189 lines
29 KiB
XML
189 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.84.68709" ID-GBIF-Dataset="b9e61bc3-4158-4bee-8d57-a65655588f70" ID-Pensoft-Pub="1314-2607-84-145" ID-Pensoft-UUID="AEDB6967BB17574BB6988187C56EE6CA" ID-Zenodo-Dep="5349694" ID-ZooBank="5F83EC637C7947FDBD29E7A2B9328518" ModsDocID="1314-2607-84-145" checkinTime="1629847073322" checkinUser="pensoft" docAuthor="Okayasu, Juriya" docDate="2021" docId="2F91B07729FE5D91ABB3599BFDB59905" docLanguage="en" docName="JourHymenoptRes 84: 145-168" docOrigin="Journal of Hymenoptera Research 84" docPubDate="2021-08-24" docSource="http://dx.doi.org/10.3897/jhr.84.68709" docTitle="Arkaditilla frim Okayasu 2021, sp. nov." docType="treatment" docUuid="E5FFF6B3-52AC-49E9-8B78-B07E69973CFF" docUuidSource="ZooBank" docVersion="4" id="AEDB6967BB17574BB6988187C56EE6CA" lastPageNumber="145" masterDocId="AEDB6967BB17574BB6988187C56EE6CA" masterDocTitle="A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species" masterLastPageNumber="168" masterPageNumber="145" pageNumber="145" updateTime="1678756233000" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Okayasu, Juriya</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6307-2939</mods:nameIdentifier>
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<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566, Japan & Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060 - 8589, Japan</mods:affiliation>
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<mods:nameIdentifier type="email">mutiphiidae@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-08-24</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>84</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>145</mods:start>
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<mods:end>168</mods:end>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.84.68709</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.84.68709</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-84-145</mods:identifier>
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<mods:identifier type="ZooBank">5F83EC637C7947FDBD29E7A2B9328518</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5349694</mods:identifier>
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<treatment ID-GBIF-Taxon="184941297" LSID="urn:lsid:zoobank.org:act:E5FFF6B3-52AC-49E9-8B78-B07E69973CFF" httpUri="http://treatment.plazi.org/id/2F91B07729FE5D91ABB3599BFDB59905" lastPageNumber="145" pageId="0" pageNumber="145">
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<subSubSection pageId="0" pageNumber="145" type="nomenclature">
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName LSID="http://zoobank.org/E5FFF6B3-52AC-49E9-8B78-B07E69973CFF" authority="Okayasu, 2021" authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla frim" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="frim" status="sp. nov.">Arkaditilla frim</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="description">
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<paragraph pageId="0" pageNumber="145">
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145">Figures 5</figureCitation>
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145">, 6</figureCitation>
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<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145">, 15</figureCitation>
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 21</figureCitation>
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 29</figureCitation>
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 30</figureCitation>
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<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">, 41</figureCitation>
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<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">, 42</figureCitation>
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<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">, 55-57</figureCitation>
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<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">, 69</figureCitation>
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<figureCitation captionStart="Figure 73–76" captionStartId="F8" captionText="Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle)." figureDoi="10.3897/jhr.84.68709.figure73-76" httpUri="https://binary.pensoft.net/fig/579487" pageId="0" pageNumber="145">, 75</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="diagnosis">
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<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
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Clypeus dorsally elevated, sloping ventrally. Mandible with weak subapical and subbasal teeth (Fig.
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<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145">15</figureCitation>
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), ventrally deeply excised with large basal tooth (Fig.
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">21</figureCitation>
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). Scape ventral carinae divergent apically (Fig.
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">29</figureCitation>
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), densely punctate between them; F1 apical 1/3, F2, F3 basal half whitish yellow on ventral half, F5-F11 with ventral yellow to reddish brown patch; F1 strongly depressed, F1 height ~0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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pedicel diameter (Fig.
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<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">30</figureCitation>
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). T3 orange (Fig.
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures1-6" httpUri="https://binary.pensoft.net/fig/579480" pageId="0" pageNumber="145">6</figureCitation>
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); cuspis weakly widened on posterior 1/3 with rounded apex (Figs
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<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">55</figureCitation>
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,
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<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">69</figureCitation>
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); digitus posteriorly expanded, forming triangular projection (Fig.
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<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">41</figureCitation>
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); paracuspis slightly tuberculate (Fig.
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<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">69</figureCitation>
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); penis valve weakly expanded ventrally (Figs
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<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">56</figureCitation>
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,
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<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">57</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="description">
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<paragraph pageId="0" pageNumber="145">Description.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
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Body length: 11.2-15.0 mm (holotype: 15.0 mm; mean: 13.1 mm); forewing length: 8.09-10.7 mm (holotype: 10.7 mm; mean: 9.37 mm).
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" pageId="0" pageNumber="145">
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<emphasis italics="true" pageId="0" pageNumber="145">Color and setae</emphasis>
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.
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</emphasis>
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Frons, vertex, gena, clypeus, pedicel, flagellum mostly, prementum, stipes, mesosoma, T4-T7, S4-S8 black; mandible brown to black, with apex always black and medial 1/5-1/2 yellowish brown; labrum, protibial spur yellow; scape, tegula brownish black; F1 apical 1/3, F2, F3 basal half whitish yellow on ventral half; F5-F11 yellow to reddish brown ventrally; trochanters brown to dark brown; coxae, femora, tibiae, tarsi dark brown to brownish black; meso- and metatibial spurs pale yellow; T1-T3, S1-S3 orange; wings pale brown on proximal 1/3, brown on distal 2/3, with veins dark brown. Upper frons, vertex, trochanters, T1, S1 with sparse long erect pale golden setae; lower frons, dorsal propodeal face, dorsal half of lateral propodeal face with dense short appressed and sparse long erect pale golden setae; eye dorsal margin with long erect black setae; gena, pronotal dorsum, metascutellum with sparse short recumbent and sparse long erect pale golden setae; clypeus with dense long recumbent pale golden setae; mandible, tibiae, tarsi, T2-T5 with sparse long recumbent pale golden setae; scape, femora, S2 with sparse long recumbent and sparse long erect pale golden setae; pedicel, F1, tegula anterior portion with sparse short recumbent pale golden setae; F2-F11 with sparse erect pale golden microsetae; pronotal collar anteriorly, T2 disc, T3-T5 medially, S3-S6 with sparse short erect pale golden setae; pronotal collar posteriorly glabrous; pronotal lateral face meso- and metacoxae with sparse long appressed pale golden setae; propleuron with sparse long appressed and sparse long erect pale golden setae; mesoscutum with sparse short recumbent black and sparse short erect pale golden to black setae; tegula inner margin with sparse short recumbent pale golden and sparse short erect pale golden to black setae; mesoscutellum with sparse short recumbent pale golden, sparse long erect black, sparse long erect pale golden setae; anterior and posterior parts of mesopleuron, metapleuron, ventral half of lateral propodeal face with sparse short appressed pale golden setae; axilla, metanotal trough, medial and ventral parts of mesopleuron, procoxa with dense long appressed and sparse long erect pale golden setae; T6-T7, S7-S8 with sparse long recumbent and sparse long erect black setae; T2 with lateral golden felt line; T1-T5, S2-S5 with posterior fringe of sparse pale golden setae; S6-S8 with posterior fringe of sparse black setae.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" pageId="0" pageNumber="145">
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<emphasis italics="true" pageId="0" pageNumber="145">Structure</emphasis>
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.
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</emphasis>
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Head 1.82-2.00
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<normalizedToken originalValue="×">x</normalizedToken>
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as wide as long (holotype: 2.00); lateral margins strongly convergent behind eye; vertex rounded posteriorly; POD/OOD = 0.56-0.58 (holotype: 0.56); OD:MOD:LOD = 1.00:1.08-1.12:0.94-1.00 (holotype: 1.00:1.12:0.94); ocellar region slightly convex, ocelli without posterior carina; antennal scrobe with weak inverted V-shaped dorsal carina not reaching inner eye margin; genal carina absent; postgenal bridge flattened; hypostomal carina weakly developed; clypeus dorsally elevated, sloping ventrally, with subventral transverse groove laterally not defined by carina; anterior clypeal margin projecting medially; mandible robust, narrowed to apex with inner margin expanded forming small subbasal tooth; mandible apex bidentate; mandible ventral margin excised into large basal tooth; mandible dorsal carina sharp; prementum flattened; scape curved medially, with two longitudinal ventral carinae divergent apically; F1 depressed, F1 height 0.67-0.80
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
pedicel diameter (holotype: 0.67); relative length of pedicel, F1 and F2 = 1.00:2.85-3.06:1.92-2.29 (holotype: 1.00:3.06:2.29); F2-F11 almost equal in width; F11 as long as F2, conical.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
Mesosoma widest at mesonotum including tegulae; head width:humeral width:pronotal width:mesonotal width = 88:62:91:100; humeral angle rounded, with sharp humeral carina on ventral half; mesoscutum strongly convex; mesoscutal width 1.12
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
its length; tegula short rounded, posteriorly reaching mesoscuto-scutellar articulation, 1.44-1.53
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as wide (holotype: 1.53); notaulus evenly wide, reaching anterior 1/2 of mesoscutum, subparallel; parapsidal line length 0.25
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
mesoscutal length; parascutal carina forming rounded, weakly elevated process on postero-lateral angle of mesoscutum; mesoscutellum convex without medial carina, tubercle or impunctate line; mesopleuron divided into dorsal and ventral convexities by distinct transverse mesopleural groove, ventrally without transverse precoxal ridge, longitudinal impunctate line, or precoxal swelling; dorsal propodeal face without distinct dorsum and sublateral carina, with anterior inverted triangular cell and semicircular sublateral cell; lateral margins of propodeum gradually convergent posteriorly; dorsal and lateral propodeal faces not separated by carina.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Wings well developed; distance between origin of RS on vein SC and base of stigmatic cell equal to stigmatic cell length and first abscissa of RS length; cell 2RS present.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Meso- and metacoxae evenly convex; metacoxa with weak complete inner longitudinal carina; protibial spur slender; tibiae without outer spines.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
T1 without distinct dorsum; T1 spiracle weakly tuberculate; T2 evenly convex, 1.28
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long; S1 with straight medial longitudinal carina with its ventral margin slightly elevated posteriorly; S2 swollen anteriorly; T7 medially with longitudinal impunctate convex line, posteriorly not forming tubercle; S6-S8 without lateral carina or tubercle; S7 entirely exposed, 0.75
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as S8; S8 mostly sclerotized with posterior membranous area evenly convex posteriorly. Paramere down-curved posteriorly, with weak inner protuberance on posterior 1/2 of dorso-inner margin; inner margin of paramere dorsally with short erect golden setae; ventral lobe posteriorly projecting, pointed; parapenial lobe large, with posterior margin slightly concave; cuspis long slender, weakly widened on posterior 1/3 with rounded apex, slightly surpassing apex of penis valve, with dense long erect setae ventrally and posteriorly; digitus straight, posteriorly expanded, forming triangular projection, with dense erect golden setae posteriorly; paracuspis slightly tuberculate with few erect golden setae; penis valves symmetrical slender, weakly expanded ventrally, with posterior ventral hook, without setae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Frons, vertex, gena, pronotal dorsum, mesoscutum, medial part of mesopleuron with large dense punctures; postgenal bridge, clypeus lateral portion, mandible outer face, scape between ventral carinae, flagellum, pronotal lateral face, propleuron, axilla, metanotal trough, anterior and posterior parts of mesopleuron, metapleuron, ventral half of lateral propodeal face, procoxa, tarsi with minute dense punctures; clypeus medial convexity with large sparse punctures; mandible mostly, pronotal collar posteriorly smooth; scape mostly, trochanters, femora, tibiae, T1 medially, T2 disc, T3-T6, S1-S2 with small sparse punctures; pedicel, pronotal collar anteriorly, meso- and metacoxa with minute sparse punctures; mesoscutellum with large confluent punctures; metascutellum with small confluent punctures; dorsal propodeal face, dorsal half of lateral propodeal face reticulate; T1-T2 laterally, T7, S3-S8 with small dense punctures; tegula mostly microreticulate with few small punctures anteriorly, postero-inner portion with minute sparse punctures.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="type material">
|
||
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
2♂.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
|
||
♂:
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Malaysia</emphasis>
|
||
: Negeri Sembilan: Pasoh Forest Reserve, 3-10.XII.1993, K. Maeto leg., coarse Malaise trap, plot 6 [FRIM].
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Paratype</emphasis>
|
||
: Malaysia
|
||
</emphasis>
|
||
: Negeri Sembilan: Pasoh Forest Reserve, 25.III-2.IV.1993, K. Konishi & K. Maeto leg., fine Malaise trap, plot 2 [1♂ NARO].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="distribution">
|
||
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Malaysia: Negeri Sembilan.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="etymology">
|
||
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">This new species is named after FRIM, Forest Research Institute Malaysia, which was the Malaysian counterpart of the Japan-Malaysia joint research project in 1993 funded by the Ministry of the Environment of Japan. Treat as a noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="remarks">
|
||
<paragraph pageId="0" pageNumber="145">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
This species occurs sympatrically with
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
|
||
</taxonomicName>
|
||
, but this new species differs from it by having the mandible with weakly developed subbasal and subapical teeth (subbasal and subapical teeth large in
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
|
||
</taxonomicName>
|
||
), the digitus posteriorly expanded (digitus slender in
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
|
||
</taxonomicName>
|
||
), and the paracuspis tuberculate (paracuspis absent in
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |