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<mods:title id="6435E19F9854D8BAF7D60DDFC9A2A074">A new species of Caloptilia (Lepidoptera: Gracillariidae) from Japan and Korea</mods:title>
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<mods:namePart id="8966201B2A1DE0F7045E434DAE543981">Yagi, Sadahisa</mods:namePart>
<mods:affiliation id="286450B57979CF7C0E9102830258F838">Entomological Laboratory, Faculty of Agriculture, Kyushu University, 744 Motooka Nishiku, Fukuoka 819 - 0395, Japan. &amp; Insect Science and Creative Entomology Center, Kyushu University, 744 Motooka Nishiku, Fukuoka 819 - 0395, Japan</mods:affiliation>
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<mods:namePart id="ED8BF681CAA4D0261D41C800107CA52C">Goto, Kiyoshiro</mods:namePart>
<mods:affiliation id="A71C344A715408F376E4EBC2707773BA">Fruit Tree Research Institute, Kumamoto Prefectural Agricultural Research Center, 2566 Toyofuku, Matsubase, Uki 869 - 0524, Japan.</mods:affiliation>
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<mods:namePart id="DB283C025D0C599A9E9189533B16FA42">Sohn, Jae-Cheon</mods:namePart>
<mods:affiliation id="2638E64F8FC3A3CE090C361CE3B8784D">Department of Science Education, Gongju National University of Education, Woongjinro 27, Gongju, Chungnam 32553, Republic of Korea.</mods:affiliation>
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<heading id="E32181F9461F383882AD0F3CFE3DA2F0" bold="true" box="[151,500,1653,1680]" fontSize="11" level="1" pageId="2" pageNumber="422" reason="1">
<emphasis id="8AA2EA87461F383882AD0F3CFE3DA2F0" bold="true" box="[151,500,1653,1680]" pageId="2" pageNumber="422">
<taxonomicName id="7FD64D16461F383882AD0F3CFE63A2EF" authority="Yagi &amp; Goto &amp; Sohn, 2024" authorityName="Yagi &amp; Goto &amp; Sohn" authorityYear="2024" box="[151,426,1653,1680]" class="Insecta" family="Gracillariidae" genus="Caloptilia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="rhynchosiae" status="sp. nov.">
<emphasis id="8AA2EA87461F383882AD0F3CFE63A2EF" bold="true" box="[151,426,1653,1680]" italics="true" pageId="2" pageNumber="422">Caloptilia rhynchosiae</emphasis>
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<taxonomicNameLabel id="919157FC461F3838838B0F3CFE3DA2F0" box="[433,500,1653,1679]" pageId="2" pageNumber="422" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="B8693695461F383882AD0FD0FED0A2CC" blockId="2.[151,500,1653,1715]" box="[151,281,1689,1715]" pageId="2" pageNumber="422">
(
<figureCitation id="20ED2A10461F383882A50FD0FED9A2CC" box="[159,272,1689,1715]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 47" captionStart-2="FIGURES 810" captionStart-3="FIGURES 1118" captionStartId-0="3.[151,264,1082,1106]" captionStartId-1="4.[151,264,1388,1412]" captionStartId-2="5.[151,264,1623,1647]" captionStartId-3="6.[151,264,1918,1942]" captionTargetBox-0="[156,1436,179,1045]" captionTargetBox-1="[153,1394,199,1315]" captionTargetBox-2="[211,1376,395,1548]" captionTargetBox-3="[151,1435,181,1894]" captionTargetId-0="figure-28@3.[150,1437,179,1057]" captionTargetId-1="figure-19@4.[153,1434,181,1364]" captionTargetId-2="figure-79@5.[211,1376,345,1599]" captionTargetId-3="figure-19@6.[151,1435,181,1894]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 13. Habitus of Caloptilia rhynchosiae n. sp. 1. Holotype, male (Korea: Jeonnam); 2. Paratype, female (Japan: Kumamoto); 3. Paratype, male (Japan: Fukuoka). Scale bars = 2.5 mm." captionText-1="FIGURES 47. Male genitalia of Caloptilia rhynchosiae n. sp. 4. Whole genitalia with saccular process spread out and phallus removed (paratype: SY1326); 5. Valvae (paratype: SY1508); 6. Phallus (paratype: SY1508); 7. Genital capsule (paratype: SY1508). Scale bar = 0.5 mm." captionText-2="FIGURES 810. Female genitalia of Caloptilia rhynchosiae n. sp. (paratype: SY1506). 8. Whole genitalia; 9. Ostium bursae; 10. Dentate signa. Scale bars = 0.5 mm (10), 0.1 mm (11, 12)." captionText-3="FIGURES 1118. Biology of Caloptilia rhynchosiae n. sp. 11. Mine on the upper side of host leaf; 12. Mine on the underside of host leaf; 13. Young larva in the mine; 14, 15. Leaf rolls; 16. Mature larva; 17. Cocoon; 17. Resting adult." figureDoi-0="http://doi.org/10.5281/zenodo.11101969" figureDoi-1="http://doi.org/10.5281/zenodo.11101971" figureDoi-2="http://doi.org/10.5281/zenodo.11101977" figureDoi-3="http://doi.org/10.5281/zenodo.11101979" httpUri-0="https://zenodo.org/record/11101969/files/figure.png" httpUri-1="https://zenodo.org/record/11101971/files/figure.png" httpUri-2="https://zenodo.org/record/11101977/files/figure.png" httpUri-3="https://zenodo.org/record/11101979/files/figure.png" pageId="2" pageNumber="422">Figs 118</figureCitation>
)
</paragraph>
<paragraph id="B8693695461F383882AD0FABFD0DA3D0" blockId="2.[151,1437,1761,2004]" pageId="2" pageNumber="422">
<emphasis id="8AA2EA87461F383882AD0FABFEC0A284" box="[151,265,1762,1787]" italics="true" pageId="2" pageNumber="422">Diagnosis</emphasis>
. This species is similar to
<taxonomicName id="7FD64D16461F3838800F0FA8FC05A284" authority="Kumata, 1982" authorityName="Kumata" authorityYear="1982" box="[565,972,1761,1788]" class="Insecta" family="Gracillariidae" genus="Caloptilia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="leucothoes">
<emphasis id="8AA2EA87461F3838800F0FA8FCEFA284" box="[565,806,1761,1787]" italics="true" pageId="2" pageNumber="422">Caloptilia leucothoes</emphasis>
<bibRefCitation id="DC474B64461F383881170FABFC05A284" author="Kumata, T." box="[813,972,1761,1788]" pageId="2" pageNumber="422" pagination="1 - 186" refId="ref5027" refString="Kumata, T. (1982) A taxonomic revision of the Gracillaria group occurring in Japan (Lepidoptera: Gracillariidae). Insecta Matsumurana, New Series, 26, 1 - 186." type="journal article" year="1982">
Kumata,
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in the external appearance and also to
<taxonomicName id="7FD64D16461F383887B90FABFE24A360" authority="(Matsumura, 1931)" baseAuthorityName="Matsumura" baseAuthorityYear="1931" class="Insecta" family="Gracillariidae" genus="Caloptilia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="sapporella">
<emphasis id="8AA2EA87461F383887B90FABFEC7A360" italics="true" pageId="2" pageNumber="422">C. sapporella</emphasis>
(
<bibRefCitation id="DC474B64461F383883270E4CFE2CA360" author="Matsumura, S." box="[285,485,1797,1823]" pageId="2" pageNumber="422" refId="ref5241" refString="Matsumura, S. (1931) 6000 Illustrated Insects of the Japan-Empire. Toko-Shoin, Tokyo, 1689 pp." type="book" year="1931">Matsumura, 1931</bibRefCitation>
)
</taxonomicName>
in having the cucullus through a neck in the male genitalia. It differs from the latter two in the presence of a saccular process on the valva of the male genitalia. In the female genitalia,
<taxonomicName id="7FD64D16461F383886C90E63FA55A33C" authority="Yagi &amp; Goto &amp; Sohn, 2024" authorityName="Yagi &amp; Goto &amp; Sohn" authorityYear="2024" box="[1267,1436,1833,1860]" class="Insecta" family="Gracillariidae" genus="Caloptilia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="rhynchosiae" status="n. sp.">
<emphasis id="8AA2EA87461F383886C90E63FA55A33C" box="[1267,1436,1833,1860]" italics="true" pageId="2" pageNumber="422">C. rhynchosiae</emphasis>
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<emphasis id="8AA2EA87461F383882AD0E04FF1FA318" bold="true" box="[151,214,1869,1895]" pageId="2" pageNumber="422">
<taxonomicNameLabel id="919157FC461F383882AD0E04FF1FA318" box="[151,214,1869,1895]" pageId="2" pageNumber="422" rank="species">n. sp.</taxonomicNameLabel>
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is similar to
<taxonomicName id="7FD64D16461F383883510E07FE12A318" baseAuthorityName="van Deventer" baseAuthorityYear="1904" box="[363,475,1869,1896]" class="Insecta" family="Gracillariidae" genus="Caloptilia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="soyella">
<emphasis id="8AA2EA87461F383883510E07FE12A318" box="[363,475,1869,1896]" italics="true" pageId="2" pageNumber="422">C. soyella</emphasis>
</taxonomicName>
in having the entirely-membranous ductus bursae. It, however, differs from the latter in the presence of the roundly-bulged basal area of ductus seminalis (narrow and then bulged in
<taxonomicName id="7FD64D16461F383886E70E3BFA87A3F4" baseAuthorityName="van Deventer" baseAuthorityYear="1904" box="[1245,1358,1905,1932]" class="Insecta" family="Gracillariidae" genus="Caloptilia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="soyella">
<emphasis id="8AA2EA87461F383886E70E3BFA87A3F4" box="[1245,1358,1905,1932]" italics="true" pageId="2" pageNumber="422">C. soyella</emphasis>
</taxonomicName>
) and a mortar-shaped sterigma (M-shaped in
<taxonomicName id="7FD64D16461F3838807E0EDFFD7AA3D0" baseAuthorityName="van Deventer" baseAuthorityYear="1904" box="[580,691,1941,1968]" class="Insecta" family="Gracillariidae" genus="Caloptilia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="soyella">
<emphasis id="8AA2EA87461F3838807E0EDFFD7AA3D0" box="[580,691,1941,1968]" italics="true" pageId="2" pageNumber="422">C. soyella</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="B8693695461F383882FD0EF3FC74A3AC" blockId="2.[151,1437,1761,2004]" box="[199,957,1977,2004]" pageId="2" pageNumber="422">
<emphasis id="8AA2EA87461F383882FD0EF3FE82A3AC" box="[199,331,1978,2003]" italics="true" pageId="2" pageNumber="422">Description</emphasis>
. Habitus (
<figureCitation id="20ED2A10461F383883850EF0FDD4A3AC" box="[447,541,1977,2003]" captionStart="FIGURES 13" captionStartId="3.[151,264,1082,1106]" captionTargetBox="[156,1436,179,1045]" captionTargetId="figure-28@3.[150,1437,179,1057]" captionTargetPageId="3" captionText="FIGURES 13. Habitus of Caloptilia rhynchosiae n. sp. 1. Holotype, male (Korea: Jeonnam); 2. Paratype, female (Japan: Kumamoto); 3. Paratype, male (Japan: Fukuoka). Scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11101969" httpUri="https://zenodo.org/record/11101969/files/figure.png" pageId="2" pageNumber="422">Figs 13</figureCitation>
)—No sexual dimorphism exhibited.
</paragraph>
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<paragraph id="B8693695461E383982AD0D73FCA3A009" blockId="3.[151,1437,1082,1143]" pageId="3" pageNumber="423">
<emphasis id="8AA2EA87461E383982AD0D73FEF4A02C" bold="true" box="[151,317,1082,1107]" pageId="3" pageNumber="423">FIGURES 13.</emphasis>
Habitus of
<taxonomicName id="7FD64D16461E383983860D72FD6EA02C" authority="Yagi &amp; Goto &amp; Sohn, 2024" authorityName="Yagi &amp; Goto &amp; Sohn" authorityYear="2024" box="[444,679,1083,1107]" class="Insecta" family="Gracillariidae" genus="Caloptilia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="423" phylum="Arthropoda" rank="species" species="rhynchosiae" status="sp. nov.">
<emphasis id="8AA2EA87461E383983860D72FD6EA02C" box="[444,679,1083,1107]" italics="true" pageId="3" pageNumber="423">Caloptilia rhynchosiae</emphasis>
</taxonomicName>
<emphasis id="8AA2EA87461E3839808A0D72FD25A02C" bold="true" box="[688,748,1083,1107]" pageId="3" pageNumber="423">
<taxonomicNameLabel id="919157FC461E3839808A0D72FD25A02C" box="[688,748,1083,1107]" pageId="3" pageNumber="423" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
1. Holotype, male (Korea: Jeonnam); 2. Paratype, female (Japan: Kumamoto); 3. Paratype, male (Japan: Fukuoka). Scale bars = 2.5 mm.
</paragraph>
</caption>
<paragraph id="B8693695461E383982FD0D80FE79A10C" blockId="3.[151,1437,1225,1900]" pageId="3" pageNumber="423">
Head: Vertex brownish gray, with purple luster, tinged with white anteriorly; frons creamy white. Antenna ca. 1.1x longer than forewing; scape brownish gray; flagellum brownish gray with pale brownish gray annulations. Labial palpus upcurved; 1
<superScript id="4FA39BDD461E383983860C58FE0EA160" attach="left" box="[444,455,1297,1311]" fontSize="6" pageId="3" pageNumber="423">st</superScript>
segment 1/4 as long as 2
<superScript id="4FA39BDD461E383980D80C58FD3BA160" attach="left" box="[738,754,1297,1311]" fontSize="6" pageId="3" pageNumber="423">nd</superScript>
segment, white; 2
<superScript id="4FA39BDD461E383981FA0C58FC19A160" attach="left" box="[960,976,1297,1311]" fontSize="6" pageId="3" pageNumber="423">nd</superScript>
segment as long as 3
<superScript id="4FA39BDD461E383986FB0C58FB06A160" attach="left" box="[1217,1231,1297,1311]" fontSize="6" pageId="3" pageNumber="423">rd</superScript>
segment, white to cream, intermixed with dark brown scales ventrally; 3
<superScript id="4FA39BDD461E383980CD0C7CFCCCA13C" attach="left" box="[759,773,1333,1347]" fontSize="6" pageId="3" pageNumber="423">rd</superScript>
segment acuminating apically, white to cream, tinged with dark brown on distal 1/3.
</paragraph>
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Thorax: Patagium brownish gray; tegula and mesonotum pale-purplish gray, intermixed with pale grayish orange scales. Forewing length
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, narrow, yellow, intermixed with lustrous, purplish brown scales on dorsal and distal areas; costa with 614 black strigulae, tinged with dark brownish gray basally; three apical strigulae black in some individuals; often one or two spots present at basal 1/4 and 1/2 of forewing; cilia dark brownish gray, peppered with pale gray on apical area and termen. Hindwing dark brownish gray, paler to base; cilia dark brownish gray. Fore- and midlegs with coxa, femur and tibia dark brown, sparsely irrorated with pale brownish gray; tarsomere white, with black ring distally. Hindleg with coxa cream, sparsely peppered with dark brown; femur pale gray with dark brown band at basal 3/5 laterally; tibia pale gray dorsally, white ventrally, intermixed with dark brown scales distally; tarsomeres pale brownish gray.
</paragraph>
<paragraph id="B8693695461E383982FD0F88FC8CA2A3" blockId="3.[151,1437,1225,1900]" box="[199,837,1729,1756]" pageId="3" pageNumber="423">Abdomen: terga lustrous, gray; sterna white or pale gray.</paragraph>
<paragraph id="B8693695461E383982FD0FACFD33A314" blockId="3.[151,1437,1225,1900]" pageId="3" pageNumber="423">
Male genitalia (
<figureCitation id="20ED2A10461E3839834D0FACFE1EA280" box="[375,471,1765,1791]" captionStart="FIGURES 47" captionStartId="4.[151,264,1388,1412]" captionTargetBox="[153,1394,199,1315]" captionTargetId="figure-19@4.[153,1434,181,1364]" captionTargetPageId="4" captionText="FIGURES 47. Male genitalia of Caloptilia rhynchosiae n. sp. 4. Whole genitalia with saccular process spread out and phallus removed (paratype: SY1326); 5. Valvae (paratype: SY1508); 6. Phallus (paratype: SY1508); 7. Genital capsule (paratype: SY1508). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11101971" httpUri="https://zenodo.org/record/11101971/files/figure.png" pageId="3" pageNumber="423">Figs 47</figureCitation>
)—Tegumen linguiform, narrowed basally; subscaphium of even width. Valva narrow, of even width in basal half; cucullus subtriangular, long-hairy except distal area with short setae; sacculus broad, 2/5 as long as valva, with large horn-like process at distal end. Vinculum linguiform, narrowly-round apically. Phallus narrow, slightly narrowed distally, with 1420 cornuti.
</paragraph>
<caption id="ECA9661D4619383E82AD0C25FE04A1B3" ID-DOI="http://doi.org/10.5281/zenodo.11101971" ID-Zenodo-Dep="11101971" httpUri="https://zenodo.org/record/11101971/files/figure.png" pageId="4" pageNumber="424" startId="4.[151,264,1388,1412]" targetBox="[153,1394,199,1315]" targetPageId="4" targetType="figure">
<paragraph id="B86936954619383E82AD0C25FE04A1B3" blockId="4.[151,1437,1388,1484]" pageId="4" pageNumber="424">
<emphasis id="8AA2EA874619383E82AD0C25FEF3A1FB" bold="true" box="[151,314,1388,1412]" pageId="4" pageNumber="424">FIGURES 47.</emphasis>
Male genitalia of
<taxonomicName id="7FD64D164619383E83C80C25FD10A1FB" authority="Yagi &amp; Goto &amp; Sohn, 2024" authorityName="Yagi &amp; Goto &amp; Sohn" authorityYear="2024" box="[498,729,1388,1412]" class="Insecta" family="Gracillariidae" genus="Caloptilia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="424" phylum="Arthropoda" rank="species" species="rhynchosiae" status="sp. nov.">
<emphasis id="8AA2EA874619383E83C80C25FD10A1FB" box="[498,729,1388,1412]" italics="true" pageId="4" pageNumber="424">Caloptilia rhynchosiae</emphasis>
</taxonomicName>
<emphasis id="8AA2EA874619383E80DA0C25FCD1A1FB" bold="true" box="[736,792,1388,1412]" pageId="4" pageNumber="424">
<taxonomicNameLabel id="919157FC4619383E80DA0C25FCD1A1FB" box="[736,792,1388,1412]" pageId="4" pageNumber="424" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
4. Whole genitalia with saccular process spread out and phallus removed (paratype: SY1326); 5. Valvae (paratype: SY1508); 6. Phallus (paratype: SY1508); 7. Genital capsule (paratype: SY1508). Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="B86936954619383E82FD0CB3FDECA2B7" blockId="4.[151,1437,1530,2025]" pageId="4" pageNumber="424">
Female genitalia (
<figureCitation id="20ED2A104619383E83A10CB3FDC4A26B" box="[411,525,1530,1556]" captionStart="FIGURES 810" captionStartId="5.[151,264,1623,1647]" captionTargetBox="[211,1376,395,1548]" captionTargetId="figure-79@5.[211,1376,345,1599]" captionTargetPageId="5" captionText="FIGURES 810. Female genitalia of Caloptilia rhynchosiae n. sp. (paratype: SY1506). 8. Whole genitalia; 9. Ostium bursae; 10. Dentate signa. Scale bars = 0.5 mm (10), 0.1 mm (11, 12)." figureDoi="http://doi.org/10.5281/zenodo.11101977" httpUri="https://zenodo.org/record/11101977/files/figure.png" pageId="4" pageNumber="424">Figs 810</figureCitation>
)—Papilla analis pad-shaped. Apophysis posterioris broad basally, as long as apophysis anterioris; apophysis anterioris relatively broad, tapering distally. Ostium bursae present between sternite VII and VIII. Sterigma well-sclerotized, mortar-shaped with minute granules. Ductus bursae entirely membranous, with minute granules between ostium bursae and opening of ductus seminalis. Ductus seminalis arising from the posterior 4/5 of ductus bursae, roundly-bulged basally. Corpus bursae ovate, wrinkled; a pair of signa curved, sickle-shaped, with dentate inner margins.
</paragraph>
<paragraph id="B86936954619383E82FD0F9AFC60A36F" blockId="4.[151,1437,1530,2025]" pageId="4" pageNumber="424">
<emphasis id="8AA2EA874619383E82FD0F9AFECFA293" box="[199,262,1747,1772]" italics="true" pageId="4" pageNumber="424">Types</emphasis>
.
<materialsCitation id="08BE3CC84619383E832C0F9BFC60A36F" collectingDate="2017-07-16" collectionCode="GJUE" collectorName="Is. Jaeundo &amp; JC Sohn" country="South Korea" county="Sinan-gun" latitude="34.895695" location="Gojang-ri" longLatPrecision="1" longitude="126.007614" municipality="Jaeun-myeon" pageId="4" pageNumber="424" specimenCount="1" stateProvince="Jeonnam" typeStatus="holotype">
<typeStatus id="676D88374619383E832C0F9BFEB6A293" box="[278,383,1746,1772]" pageId="4" pageNumber="424" type="holotype">Holotype</typeStatus>
—male,
<collectingCountry id="C0C176054619383E83DB0F9AFDE3A292" box="[481,554,1747,1773]" name="South Korea" pageId="4" pageNumber="424">Korea</collectingCountry>
:
<collectingRegion id="7A12F8774619383E800C0F9AFD16A292" box="[566,735,1747,1773]" country="South Korea" name="Jeonnam" pageId="4" pageNumber="424">Jeonnam Prov.</collectingRegion>
,
<collectingCounty id="51084E194619383E80D60F9BFCABA293" box="[748,866,1746,1772]" pageId="4" pageNumber="424">Sinan-gun</collectingCounty>
,
<collectorName id="152353434619383E81550F9AFC3BA293" box="[879,1010,1746,1773]" pageId="4" pageNumber="424">Is. Jaeundo</collectorName>
,
<collectingMunicipality id="580DACEF4619383E81C40F9AFB5FA292" box="[1022,1174,1747,1773]" pageId="4" pageNumber="424">Jaeun-myeon</collectingMunicipality>
,
<location id="BD09604E4619383E869E0F9BFADDA293" LSID="urn:lsid:plazi:treatment:307F8783461F383F82AD0F3CFC27A383:BD09604E4619383E869E0F9BFADDA293" box="[1188,1300,1746,1772]" country="South Korea" county="Sinan-gun" latitude="34.895695" longLatPrecision="1" longitude="126.007614" municipality="Jaeun-myeon" name="Gojang-ri" pageId="4" pageNumber="424" stateProvince="Jeonnam">Gojang-ri</location>
,
<location id="BD09604E4619383E871A0F9BFA9FA293" LSID="urn:lsid:plazi:treatment:307F8783461F383F82AD0F3CFC27A383:BD09604E4619383E871A0F9BFA9FA293" box="[1312,1366,1746,1772]" country="South Korea" county="Sinan-gun" latitude="34.895695" longLatPrecision="1" longitude="126.007614" municipality="Jaeun-myeon" name="Oegi" pageId="4" pageNumber="424" stateProvince="Jeonnam">Oegi</location>
sand-dunes (
<geoCoordinate id="DDE250524619383E82DD0FB1FE4AA36C" box="[231,387,1784,1811]" degrees="34" direction="north" minutes="53" orientation="latitude" pageId="4" pageNumber="424" precision="1" seconds="44.5" value="34.895695">34°5344.5″N</geoCoordinate>
<geoCoordinate id="DDE250524619383E83B00FB1FDFAA36C" box="[394,563,1784,1811]" degrees="126" direction="east" minutes="00" orientation="longitude" pageId="4" pageNumber="424" precision="1" seconds="27.4" value="126.007614">126°0027.4″E</geoCoordinate>
),
<date id="CC6810554619383E807C0FBFFD1EA36F" box="[582,727,1782,1808]" pageId="4" pageNumber="424" value="2017-07-16">
<collectingDate id="DC2CE9BD4619383E807C0FBFFD1EA36F" box="[582,727,1782,1808]" pageId="4" pageNumber="424" value="2017-07-16">16 July 2017</collectingDate>
</date>
(
<collectorName id="152353434619383E80DC0FBFFC83A36F" box="[742,842,1782,1808]" pageId="4" pageNumber="424">JC Sohn</collectorName>
),
<collectionCode id="DEC7AE504619383E81670FBFFC6CA36F" box="[861,933,1782,1808]" pageId="4" pageNumber="424">GJUE</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B86936954619383E82FD0E52FA54A397" blockId="4.[151,1437,1530,2025]" pageId="4" pageNumber="424">
<materialsCitation id="08BE3CC84619383E82FD0E52FDCFA34B" box="[199,518,1818,1845]" country="Japan" location="Japan" pageId="4" pageNumber="424" specimenCount="15" specimenCount-female="7" specimenCount-male="8" typeStatus="paratype">
<typeStatus id="676D88374619383E82FD0E52FEFDA34A" box="[199,308,1819,1845]" pageId="4" pageNumber="424" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="AED0FD1C4619383E837D0E53FEA0A34B" box="[327,361,1818,1844]" count="8" pageId="4" pageNumber="424" type="male">8♂</specimenCount>
<specimenCount id="AED0FD1C4619383E83520E53FE4DA34B" box="[360,388,1818,1844]" count="7" pageId="4" pageNumber="424" type="female">7♀</specimenCount>
)—[
<collectingCountry id="C0C176054619383E838F0E53FE37A34B" box="[437,510,1818,1844]" name="Japan" pageId="4" pageNumber="424">Japan</collectingCountry>
]
</materialsCitation>
<materialsCitation id="08BE3CC84619383E802A0E53FD33A302" collectingDate="2020-06-02" collectingDateMax="2020-06-10" collectingDateMin="2020-06-02" collectionCode="ELKU" collectorName="S Yagi" country="Japan" county="Kyushu" latitude="33.598" location="Nishi-ku" longLatPrecision="1" longitude="130.2168" municipality="Fukuoka-shi" pageId="4" pageNumber="424" specimenCount="1" specimenCount-male="1" stateProvince="Fukuoka" typeStatus="paratype">
<specimenCount id="AED0FD1C4619383E802A0E53FDFDA34B" box="[528,564,1818,1844]" count="1" pageId="4" pageNumber="424" type="male">1♂</specimenCount>
,
<collectingCounty id="51084E194619383E807E0E53FD57A34B" box="[580,670,1818,1844]" pageId="4" pageNumber="424">Kyushu</collectingCounty>
,
<collectingRegion id="7A12F8774619383E80970E53FC9BA34B" box="[685,850,1818,1844]" country="Japan" name="Fukuoka" pageId="4" pageNumber="424">Fukuoka Pref.</collectingRegion>
,
<collectingMunicipality id="580DACEF4619383E815A0E53FC26A34B" box="[864,1007,1818,1844]" pageId="4" pageNumber="424">Fukuoka-shi</collectingMunicipality>
,
<location id="BD09604E4619383E81C70E53FBABA34B" LSID="urn:lsid:plazi:treatment:307F8783461F383F82AD0F3CFC27A383:BD09604E4619383E81C70E53FBABA34B" box="[1021,1122,1818,1844]" country="Japan" county="Kyushu" latitude="33.598" longLatPrecision="1" longitude="130.2168" municipality="Fukuoka-shi" name="Nishi-ku" pageId="4" pageNumber="424" stateProvince="Fukuoka">Nishi-ku</location>
,
<location id="BD09604E4619383E864B0E53FB2EA34B" LSID="urn:lsid:plazi:treatment:307F8783461F383F82AD0F3CFC27A383:BD09604E4619383E864B0E53FB2EA34B" box="[1137,1255,1818,1844]" country="Japan" county="Kyushu" latitude="33.598" longLatPrecision="1" longitude="130.2168" municipality="Fukuoka-shi" name="Kuwabara" pageId="4" pageNumber="424" stateProvince="Fukuoka">Kuwabara</location>
,
<location id="BD09604E4619383E86CF0E52FEA1A327" LSID="urn:lsid:plazi:treatment:307F8783461F383F82AD0F3CFC27A383:BD09604E4619383E86CF0E52FEA1A327" country="Japan" county="Kyushu" latitude="33.598" longLatPrecision="1" longitude="130.2168" municipality="Fukuoka-shi" name="Ito Campus of Kyushu University" pageId="4" pageNumber="424" stateProvince="Fukuoka">Ito Campus of Kyushu University</location>
(
<geoCoordinate id="DDE250524619383E834F0E77FDDCA326" box="[373,533,1854,1881]" degrees="33" direction="north" minutes="35" orientation="latitude" pageId="4" pageNumber="424" precision="1" seconds="52.8" value="33.598">33°3552.8”N</geoCoordinate>
<geoCoordinate id="DDE250524619383E80230E77FD0DA326" box="[537,708,1854,1881]" degrees="130" direction="east" minutes="13" orientation="longitude" pageId="4" pageNumber="424" precision="1" seconds="00.5" value="130.2168">130°1300.5”E</geoCoordinate>
),
<date id="CC6810554619383E80EC0E77FC92A327" box="[726,859,1854,1881]" pageId="4" pageNumber="424" value="2020-06-02">
<collectingDate id="DC2CE9BD4619383E80EC0E77FC92A327" box="[726,859,1854,1881]" pageId="4" pageNumber="424" value="2020-06-02">2 June 2020</collectingDate>
</date>
, larva
<collectedFrom id="0BCA75684619383E81A50E76FB70A327" box="[927,1209,1854,1881]" pageId="4" pageNumber="424">
on
<taxonomicName id="7FD64D164619383E81850E77FB70A327" box="[959,1209,1854,1880]" class="Magnoliopsida" family="Fabaceae" genus="Rhynchosia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="424" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="8AA2EA874619383E81850E77FB70A327" box="[959,1209,1854,1880]" italics="true" pageId="4" pageNumber="424">Rhynchosia tomentosa</emphasis>
</taxonomicName>
</collectedFrom>
, emerged on
<date id="CC6810554619383E87730E77FF06A303" pageId="4" pageNumber="424" value="2020-06-10">
<collectingDate id="DC2CE9BD4619383E87730E77FF06A303" pageId="4" pageNumber="424" value="2020-06-10">10 June 2020</collectingDate>
</date>
(
<collectorName id="152353434619383E82E50E2BFEE4A303" box="[223,301,1890,1916]" pageId="4" pageNumber="424">S Yagi</collectorName>
), [GSN] SY1326, [COI] SaY777,
<collectionCode id="DEC7AE504619383E80960E2AFD33A302" box="[684,762,1891,1917]" pageId="4" pageNumber="424">ELKU</collectionCode>
</materialsCitation>
;
<materialsCitation id="08BE3CC84619383E813C0E2BFBFEA3DE" collectingDate="2023-07-03" collectingDateMax="2023-08-29" collectingDateMin="2023-07-03" collectionCode="ELKU" collectorName="S Yagi" country="Japan" county="Kyushu" latitude="33.598" location="Nishi-ku" longLatPrecision="1" longitude="130.2168" municipality="Fukuoka-shi" pageId="4" pageNumber="424" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Fukuoka" typeStatus="paratype">
<specimenCount id="AED0FD1C4619383E813C0E2BFCE4A303" box="[774,813,1890,1916]" count="3" pageId="4" pageNumber="424" type="male">3♂</specimenCount>
<specimenCount id="AED0FD1C4619383E81170E2BFC84A303" box="[813,845,1890,1916]" count="3" pageId="4" pageNumber="424" type="female">3♀</specimenCount>
, ditto,
<date id="CC6810554619383E81A10E2BFBE9A303" box="[923,1056,1890,1916]" pageId="4" pageNumber="424" value="2023-07-03">
<collectingDate id="DC2CE9BD4619383E81A10E2BFBE9A303" box="[923,1056,1890,1916]" pageId="4" pageNumber="424" value="2023-07-03">3 July 2023</collectingDate>
</date>
, larvae
<collectedFrom id="0BCA75684619383E864C0E2AFA5CA303" box="[1142,1429,1890,1917]" pageId="4" pageNumber="424">
on
<taxonomicName id="7FD64D164619383E86A30E2BFA5CA303" box="[1177,1429,1890,1916]" class="Magnoliopsida" family="Fabaceae" genus="Rhynchosia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="424" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="8AA2EA874619383E86A30E2BFA5CA303" box="[1177,1429,1890,1916]" italics="true" pageId="4" pageNumber="424">Rhynchosia tomentosa</emphasis>
</taxonomicName>
</collectedFrom>
, emerged on
<date id="CC6810554619383E83180ECFFE38A3DF" box="[290,497,1926,1953]" pageId="4" pageNumber="424" value="2023-08-06" valueMax="2023-08-29" valueMin="2023-08-06">
<collectingDate id="DC2CE9BD4619383E83180ECFFE38A3DF" box="[290,497,1926,1953]" pageId="4" pageNumber="424" value="2023-08-06" valueMax="2023-08-29" valueMin="2023-08-06">629 August 2023</collectingDate>
</date>
(
<collectorName id="152353434619383E803B0ECFFD99A3DF" box="[513,592,1926,1952]" pageId="4" pageNumber="424">S Yagi</collectorName>
), [GSN] SY1506 (
<specimenCount id="AED0FD1C4619383E811E0ECFFCFEA3DF" box="[804,823,1926,1952]" count="1" pageId="4" pageNumber="424" type="female"></specimenCount>
), SY1508 (
<specimenCount id="AED0FD1C4619383E81860ECFFC1AA3DF" box="[956,979,1926,1952]" count="1" pageId="4" pageNumber="424" type="male"></specimenCount>
),
<collectionCode id="DEC7AE504619383E81D30ECEFBFEA3DE" box="[1001,1079,1927,1953]" pageId="4" pageNumber="424">ELKU</collectionCode>
</materialsCitation>
;
<materialsCitation id="08BE3CC84619383E86790ECFFE93A396" collectingDate="2023-06-28" collectingDateMax="2023-08-01" collectingDateMin="2023-06-28" collectionCode="ELKU" collectorName="K Goto" country="Japan" county="Kyushu" location="Matsubase-machi" municipality="Uki-shi" pageId="4" pageNumber="424" specimenCount="1" specimenCount-female="1" stateProvince="Kumamoto" typeStatus="paratype">
<specimenCount id="AED0FD1C4619383E86790ECFFBAFA3DF" box="[1091,1126,1926,1952]" count="1" pageId="4" pageNumber="424" type="female">1♀</specimenCount>
,
<collectingCounty id="51084E194619383E864E0ECFFB07A3DF" box="[1140,1230,1926,1952]" pageId="4" pageNumber="424">Kyushu</collectingCounty>
,
<collectingRegion id="7A12F8774619383E86E00ECEFA50A3DF" box="[1242,1433,1926,1953]" country="Japan" name="Kumamoto" pageId="4" pageNumber="424">Kumamoto Pref.</collectingRegion>
,
<collectingMunicipality id="580DACEF4619383E82AD0EE3FF27A3BB" box="[151,238,1962,1988]" pageId="4" pageNumber="424">Uki-shi</collectingMunicipality>
,
<location id="BD09604E4619383E82C10EE3FE0AA3BB" LSID="urn:lsid:plazi:treatment:307F8783461F383F82AD0F3CFC27A383:BD09604E4619383E82C10EE3FE0AA3BB" box="[251,451,1962,1988]" country="Japan" county="Kyushu" municipality="Uki-shi" name="Matsubase-machi" pageId="4" pageNumber="424" stateProvince="Kumamoto">Matsubase-machi</location>
,
<location id="BD09604E4619383E83EA0EE3FDF6A3BB" LSID="urn:lsid:plazi:treatment:307F8783461F383F82AD0F3CFC27A383:BD09604E4619383E83EA0EE3FDF6A3BB" box="[464,575,1962,1988]" country="Japan" county="Kyushu" municipality="Uki-shi" name="Toyofuku" pageId="4" pageNumber="424" stateProvince="Kumamoto">Toyofuku</location>
,
<date id="CC6810554619383E80760EE3FD21A3BB" box="[588,744,1962,1989]" pageId="4" pageNumber="424" value="2023-06-28">
<collectingDate id="DC2CE9BD4619383E80760EE3FD21A3BB" box="[588,744,1962,1989]" pageId="4" pageNumber="424" value="2023-06-28">28 June 2023</collectingDate>
</date>
, larva
<collectedFrom id="0BCA75684619383E810F0EE2FB91A3BB" box="[821,1112,1962,1989]" pageId="4" pageNumber="424">
on
<taxonomicName id="7FD64D164619383E81600EE3FB91A3BB" box="[858,1112,1962,1988]" class="Magnoliopsida" family="Fabaceae" genus="Rhynchosia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="424" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="8AA2EA874619383E81600EE3FB91A3BB" box="[858,1112,1962,1988]" italics="true" pageId="4" pageNumber="424">Rhynchosia tomentosa</emphasis>
</taxonomicName>
</collectedFrom>
, emerged on
<date id="CC6810554619383E86CF0EE3FA55A3BB" box="[1269,1436,1962,1989]" pageId="4" pageNumber="424" value="2023-08-01">
<collectingDate id="DC2CE9BD4619383E86CF0EE3FA55A3BB" box="[1269,1436,1962,1989]" pageId="4" pageNumber="424" value="2023-08-01">1 August 2023</collectingDate>
</date>
(
<collectorName id="152353434619383E82A50E87FF31A397" box="[159,248,1998,2024]" pageId="4" pageNumber="424">K Goto</collectorName>
),
<collectionCode id="DEC7AE504619383E83360E86FE93A396" box="[268,346,1999,2025]" pageId="4" pageNumber="424">ELKU</collectionCode>
</materialsCitation>
;
<materialsCitation id="08BE3CC84619383E835C0E87FA54A397" box="[358,1437,1998,2025]" collectingDate="2023-07-13" collectingDateMax="2023-07-26" collectingDateMin="2023-07-13" collectionCode="ELKU" collectorName="K Goto" country="Japan" county="Kyushu" location="Matsubase-machi" municipality="Uki-shi" pageId="4" pageNumber="424" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kumamoto" typeStatus="paratype">
<specimenCount id="AED0FD1C4619383E835C0E87FE47A397" box="[358,398,1998,2024]" count="1" pageId="4" pageNumber="424" type="male">1♂</specimenCount>
<specimenCount id="AED0FD1C4619383E83B40E87FE64A397" box="[398,429,1998,2024]" count="1" pageId="4" pageNumber="424" type="female">1♀</specimenCount>
, ditto,
<date id="CC6810554619383E83C70E87FD5BA397" box="[509,658,1998,2024]" pageId="4" pageNumber="424" value="2023-07-13">
<collectingDate id="DC2CE9BD4619383E83C70E87FD5BA397" box="[509,658,1998,2024]" pageId="4" pageNumber="424" value="2023-07-13">13 July 2023</collectingDate>
</date>
, larvae
<collectedFrom id="0BCA75684619383E80D30E86FBC3A397" box="[745,1034,1998,2025]" pageId="4" pageNumber="424">
on
<taxonomicName id="7FD64D164619383E81370E87FBC3A397" box="[781,1034,1998,2024]" class="Magnoliopsida" family="Fabaceae" genus="Rhynchosia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="424" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="8AA2EA874619383E81370E87FBC3A397" box="[781,1034,1998,2024]" italics="true" pageId="4" pageNumber="424">Rhynchosia tomentosa</emphasis>
</taxonomicName>
</collectedFrom>
, emerged on 24 &amp;
<date id="CC6810554619383E86DF0E87FABEA397" box="[1253,1399,1998,2024]" pageId="4" pageNumber="424" value="2023-07-26">
<collectingDate id="DC2CE9BD4619383E86DF0E87FABEA397" box="[1253,1399,1998,2024]" pageId="4" pageNumber="424" value="2023-07-26">26 July 2023</collectingDate>
</date>
(K
</materialsCitation>
</paragraph>
<paragraph id="B86936954618383F82AD09DEFE9BA562" blockId="5.[151,1437,151,285]" pageId="5" pageNumber="425">
Goto), ELKU;
<materialsCitation id="08BE3CC84618383F837A09DEFE69A4AA" collectingDate="2023-08-16" collectingDateMax="2023-09-01" collectingDateMin="2023-08-16" collectionCode="ELKU" collectorName="August &amp; K Goto" pageId="5" pageNumber="425" specimenCount="2" specimenCount-female="2">
<specimenCount id="AED0FD1C4618383F837A09DEFEACA4CE" box="[320,357,151,177]" count="2" pageId="5" pageNumber="425" type="female">2♀</specimenCount>
, ditto,
<date id="CC6810554618383F838909DEFDA1A4CE" box="[435,616,151,177]" pageId="5" pageNumber="425" value="2023-08-16">
<collectingDate id="DC2CE9BD4618383F838909DEFDA1A4CE" box="[435,616,151,177]" pageId="5" pageNumber="425" value="2023-08-16">16 August 2023</collectingDate>
</date>
, larvae
<collectedFrom id="0BCA75684618383F808509DEFC16A4CF" box="[703,991,151,177]" pageId="5" pageNumber="425">
on
<taxonomicName id="7FD64D164618383F80D909DEFC16A4CF" box="[739,991,151,177]" class="Magnoliopsida" family="Fabaceae" genus="Rhynchosia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="425" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="8AA2EA874618383F80D909DEFC16A4CF" box="[739,991,151,177]" italics="true" pageId="5" pageNumber="425">Rhynchosia tomentosa</emphasis>
</taxonomicName>
</collectedFrom>
, emerged on 28
<collectorName id="152353434618383F86A009DEFB22A4CE" box="[1178,1259,151,177]" pageId="5" pageNumber="425">August</collectorName>
&amp;
<date id="CC6810554618383F873509DEFF06A4AA" pageId="5" pageNumber="425" value="2023-09-01">
<collectingDate id="DC2CE9BD4618383F873509DEFF06A4AA" pageId="5" pageNumber="425" value="2023-09-01">1 September 2023</collectingDate>
</date>
(
<collectorName id="152353434618383F82DB09F2FEF5A4AA" box="[225,316,187,213]" pageId="5" pageNumber="425">K Goto</collectorName>
),
<collectionCode id="DEC7AE504618383F836809F2FE69A4AA" box="[338,416,187,213]" pageId="5" pageNumber="425">ELKU</collectionCode>
</materialsCitation>
.
<materialsCitation id="08BE3CC84618383F839709F2FF14A486" collectingDate="2017-07-16" collectionCode="GJUE, NIBR" collectorName="Is. Jaeundo &amp; JC Sohn" country="South Korea" county="Sinan-gun" latitude="34.895695" location="Gojang-ri" longLatPrecision="1" longitude="126.007614" municipality="Jaeun-myeon" pageId="5" pageNumber="425" specimenCount="2" specimenCount-male="2" stateProvince="Jeonnam">
[
<collectingCountry id="C0C176054618383F838E09F2FE37A4AA" box="[436,510,187,213]" name="South Korea" pageId="5" pageNumber="425">Korea</collectingCountry>
]
<specimenCount id="AED0FD1C4618383F803409F2FDFAA4AA" box="[526,563,187,213]" count="2" pageId="5" pageNumber="425" type="male">2♂</specimenCount>
, same data as holotype, [GSN] SJC-1253, [COI] JCS-COI D-325,
<collectionCode id="DEC7AE504618383F870209F2FAB5A4AA" box="[1336,1404,187,213]" pageId="5" pageNumber="425">GJUE</collectionCode>
&amp;
<collectionCode id="DEC7AE504618383F82AD0996FF14A486" box="[151,221,223,249]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="5" pageNumber="425">NIBR</collectionCode>
</materialsCitation>
;
<materialsCitation id="08BE3CC84618383F82D20996FE87A562" collectingDate="2004-06-18" collectionCode="GJUE" collectorName="Is. Geoje-do &amp; Jangmok-myeon &amp; Jangmunpo-waeseong" country="South Korea" location="Gyeongnam Prov." pageId="5" pageNumber="425" specimenCount="1" specimenCount-male="1" stateProvince="Gyeongnam">
<specimenCount id="AED0FD1C4618383F82D20996FEC4A486" box="[232,269,223,249]" count="1" pageId="5" pageNumber="425" type="male">1♂</specimenCount>
,
<collectingRegion id="7A12F8774618383F83230996FE2FA486" box="[281,486,223,249]" country="South Korea" name="Gyeongnam" pageId="5" pageNumber="425">Gyeongnam Prov.</collectingRegion>
,
<collectorName id="152353434618383F83CA0996FDB4A486" box="[496,637,223,249]" pageId="5" pageNumber="425">Is. Geoje-do</collectorName>
,
<collectorName id="152353434618383F80B20996FC8FA486" box="[648,838,223,249]" pageId="5" pageNumber="425">Jangmok-myeon</collectorName>
,
<collectorName id="152353434618383F816A0996FB80A486" box="[848,1097,223,249]" pageId="5" pageNumber="425">Jangmunpo-waeseong</collectorName>
(castle),
<date id="CC6810554618383F86950996FA8EA486" box="[1199,1351,223,249]" pageId="5" pageNumber="425" value="2004-06-18">
<collectingDate id="DC2CE9BD4618383F86950996FA8EA486" box="[1199,1351,223,249]" pageId="5" pageNumber="425" value="2004-06-18">18 June 2004</collectingDate>
</date>
, [GSN] SJC-792,
<collectionCode id="DEC7AE504618383F833C084AFE87A562" box="[262,334,259,285]" pageId="5" pageNumber="425">GJUE</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="ECA9661D4618383F82AD0F1EFCC2A2EC" ID-DOI="http://doi.org/10.5281/zenodo.11101977" ID-Zenodo-Dep="11101977" httpUri="https://zenodo.org/record/11101977/files/figure.png" pageId="5" pageNumber="425" startId="5.[151,264,1623,1647]" targetBox="[211,1376,395,1548]" targetPageId="5" targetType="figure">
<paragraph id="B86936954618383F82AD0F1EFCC2A2EC" blockId="5.[151,1436,1623,1683]" pageId="5" pageNumber="425">
<emphasis id="8AA2EA874618383F82AD0F1EFE81A210" bold="true" box="[151,328,1623,1647]" pageId="5" pageNumber="425">FIGURES 810.</emphasis>
Female genitalia of
<taxonomicName id="7FD64D164618383F80230F1EFCCBA210" authority="Yagi &amp; Goto &amp; Sohn, 2024" authorityName="Yagi &amp; Goto &amp; Sohn" authorityYear="2024" box="[537,770,1623,1647]" class="Insecta" family="Gracillariidae" genus="Caloptilia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="425" phylum="Arthropoda" rank="species" species="rhynchosiae" status="sp. nov.">
<emphasis id="8AA2EA874618383F80230F1EFCCBA210" box="[537,770,1623,1647]" italics="true" pageId="5" pageNumber="425">Caloptilia rhynchosiae</emphasis>
</taxonomicName>
<emphasis id="8AA2EA874618383F81330F1EFC8AA210" bold="true" box="[777,835,1623,1647]" pageId="5" pageNumber="425">
<taxonomicNameLabel id="919157FC4618383F81330F1EFC8AA210" box="[777,835,1623,1647]" pageId="5" pageNumber="425" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(paratype: SY1506). 8. Whole genitalia; 9. Ostium bursae; 10. Dentate signa. Scale bars = 0.5 mm (10), 0.1 mm (11, 12).
</paragraph>
</caption>
<paragraph id="B86936954618383F82FD0F88FD10A2A3" blockId="5.[151,1436,1729,2044]" box="[199,729,1729,1756]" pageId="5" pageNumber="425">
<emphasis id="8AA2EA874618383F82FD0F88FE99A2A4" box="[199,336,1729,1755]" italics="true" pageId="5" pageNumber="425">Distribution</emphasis>
.
<collectingCountry id="C0C176054618383F83640F8BFE57A2A3" box="[350,414,1730,1756]" name="Japan" pageId="5" pageNumber="425">Japan</collectingCountry>
(Kyushu) and
<collectingCountry id="C0C176054618383F807F0F88FD1CA2A3" box="[581,725,1729,1756]" name="South Korea" pageId="5" pageNumber="425">South Korea</collectingCountry>
.
</paragraph>
<paragraph id="B86936954618383F82FD0FACFB96A280" blockId="5.[151,1436,1729,2044]" box="[199,1119,1765,1792]" pageId="5" pageNumber="425">
<emphasis id="8AA2EA874618383F82FD0FACFEF6A280" box="[199,319,1765,1791]" italics="true" pageId="5" pageNumber="425">Etymology</emphasis>
. The species epithet is derived from the generic name of the host plant.
</paragraph>
<paragraph id="B86936954618383F82FD0E43FB9DA35C" blockId="5.[151,1436,1729,2044]" box="[199,1108,1801,1828]" pageId="5" pageNumber="425">
<emphasis id="8AA2EA874618383F82FD0E43FE8EA35C" box="[199,327,1801,1827]" italics="true" pageId="5" pageNumber="425">Host plants</emphasis>
.
<taxonomicName id="7FD64D164618383F836C0E40FE77A35C" box="[342,446,1801,1827]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="425" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
<taxonomicName id="7FD64D164618383F83E00E40FC48A35C" authority="var. tomentosa" box="[474,897,1801,1828]" class="Magnoliopsida" family="Fabaceae" genus="Rhynchosia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="425" phylum="Tracheophyta" rank="variety" species="tomentosa" variety="tomentosa">
<emphasis id="8AA2EA874618383F83E00E40FD1FA35C" box="[474,726,1801,1827]" italics="true" pageId="5" pageNumber="425">Rhynchosia tomentosa</emphasis>
var.
<emphasis id="8AA2EA874618383F81340E43FC48A35C" box="[782,897,1802,1827]" italics="true" pageId="5" pageNumber="425">tomentosa</emphasis>
</taxonomicName>
(L.) Hook, &amp; Arn.
</paragraph>
<paragraph id="B86936954618383F82FD0E64FC27A383" blockId="5.[151,1436,1729,2044]" pageId="5" pageNumber="425">
<emphasis id="8AA2EA874618383F82FD0E64FED7A338" box="[199,286,1837,1863]" italics="true" pageId="5" pageNumber="425">Biology</emphasis>
. This species has been collected from the sand-dunes and coastal forests in
<collectingCountry id="C0C176054618383F865B0E67FB6CA337" box="[1121,1189,1838,1864]" name="South Korea" pageId="5" pageNumber="425">Korea</collectingCountry>
and the low mountain areas in
<collectingCountry id="C0C176054618383F82CE0E1BFEFEA313" box="[244,311,1874,1900]" name="Japan" pageId="5" pageNumber="425">Japan</collectingCountry>
. These habitats are matching with where its host plants occur. The young larvae made a linear-blotch mine on the underside of a leaf or a trumpet-shaped mine on the upper side of a leaf (
<figureCitation id="20ED2A104618383F86690E3CFB1DA3F0" box="[1107,1236,1909,1935]" captionStart="FIGURES 1118" captionStartId="6.[151,264,1918,1942]" captionTargetBox="[151,1435,181,1894]" captionTargetId="figure-19@6.[151,1435,181,1894]" captionTargetPageId="6" captionText="FIGURES 1118. Biology of Caloptilia rhynchosiae n. sp. 11. Mine on the upper side of host leaf; 12. Mine on the underside of host leaf; 13. Young larva in the mine; 14, 15. Leaf rolls; 16. Mature larva; 17. Cocoon; 17. Resting adult." figureDoi="http://doi.org/10.5281/zenodo.11101979" httpUri="https://zenodo.org/record/11101979/files/figure.png" pageId="5" pageNumber="425">Figs 1112</figureCitation>
). The later instars of larvae made a cigarette-like leaf roll. The spindle-shaped cocoons (
<figureCitation id="20ED2A104618383F81820ED0FBC2A3CC" box="[952,1035,1945,1971]" captionStart="FIGURES 1118" captionStartId="6.[151,264,1918,1942]" captionTargetBox="[151,1435,181,1894]" captionTargetId="figure-19@6.[151,1435,181,1894]" captionTargetPageId="6" captionText="FIGURES 1118. Biology of Caloptilia rhynchosiae n. sp. 11. Mine on the upper side of host leaf; 12. Mine on the underside of host leaf; 13. Young larva in the mine; 14, 15. Leaf rolls; 16. Mature larva; 17. Cocoon; 17. Resting adult." figureDoi="http://doi.org/10.5281/zenodo.11101979" httpUri="https://zenodo.org/record/11101979/files/figure.png" pageId="5" pageNumber="425">Fig. 17</figureCitation>
) were formed inside/outside of the rolls. The larvae were collected mainly from the places that get a lot of sunshine from June to August and they were emerged in June to September, suggesting two or more generations per year.
</paragraph>
</subSubSection>
</treatment>
</document>