706 lines
106 KiB
XML
706 lines
106 KiB
XML
<document id="056E535E88B4754EBBB86BE03A12C969" ID-CLB-Dataset="56039" ID-DOI="10.5281/zenodo.5306071" ID-GBIF-Dataset="6f1e77a4-b2ef-4a36-bbb5-71920341e9ad" ID-ISSN="0374-1036" ID-Zenodo-Dep="5306071" ID-ZooBank="7EFF344A-B2FF-4A14-80AC-3B047298201D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630112253027" checkinUser="marcus" docAuthor="Hillert, Oliver, Arnone, Marcello, Král, David & Massa, Bruno" docDate="2016" docId="2F75BE40FFA6BE1CFEF9FAB2FBBEF16E" docLanguage="en" docName="ActaEntMusNatPra.2016.56.1.211-254.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 56 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Bolbelasmus Boucomont 1911" docType="key" docVersion="5" lastPageNumber="244" masterDocId="D34CC638FFB8BE3DFFECFF96FFFEF51D" masterDocTitle="The genusBolbelasmus in the western and southern regions of the Mediterranean Basin (Coleoptera: Geotrupidae: Bolboceratinae)" masterLastPageNumber="254" masterPageNumber="211" pageNumber="241" updateTime="1699159846572" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="15129E8580E4D3C94C190C987E29DCAA">The genusBolbelasmus in the western and southern regions of the Mediterranean Basin (Coleoptera: Geotrupidae: Bolboceratinae)</mods:title>
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<mods:name id="DEA6132DF9A29A9BC667ECCDE2D3BE83" type="personal">
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<mods:namePart id="D55B1BD4B65F158802385A827B6D6D0C">Hillert, Oliver</mods:namePart>
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<mods:affiliation id="90C3A19A8D82383210D293FFE96B810B">Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany; e-mail: ohillert @ yahoo. de</mods:affiliation>
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<mods:nameIdentifier id="34D659E02950DB2CC5A905EE898B365C" type="email">ohillert@yahoo.de</mods:nameIdentifier>
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<mods:namePart id="6964F2D3CB292D093EC0E4B9F552C6C6">Arnone, Marcello</mods:namePart>
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<mods:affiliation id="B183BFAA9D396BA8FE282189F15FABBE">Via San Michele 8, I- 90010 Altavilla Milicia (Palermo), Italy; e-mail: marcel. arnone @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="F64712A36F0496A09181AB37848D2D15" type="email">marcel.arnone@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0650164FA000168CCF6B3F2DFC89E042">Král, David</mods:namePart>
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<mods:affiliation id="7956EC83E9933FF92E5E03CE99B2CA24">Charles University in Prague, Faculty of Science, Department of Zoology, Viničná 7, CZ- 128 43 Praha 2, Czech Republic; e-mail: kral @ natur. cuni. cz</mods:affiliation>
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<mods:nameIdentifier id="1CD70495B22DAED7BAF29D9EB06D7B63" type="email">kral@natur.cuni.cz</mods:nameIdentifier>
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<mods:namePart id="D30004AD89C1A3A64D2CC9C209306424">Massa, Bruno</mods:namePart>
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<mods:affiliation id="37EA60FF737A789A7680113046942D7D">University of Palermo, Department of Agricultural and Forest Sciences, Viale Scienze 13, I- 90128 Palermo, Italy; e-mail: bruno. massa @ unipa. it</mods:affiliation>
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<mods:nameIdentifier id="E8EE8D566851379496D1BEA3504EBBB7" type="email">bruno.massa@unipa.it</mods:nameIdentifier>
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<mods:title id="1173E86352BBB4CDF0696BE23B3A1A99">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="4F9200897F93FD2D07AED717C9281303">2016</mods:date>
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<mods:number id="29D5F304CE3A7C82F9DB67040F692F64">2016-07-15</mods:number>
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<treatment id="2F75BE40FFA6BE1CFEF9FAB2FBBEF16E" ID-DOI="http://doi.org/10.5281/zenodo.6454369" ID-GBIF-Taxon="194587790" ID-Zenodo-Dep="6454369" LSID="urn:lsid:plazi:treatment:2F75BE40FFA6BE1CFEF9FAB2FBBEF16E" httpUri="http://treatment.plazi.org/id/2F75BE40FFA6BE1CFEF9FAB2FBBEF16E" lastPageId="33" lastPageNumber="244" pageId="30" pageNumber="241">
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<subSubSection id="EFC65CDDFFA6BE23FEF9FAB2FC17F023" box="[277,1001,1316,1342]" pageId="30" pageNumber="241" type="nomenclature">
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<paragraph id="A7630F56FFA6BE23FEF9FAB2FC17F023" blockId="30.[277,1001,1316,1342]" box="[277,1001,1316,1342]" pageId="30" pageNumber="241">
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<heading id="FC2BB83AFFA6BE23FEF9FAB2FC17F023" bold="true" box="[277,1001,1316,1342]" centered="true" fontSize="11" level="1" pageId="30" pageNumber="241" reason="2">
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<emphasis id="95A8D344FFA6BE23FEF9FAB2FC17F023" bold="true" box="[277,1001,1316,1342]" pageId="30" pageNumber="241">
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Identification key for
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<taxonomicName id="60DC74D5FFA6BE23FDC3FAB2FCF0F023" authorityName="Boucomont" authorityYear="1911" box="[559,782,1316,1342]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="241" phylum="Arthropoda" rank="genus">well and medium</taxonomicName>
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developed males
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="EFC65CDDFFA6BE22FF7BFACAFBBEF0EE" lastPageId="31" lastPageNumber="242" pageId="30" pageNumber="241" type="key">
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<key id="D34D5867FFA6BE22FF7BFACAFBBEF0EE" lastPageId="31" lastPageNumber="242" pageId="30" pageNumber="241">
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<paragraph id="A7630F56FFA6BE23FF7BFACAFE6BF08E" blockId="30.[151,1126,1372,1555]" pageId="30" pageNumber="241">
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<keyLead id="1C2DAC63FFA6BE23FF7BFACAFE6BF08E" pageId="30" pageNumber="241">
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<keyStep id="1C2817F3FFA6BE23FF7BFACAFE6BF08E" pageId="30" pageNumber="241">1 (10) Scutellar shield impunctate or indistinctly shallowly punctate, frontal horn simple or furcate apically.</keyStep>
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</keyLead>
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</paragraph>
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<paragraph id="A7630F56FFA6BE23FF7BFA0AFD2AF0CE" blockId="30.[151,1126,1372,1555]" pageId="30" pageNumber="241">
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<keyLead id="1C2DAC63FFA6BE23FF7BFA0AFD2AF0CE" pageId="30" pageNumber="241">
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<keyStep id="1C2817F3FFA6BE23FF7BFA0AFD2AF0CE" pageId="30" pageNumber="241">2 (5) Frontal horn simple; genae evenly rounded; oblique keel above eyes over whole eyes length; head strongly constricted behind eyes.</keyStep>
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</keyLead>
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||
</paragraph>
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<paragraph id="A7630F56FFA6BE22FF7BFA4AFBBFF42E" blockId="30.[151,1126,1372,1555]" lastBlockId="31.[113,1089,220,1523]" lastPageId="31" lastPageNumber="242" pageId="30" pageNumber="241">
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<keyLead id="1C2DAC63FFA6BE22FF7BFA4AFBBFF42E" lastPageId="31" lastPageNumber="242" pageId="30" pageNumber="241">
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<keyStep id="1C2817F3FFA6BE22FF7BFA4AFBBFF42E" lastPageId="31" lastPageNumber="242" pageId="30" pageNumber="241">
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3 (4) Frontal horn directed forwards and upwards in lateral view; clypeus with short transversal carina; scutellar shield triangular, approximately as long as wide; body surface brownish, maximal body length
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<quantity id="6024A2B3FFA7BE22FD95FF4AFD37F5EE" box="[633,713,220,243]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="31" pageNumber="242" unit="mm" value="14.0">14 mm</quantity>
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. Southern parts of Central Europe, Alsace,
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<collectingCountry id="DFCB4FC6FFA7BE22FEF6FF6AFEB2F40E" box="[282,332,252,275]" name="Italy" pageId="31" pageNumber="242">Italy</collectingCountry>
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, Balkan Peninsula,
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<collectingCountry id="DFCB4FC6FFA7BE22FDF2FF6AFD95F40E" box="[542,619,252,275]" name="Poland" pageId="31" pageNumber="242">Poland</collectingCountry>
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,
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<collectingCountry id="DFCB4FC6FFA7BE22FD9AFF6AFD43F40E" box="[630,701,252,275]" name="Russia" pageId="31" pageNumber="242">Russia</collectingCountry>
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and
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||
<collectingCountry id="DFCB4FC6FFA7BE22FD1CFF6AFCB7F40E" box="[752,841,252,275]" name="Ukraine" pageId="31" pageNumber="242">Ukraine</collectingCountry>
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. ................................... ....................................................................
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||
<taxonomicName id="60DC74D5FFA7BE22FD79FE8AFBBFF42E" authority="(Schrank von Paula, 1789)" baseAuthorityName="Schrank von Paula" baseAuthorityYear="1789" box="[661,1089,283,307]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="242" phylum="Arthropoda" rank="species" species="unicornis">
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||
<emphasis id="95A8D344FFA7BE22FD79FE8AFCE3F42E" bold="true" box="[661,797,283,307]" italics="true" pageId="31" pageNumber="242">B. unicornis</emphasis>
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||
(
|
||
<bibRefCitation id="C34D72A7FFA7BE22FCC7FE8AFBC7F42E" author="Schrank von Paula" box="[811,1081,284,307]" firstAuthor="Schrank von Paula" pageId="31" pageNumber="242" pagination="60 - 90" refId="ref28580" refString="SCHRANK VON PAULA F. 1789: Entomologische Beobachtungen. Der Naturforscher 24: 60 - 90." type="journal article" year="1789">Schrank von Paula, 1789</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
</keyStep>
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||
</keyLead>
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</paragraph>
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||
<paragraph id="A7630F56FFA7BE22FF9DFEAAFBBEF4AE" blockId="31.[113,1089,220,1523]" pageId="31" pageNumber="242">
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<keyLead id="1C2DAC63FFA7BE22FF9DFEAAFBBEF4AE" pageId="31" pageNumber="242">
|
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<keyStep id="1C2817F3FFA7BE22FF9DFEAAFBBEF4AE" pageId="31" pageNumber="242">
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||
4 (3) Frontal horn bending not forwards (
|
||
<figureCitation id="3FE713D3FFA7BE22FDA5FEAAFD39F44E" box="[585,711,316,339]" captionStart="Figs 14–20" captionStartId="14.[151,188,1475,1494]" captionTargetBox="[165,1111,442,1454]" captionTargetId="figure-10@14.[156,1123,435,1471]" captionTargetPageId="14" captionText="Figs 14–20. Male habitus in lateral aspect. 14 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length:14.0 mm, NMPC); 15 – B. brancoi Hillert & Král sp.nov.(holotype); 16 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône,Barbentane, 13.8 mm, DJCP); 17 – B. howdeni Hillert & Král sp. nov.(holotype); 18 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 19 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, body length: 11.8 mm, DKCP); 20 – B. vaulogeri (Italy: Sicily, Menfi (Agrigento), Cinquanta, body length: 11.6 mm, MAPC)." figureDoi="http://doi.org/10.5281/zenodo.5306081" httpUri="https://zenodo.org/record/5306081/files/figure.png" pageId="31" pageNumber="242">Figs 19–20</figureCitation>
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); transversal clypeal carina absent; scutellar shield triangular longer than wide; body surface blackish, maximal body length
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||
<quantity id="6024A2B3FFA7BE22FEE2FEEAFEA7F48E" box="[270,345,380,403]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="31" pageNumber="242" unit="mm" value="13.0">13 mm</quantity>
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(
|
||
<figureCitation id="3FE713D3FFA7BE22FE84FEEAFE36F48E" box="[360,456,380,403]" captionStart="Figs 5–7" captionStartId="8.[151,187,1490,1509]" captionTargetBox="[217,1057,222,1460]" captionTargetId="figure-10@8.[211,1067,219,1489]" captionTargetPageId="8" captionText="Figs 5–7.Male habitus in dorsal aspect.5 – Bolbelasmus nikolajevi Hillert,Arnone, Král & Massa sp. nov. (holotype); 6 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, 11.8 mm, DKCP); 7 – B. vaulogeri (Italy: Sicily, Menfi (Agrigento), Cinquanta, body length: 11.6 mm, MAPC)." figureDoi="http://doi.org/10.5281/zenodo.5306075" httpUri="https://zenodo.org/record/5306075/files/figure.png" pageId="31" pageNumber="242">Figs 6–7</figureCitation>
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||
,
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||
<figureCitation id="3FE713D3FFA7BE22FE38FEEAFE0CF48E" box="[468,498,380,403]" captionStart="Figs 33–44" captionStartId="17.[113,150,1415,1434]" captionTargetBox="[183,1019,223,1400]" captionTargetId="figure-14@17.[183,1019,219,1411]" captionTargetPageId="17" captionText="Figs 33–44. Head in dorsal aspect, 33–38 –♂♂; 39–44 – ♀♀. 33 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, NMPC); 34 – B. brancoi Hillert & Král sp. nov. (holotype); 35 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 36 – B. howdeni Hillert & Král sp. nov. (holotype); 37 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 38 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP); 39 – B. bocchus (Morocco: Gaada de Debdou plateau., E of Rchida, DKCP); 40 – B. brancoi (allotype); 41 – B. gallicus (Spain: Andalusia, Aljaraque vill. env., JSCP); 42 – B. howdeni (allotype); 43 – B. nikolajevi (allotype); 44 – B. vaulogeri (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306091" httpUri="https://zenodo.org/record/5306091/files/figure.png" pageId="31" pageNumber="242">38</figureCitation>
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); aedeagus as in
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||
<figureCitation id="3FE713D3FFA7BE22FD44FEEAFD09F48E" box="[680,759,380,403]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="31" pageNumber="242">Fig. 65</figureCitation>
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||
;
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FCEDFEEAFCAAF48E" box="[769,852,380,403]" name="Algeria" pageId="31" pageNumber="242">Algeria</collectingCountry>
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||
,
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FCB3FEEAFC6EF48E" box="[863,912,380,403]" name="Italy" pageId="31" pageNumber="242">Italy</collectingCountry>
|
||
(
|
||
<collectingRegion id="6518C1B4FFA7BE22FC72FEEAFC21F48E" box="[926,991,380,403]" country="Italy" name="Sicilia" pageId="31" pageNumber="242">Sicily</collectingRegion>
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||
) and
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||
<collectingCountry id="DFCB4FC6FFA7BE22FBF7FEEAFF08F4AE" name="Tunisia" pageId="31" pageNumber="242">Tunisia</collectingCountry>
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||
(Fig. 59). ...............................................
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<taxonomicName id="60DC74D5FFA7BE22FD44FE0AFBBEF4AE" authority="(Abeille de Perrin, 1898)" baseAuthorityName="Abeille de Perrin" baseAuthorityYear="1898" box="[680,1088,411,435]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="242" phylum="Arthropoda" rank="species" species="vaulogeri">
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||
<emphasis id="95A8D344FFA7BE22FD44FE0AFCD1F4AE" bold="true" box="[680,815,411,435]" italics="true" pageId="31" pageNumber="242">B. vaulogeri</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FFA7BE22FCD1FE0AFBC7F4AE" author="Abeille de Perrin" box="[829,1081,412,435]" firstAuthor="Abeille de Perrin" pageId="31" pageNumber="242" pagination="254 - 258" refId="ref25891" refString="ABEILLE DE PERRIN E. 1898: Descriptions de Coleopteres du Nord de l'Afrique. Bulletin de la Societe Entomologique de France 1898: 254 - 258." type="journal article" year="1898">Abeille de Perrin, 1898</bibRefCitation>
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||
)
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</taxonomicName>
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||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FFA7BE22FF9DFE2AFEB4F70E" blockId="31.[113,1089,220,1523]" pageId="31" pageNumber="242">
|
||
<keyLead id="1C2DAC63FFA7BE22FF9DFE2AFEB4F70E" pageId="31" pageNumber="242">
|
||
<keyStep id="1C2817F3FFA7BE22FF9DFE2AFEB4F70E" pageId="31" pageNumber="242">5 (2) Frontal horn furcate apically; genae more or less angulate anterolaterally; oblique keel above eyes reaching only to anterior part of eyes; head not strongly constricted behind eyes.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FFA7BE22FF9DFD8AFBBEF7CE" blockId="31.[113,1089,220,1523]" pageId="31" pageNumber="242">
|
||
<keyLead id="1C2DAC63FFA7BE22FF9DFD8AFBBEF7CE" pageId="31" pageNumber="242">
|
||
<keyStep id="1C2817F3FFA7BE22FF9DFD8AFBBEF7CE" pageId="31" pageNumber="242">
|
||
6 (7) Frontal horn short and not longer than broad, horn apex approximately as wide as horn base (
|
||
<figureCitation id="3FE713D3FFA7BE22FED7FDAAFE7FF74E" box="[315,385,572,595]" captionStart="Figs 1–4" captionStartId="7.[113,150,1490,1509]" captionTargetBox="[122,1077,229,1442]" captionTargetId="figure-14@7.[112,1092,216,1462]" captionTargetPageId="7" captionText="Figs 1–4. Male habitus in dorsal aspect. 1 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length: 14.0 mm, NMPC); 2 – B. brancoi Hillert & Král sp. nov. (holotype); 3 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, body length 13.7 mm, DJCP); 4 – B. howdeni Hillert & Král sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5306073" httpUri="https://zenodo.org/record/5306073/files/figure.png" pageId="31" pageNumber="242">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FE62FDAAFE55F74E" box="[398,427,572,595]" captionStart="Figs 14–20" captionStartId="14.[151,188,1475,1494]" captionTargetBox="[165,1111,442,1454]" captionTargetId="figure-10@14.[156,1123,435,1471]" captionTargetPageId="14" captionText="Figs 14–20. Male habitus in lateral aspect. 14 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length:14.0 mm, NMPC); 15 – B. brancoi Hillert & Král sp.nov.(holotype); 16 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône,Barbentane, 13.8 mm, DJCP); 17 – B. howdeni Hillert & Král sp. nov.(holotype); 18 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 19 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, body length: 11.8 mm, DKCP); 20 – B. vaulogeri (Italy: Sicily, Menfi (Agrigento), Cinquanta, body length: 11.6 mm, MAPC)." figureDoi="http://doi.org/10.5281/zenodo.5306081" httpUri="https://zenodo.org/record/5306081/files/figure.png" pageId="31" pageNumber="242">15</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FE5AFDAAFE2AF74E" box="[438,468,572,595]" captionStart="Figs 21–26" captionStartId="15.[113,150,1491,1510]" captionTargetBox="[157,1030,223,1437]" captionTargetId="figure-14@15.[152,1050,219,1464]" captionTargetPageId="15" captionText="Figs 21–26. Pronotum of male in dorsal aspect.21 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 22 – B. brancoi Hillert & Král sp. nov. (holotype); 23 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 24 – B. howdeni Hillert & Král sp. nov. (holotype); 25 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 26 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306085" httpUri="https://zenodo.org/record/5306085/files/figure.png" pageId="31" pageNumber="242">22</figureCitation>
|
||
); pronotal punctation denser with exception of basal part, punctures separated approximately by distance equal to their diameters; pronotal base with distinct visible but fine, regularly pinned points; elytral striae well impressed, visible as complete lines; elytral intervals moderately convex; aedeagus as in
|
||
<figureCitation id="3FE713D3FFA7BE22FF0CFD2AFED0F7CE" box="[224,302,700,723]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="31" pageNumber="242">Fig. 61</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FED6FD2AFE89F7CE" box="[314,375,700,723]" name="Spain" pageId="31" pageNumber="242">Spain</collectingCountry>
|
||
(Fig. 59). .................................
|
||
<taxonomicName id="60DC74D5FFA7BE22FD3FFD2AFC1AF7CE" authority="Hillert & Kral" authorityName="Hillert & Kral" authorityYear="2016" box="[723,996,699,723]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="242" phylum="Arthropoda" rank="species" species="brancoi" status="sp. nov.">
|
||
<emphasis id="95A8D344FFA7BE22FD3FFD2AFCB8F7CE" bold="true" box="[723,838,699,723]" italics="true" pageId="31" pageNumber="242">B. brancoi</emphasis>
|
||
Hillert & Král
|
||
</taxonomicName>
|
||
<emphasis id="95A8D344FFA7BE22FC07FD2AFBBEF7CE" bold="true" box="[1003,1088,700,723]" pageId="31" pageNumber="242">
|
||
<taxonomicNameLabel id="8E9B6E3FFFA7BE22FC07FD2AFBBEF7CE" box="[1003,1088,700,723]" pageId="31" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FFA7BE22FF9DFD4AFC9EF66E" blockId="31.[113,1089,220,1523]" pageId="31" pageNumber="242">
|
||
<keyLead id="1C2DAC63FFA7BE22FF9DFD4AFC9EF66E" pageId="31" pageNumber="242">
|
||
<keyStep id="1C2817F3FFA7BE22FF9DFD4AFC9EF66E" pageId="31" pageNumber="242">
|
||
7 (6) Frontal horn long, longer than broad, horn becoming gradually thinner from base to apex (
|
||
<figureCitation id="3FE713D3FFA7BE22FEE8FD6AFEB8F60E" box="[260,326,764,787]" captionStart="Figs 1–4" captionStartId="7.[113,150,1490,1509]" captionTargetBox="[122,1077,229,1442]" captionTargetId="figure-14@7.[112,1092,216,1462]" captionTargetPageId="7" captionText="Figs 1–4. Male habitus in dorsal aspect. 1 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length: 14.0 mm, NMPC); 2 – B. brancoi Hillert & Král sp. nov. (holotype); 3 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, body length 13.7 mm, DJCP); 4 – B. howdeni Hillert & Král sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5306073" httpUri="https://zenodo.org/record/5306073/files/figure.png" pageId="31" pageNumber="242">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FEB8FD6AFE9DF60E" box="[340,355,764,787]" captionStart="Figs 5–7" captionStartId="8.[151,187,1490,1509]" captionTargetBox="[217,1057,222,1460]" captionTargetId="figure-10@8.[211,1067,219,1489]" captionTargetPageId="8" captionText="Figs 5–7.Male habitus in dorsal aspect.5 – Bolbelasmus nikolajevi Hillert,Arnone, Král & Massa sp. nov. (holotype); 6 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, 11.8 mm, DKCP); 7 – B. vaulogeri (Italy: Sicily, Menfi (Agrigento), Cinquanta, body length: 11.6 mm, MAPC)." figureDoi="http://doi.org/10.5281/zenodo.5306075" httpUri="https://zenodo.org/record/5306075/files/figure.png" pageId="31" pageNumber="242">5</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FE82FD6AFE4CF60E" box="[366,434,764,787]" captionStart="Figs 14–20" captionStartId="14.[151,188,1475,1494]" captionTargetBox="[165,1111,442,1454]" captionTargetId="figure-10@14.[156,1123,435,1471]" captionTargetPageId="14" captionText="Figs 14–20. Male habitus in lateral aspect. 14 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length:14.0 mm, NMPC); 15 – B. brancoi Hillert & Král sp.nov.(holotype); 16 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône,Barbentane, 13.8 mm, DJCP); 17 – B. howdeni Hillert & Král sp. nov.(holotype); 18 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 19 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, body length: 11.8 mm, DKCP); 20 – B. vaulogeri (Italy: Sicily, Menfi (Agrigento), Cinquanta, body length: 11.6 mm, MAPC)." figureDoi="http://doi.org/10.5281/zenodo.5306081" httpUri="https://zenodo.org/record/5306081/files/figure.png" pageId="31" pageNumber="242">14, 18</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FE52FD6AFDFFF60E" box="[446,513,764,787]" captionStart="Figs 21–26" captionStartId="15.[113,150,1491,1510]" captionTargetBox="[157,1030,223,1437]" captionTargetId="figure-14@15.[152,1050,219,1464]" captionTargetPageId="15" captionText="Figs 21–26. Pronotum of male in dorsal aspect.21 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 22 – B. brancoi Hillert & Král sp. nov. (holotype); 23 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 24 – B. howdeni Hillert & Král sp. nov. (holotype); 25 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 26 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306085" httpUri="https://zenodo.org/record/5306085/files/figure.png" pageId="31" pageNumber="242">21, 25</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FDE1FD6AFDAEF60E" box="[525,592,764,787]" captionStart="Figs 27–32" captionStartId="16.[151,188,1490,1509]" captionTargetBox="[151,1097,253,1372]" captionTargetId="figure-10@16.[151,1126,248,1394]" captionTargetPageId="16" captionText="Figs 27–32.Pronotum of male in frontal aspect. 27 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 28 – B. brancoi Hillert & Král sp. nov. (holotype); 29 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 30 – B. howdeni Hillert & Král sp. nov. (holotype); 31 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 32 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306087" httpUri="https://zenodo.org/record/5306087/files/figure.png" pageId="31" pageNumber="242">27, 31</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FDB0FD6AFD5EF60E" box="[604,672,764,787]" captionStart="Figs 33–44" captionStartId="17.[113,150,1415,1434]" captionTargetBox="[183,1019,223,1400]" captionTargetId="figure-14@17.[183,1019,219,1411]" captionTargetPageId="17" captionText="Figs 33–44. Head in dorsal aspect, 33–38 –♂♂; 39–44 – ♀♀. 33 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, NMPC); 34 – B. brancoi Hillert & Král sp. nov. (holotype); 35 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 36 – B. howdeni Hillert & Král sp. nov. (holotype); 37 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 38 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP); 39 – B. bocchus (Morocco: Gaada de Debdou plateau., E of Rchida, DKCP); 40 – B. brancoi (allotype); 41 – B. gallicus (Spain: Andalusia, Aljaraque vill. env., JSCP); 42 – B. howdeni (allotype); 43 – B. nikolajevi (allotype); 44 – B. vaulogeri (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306091" httpUri="https://zenodo.org/record/5306091/files/figure.png" pageId="31" pageNumber="242">33, 37</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FD47FD6AFD36F60E" box="[683,712,764,787]" captionStart="Figs 57–58" captionStartId="26.[151,188,1519,1538]" captionTargetBox="[180,1097,219,1493]" captionTargetId="figure-10@26.[180,1098,219,1493]" captionTargetPageId="26" captionText="Figs 57–58. Living specimens. 57 – Bolbelasmus nikolajevi Hillert, Arnone, Král & Massa sp. nov. (paratype, ♂, Tunisia, Cap Bon peninsula, 6 km W of El Hawariah); 58 – B. vaulogeri (Abeille de Perrin, 1898) (♂, Tunisia, Cap Bon peninsula, 2 km NW of Douela). Photos by Petr Šípek." figureDoi="http://doi.org/10.5281/zenodo.5306099" httpUri="https://zenodo.org/record/5306099/files/figure.png" pageId="31" pageNumber="242">57</figureCitation>
|
||
); pronotal punctation sparser, punctures separated by approximately three times their diameters; pronotal base with very fine, weakly visible and well separated pinned points; elytral striae not impressed, visible as row of punctures; elytral intervals flat; nothern Africa.
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FFA7BE22FF9DFCEAFBBEF12E" blockId="31.[113,1089,220,1523]" pageId="31" pageNumber="242">
|
||
<keyLead id="1C2DAC63FFA7BE22FF9DFCEAFBBEF12E" pageId="31" pageNumber="242">
|
||
<keyStep id="1C2817F3FFA7BE22FF9DFCEAFBBEF12E" pageId="31" pageNumber="242">
|
||
8 (9) Base of frontal horn situated at level of anterior part of eyes; horn sharply keeled from base to half of its length in dorsal aspect (
|
||
<figureCitation id="3FE713D3FFA7BE22FD50FC0AFD01F6AE" box="[700,767,924,947]" captionStart="Figs 1–4" captionStartId="7.[113,150,1490,1509]" captionTargetBox="[122,1077,229,1442]" captionTargetId="figure-14@7.[112,1092,216,1462]" captionTargetPageId="7" captionText="Figs 1–4. Male habitus in dorsal aspect. 1 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length: 14.0 mm, NMPC); 2 – B. brancoi Hillert & Král sp. nov. (holotype); 3 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, body length 13.7 mm, DJCP); 4 – B. howdeni Hillert & Král sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5306073" httpUri="https://zenodo.org/record/5306073/files/figure.png" pageId="31" pageNumber="242">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FCE1FC0AFCD4F6AE" box="[781,810,924,947]" captionStart="Figs 14–20" captionStartId="14.[151,188,1475,1494]" captionTargetBox="[165,1111,442,1454]" captionTargetId="figure-10@14.[156,1123,435,1471]" captionTargetPageId="14" captionText="Figs 14–20. Male habitus in lateral aspect. 14 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length:14.0 mm, NMPC); 15 – B. brancoi Hillert & Král sp.nov.(holotype); 16 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône,Barbentane, 13.8 mm, DJCP); 17 – B. howdeni Hillert & Král sp. nov.(holotype); 18 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 19 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, body length: 11.8 mm, DKCP); 20 – B. vaulogeri (Italy: Sicily, Menfi (Agrigento), Cinquanta, body length: 11.6 mm, MAPC)." figureDoi="http://doi.org/10.5281/zenodo.5306081" httpUri="https://zenodo.org/record/5306081/files/figure.png" pageId="31" pageNumber="242">14</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FCD9FC0AFCACF6AE" box="[821,850,924,947]" captionStart="Figs 21–26" captionStartId="15.[113,150,1491,1510]" captionTargetBox="[157,1030,223,1437]" captionTargetId="figure-14@15.[152,1050,219,1464]" captionTargetPageId="15" captionText="Figs 21–26. Pronotum of male in dorsal aspect.21 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 22 – B. brancoi Hillert & Král sp. nov. (holotype); 23 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 24 – B. howdeni Hillert & Král sp. nov. (holotype); 25 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 26 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306085" httpUri="https://zenodo.org/record/5306085/files/figure.png" pageId="31" pageNumber="242">21</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FCB2FC0AFC85F6AE" box="[862,891,924,947]" captionStart="Figs 27–32" captionStartId="16.[151,188,1490,1509]" captionTargetBox="[151,1097,253,1372]" captionTargetId="figure-10@16.[151,1126,248,1394]" captionTargetPageId="16" captionText="Figs 27–32.Pronotum of male in frontal aspect. 27 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 28 – B. brancoi Hillert & Král sp. nov. (holotype); 29 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 30 – B. howdeni Hillert & Král sp. nov. (holotype); 31 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 32 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306087" httpUri="https://zenodo.org/record/5306087/files/figure.png" pageId="31" pageNumber="242">27</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FC6AFC0AFC5AF6AE" box="[902,932,924,947]" captionStart="Figs 33–44" captionStartId="17.[113,150,1415,1434]" captionTargetBox="[183,1019,223,1400]" captionTargetId="figure-14@17.[183,1019,219,1411]" captionTargetPageId="17" captionText="Figs 33–44. Head in dorsal aspect, 33–38 –♂♂; 39–44 – ♀♀. 33 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, NMPC); 34 – B. brancoi Hillert & Král sp. nov. (holotype); 35 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 36 – B. howdeni Hillert & Král sp. nov. (holotype); 37 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 38 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP); 39 – B. bocchus (Morocco: Gaada de Debdou plateau., E of Rchida, DKCP); 40 – B. brancoi (allotype); 41 – B. gallicus (Spain: Andalusia, Aljaraque vill. env., JSCP); 42 – B. howdeni (allotype); 43 – B. nikolajevi (allotype); 44 – B. vaulogeri (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306091" httpUri="https://zenodo.org/record/5306091/files/figure.png" pageId="31" pageNumber="242">33</figureCitation>
|
||
); medial hornlike apophysis of pronotum directed vertically upwards in lateral aspect, subapical protrusion situated basally; lateral hornlike apophyses of pronotum directed forwards and weakly divergent; aedeagus as in
|
||
<figureCitation id="3FE713D3FFA7BE22FD54FC6AFCF5F10E" box="[696,779,1020,1043]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="31" pageNumber="242">Fig. 60</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FCF4FC6AFC96F10E" box="[792,872,1020,1043]" name="Algeria" pageId="31" pageNumber="242">Algeria</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FC4FFC6AFBFDF10E" box="[931,1027,1020,1043]" name="Morocco" pageId="31" pageNumber="242">Morocco</collectingCountry>
|
||
(Fig. 59). ...............................................................................
|
||
<taxonomicName id="60DC74D5FFA7BE22FCE1FB8AFBBEF12E" authority="(Erichson, 1841)" baseAuthorityName="Erichson" baseAuthorityYear="1841" box="[781,1088,1051,1075]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="242" phylum="Arthropoda" rank="species" species="bocchus">
|
||
<emphasis id="95A8D344FFA7BE22FCE1FB8AFC78F12E" bold="true" box="[781,902,1051,1075]" italics="true" pageId="31" pageNumber="242">B. bocchus</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FFA7BE22FC78FB8AFBC6F12E" author="Erichson" box="[916,1080,1052,1075]" firstAuthor="Erichson" pageId="31" pageNumber="242" refId="ref26768" refString="ERICHSON W. F. 1841: Ueber die Insecten von Algier mit besonderer Berucksichtigung ihrer Verbreitung. Pp. 140 - 194. In: WAGNER M. F.: Reisen in der Regentschaft Algier in den Jahren 1836, 1837 und 1838 nebst einem naturhistorischen Anhang und einem Kupferatlas. Dritter Band. L. Voss, Leipzig, xviii + 297 pp. + 17 pls." type="book" year="1841">Erichson, 1841</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FFA7BE22FF9DFBAAFBBEF1EE" blockId="31.[113,1089,220,1523]" pageId="31" pageNumber="242">
|
||
<keyLead id="1C2DAC63FFA7BE22FF9DFBAAFBBEF1EE" pageId="31" pageNumber="242">
|
||
<keyStep id="1C2817F3FFA7BE22FF9DFBAAFBBEF1EE" pageId="31" pageNumber="242">
|
||
9 (8) Base of frontal horn situated at level of posterior part of eyes; horn broadly keeled from base to apex in dorsal view (
|
||
<figureCitation id="3FE713D3FFA7BE22FDC8FBCAFD96F16E" box="[548,616,1116,1139]" captionStart="Figs 5–7" captionStartId="8.[151,187,1490,1509]" captionTargetBox="[217,1057,222,1460]" captionTargetId="figure-10@8.[211,1067,219,1489]" captionTargetPageId="8" captionText="Figs 5–7.Male habitus in dorsal aspect.5 – Bolbelasmus nikolajevi Hillert,Arnone, Král & Massa sp. nov. (holotype); 6 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, 11.8 mm, DKCP); 7 – B. vaulogeri (Italy: Sicily, Menfi (Agrigento), Cinquanta, body length: 11.6 mm, MAPC)." figureDoi="http://doi.org/10.5281/zenodo.5306075" httpUri="https://zenodo.org/record/5306075/files/figure.png" pageId="31" pageNumber="242">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FD9EFBCAFD71F16E" box="[626,655,1116,1139]" captionStart="Figs 14–20" captionStartId="14.[151,188,1475,1494]" captionTargetBox="[165,1111,442,1454]" captionTargetId="figure-10@14.[156,1123,435,1471]" captionTargetPageId="14" captionText="Figs 14–20. Male habitus in lateral aspect. 14 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length:14.0 mm, NMPC); 15 – B. brancoi Hillert & Král sp.nov.(holotype); 16 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône,Barbentane, 13.8 mm, DJCP); 17 – B. howdeni Hillert & Král sp. nov.(holotype); 18 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 19 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, body length: 11.8 mm, DKCP); 20 – B. vaulogeri (Italy: Sicily, Menfi (Agrigento), Cinquanta, body length: 11.6 mm, MAPC)." figureDoi="http://doi.org/10.5281/zenodo.5306081" httpUri="https://zenodo.org/record/5306081/files/figure.png" pageId="31" pageNumber="242">18</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FD74FBCAFD4BF16E" box="[664,693,1116,1139]" captionStart="Figs 21–26" captionStartId="15.[113,150,1491,1510]" captionTargetBox="[157,1030,223,1437]" captionTargetId="figure-14@15.[152,1050,219,1464]" captionTargetPageId="15" captionText="Figs 21–26. Pronotum of male in dorsal aspect.21 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 22 – B. brancoi Hillert & Král sp. nov. (holotype); 23 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 24 – B. howdeni Hillert & Král sp. nov. (holotype); 25 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 26 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306085" httpUri="https://zenodo.org/record/5306085/files/figure.png" pageId="31" pageNumber="242">25</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FD53FBCAFD22F16E" box="[703,732,1116,1139]" captionStart="Figs 27–32" captionStartId="16.[151,188,1490,1509]" captionTargetBox="[151,1097,253,1372]" captionTargetId="figure-10@16.[151,1126,248,1394]" captionTargetPageId="16" captionText="Figs 27–32.Pronotum of male in frontal aspect. 27 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 28 – B. brancoi Hillert & Král sp. nov. (holotype); 29 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 30 – B. howdeni Hillert & Král sp. nov. (holotype); 31 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 32 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306087" httpUri="https://zenodo.org/record/5306087/files/figure.png" pageId="31" pageNumber="242">31</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FD09FBCAFCFCF16E" box="[741,770,1116,1139]" captionStart="Figs 33–44" captionStartId="17.[113,150,1415,1434]" captionTargetBox="[183,1019,223,1400]" captionTargetId="figure-14@17.[183,1019,219,1411]" captionTargetPageId="17" captionText="Figs 33–44. Head in dorsal aspect, 33–38 –♂♂; 39–44 – ♀♀. 33 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, NMPC); 34 – B. brancoi Hillert & Král sp. nov. (holotype); 35 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 36 – B. howdeni Hillert & Král sp. nov. (holotype); 37 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 38 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP); 39 – B. bocchus (Morocco: Gaada de Debdou plateau., E of Rchida, DKCP); 40 – B. brancoi (allotype); 41 – B. gallicus (Spain: Andalusia, Aljaraque vill. env., JSCP); 42 – B. howdeni (allotype); 43 – B. nikolajevi (allotype); 44 – B. vaulogeri (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306091" httpUri="https://zenodo.org/record/5306091/files/figure.png" pageId="31" pageNumber="242">37</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FCE0FBCAFCD4F16E" box="[780,810,1116,1139]" captionStart="Figs 57–58" captionStartId="26.[151,188,1519,1538]" captionTargetBox="[180,1097,219,1493]" captionTargetId="figure-10@26.[180,1098,219,1493]" captionTargetPageId="26" captionText="Figs 57–58. Living specimens. 57 – Bolbelasmus nikolajevi Hillert, Arnone, Král & Massa sp. nov. (paratype, ♂, Tunisia, Cap Bon peninsula, 6 km W of El Hawariah); 58 – B. vaulogeri (Abeille de Perrin, 1898) (♂, Tunisia, Cap Bon peninsula, 2 km NW of Douela). Photos by Petr Šípek." figureDoi="http://doi.org/10.5281/zenodo.5306099" httpUri="https://zenodo.org/record/5306099/files/figure.png" pageId="31" pageNumber="242">57</figureCitation>
|
||
); medial hornlike apophysis of pronotum directed obliquely forwards in lateral aspect, subapical protrusion situated near apex; lateral hornlike apophyses of pronotum directed forwards; aedeagus as in
|
||
<figureCitation id="3FE713D3FFA7BE22FEC4FB2AFE8FF1CE" box="[296,369,1212,1235]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="31" pageNumber="242">Fig 64</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FE97FB2AFE43F1CE" box="[379,445,1212,1235]" name="Egypt" pageId="31" pageNumber="242">Egypt</collectingCountry>
|
||
,
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FE25FB2AFDF9F1CE" box="[457,519,1212,1235]" name="Libya" pageId="31" pageNumber="242">Libya</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FDD6FB2AFD77F1CE" box="[570,649,1212,1235]" name="Tunisia" pageId="31" pageNumber="242">Tunisia</collectingCountry>
|
||
(Fig. 59). ................................................ ...............................................
|
||
<taxonomicName id="60DC74D5FFA7BE22FDE2FB4AFC1AF1EE" authority="Hillert, Arnone, Kral & Massa" authorityName="Hillert, Arnone, Kral & Massa" authorityYear="2016" box="[526,996,1243,1267]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="242" phylum="Arthropoda" rank="species" species="nikolajevi" status="sp. nov.">
|
||
<emphasis id="95A8D344FFA7BE22FDE2FB4AFD67F1EE" bold="true" box="[526,665,1243,1267]" italics="true" pageId="31" pageNumber="242">B. nikolajevi</emphasis>
|
||
Hillert, Arnone, Král & Massa
|
||
</taxonomicName>
|
||
<emphasis id="95A8D344FFA7BE22FC07FB4AFBBEF1EE" bold="true" box="[1003,1088,1244,1267]" pageId="31" pageNumber="242">
|
||
<taxonomicNameLabel id="8E9B6E3FFFA7BE22FC07FB4AFBBEF1EE" box="[1003,1088,1244,1267]" pageId="31" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FFA7BE22FF9DFB6AFCA0F00E" blockId="31.[113,1089,220,1523]" box="[113,862,1276,1299]" pageId="31" pageNumber="242">
|
||
<keyLead id="1C2DAC63FFA7BE22FF9DFB6AFCA0F00E" box="[113,862,1276,1299]" pageId="31" pageNumber="242">
|
||
<keyStep id="1C2817F3FFA7BE22FF9DFB6AFCA0F00E" box="[113,862,1276,1299]" pageId="31" pageNumber="242">10 (1) Scutellar shield densely punctate; frontal horn simple apically.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FFA7BE22FF9DFA8AFBBFF08E" blockId="31.[113,1089,220,1523]" pageId="31" pageNumber="242">
|
||
<keyLead id="1C2DAC63FFA7BE22FF9DFA8AFBBFF08E" pageId="31" pageNumber="242">
|
||
<keyStep id="1C2817F3FFA7BE22FF9DFA8AFBBFF08E" pageId="31" pageNumber="242">
|
||
11 (12) Apex of frontal horn approximately as wide as horn base, apex broadly acute-angled, strongly flattened on front part of apex (
|
||
<figureCitation id="3FE713D3FFA7BE22FD98FAAAFD44F04E" box="[628,698,1340,1363]" captionStart="Figs 1–4" captionStartId="7.[113,150,1490,1509]" captionTargetBox="[122,1077,229,1442]" captionTargetId="figure-14@7.[112,1092,216,1462]" captionTargetPageId="7" captionText="Figs 1–4. Male habitus in dorsal aspect. 1 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length: 14.0 mm, NMPC); 2 – B. brancoi Hillert & Král sp. nov. (holotype); 3 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, body length 13.7 mm, DJCP); 4 – B. howdeni Hillert & Král sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5306073" httpUri="https://zenodo.org/record/5306073/files/figure.png" pageId="31" pageNumber="242">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FD2AFAAAFD1DF04E" box="[710,739,1340,1363]" captionStart="Figs 14–20" captionStartId="14.[151,188,1475,1494]" captionTargetBox="[165,1111,442,1454]" captionTargetId="figure-10@14.[156,1123,435,1471]" captionTargetPageId="14" captionText="Figs 14–20. Male habitus in lateral aspect. 14 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length:14.0 mm, NMPC); 15 – B. brancoi Hillert & Král sp.nov.(holotype); 16 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône,Barbentane, 13.8 mm, DJCP); 17 – B. howdeni Hillert & Král sp. nov.(holotype); 18 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 19 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, body length: 11.8 mm, DKCP); 20 – B. vaulogeri (Italy: Sicily, Menfi (Agrigento), Cinquanta, body length: 11.6 mm, MAPC)." figureDoi="http://doi.org/10.5281/zenodo.5306081" httpUri="https://zenodo.org/record/5306081/files/figure.png" pageId="31" pageNumber="242">16</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FD03FAAAFCF2F04E" box="[751,780,1340,1363]" captionStart="Figs 21–26" captionStartId="15.[113,150,1491,1510]" captionTargetBox="[157,1030,223,1437]" captionTargetId="figure-14@15.[152,1050,219,1464]" captionTargetPageId="15" captionText="Figs 21–26. Pronotum of male in dorsal aspect.21 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 22 – B. brancoi Hillert & Král sp. nov. (holotype); 23 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 24 – B. howdeni Hillert & Král sp. nov. (holotype); 25 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 26 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306085" httpUri="https://zenodo.org/record/5306085/files/figure.png" pageId="31" pageNumber="242">23</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FCF4FAAAFCCBF04E" box="[792,821,1340,1363]" captionStart="Figs 27–32" captionStartId="16.[151,188,1490,1509]" captionTargetBox="[151,1097,253,1372]" captionTargetId="figure-10@16.[151,1126,248,1394]" captionTargetPageId="16" captionText="Figs 27–32.Pronotum of male in frontal aspect. 27 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 28 – B. brancoi Hillert & Král sp. nov. (holotype); 29 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 30 – B. howdeni Hillert & Král sp. nov. (holotype); 31 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 32 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306087" httpUri="https://zenodo.org/record/5306087/files/figure.png" pageId="31" pageNumber="242">29</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FCADFAAAFCA1F04E" box="[833,863,1340,1363]" captionStart="Figs 33–44" captionStartId="17.[113,150,1415,1434]" captionTargetBox="[183,1019,223,1400]" captionTargetId="figure-14@17.[183,1019,219,1411]" captionTargetPageId="17" captionText="Figs 33–44. Head in dorsal aspect, 33–38 –♂♂; 39–44 – ♀♀. 33 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, NMPC); 34 – B. brancoi Hillert & Král sp. nov. (holotype); 35 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 36 – B. howdeni Hillert & Král sp. nov. (holotype); 37 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 38 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP); 39 – B. bocchus (Morocco: Gaada de Debdou plateau., E of Rchida, DKCP); 40 – B. brancoi (allotype); 41 – B. gallicus (Spain: Andalusia, Aljaraque vill. env., JSCP); 42 – B. howdeni (allotype); 43 – B. nikolajevi (allotype); 44 – B. vaulogeri (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306091" httpUri="https://zenodo.org/record/5306091/files/figure.png" pageId="31" pageNumber="242">35</figureCitation>
|
||
); aedeagus as in
|
||
<figureCitation id="3FE713D3FFA7BE22FBFAFAAAFF1CF06E" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="31" pageNumber="242">Fig. 62</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FF01FACAFEC8F06E" box="[237,310,1372,1395]" name="France" pageId="31" pageNumber="242">France</collectingCountry>
|
||
(including
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FE5DFACAFDF9F06E" box="[433,519,1372,1395]" name="France" pageId="31" pageNumber="242">Corsica</collectingCountry>
|
||
),
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FDF4FACAFD8CF06E" box="[536,626,1372,1395]" name="Portugal" pageId="31" pageNumber="242">Portugal</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FD4BFACAFD1AF06E" box="[679,740,1372,1395]" name="Spain" pageId="31" pageNumber="242">Spain</collectingCountry>
|
||
(Fig. 59). ................................... .......................................................................................
|
||
<taxonomicName id="60DC74D5FFA7BE22FCF5FAEAFBBFF08E" authority="(Mulsant, 1842)" baseAuthorityName="Mulsant" baseAuthorityYear="1842" box="[793,1089,1403,1427]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="242" phylum="Arthropoda" rank="species" species="gallicus">
|
||
<emphasis id="95A8D344FFA7BE22FCF5FAEAFC70F08E" bold="true" box="[793,910,1403,1427]" italics="true" pageId="31" pageNumber="242">B. gallicus</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FFA7BE22FC70FAEAFBC7F08E" author="Mulsant" box="[924,1081,1404,1427]" firstAuthor="Mulsant" pageId="31" pageNumber="242" refId="ref27806" refString="MULSANT E. 1842: Histoire naturelle des Coleopteres de France. Lamellicornes. Maison Libraire, Paris & Imprimerie de Dumoulin, Ronet et Sibuet, Lyon, viii + 626 pp. + 3 pls." type="book" year="1842">Mulsant, 1842</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FFA7BE22FF9DFA0AFBBEF0EE" blockId="31.[113,1089,220,1523]" pageId="31" pageNumber="242">
|
||
<keyLead id="1C2DAC63FFA7BE22FF9DFA0AFBBEF0EE" pageId="31" pageNumber="242">
|
||
<keyStep id="1C2817F3FFA7BE22FF9DFA0AFBBEF0EE" pageId="31" pageNumber="242">
|
||
12 (11) Apex of frontal horn becoming gradually thinner from base to apex, apex acute-angled, only weakly flattened on front part of apex (
|
||
<figureCitation id="3FE713D3FFA7BE22FD22FA2AFCECF0CE" box="[718,786,1468,1491]" captionStart="Figs 1–4" captionStartId="7.[113,150,1490,1509]" captionTargetBox="[122,1077,229,1442]" captionTargetId="figure-14@7.[112,1092,216,1462]" captionTargetPageId="7" captionText="Figs 1–4. Male habitus in dorsal aspect. 1 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length: 14.0 mm, NMPC); 2 – B. brancoi Hillert & Král sp. nov. (holotype); 3 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, body length 13.7 mm, DJCP); 4 – B. howdeni Hillert & Král sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5306073" httpUri="https://zenodo.org/record/5306073/files/figure.png" pageId="31" pageNumber="242">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FCF1FA2AFCC4F0CE" box="[797,826,1468,1491]" captionStart="Figs 14–20" captionStartId="14.[151,188,1475,1494]" captionTargetBox="[165,1111,442,1454]" captionTargetId="figure-10@14.[156,1123,435,1471]" captionTargetPageId="14" captionText="Figs 14–20. Male habitus in lateral aspect. 14 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, body length:14.0 mm, NMPC); 15 – B. brancoi Hillert & Král sp.nov.(holotype); 16 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône,Barbentane, 13.8 mm, DJCP); 17 – B. howdeni Hillert & Král sp. nov.(holotype); 18 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 19 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, body length: 11.8 mm, DKCP); 20 – B. vaulogeri (Italy: Sicily, Menfi (Agrigento), Cinquanta, body length: 11.6 mm, MAPC)." figureDoi="http://doi.org/10.5281/zenodo.5306081" httpUri="https://zenodo.org/record/5306081/files/figure.png" pageId="31" pageNumber="242">17</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FCA8FA2AFC9FF0CE" box="[836,865,1468,1491]" captionStart="Figs 21–26" captionStartId="15.[113,150,1491,1510]" captionTargetBox="[157,1030,223,1437]" captionTargetId="figure-14@15.[152,1050,219,1464]" captionTargetPageId="15" captionText="Figs 21–26. Pronotum of male in dorsal aspect.21 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 22 – B. brancoi Hillert & Král sp. nov. (holotype); 23 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 24 – B. howdeni Hillert & Král sp. nov. (holotype); 25 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 26 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306085" httpUri="https://zenodo.org/record/5306085/files/figure.png" pageId="31" pageNumber="242">24</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FC87FA2AFC76F0CE" box="[875,904,1468,1491]" captionStart="Figs 27–32" captionStartId="16.[151,188,1490,1509]" captionTargetBox="[151,1097,253,1372]" captionTargetId="figure-10@16.[151,1126,248,1394]" captionTargetPageId="16" captionText="Figs 27–32.Pronotum of male in frontal aspect. 27 – Bolbelasmus bocchus (Erichson, 1841) (Morocco:Azrou, Foret de Cedres, NMPC); 28 – B. brancoi Hillert & Král sp. nov. (holotype); 29 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 30 – B. howdeni Hillert & Král sp. nov. (holotype); 31 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 32 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306087" httpUri="https://zenodo.org/record/5306087/files/figure.png" pageId="31" pageNumber="242">30</figureCitation>
|
||
,
|
||
<figureCitation id="3FE713D3FFA7BE22FC7EFA2AFC4EF0CE" box="[914,944,1468,1491]" captionStart="Figs 33–44" captionStartId="17.[113,150,1415,1434]" captionTargetBox="[183,1019,223,1400]" captionTargetId="figure-14@17.[183,1019,219,1411]" captionTargetPageId="17" captionText="Figs 33–44. Head in dorsal aspect, 33–38 –♂♂; 39–44 – ♀♀. 33 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, NMPC); 34 – B. brancoi Hillert & Král sp. nov. (holotype); 35 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 36 – B. howdeni Hillert & Král sp. nov. (holotype); 37 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 38 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP); 39 – B. bocchus (Morocco: Gaada de Debdou plateau., E of Rchida, DKCP); 40 – B. brancoi (allotype); 41 – B. gallicus (Spain: Andalusia, Aljaraque vill. env., JSCP); 42 – B. howdeni (allotype); 43 – B. nikolajevi (allotype); 44 – B. vaulogeri (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306091" httpUri="https://zenodo.org/record/5306091/files/figure.png" pageId="31" pageNumber="242">36</figureCitation>
|
||
); aedeagus as in
|
||
<figureCitation id="3FE713D3FFA7BE22FF0CFA4AFED1F0EE" box="[224,303,1500,1523]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="31" pageNumber="242">Fig. 63</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FED6FA4AFE87F0EE" box="[314,377,1500,1523]" name="Spain" pageId="31" pageNumber="242">Spain</collectingCountry>
|
||
,
|
||
<collectingCountry id="DFCB4FC6FFA7BE22FE68FA4AFE1AF0EE" box="[388,484,1500,1523]" name="Gibraltar" pageId="31" pageNumber="242">Gibraltar</collectingCountry>
|
||
(Fig. 59). ................
|
||
<taxonomicName id="60DC74D5FFA7BE22FD26FA4AFC1AF0EE" authority="Hillert & Kral" authorityName="Hillert & Kral" authorityYear="2016" box="[714,996,1499,1523]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="242" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
|
||
<emphasis id="95A8D344FFA7BE22FD26FA4AFCB8F0EE" bold="true" box="[714,838,1499,1523]" italics="true" pageId="31" pageNumber="242">B. howdeni</emphasis>
|
||
Hillert & Král
|
||
</taxonomicName>
|
||
<emphasis id="95A8D344FFA7BE22FC07FA4AFBBEF0EE" bold="true" box="[1003,1088,1500,1523]" pageId="31" pageNumber="242">
|
||
<taxonomicNameLabel id="8E9B6E3FFFA7BE22FC07FA4AFBBEF0EE" box="[1003,1088,1500,1523]" pageId="31" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</subSubSection>
|
||
<subSubSection id="EFC65CDDFF98BE1DFE82FF4AFC6EF5EB" box="[366,912,220,246]" pageId="32" pageNumber="243" type="description">
|
||
<paragraph id="A7630F56FF98BE1DFE82FF4AFC6EF5EB" blockId="32.[366,912,220,246]" box="[366,912,220,246]" pageId="32" pageNumber="243">
|
||
<heading id="FC2BB83AFF98BE1DFE82FF4AFC6EF5EB" bold="true" box="[366,912,220,246]" centered="true" fontSize="11" level="1" pageId="32" pageNumber="243" reason="2">
|
||
<emphasis id="95A8D344FF98BE1DFE82FF4AFC6EF5EB" bold="true" box="[366,912,220,246]" pageId="32" pageNumber="243">Identification key for less developed males</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFC65CDDFF98BE1DFF7BFE82FB98F036" pageId="32" pageNumber="243" type="key">
|
||
<key id="D34D5867FF98BE1DFF7BFE82FB98F036" pageId="32" pageNumber="243">
|
||
<paragraph id="A7630F56FF98BE1DFF7BFE82FE6BF456" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFE82FE6BF456" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFE82FE6BF456" pageId="32" pageNumber="243">1 (10) Scutellar shield impunctate or weakly shallowly punctate, frontal horn simple or furcate apically.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFEC2FC76F496" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFEC2FC76F496" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFEC2FC76F496" pageId="32" pageNumber="243">2 (5) Frontal horn simple; genae regularly rounded; oblique keel above eyes extending over whole eyes length; head strongly constricted behind eyes.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFE02FB98F4F6" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFE02FB98F4F6" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFE02FB98F4F6" pageId="32" pageNumber="243">
|
||
3 (4) Scutellar shield triangular, approximately as long as wide; body surface brownish, maximal body length
|
||
<quantity id="6024A2B3FF98BE1DFE3CFE22FDE3F4D6" box="[464,541,436,459]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="243" unit="mm" value="12.0">12 mm</quantity>
|
||
. Southern parts of Central Europe, Alsace,
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFC0CFE22FBECF4D6" box="[992,1042,436,459]" name="Italy" pageId="32" pageNumber="243">Italy</collectingCountry>
|
||
, Balkan Peninsula,
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFEB3FE42FE52F4F6" box="[351,428,468,491]" name="Poland" pageId="32" pageNumber="243">Poland</collectingCountry>
|
||
,
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFE5BFE42FE00F4F6" box="[439,510,468,491]" name="Russia" pageId="32" pageNumber="243">Russia</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFDDEFE42FD75F4F6" box="[562,651,468,491]" name="Ukraine" pageId="32" pageNumber="243">Ukraine</collectingCountry>
|
||
. ....
|
||
<taxonomicName id="60DC74D5FF98BE1DFD57FE42FB98F4F6" authority="(Schrank von Paula, 1789)" baseAuthorityName="Schrank von Paula" baseAuthorityYear="1789" box="[699,1126,467,491]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="243" phylum="Arthropoda" rank="species" species="unicornis">
|
||
<emphasis id="95A8D344FF98BE1DFD57FE42FCBDF4F6" bold="true" box="[699,835,467,491]" italics="true" pageId="32" pageNumber="243">B. unicornis</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FF98BE1DFCBDFE42FBA1F4F6" author="Schrank von Paula" box="[849,1119,468,491]" firstAuthor="Schrank von Paula" pageId="32" pageNumber="243" pagination="60 - 90" refId="ref28580" refString="SCHRANK VON PAULA F. 1789: Entomologische Beobachtungen. Der Naturforscher 24: 60 - 90." type="journal article" year="1789">Schrank von Paula, 1789</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFE62FB98F756" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFE62FB98F756" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFE62FB98F756" pageId="32" pageNumber="243">
|
||
4 (3) Scutellar shield triangular, longer than wide; body surface blackish, maximal body length
|
||
<quantity id="6024A2B3FF98BE1DFED9FD82FE7DF736" box="[309,387,532,555]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="32" pageNumber="243" unit="mm" value="11.0">11 mm</quantity>
|
||
; aedeagus as in
|
||
<figureCitation id="3FE713D3FF98BE1DFDDFFD82FD7DF736" box="[563,643,532,555]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="32" pageNumber="243">Fig. 65</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFD62FD82FD1FF736" box="[654,737,532,555]" name="Algeria" pageId="32" pageNumber="243">Algeria</collectingCountry>
|
||
,
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFD00FD82FCE3F736" box="[748,797,532,555]" name="Italy" pageId="32" pageNumber="243">Italy</collectingCountry>
|
||
(
|
||
<collectingRegion id="6518C1B4FF98BE1DFCC0FD82FC93F736" box="[812,877,532,555]" country="Italy" name="Sicilia" pageId="32" pageNumber="243">Sicily</collectingRegion>
|
||
) and
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFC46FD82FC07F736" box="[938,1017,532,555]" name="Tunisia" pageId="32" pageNumber="243">Tunisia</collectingCountry>
|
||
(Fig. 59). .......................................................................
|
||
<taxonomicName id="60DC74D5FF98BE1DFD22FDA2FB98F756" authority="(Abeille de Perrin, 1898)" baseAuthorityName="Abeille de Perrin" baseAuthorityYear="1898" box="[718,1126,563,587]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="243" phylum="Arthropoda" rank="species" species="vaulogeri">
|
||
<emphasis id="95A8D344FF98BE1DFD22FDA2FCABF756" bold="true" box="[718,853,563,587]" italics="true" pageId="32" pageNumber="243">B. vaulogeri</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FF98BE1DFC8FFDA2FBA1F756" author="Abeille de Perrin" box="[867,1119,564,587]" firstAuthor="Abeille de Perrin" pageId="32" pageNumber="243" pagination="254 - 258" refId="ref25891" refString="ABEILLE DE PERRIN E. 1898: Descriptions de Coleopteres du Nord de l'Afrique. Bulletin de la Societe Entomologique de France 1898: 254 - 258." type="journal article" year="1898">Abeille de Perrin, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFDC2FE8EF7B6" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFDC2FE8EF7B6" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFDC2FE8EF7B6" pageId="32" pageNumber="243">5 (2) Frontal horn furcate apically; genae more or less angulate anterolaterally; oblique keel above eyes reaching only to anterior part of eyes; head not strongly constricted behind eyes.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFD22FB98F656" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFD22FB98F656" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFD22FB98F656" pageId="32" pageNumber="243">
|
||
6 (7) Pronotal punctation denser with exception of basal part, punctures separated approximately by distance equal to their diameters; pronotal base with distinct visible but fine, regularly spaced pinned points, elytral striae well impressed, visible as complete lines; elytral intervals moderately convex; aedeagus as in
|
||
<figureCitation id="3FE713D3FF98BE1DFC78FC82FC1DF636" box="[916,995,788,811]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="32" pageNumber="243">Fig. 61</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFC03FC82FBD2F636" box="[1007,1068,788,811]" name="Spain" pageId="32" pageNumber="243">Spain</collectingCountry>
|
||
(Fig. 59). ......................................................................
|
||
<taxonomicName id="60DC74D5FF98BE1DFD15FCA2FBF4F656" authority="Hillert & Kral" authorityName="Hillert & Kral" authorityYear="2016" box="[761,1034,819,843]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="243" phylum="Arthropoda" rank="species" species="brancoi" status="sp. nov.">
|
||
<emphasis id="95A8D344FF98BE1DFD15FCA2FC92F656" bold="true" box="[761,876,819,843]" italics="true" pageId="32" pageNumber="243">B. brancoi</emphasis>
|
||
Hillert & Král
|
||
</taxonomicName>
|
||
<emphasis id="95A8D344FF98BE1DFBFDFCA2FB98F656" bold="true" box="[1041,1126,820,843]" pageId="32" pageNumber="243">
|
||
<taxonomicNameLabel id="8E9B6E3FFF98BE1DFBFDFCA2FB98F656" box="[1041,1126,820,843]" pageId="32" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFCC2FE68F6D6" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFCC2FE68F6D6" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFCC2FE68F6D6" pageId="32" pageNumber="243">7 (6) Pronotal punctation sparser, punctures separated by approximately three times their diameters pronotal base with very fine, weakly visible and well separated pinned points; elytral striae not impressed, visible as row of punctures; elytral intervals flat; northern Africa.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFC42FB98F136" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFC42FB98F136" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFC42FB98F136" pageId="32" pageNumber="243">
|
||
8 (9) Base of frontal horn situated at level of anterior part of eyes; horn sharply keeled basally only; aedeagus as in
|
||
<figureCitation id="3FE713D3FF98BE1DFDF7FC62FD94F116" box="[539,618,1012,1035]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="32" pageNumber="243">Fig. 60</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFD9FFC62FD3DF116" box="[627,707,1012,1035]" name="Algeria" pageId="32" pageNumber="243">Algeria</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFD1BFC62FCA9F116" box="[759,855,1012,1035]" name="Morocco" pageId="32" pageNumber="243">Morocco</collectingCountry>
|
||
(Fig. 59). ....................... ......................................................................................
|
||
<taxonomicName id="60DC74D5FF98BE1DFCDFFB82FB98F136" authority="(Erichson, 1841)" baseAuthorityName="Erichson" baseAuthorityYear="1841" box="[819,1126,1043,1067]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="243" phylum="Arthropoda" rank="species" species="bocchus">
|
||
<emphasis id="95A8D344FF98BE1DFCDFFB82FC52F136" bold="true" box="[819,940,1043,1067]" italics="true" pageId="32" pageNumber="243">B. bocchus</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FF98BE1DFC56FB82FBA0F136" author="Erichson" box="[954,1118,1044,1067]" firstAuthor="Erichson" pageId="32" pageNumber="243" refId="ref26768" refString="ERICHSON W. F. 1841: Ueber die Insecten von Algier mit besonderer Berucksichtigung ihrer Verbreitung. Pp. 140 - 194. In: WAGNER M. F.: Reisen in der Regentschaft Algier in den Jahren 1836, 1837 und 1838 nebst einem naturhistorischen Anhang und einem Kupferatlas. Dritter Band. L. Voss, Leipzig, xviii + 297 pp. + 17 pls." type="book" year="1841">Erichson, 1841</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFBA2FB98F196" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFBA2FB98F196" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFBA2FB98F196" pageId="32" pageNumber="243">
|
||
9 (8) Base of frontal horn situated at level of posterior part of eyes; horn broadly keeled from base to apex in dorsal view; aedeagus as in
|
||
<figureCitation id="3FE713D3FF98BE1DFD10FBC2FCB5F176" box="[764,843,1108,1131]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="32" pageNumber="243">Fig. 64</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFCBBFBC2FC67F176" box="[855,921,1108,1131]" name="Egypt" pageId="32" pageNumber="243">Egypt</collectingCountry>
|
||
,
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFC49FBC2FC1DF176" box="[933,995,1108,1131]" name="Libya" pageId="32" pageNumber="243">Libya</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFBFBFBC2FB98F176" box="[1047,1126,1108,1131]" name="Tunisia" pageId="32" pageNumber="243">Tunisia</collectingCountry>
|
||
(Fig. 59). ...............................
|
||
<taxonomicName id="60DC74D5FF98BE1DFDD8FBE2FBF4F196" authority="Hillert, Arnone, Kral & Massa" authorityName="Hillert, Arnone, Kral & Massa" authorityYear="2016" box="[564,1034,1139,1163]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="243" phylum="Arthropoda" rank="species" species="nikolajevi" status="sp. nov.">
|
||
<emphasis id="95A8D344FF98BE1DFDD8FBE2FD41F196" bold="true" box="[564,703,1139,1163]" italics="true" pageId="32" pageNumber="243">B. nikolajevi</emphasis>
|
||
Hillert, Arnone, Král & Massa
|
||
</taxonomicName>
|
||
<emphasis id="95A8D344FF98BE1DFBFDFBE2FB98F196" bold="true" box="[1041,1126,1140,1163]" pageId="32" pageNumber="243">
|
||
<taxonomicNameLabel id="8E9B6E3FFF98BE1DFBFDFBE2FB98F196" box="[1041,1126,1140,1163]" pageId="32" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFB02FC7AF1B6" blockId="32.[151,1127,276,1323]" box="[151,900,1172,1195]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFB02FC7AF1B6" box="[151,900,1172,1195]" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFB02FC7AF1B6" box="[151,900,1172,1195]" pageId="32" pageNumber="243">10 (1) Scutellar shield densely punctate; frontal horn simple apically.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFB22FB98F1F6" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFB22FB98F1F6" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFB22FB98F1F6" pageId="32" pageNumber="243">
|
||
11 (12) Apex of frontal horn flattened on front part of apex; aedeagus as in
|
||
<figureCitation id="3FE713D3FF98BE1DFC25FB22FBECF1D6" box="[969,1042,1204,1227]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="32" pageNumber="243">Fig 62</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFBF2FB22FB99F1D6" box="[1054,1127,1204,1227]" name="France" pageId="32" pageNumber="243">France</collectingCountry>
|
||
(including
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFEB3FB42FE4BF1F6" box="[351,437,1236,1259]" name="France" pageId="32" pageNumber="243">Corsica</collectingCountry>
|
||
),
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFE2AFB42FDDEF1F6" box="[454,544,1236,1259]" name="Portugal" pageId="32" pageNumber="243">Portugal</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFDB9FB42FD6CF1F6" box="[597,658,1236,1259]" name="Spain" pageId="32" pageNumber="243">Spain</collectingCountry>
|
||
(Fig. 59). .......
|
||
<taxonomicName id="60DC74D5FF98BE1DFCD2FB42FB98F1F6" authority="(Mulsant, 1842)" baseAuthorityName="Mulsant" baseAuthorityYear="1842" box="[830,1126,1235,1259]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="243" phylum="Arthropoda" rank="species" species="gallicus">
|
||
<emphasis id="95A8D344FF98BE1DFCD2FB42FC4AF1F6" bold="true" box="[830,948,1235,1259]" italics="true" pageId="32" pageNumber="243">B. gallicus</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FF98BE1DFC2DFB42FBA1F1F6" author="Mulsant" box="[961,1119,1236,1259]" firstAuthor="Mulsant" pageId="32" pageNumber="243" refId="ref27806" refString="MULSANT E. 1842: Histoire naturelle des Coleopteres de France. Lamellicornes. Maison Libraire, Paris & Imprimerie de Dumoulin, Ronet et Sibuet, Lyon, viii + 626 pp. + 3 pls." type="book" year="1842">Mulsant, 1842</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFB62FB98F036" blockId="32.[151,1127,276,1323]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFB62FB98F036" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFB62FB98F036" pageId="32" pageNumber="243">
|
||
12 (11) Apex of frontal horn regularly arcuate, not flattened on front part of apex; aedeagus as in
|
||
<figureCitation id="3FE713D3FF98BE1DFECFFA82FE8CF036" box="[291,370,1300,1323]" captionStart="Figs 60–65" captionStartId="29.[113,150,1482,1501]" captionTargetBox="[200,960,228,1448]" captionTargetId="figure-14@29.[192,1010,219,1474]" captionTargetPageId="29" captionText="Figs 60–65. Aedeagus of Bolbelasmus species, ventral aspect, schematically. 60 – B. bocchus (Erichson, 1841), 61 – B. brancoi Hillert & Král sp. nov., 62 – B. gallicus (Mulsant, 1842), 63 – B. howdeni Hillert & Král sp. nov., 64 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov., 65 – B. vaulogeri (Abeille de Perrin, 1898)." figureDoi="http://doi.org/10.5281/zenodo.5306101" httpUri="https://zenodo.org/record/5306101/files/figure.png" pageId="32" pageNumber="243">Fig. 63</figureCitation>
|
||
;
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFE90FA82FE45F036" box="[380,443,1300,1323]" name="Spain" pageId="32" pageNumber="243">Spain</collectingCountry>
|
||
,
|
||
<collectingCountry id="DFCB4FC6FF98BE1DFE2AFA82FDD8F036" box="[454,550,1300,1323]" name="Gibraltar" pageId="32" pageNumber="243">Gibraltar</collectingCountry>
|
||
(Fig. 59). ............
|
||
<taxonomicName id="60DC74D5FF98BE1DFD1CFA82FBF4F036" authority="Hillert & Kral" authorityName="Hillert & Kral" authorityYear="2016" box="[752,1034,1299,1323]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="243" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
|
||
<emphasis id="95A8D344FF98BE1DFD1CFA82FC92F036" bold="true" box="[752,876,1299,1323]" italics="true" pageId="32" pageNumber="243">B. howdeni</emphasis>
|
||
Hillert & Král
|
||
</taxonomicName>
|
||
<emphasis id="95A8D344FF98BE1DFBFDFA82FB98F036" bold="true" box="[1041,1126,1300,1323]" pageId="32" pageNumber="243">
|
||
<taxonomicNameLabel id="8E9B6E3FFF98BE1DFBFDFA82FB98F036" box="[1041,1126,1300,1323]" pageId="32" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</subSubSection>
|
||
<subSubSection id="EFC65CDDFF98BE1DFE2FFACDFCC4F068" box="[451,826,1371,1397]" pageId="32" pageNumber="243" type="description">
|
||
<paragraph id="A7630F56FF98BE1DFE2FFACDFCC4F068" blockId="32.[451,826,1371,1397]" box="[451,826,1371,1397]" pageId="32" pageNumber="243">
|
||
<heading id="FC2BB83AFF98BE1DFE2FFACDFCC4F068" bold="true" box="[451,826,1371,1397]" centered="true" fontSize="11" level="1" pageId="32" pageNumber="243" reason="2">
|
||
<emphasis id="95A8D344FF98BE1DFE2FFACDFCC4F068" bold="true" box="[451,826,1371,1397]" pageId="32" pageNumber="243">Identification key for females</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFC65CDDFF98BE1CFF7BFA05FBBEF16E" lastPageId="33" lastPageNumber="244" pageId="32" pageNumber="243" type="key">
|
||
<key id="D34D5867FF98BE1CFF7BFA05FBBEF16E" lastPageId="33" lastPageNumber="244" pageId="32" pageNumber="243">
|
||
<paragraph id="A7630F56FF98BE1DFF7BFA05FCA5F0B7" blockId="32.[151,1126,1427,1514]" box="[151,859,1427,1450]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFA05FCA5F0B7" box="[151,859,1427,1450]" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFA05FCA5F0B7" box="[151,859,1427,1450]" pageId="32" pageNumber="243">1 (10) Scutellar shield impunctate, or weakly shallowly punctate.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF98BE1DFF7BFA25FBDFF0F7" blockId="32.[151,1126,1427,1514]" pageId="32" pageNumber="243">
|
||
<keyLead id="1C2DAC63FF98BE1DFF7BFA25FBDFF0F7" pageId="32" pageNumber="243">
|
||
<keyStep id="1C2817F3FF98BE1DFF7BFA25FBDFF0F7" pageId="32" pageNumber="243">
|
||
2 (5) Frontal tubercle simple (
|
||
<figureCitation id="3FE713D3FF98BE1DFE17FA25FD55F0D7" box="[507,683,1459,1482]" captionStart="Figs 33–44" captionStartId="17.[113,150,1415,1434]" captionTargetBox="[183,1019,223,1400]" captionTargetId="figure-14@17.[183,1019,219,1411]" captionTargetPageId="17" captionText="Figs 33–44. Head in dorsal aspect, 33–38 –♂♂; 39–44 – ♀♀. 33 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, NMPC); 34 – B. brancoi Hillert & Král sp. nov. (holotype); 35 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 36 – B. howdeni Hillert & Král sp. nov. (holotype); 37 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 38 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP); 39 – B. bocchus (Morocco: Gaada de Debdou plateau., E of Rchida, DKCP); 40 – B. brancoi (allotype); 41 – B. gallicus (Spain: Andalusia, Aljaraque vill. env., JSCP); 42 – B. howdeni (allotype); 43 – B. nikolajevi (allotype); 44 – B. vaulogeri (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306091" httpUri="https://zenodo.org/record/5306091/files/figure.png" pageId="32" pageNumber="243">Figs 41, 42, 44</figureCitation>
|
||
); genae regularly rounded; oblique keel above eyes over whole length of eyes; head strongly constricted behind eyes.
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF99BE1CFF9DFF4AFBBFF42E" blockId="33.[113,1089,220,1139]" pageId="33" pageNumber="244">
|
||
<keyLead id="1C2DAC63FF99BE1CFF9DFF4AFBBFF42E" pageId="33" pageNumber="244">
|
||
<keyStep id="1C2817F3FF99BE1CFF9DFF4AFBBFF42E" pageId="33" pageNumber="244">
|
||
3 (4) Scutellar shield triangular, approximately as long as wide; body surface brownish, minimal body length
|
||
<quantity id="6024A2B3FF99BE1CFE4BFF6AFE0AF40E" box="[423,500,252,275]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="244" unit="mm" value="12.0">12 mm</quantity>
|
||
. Southern parts of Central Europe, Alsace,
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFC56FF6AFC12F40E" box="[954,1004,252,275]" name="Italy" pageId="33" pageNumber="244">Italy</collectingCountry>
|
||
, Balkan Peninsula,
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFED6FE8AFE79F42E" box="[314,391,284,307]" name="Poland" pageId="33" pageNumber="244">Poland</collectingCountry>
|
||
,
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFE7DFE8AFE26F42E" box="[401,472,284,307]" name="Russia" pageId="33" pageNumber="244">Russia</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFDE0FE8AFD9BF42E" box="[524,613,284,307]" name="Ukraine" pageId="33" pageNumber="244">Ukraine</collectingCountry>
|
||
. ....
|
||
<taxonomicName id="60DC74D5FF99BE1CFD79FE8AFBBFF42E" authority="(Schrank von Paula, 1789)" baseAuthorityName="Schrank von Paula" baseAuthorityYear="1789" box="[661,1089,283,307]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="unicornis">
|
||
<emphasis id="95A8D344FF99BE1CFD79FE8AFCE3F42E" bold="true" box="[661,797,283,307]" italics="true" pageId="33" pageNumber="244">B. unicornis</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FF99BE1CFCC7FE8AFBC7F42E" author="Schrank von Paula" box="[811,1081,284,307]" firstAuthor="Schrank von Paula" pageId="33" pageNumber="244" pagination="60 - 90" refId="ref28580" refString="SCHRANK VON PAULA F. 1789: Entomologische Beobachtungen. Der Naturforscher 24: 60 - 90." type="journal article" year="1789">Schrank von Paula, 1789</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF99BE1CFF9DFEAAFBBEF48E" blockId="33.[113,1089,220,1139]" pageId="33" pageNumber="244">
|
||
<keyLead id="1C2DAC63FF99BE1CFF9DFEAAFBBEF48E" pageId="33" pageNumber="244">
|
||
<keyStep id="1C2817F3FF99BE1CFF9DFEAAFBBEF48E" pageId="33" pageNumber="244">
|
||
4 (3) Scutellar shield triangular longer than wide; body surface blackish, minimal body length
|
||
<quantity id="6024A2B3FF99BE1CFEE2FECAFEA9F46E" box="[270,343,348,371]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="33" pageNumber="244" unit="mm" value="11.0">11 mm</quantity>
|
||
(
|
||
<figureCitation id="3FE713D3FF99BE1CFE8AFECAFE50F46E" box="[358,430,348,371]" captionStart="Figs 12–13" captionStartId="10.[151,188,914,933]" captionTargetBox="[177,1107,239,881]" captionTargetId="figure-10@10.[167,1111,232,885]" captionTargetPageId="10" captionText="Figs 12–13. Female habitus in dorsal aspect. 12 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (allotype); 13 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, 12.0 mm, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306079" httpUri="https://zenodo.org/record/5306079/files/figure.png" pageId="33" pageNumber="244">Fig 13</figureCitation>
|
||
);
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFE2EFECAFDEBF46E" box="[450,533,348,371]" name="Algeria" pageId="33" pageNumber="244">Algeria</collectingCountry>
|
||
,
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFDF3FECAFDAEF46E" box="[543,592,348,371]" name="Italy" pageId="33" pageNumber="244">Italy</collectingCountry>
|
||
(
|
||
<collectingRegion id="6518C1B4FF99BE1CFDB2FECAFD61F46E" box="[606,671,348,371]" country="Italy" name="Sicilia" pageId="33" pageNumber="244">Sicily</collectingRegion>
|
||
) and
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFD36FECAFCD7F46E" box="[730,809,348,371]" name="Tunisia" pageId="33" pageNumber="244">Tunisia</collectingCountry>
|
||
(Fig. 59). ........................ .......................................................................
|
||
<taxonomicName id="60DC74D5FF99BE1CFD44FEEAFBBEF48E" authority="(Abeille de Perrin, 1898)" baseAuthorityName="Abeille de Perrin" baseAuthorityYear="1898" box="[680,1088,379,403]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="vaulogeri">
|
||
<emphasis id="95A8D344FF99BE1CFD44FEEAFCD1F48E" bold="true" box="[680,815,379,403]" italics="true" pageId="33" pageNumber="244">B. vaulogeri</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FF99BE1CFCD1FEEAFBC7F48E" author="Abeille de Perrin" box="[829,1081,380,403]" firstAuthor="Abeille de Perrin" pageId="33" pageNumber="244" pagination="254 - 258" refId="ref25891" refString="ABEILLE DE PERRIN E. 1898: Descriptions de Coleopteres du Nord de l'Afrique. Bulletin de la Societe Entomologique de France 1898: 254 - 258." type="journal article" year="1898">Abeille de Perrin, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF99BE1CFF9DFE0AFDDAF4EE" blockId="33.[113,1089,220,1139]" pageId="33" pageNumber="244">
|
||
<keyLead id="1C2DAC63FF99BE1CFF9DFE0AFDDAF4EE" pageId="33" pageNumber="244">
|
||
<keyStep id="1C2817F3FF99BE1CFF9DFE0AFDDAF4EE" pageId="33" pageNumber="244">
|
||
5 (2) Frontal tubercle furcate apically (
|
||
<figureCitation id="3FE713D3FF99BE1CFDCBFE0AFD34F4AE" box="[551,714,412,435]" captionStart="Figs 33–44" captionStartId="17.[113,150,1415,1434]" captionTargetBox="[183,1019,223,1400]" captionTargetId="figure-14@17.[183,1019,219,1411]" captionTargetPageId="17" captionText="Figs 33–44. Head in dorsal aspect, 33–38 –♂♂; 39–44 – ♀♀. 33 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Azrou, Foret de Cedres, NMPC); 34 – B. brancoi Hillert & Král sp. nov. (holotype); 35 – B. gallicus (Mulsant, 1842) (France: Bouches-du-Rhône, Barbentane, DJCP); 36 – B. howdeni Hillert & Král sp. nov. (holotype); 37 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (holotype); 38 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP); 39 – B. bocchus (Morocco: Gaada de Debdou plateau., E of Rchida, DKCP); 40 – B. brancoi (allotype); 41 – B. gallicus (Spain: Andalusia, Aljaraque vill. env., JSCP); 42 – B. howdeni (allotype); 43 – B. nikolajevi (allotype); 44 – B. vaulogeri (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306091" httpUri="https://zenodo.org/record/5306091/files/figure.png" pageId="33" pageNumber="244">Figs 39–40, 43</figureCitation>
|
||
); genae more or less angulate anterolaterally; oblique keel above eyes reaching only to anterior part of eyes; head not strongly constricted behind eyes.
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF99BE1CFF9DFE6AFBBEF78E" blockId="33.[113,1089,220,1139]" pageId="33" pageNumber="244">
|
||
<keyLead id="1C2DAC63FF99BE1CFF9DFE6AFBBEF78E" pageId="33" pageNumber="244">
|
||
<keyStep id="1C2817F3FF99BE1CFF9DFE6AFBBEF78E" pageId="33" pageNumber="244">
|
||
6 (7) Pronotal punctation denser with exception of basal part, punctures separated approximately by distance equal to their diameters; pronotal base with distinct visible but fine, regularly pinned points; elytral striae well impressed, visible as complete lines; elytral intervals moderately convex (
|
||
<figureCitation id="3FE713D3FF99BE1CFDA0FDCAFD73F76E" box="[588,653,604,627]" captionStart="Figs 8–11" captionStartId="9.[113,150,1520,1539]" captionTargetBox="[136,1072,224,1492]" captionTargetId="figure-14@9.[129,1073,219,1523]" captionTargetPageId="9" captionText="Figs 8–11. Female habitus in dorsal aspect. 8 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Gaada de Debdou plateau., E of Rchida, 13.5 mm, DKCP); 9 – B. brancoi Hillert & Král sp. nov. (allotype); 10 – B. gallicus (Mulsant, 1842) (Spain: Andalusia, Aljaraque vill. env., 13.6 mm, JSCP); 11 – B. howdeni Hillert & Král sp. nov. (allotype)." figureDoi="http://doi.org/10.5281/zenodo.5306077" httpUri="https://zenodo.org/record/5306077/files/figure.png" pageId="33" pageNumber="244">Fig. 9</figureCitation>
|
||
);
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFD4EFDCAFD21F76E" box="[674,735,604,627]" name="Spain" pageId="33" pageNumber="244">Spain</collectingCountry>
|
||
(Fig. 59). ................................... .............................................................................
|
||
<taxonomicName id="60DC74D5FF99BE1CFD3FFDEAFC1AF78E" authority="Hillert & Kral" authorityName="Hillert & Kral" authorityYear="2016" box="[723,996,635,659]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="brancoi" status="sp. nov.">
|
||
<emphasis id="95A8D344FF99BE1CFD3FFDEAFCB8F78E" bold="true" box="[723,838,635,659]" italics="true" pageId="33" pageNumber="244">B. brancoi</emphasis>
|
||
Hillert & Král
|
||
</taxonomicName>
|
||
<emphasis id="95A8D344FF99BE1CFC07FDEAFBBEF78E" bold="true" box="[1003,1088,636,659]" pageId="33" pageNumber="244">
|
||
<taxonomicNameLabel id="8E9B6E3FFF99BE1CFC07FDEAFBBEF78E" box="[1003,1088,636,659]" pageId="33" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF99BE1CFF9DFD0AFE8EF60E" blockId="33.[113,1089,220,1139]" pageId="33" pageNumber="244">
|
||
<keyLead id="1C2DAC63FF99BE1CFF9DFD0AFE8EF60E" pageId="33" pageNumber="244">
|
||
<keyStep id="1C2817F3FF99BE1CFF9DFD0AFE8EF60E" pageId="33" pageNumber="244">7 (6) Pronotal punctation sparser, punctures separated by approximately three times their diameters pronotal base with very fine weak visible and well separated pinned points; elytral striae not impressed, visible as row of punctures; elytral intervals flat; northern Africa.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF99BE1CFF9DFC8AFBBEF66E" blockId="33.[113,1089,220,1139]" pageId="33" pageNumber="244">
|
||
<keyLead id="1C2DAC63FF99BE1CFF9DFC8AFBBEF66E" pageId="33" pageNumber="244">
|
||
<keyStep id="1C2817F3FF99BE1CFF9DFC8AFBBEF66E" pageId="33" pageNumber="244">
|
||
8 (9) Base of frontal tubercle situated at level of anterior part of eyes; tubercle sharply keeled basally only in dorsal view (
|
||
<figureCitation id="3FE713D3FF99BE1CFDD3FCAAFD7EF64E" box="[575,640,828,851]" captionStart="Figs 8–11" captionStartId="9.[113,150,1520,1539]" captionTargetBox="[136,1072,224,1492]" captionTargetId="figure-14@9.[129,1073,219,1523]" captionTargetPageId="9" captionText="Figs 8–11. Female habitus in dorsal aspect. 8 – Bolbelasmus bocchus (Erichson, 1841) (Morocco: Gaada de Debdou plateau., E of Rchida, 13.5 mm, DKCP); 9 – B. brancoi Hillert & Král sp. nov. (allotype); 10 – B. gallicus (Mulsant, 1842) (Spain: Andalusia, Aljaraque vill. env., 13.6 mm, JSCP); 11 – B. howdeni Hillert & Král sp. nov. (allotype)." figureDoi="http://doi.org/10.5281/zenodo.5306077" httpUri="https://zenodo.org/record/5306077/files/figure.png" pageId="33" pageNumber="244">Fig. 8</figureCitation>
|
||
);
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFD7FFCAAFD1DF64E" box="[659,739,828,851]" name="Algeria" pageId="33" pageNumber="244">Algeria</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFCFBFCAAFC89F64E" box="[791,887,828,851]" name="Morocco" pageId="33" pageNumber="244">Morocco</collectingCountry>
|
||
(Fig. 59). ............ ......................................................................................
|
||
<taxonomicName id="60DC74D5FF99BE1CFCE1FCCAFBBEF66E" authority="(Erichson, 1841)" baseAuthorityName="Erichson" baseAuthorityYear="1841" box="[781,1088,859,883]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="bocchus">
|
||
<emphasis id="95A8D344FF99BE1CFCE1FCCAFC78F66E" bold="true" box="[781,902,859,883]" italics="true" pageId="33" pageNumber="244">B. bocchus</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FF99BE1CFC78FCCAFBC6F66E" author="Erichson" box="[916,1080,860,883]" firstAuthor="Erichson" pageId="33" pageNumber="244" refId="ref26768" refString="ERICHSON W. F. 1841: Ueber die Insecten von Algier mit besonderer Berucksichtigung ihrer Verbreitung. Pp. 140 - 194. In: WAGNER M. F.: Reisen in der Regentschaft Algier in den Jahren 1836, 1837 und 1838 nebst einem naturhistorischen Anhang und einem Kupferatlas. Dritter Band. L. Voss, Leipzig, xviii + 297 pp. + 17 pls." type="book" year="1841">Erichson, 1841</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF99BE1CFF9DFCEAFBBEF6CE" blockId="33.[113,1089,220,1139]" pageId="33" pageNumber="244">
|
||
<keyLead id="1C2DAC63FF99BE1CFF9DFCEAFBBEF6CE" pageId="33" pageNumber="244">
|
||
<keyStep id="1C2817F3FF99BE1CFF9DFCEAFBBEF6CE" pageId="33" pageNumber="244">
|
||
9 (8) Base of frontal tubercle situated at level of posterior part of eyes; horn broadly keeled from base to apex in dorsal view (
|
||
<figureCitation id="3FE713D3FF99BE1CFDC3FC0AFD80F6AE" box="[559,638,924,947]" captionStart="Figs 12–13" captionStartId="10.[151,188,914,933]" captionTargetBox="[177,1107,239,881]" captionTargetId="figure-10@10.[167,1111,232,885]" captionTargetPageId="10" captionText="Figs 12–13. Female habitus in dorsal aspect. 12 – B. nikolajevi Hillert, Arnone, Král & Massa sp. nov. (allotype); 13 – B. vaulogeri (Abeille de Perrin, 1898) (Tunisia: Cap Bon peninsula, 6 km W of El Hawariah, 12.0 mm, DKCP)." figureDoi="http://doi.org/10.5281/zenodo.5306079" httpUri="https://zenodo.org/record/5306079/files/figure.png" pageId="33" pageNumber="244">Fig. 12</figureCitation>
|
||
);
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFD7FFC0AFD2BF6AE" box="[659,725,924,947]" name="Egypt" pageId="33" pageNumber="244">Egypt</collectingCountry>
|
||
,
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFD0CFC0AFCE0F6AE" box="[736,798,924,947]" name="Libya" pageId="33" pageNumber="244">Libya</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFCBDFC0AFC5EF6AE" box="[849,928,924,947]" name="Tunisia" pageId="33" pageNumber="244">Tunisia</collectingCountry>
|
||
(Fig. 59). ...... ...............................................
|
||
<taxonomicName id="60DC74D5FF99BE1CFDE2FC2AFC1AF6CE" authority="Hillert, Arnone, Kral & Massa" authorityName="Hillert, Arnone, Kral & Massa" authorityYear="2016" box="[526,996,955,979]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="nikolajevi" status="sp. nov.">
|
||
<emphasis id="95A8D344FF99BE1CFDE2FC2AFD67F6CE" bold="true" box="[526,665,955,979]" italics="true" pageId="33" pageNumber="244">B. nikolajevi</emphasis>
|
||
Hillert, Arnone, Král & Massa
|
||
</taxonomicName>
|
||
<emphasis id="95A8D344FF99BE1CFC07FC2AFBBEF6CE" bold="true" box="[1003,1088,956,979]" pageId="33" pageNumber="244">
|
||
<taxonomicNameLabel id="8E9B6E3FFF99BE1CFC07FC2AFBBEF6CE" box="[1003,1088,956,979]" pageId="33" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF99BE1CFF9DFC4AFDD4F6EE" blockId="33.[113,1089,220,1139]" box="[113,554,988,1011]" pageId="33" pageNumber="244">
|
||
<keyLead id="1C2DAC63FF99BE1CFF9DFC4AFDD4F6EE" box="[113,554,988,1011]" pageId="33" pageNumber="244">
|
||
<keyStep id="1C2817F3FF99BE1CFF9DFC4AFDD4F6EE" box="[113,554,988,1011]" pageId="33" pageNumber="244">10 (1) Scutellar shield densely punctate.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF99BE1CFF9DFC6AFBBFF12E" blockId="33.[113,1089,220,1139]" pageId="33" pageNumber="244">
|
||
<keyLead id="1C2DAC63FF99BE1CFF9DFC6AFBBFF12E" pageId="33" pageNumber="244">
|
||
<keyStep id="1C2817F3FF99BE1CFF9DFC6AFBBFF12E" pageId="33" pageNumber="244">
|
||
11 (12)
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFF28FC6AFEF3F10E" box="[196,269,1020,1043]" name="France" pageId="33" pageNumber="244">France</collectingCountry>
|
||
(including
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFE60FC6AFE30F10E" box="[396,462,1020,1043]" name="France" pageId="33" pageNumber="244">Corse</collectingCountry>
|
||
),
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFE0EFC6AFDC2F10E" box="[482,572,1020,1043]" name="Portugal" pageId="33" pageNumber="244">Portugal</collectingCountry>
|
||
and
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFD99FC6AFD4CF10E" box="[629,690,1020,1043]" name="Spain" pageId="33" pageNumber="244">Spain</collectingCountry>
|
||
(see distribution map on Fig. 59 for details). .........................................................................
|
||
<taxonomicName id="60DC74D5FF99BE1CFCF5FB8AFBBFF12E" authority="(Mulsant, 1842)" baseAuthorityName="Mulsant" baseAuthorityYear="1842" box="[793,1089,1051,1075]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="gallicus">
|
||
<emphasis id="95A8D344FF99BE1CFCF5FB8AFC70F12E" bold="true" box="[793,910,1051,1075]" italics="true" pageId="33" pageNumber="244">B. gallicus</emphasis>
|
||
(
|
||
<bibRefCitation id="C34D72A7FF99BE1CFC70FB8AFBC7F12E" author="Mulsant" box="[924,1081,1052,1075]" firstAuthor="Mulsant" pageId="33" pageNumber="244" refId="ref27806" refString="MULSANT E. 1842: Histoire naturelle des Coleopteres de France. Lamellicornes. Maison Libraire, Paris & Imprimerie de Dumoulin, Ronet et Sibuet, Lyon, viii + 626 pp. + 3 pls." type="book" year="1842">Mulsant, 1842</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A7630F56FF99BE1CFF9DFBAAFBBEF16E" blockId="33.[113,1089,220,1139]" pageId="33" pageNumber="244">
|
||
<keyLead id="1C2DAC63FF99BE1CFF9DFBAAFBBEF16E" pageId="33" pageNumber="244">
|
||
<keyStep id="1C2817F3FF99BE1CFF9DFBAAFBBEF16E" pageId="33" pageNumber="244">
|
||
12 (11)
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFF28FBAAFEFDF14E" box="[196,259,1084,1107]" name="Spain" pageId="33" pageNumber="244">Spain</collectingCountry>
|
||
,
|
||
<collectingCountry id="DFCB4FC6FF99BE1CFEE3FBAAFE91F14E" box="[271,367,1084,1107]" name="Gibraltar" pageId="33" pageNumber="244">Gibraltar</collectingCountry>
|
||
(see distribution map on Fig. 59, for details). .................................. ............................................................................
|
||
<taxonomicName id="60DC74D5FF99BE1CFD26FBCAFC1AF16E" authority="Hillert & Kral" authorityName="Hillert & Kral" authorityYear="2016" box="[714,996,1115,1139]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
|
||
<emphasis id="95A8D344FF99BE1CFD26FBCAFCB8F16E" bold="true" box="[714,838,1115,1139]" italics="true" pageId="33" pageNumber="244">B. howdeni</emphasis>
|
||
Hillert & Král
|
||
</taxonomicName>
|
||
<emphasis id="95A8D344FF99BE1CFC07FBCAFBBEF16E" bold="true" box="[1003,1088,1116,1139]" pageId="33" pageNumber="244">
|
||
<taxonomicNameLabel id="8E9B6E3FFF99BE1CFC07FBCAFBBEF16E" box="[1003,1088,1116,1139]" pageId="33" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |