treatments-xml/data/2F/67/58/2F6758F0659B5A2FB0184BB2FD61DCAE.xml

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<mods:title id="D561504AE99E8295248C67E561BA1BEA">Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)</mods:title>
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<mods:namePart id="55FAA6A640B3743E9A4695E6F34CE6E5">Will, Kipling</mods:namePart>
<mods:nameIdentifier id="1E6EB878B7600ECDCCF87C0B92DD9399" type="ORCID">https://orcid.org/0000-0002-7056-9011</mods:nameIdentifier>
<mods:affiliation id="90A7C68EE58D10EB8DDE820E4C8AE5C3">University of California Berkeley, Essig Museum of Entomology. Berkeley, California, USA</mods:affiliation>
<mods:nameIdentifier id="9A469DE174745E107E403CC8E74B2D9A" type="email">kipwill@berkeley.edu</mods:nameIdentifier>
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<mods:namePart id="B235485ABCECAFF660965E4D2BED0C43">Gueorguiev, Borislav</mods:namePart>
<mods:affiliation id="000747D0120155FE24134F345E2DB796">National Museum of Natural History, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
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<mods:date id="7CA1B3CA464076034A711E43325F28A2">2021</mods:date>
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<mods:number id="1F5242718AF7E4F23E9B6E10ECDC7DB0">1044</mods:number>
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<mods:url id="3D60445BA2FFED886212E5DD9479F1B4">http://dx.doi.org/10.3897/zookeys.1044.63775</mods:url>
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<mods:classification id="4F705F1119A395EA7B251285B3F371B6">journal article</mods:classification>
<mods:identifier id="759C5FB37526DB6BAB91F2BA4A19EC1C" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.63775</mods:identifier>
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<treatment id="2F6758F0659B5A2FB0184BB2FD61DCAE" ID-GBIF-Taxon="182754031" LSID="urn:lsid:plazi:treatment:2F6758F0659B5A2FB0184BB2FD61DCAE" httpUri="http://treatment.plazi.org/id/2F6758F0659B5A2FB0184BB2FD61DCAE" lastPageNumber="375" pageId="0" pageNumber="375" scope_family="Carabidae" scope_order="Coleoptera" scope_subTribe="Thryptocerina" scope_tribe="Oodini">
<subSubSection id="B53641AFFB6EA25A3BF4F81FF9F78BD3" pageId="0" pageNumber="375" type="nomenclature">
<paragraph id="90DC99FAA80C4D79F74FA2072DC19A47" pageId="0" pageNumber="375">
<taxonomicName id="A2600B4454E581850A7BE63EE9827AF0" ID-CoL="8MNHL" LSID="2F6758F0-659B-5A2F-B018-4BB2FD61DCAE" authority="(Fauvel, 1882)" authorityName="Will &amp; Guéorguiev" authorityYear="2021" baseAuthorityName="Fauvel" baseAuthorityYear="1882" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus cyllodinus" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="cyllodinus" status="comb. nov.">Coptocarpus cyllodinus (Fauvel, 1882)</taxonomicName>
<taxonomicNameLabel id="43C723C830B61B7DC449961B522D77E3" pageId="0" pageNumber="375">comb. nov.</taxonomicNameLabel>
<figureCitation id="6D15053DD73E6C81CBB820A15A37F2D6" captionStart="Figures 9–12" captionStartId="F3" captionText="Figures 9 - 12. Female reproductive tract of 9 Coptocarpus grossus Erwin (dorsal view, Margaret River, Western Australia, Australia) 10 Coptocarpus sp. group 6 (ventral view, Mt. Lewis, Queensland, Australia) 11 C. erwini sp. nov. (ventral view, Mt. Humboldt, New Caledonia) 12 C. cyllodinus (Fauvel) (ventral view, Riviere Bleue, New Caledonia). Abbreviations: at, atrium; bc, bursa copulatrix; co, common oviduct; rc, receptaculum; sg, spermatheca gland; smc, seminal canal; spc, spermathecal canal. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1044.63775.figures9-12" httpUri="https://binary.pensoft.net/fig/554050" pageId="0" pageNumber="375">Figs 12</figureCitation>
<figureCitation id="1CC4FDF3A660F6BD231F6B20798D3213" captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Dorsal habitus of 18 C. erwini sp. nov. 19 holotype of Coptocarpus cyllodinus, with labels 20 C. amieuensis sp. nov. 21 C. lescheni sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.63775.figures18-21" httpUri="https://binary.pensoft.net/fig/554053" pageId="0" pageNumber="375">, 19</figureCitation>
<figureCitation id="1B1DF88708FA2C80C05C4FF9A75F1241" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">, 24</figureCitation>
<figureCitation id="F5088FFEEF313C0B5579773E483CE140" captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">, 34</figureCitation>
<figureCitation id="0A57FBB3B367901CC7CB90EE6F302C28" captionStart="Figures 46, 47" captionStartId="F12" captionText="Figures 46, 47. Humeral region of left elytron of 46 Coptocarpus erwini sp. nov. 47 C. cyllodinus (Fauvel). Arrow indicates the submarginal carina." figureDoi="10.3897/zookeys.1044.63775.figures46-47" httpUri="https://binary.pensoft.net/fig/554059" pageId="0" pageNumber="375">, 47</figureCitation>
<figureCitation id="C80A92993AB88DFA04CC884DF9359690" captionStart="Figure 48" captionStartId="F13" captionText="Figure 48. Left hind leg, anterior view of Coptocarpus cyllodinus (Fauvel) male." figureDoi="10.3897/zookeys.1044.63775.figure48" httpUri="https://binary.pensoft.net/fig/554060" pageId="0" pageNumber="375">, 48</figureCitation>
<figureCitation id="34B0E0A2FBCBF85D78DB3EC7C8FFEF2E" captionStart="Figures 49–51" captionStartId="F14" captionText="Figures 49 - 51. Head in left lateral view of 49 Coptocarpus microps sp. nov. 50 C. cyllodinus (Fauvel) 51 C. magnus sp. nov. Arrow indicates location of notch beneath the eye." figureDoi="10.3897/zookeys.1044.63775.figures49-51" httpUri="https://binary.pensoft.net/fig/554061" pageId="0" pageNumber="375">, 50</figureCitation>
<figureCitation id="F2F90A745EFC21EC0E36F22575436D04" captionStart="Figures 52, 53" captionStartId="F15" captionText="Figures 52, 53. Maps of localities of New Caledonian Oodini." figureDoi="10.3897/zookeys.1044.63775.figures52-53" httpUri="https://binary.pensoft.net/fig/554062" pageId="0" pageNumber="375">, 52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6521E4B17B44052C44DC6EFD0FFC5FC0" pageId="0" pageNumber="375" type="reference_group">
<paragraph id="A9E5B5878B626B12A63795A84954A4FF" pageId="0" pageNumber="375">
<taxonomicName id="C2041045972384A691A7852C1527E390" authorityName="Will &amp; Guéorguiev" authorityYear="2021" baseAuthorityName="Will &amp; Guéorguiev" baseAuthorityYear="2021" class="Insecta" family="Carabidae" genus="Hoplolenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplolenus cyllodinus" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="cyllodinus" status="comb. nov.">Hoplolenus cyllodinus</taxonomicName>
Fauvel, 1882: 266.
</paragraph>
</subSubSection>
<subSubSection id="84E5631394105B4962A8C110539F5469" pageId="0" pageNumber="375" type="type locality">
<paragraph id="9CB4798A1213FB5ED6B0BE439A2BE1D9" pageId="0" pageNumber="375">Type locality.</paragraph>
<paragraph id="74FF3B054F8A69F731229031BCCC85E9" pageId="0" pageNumber="375">
<normalizedToken id="14457557669413D8423A461EB7F96418" originalValue="Nouméa">Noumea</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="876BD9FF0B04FC546D3672993536FF41" pageId="0" pageNumber="375" type="materials_examined">
<paragraph id="42F18B7F2926F5B6F4FCA609842D6580" pageId="0" pageNumber="375">Material examined.</paragraph>
<paragraph id="501AD3F6563C2E3F6BECCDB033E56FAB" pageId="0" pageNumber="375">
<materialsCitation id="D32F847EA2CD95175DED8DE11380378C" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Coll. R. I. Sc. N. B. Nouvelle Caledonie, Fauvel, R. I. Sc." country="New Caledonia" determinerName="A. Fauvel Hoplolenus" location="New Caledonia" specimenCount="♂" typeStatus="Holotype">
<emphasis id="D61FB51BC98BE6E69DD4FEB942B5A030" bold="true" italics="true" pageId="0" pageNumber="375">
<typeStatus id="1AE7F5DFDEE95C0858064B33533816FB">Holotype</typeStatus>
</emphasis>
: 
<collectingCountry id="C299E79D72C46EFBC6E9203589E4C9F8" name="New Caledonia">New Caledonia</collectingCountry>
• 
<specimenCount id="5E941E53150B93E448B9B232A75C8532">♂</specimenCount>
, card-mounted specimen, original label with 
<normalizedToken id="351018043ABC98882058D887DDA85DF4" originalValue="“Nouméa”">&quot;Noumea&quot;</normalizedToken>
glued to pink label &quot; 
<collectorName id="06B318DFADDAB0FFA532330D598909FC">
Coll. R.I.Sc.N.B. Nouvelle 
<normalizedToken id="F4C0F5B713190A07A228FCC23F439690" originalValue="Calédonie">Caledonie</normalizedToken>
</collectorName>
rec Deplanche ex coll. 
<collectorName id="33A032DF31F6649C5FDB5B5BEFF18554">Fauvel</collectorName>
&quot;// 
<normalizedToken id="E760CEF0DE270E877F68FD4641C0D011" originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
// &quot;Coll.et det. 
<determinerName id="903D77DA8F26DAE1DB0322099E62FF18">
A. Fauvel 
<emphasis id="A24188D4018F75A7B2C6072F5E6639A8" italics="true" pageId="0" pageNumber="375">Hoplolenus cyllodinus</emphasis>
</determinerName>
n. sp. 
<collectorName id="7F6EF287E1EE387B66820CB52DE6BBAE">R.I.Sc.</collectorName>
N.B. 17.479&quot; [RBINS].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A4893393B1E0945E8AB78E580CE49427" pageId="0" pageNumber="375" type="materials_examined">
<paragraph id="B807511A1AFDB386CD5DB0BB204C453C" pageId="0" pageNumber="375">Other material.</paragraph>
<paragraph id="68791EA041B950BAB4998CBF0446C151" pageId="0" pageNumber="375">
<materialsCitation id="8A42D75BD14423860F2ED4589BBF03BA" collectingDate="2006-05-06" collectorName="G. B. Monteith" country="New Caledonia" elevation="500" latitude="-22.183332" location="Mt. Koghis" longLatPrecision="1261" longitude="166.51666" specimenCount="♂">
<collectingCountry id="F76E7F93EC3A8ABACB76667A7F5D40AF" name="New Caledonia">New Caledonia</collectingCountry>
• 
<specimenCount id="95D04CB7DA44842BB722513C482D6B2E">♂</specimenCount>
; 
<location id="B4E9696E0AED5146668D540DD847BD6C" LSID="urn:lsid:plazi:treatment:2F6758F0659B5A2FB0184BB2FD61DCAE:B4E9696E0AED5146668D540DD847BD6C" country="New Caledonia" latitude="-22.183332" longLatPrecision="1261" longitude="166.51666" name="Mt. Koghis">Mt. Koghis</location>
; 
<geoCoordinate id="3B9A59D7CAF1C9BA5EACE63F2D696CD5" degrees="22" direction="south" minutes="11" orientation="latitude" precision="925" value="-22.183332">22°11'S</geoCoordinate>
, 
<geoCoordinate id="842D67FB9A2F2C7C60D716C16F3418D7" degrees="166" direction="east" minutes="31" orientation="longitude" precision="925" value="166.51666">166°31'E</geoCoordinate>
; track entrance; 
<quantity id="9396B731B2831820CD49B3E9CA63927C" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<elevation id="00E07CD698AF27FCCEC632AA21F657BA" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</elevation>
</quantity>
; 
<collectingDate id="ECB7E3D7B09D7AF2C1EE4624F70CC45A" value="2006-05-06">6.v.2006</collectingDate>
; berleseate; 
<collectorName id="5BAB56037F7DB136DBF86F0968D6D5C4">G.B.Monteith</collectorName>
; [QM]
</materialsCitation>
• 
<materialsCitation id="67DACC607E065C3CF584F8F86A3C1787" collectingDate="2000-11-29" collectorName="G. B. Monteith" country="New Caledonia" elevation="750" latitude="-22.183332" location="Mt. Koghis" longLatPrecision="1261" longitude="166.0" specimenCount="♀">
<specimenCount id="4741AD9DE9C784CD684D02BCFA5BFA26">♀</specimenCount>
; 
<location id="DD444E6071F1454BDBF20B387B1D0826" LSID="urn:lsid:plazi:treatment:2F6758F0659B5A2FB0184BB2FD61DCAE:DD444E6071F1454BDBF20B387B1D0826" country="New Caledonia" latitude="-22.183332" longLatPrecision="1261" longitude="166.0" name="Mt. Koghis">Mt. Koghis</location>
; 
<geoCoordinate id="8E86A6DB1948C20E5D78F9BC0473545C" degrees="22" direction="south" minutes="11" orientation="latitude" precision="925" value="-22.183332">22°11'S</geoCoordinate>
, 
<geoCoordinate id="8014CBD9C8FE0D97AD3D8F8FCBF095A6" degrees="166" direction="east" minutes="00" orientation="longitude" precision="925" value="166.0">166°00'E</geoCoordinate>
; 
<quantity id="ECC367E0F6F9AE3E004D42F749DABBB5" metricMagnitude="2" metricUnit="m" metricValue="7.5" unit="m" value="750.0">
<elevation id="0275913D831524A236C95BF47FDD2D12" metricMagnitude="2" metricUnit="m" metricValue="7.5" unit="m" value="750.0">750 m</elevation>
</quantity>
; 
<collectingDate id="6CC5B5F12A7691DE2F978BC90F37DEAA" value="2000-11-29">29.xi.2000</collectingDate>
; berleseate, sieved litter, rainforest; 
<collectorName id="CC763CC32EC09E6E910962579DFE9B79">G.B. Monteith</collectorName>
; [QM]
</materialsCitation>
• 
<specimenCount id="64905D0C8E949C76AF5165F2E855D885">♂</specimenCount>
; 
<geoCoordinate id="5D6DE17706649AE37430E75047C23D14" degrees="22" direction="south" minutes="10" orientation="latitude" precision="15" seconds="28" value="-22.174444">22°10'28&quot;S</geoCoordinate>
, 
<geoCoordinate id="A84C29690361813A235F938CCEDB64E7" degrees="166" direction="east" minutes="30" orientation="longitude" precision="15" seconds="48" value="166.51334">166°30'48&quot;E</geoCoordinate>
, 
<quantity id="A8245F386E8EEB8117E3F0D30B0BC8F5" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">700 m</quantity>
, 
<date id="74B4449007453E618A8169BE1457376D" value="2007-03-12">12.iii.2007</date>
; K. Will; EMEC1075689; [EMEC] • 
<materialsCitation id="061AD10AB70A624382223E10C70DA963" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="G. B. Monteith" country="New Caledonia" elevation="750" latitude="-22.183332" location="Mt. Koghis" longLatPrecision="1261" longitude="166.0" specimenCount="♂">
<specimenCount id="E4EBE713858E70A44CDD6787BAE65FF2">♂</specimenCount>
; same data as for preceding; headlamp search; EMEC1075690; [EMEC]
</materialsCitation>
• 
<materialsCitation id="1A8F373C11D675C40E1DE1D8B3B1F21C" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="G. B. Monteith" country="New Caledonia" elevation="750" latitude="-22.183332" location="Mt. Koghis" longLatPrecision="1261" longitude="166.0" specimenCount="♂">
<specimenCount id="0147B4472BDDE60506E9017CA97CBE24">♂</specimenCount>
; same data as for preceding; under rocks/logs; EMEC1075691; [EMEC]
</materialsCitation>
• 
<materialsCitation id="284CE73C76473529EB8D0AA6C4FB8F48" collectingDate="1977-02-12" collectingDateMax="1977-02-13" collectingDateMin="1977-02-12" collectorName="Dr. J." country="New Caledonia" location="Noumea Mt." specimenCount="♂">
<specimenCount id="EFB6604D7E13762286DF1FBAA997A9FB">♂</specimenCount>
; 
<location id="18588F26B15B10C8210A186191A137C7" LSID="urn:lsid:plazi:treatment:2F6758F0659B5A2FB0184BB2FD61DCAE:18588F26B15B10C8210A186191A137C7" country="New Caledonia" name="Noumea Mt.">
<normalizedToken id="B238731219C5B444A313EF6097D2DF82" originalValue="Nouméa">Noumea</normalizedToken>
Mt.
</location>
Kogni; 
<collectingDate id="C2F3F250F5CF30E3F3F25C00752A041D" value="1977-02-12" valueMax="1977-02-13" valueMin="1977-02-12">12-13.II.1977</collectingDate>
; leg. 
<collectorName id="1183BE30A4C13074AA957D4E4838D63C">Dr. J.</collectorName>
BALOGH; [NHMUK]
</materialsCitation>
• 
<materialsCitation id="2FB1028FCC6BD30AAD334DF6A8968A2F" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="P. D. Montague, B. M." country="New Caledonia" location="Mt. Kouakoue" specimenCount="♀">
<specimenCount id="19BEF15919CE4AAAE2BA809E88316EFB">♀</specimenCount>
; 
<normalizedToken id="6848B5C568E515E94B6787ACC9FB632F" originalValue="“Kuakui”">&quot;Kuakui&quot;</normalizedToken>
[
<location id="42A17D674BC2DFA1B5E0ABB7AB6ECE0D" LSID="urn:lsid:plazi:treatment:2F6758F0659B5A2FB0184BB2FD61DCAE:42A17D674BC2DFA1B5E0ABB7AB6ECE0D" country="New Caledonia" name="Mt. Kouakoue">
Mt. 
<normalizedToken id="73B9579EE050F9E30080DD09CBC8A496" originalValue="Kouakoué">Kouakoue</normalizedToken>
</location>
]; 
<collectorName id="35405624BE7CEAE3BA8CD5A0A1EAD58E">P.D. Montague</collectorName>
; 
<collectorName id="89DA17C1C4EAC66D964248EA42CC3578" name="G.B. Monteith">B. M.</collectorName>
1946-210; [NHMUK]
</materialsCitation>
• 
<materialsCitation id="D4161AE809D446E7A225244806A34949" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Fauvel" country="New Caledonia" location="Yahoue" specimenCount="♂" typeStatus="syntype">
<specimenCount id="74A9FA12A162B998451F626C353F92D1">♂</specimenCount>
; 
<normalizedToken id="0DDD4B7F6448B6EACD46A61913964D4E" originalValue="Yahoué">
<location id="A525417101B5B962161EC1EEDDAF692C" LSID="urn:lsid:plazi:treatment:2F6758F0659B5A2FB0184BB2FD61DCAE:A525417101B5B962161EC1EEDDAF692C" country="New Caledonia" name="Yahoue">Yahoue</location>
</normalizedToken>
; 
<normalizedToken id="C53ED3B30061E9C2A8F16CB7EF115988" originalValue="“février”">&quot;fevrier&quot;</normalizedToken>
; [RBINS, 
<collectorName id="4A8CEBC003FC2FF7614F3D3E2ED49F0F">Fauvel</collectorName>
collection, but not a 
<typeStatus id="E7BF496282DF1F1D4BF3F94D097E4A8B">syntype</typeStatus>
]
</materialsCitation>
• 
<materialsCitation id="B6229A5EB7ABAEAA8673F212507AF235" collectingDate="2007-03-13" collectorName="K. Will" country="New Caledonia" elevation="330" latitude="-22.15" location="Riviere Bleue Park" longLatPrecision="21" longitude="166.68666" municipality="Houp Geant" specimenCount="♂" typeStatus="syntype">
<specimenCount id="7D89F76A8B8C3F354B777FD2BADF5979">♂</specimenCount>
; 
<location id="888EAF9BD90D8D7720181B00D26E9090" LSID="urn:lsid:plazi:treatment:2F6758F0659B5A2FB0184BB2FD61DCAE:888EAF9BD90D8D7720181B00D26E9090" country="New Caledonia" latitude="-22.15" longLatPrecision="21" longitude="166.68666" municipality="Houp Geant" name="Riviere Bleue Park">
<normalizedToken id="0EA96003EA9DB65EC5227FE6C7182511" originalValue="Riviére">Riviere</normalizedToken>
Bleue Park
</location>
: 
<geoCoordinate id="53D5E001E3BB8C72A4B2A9DAC138C396" degrees="22" direction="south" minutes="09" orientation="latitude" precision="15" seconds="00" value="-22.15">22°09'00&quot;S</geoCoordinate>
, 
<geoCoordinate id="7B63452EF9CB1EB9DF3398F018CCF3FC" degrees="166" direction="east" minutes="41" orientation="longitude" precision="15" seconds="12" value="166.68666">166°41'12&quot;E</geoCoordinate>
, 
<quantity id="0E90AE0C6A0EC54A50484752E509A3DE" metricMagnitude="2" metricUnit="m" metricValue="3.3" unit="m" value="330.0">
<elevation id="9BE909237AD00044C0AAE0977046D17B" metricMagnitude="2" metricUnit="m" metricValue="3.3" unit="m" value="330.0">330 m</elevation>
</quantity>
, 
<collectingMunicipality id="C87A0F0833DFFBFBE7A9BEF622A67436">
Houp 
<normalizedToken id="3272DAF1B90C3E1335795C2FAC5E5AAD" originalValue="Géant">Geant</normalizedToken>
</collectingMunicipality>
trail, 
<collectingDate id="A03CF29C9A40E9DA6B34E8372791EB3D" value="2007-03-13">13.iii.2007</collectingDate>
, headlamp search, 
<collectorName id="BA5478E3BE07E5748EB194848B070D32">K. Will</collectorName>
; EMEC1075688; [EMEC]
</materialsCitation>
• 
<materialsCitation id="B849A9FE42B3FFED0A22594A76BDC3C8" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="K. Will" country="New Caledonia" elevation="330" latitude="-22.15" location="Riviere Bleue Park" longLatPrecision="21" longitude="166.68666" municipality="Houp Geant" specimenCount="♀" typeStatus="syntype">
<specimenCount id="76E879DD2E8580D64D5BCE6AF414840B">♀</specimenCount>
; same data as for preceding; [EMCE]
</materialsCitation>
• 
<materialsCitation id="F07F6A9025077167DFBFCB09819FAD27" collectingDate="2000-12-01" collectorName="G. B. Monteith" country="New Caledonia" elevation="700" latitude="-22.05" location="Mt. Dzumac" longLatPrecision="1261" longitude="166.46666" municipality="Mt. Dzumac Road" specimenCount="♂" typeStatus="syntype">
<specimenCount id="137CE35F690E66079FB123BE527E3B34">♂</specimenCount>
; 
<location id="03077A76A6BE10D65D7F43C385C3B4AD" LSID="urn:lsid:plazi:treatment:2F6758F0659B5A2FB0184BB2FD61DCAE:03077A76A6BE10D65D7F43C385C3B4AD" country="New Caledonia" latitude="-22.05" longLatPrecision="1261" longitude="166.46666" municipality="Mt. Dzumac Road" name="Mt. Dzumac">Mt. Dzumac</location>
; 
<geoCoordinate id="CE68F854AE21C927EC871E4805A6D5EC" degrees="22" direction="south" minutes="03" orientation="latitude" precision="925" value="-22.05">22°03'S</geoCoordinate>
, 
<geoCoordinate id="B20A459EF5B3AC2A0A640979DFF9C0EF" degrees="166" direction="east" minutes="28" orientation="longitude" precision="925" value="166.46666">166°28'E</geoCoordinate>
; 
<collectingMunicipality id="5382474E9A8AB46ADAB4C676DA8783F4">Mt. Dzumac Road</collectingMunicipality>
; 
<quantity id="721BA7738D041059042D4207C3925CE8" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">
<elevation id="AED70503FBF8B514460297A2FAAE2B38" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">700 m</elevation>
</quantity>
; 
<collectingDate id="34F6E51001E666BA0C3FD81D93902279" value="2000-12-01">1.xii.2000</collectingDate>
; berleseate, sieved litter, rainforest; 
<collectorName id="AB3EF3E562991B7383C93DEA7BFEE20E">G.B. Monteith</collectorName>
; [QM]
</materialsCitation>
• 
<materialsCitation id="A85434A96196E83745D00605B29A3E54" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="G. B. Monteith" country="New Caledonia" elevation="700" latitude="-22.05" location="Mt. Dzumac" longLatPrecision="1261" longitude="166.46666" municipality="Mt. Dzumac Road" specimenCount="♂" typeStatus="syntype">
<specimenCount id="ACD4587E80E45C4102F459B1DACD0530">♂</specimenCount>
; same data as for preceding; [ZSM]
</materialsCitation>
• 
<materialsCitation id="F3085E65F9CCBFE348D64BD0CB008D99" collectingDate="2004-11-02" collectorName="CJ Burwell" country="New Caledonia" latitude="-22.166666" location="Col de Yate" longLatPrecision="1261" longitude="166.91667" specimenCount="♀" typeStatus="syntype">
<specimenCount id="58FC4D921126254350F7146E61E287D6">♀</specimenCount>
; 
<location id="7D8F53F25EFDBE32C53274DCC4C8E087" LSID="urn:lsid:plazi:treatment:2F6758F0659B5A2FB0184BB2FD61DCAE:7D8F53F25EFDBE32C53274DCC4C8E087" country="New Caledonia" latitude="-22.166666" longLatPrecision="1261" longitude="166.91667" name="Col de Yate">Col de Yate</location>
; 
<geoCoordinate id="AF9B7CA55A66EC3B0D3248CC9F08F247" degrees="22" direction="south" minutes="10" orientation="latitude" precision="925" value="-22.166666">22°10'S</geoCoordinate>
, 
<geoCoordinate id="E522C98081FCEA95DB33EFC53012F664" degrees="166" direction="east" minutes="55" orientation="longitude" precision="925" value="166.91667">166°55'E</geoCoordinate>
; 
<collectingDate id="7F2A9E520F0BB0F2F223A473A8F922FC" value="2004-11-02">2.xi.2004</collectingDate>
; hand collecting, 
<collectorName id="7E4AFD747609B6212B428783EF27B0AE">CJ Burwell</collectorName>
; [QM]
</materialsCitation>
• 
<materialsCitation id="0F53B663FDE3774F4DB89C244F24D4B2" collectingDate="2004-11-25" collectingDateMax="2004-11-26" collectingDateMin="2004-11-25" country="New Caledonia" elevation="280" latitude="-22.233334" location="Pic du Pin" longLatPrecision="1260" longitude="166.83333" specimenCount="♀" typeStatus="syntype">
<specimenCount id="48D00C388351652EED1F11CEA688A684">♀</specimenCount>
; 
<location id="E57D263C84E685DF0ED86049AA377AC9" LSID="urn:lsid:plazi:treatment:2F6758F0659B5A2FB0184BB2FD61DCAE:E57D263C84E685DF0ED86049AA377AC9" country="New Caledonia" latitude="-22.233334" longLatPrecision="1260" longitude="166.83333" name="Pic du Pin">Pic du Pin</location>
; 
<geoCoordinate id="D36952A5CFC37CBD41DE85977CC8CD74" degrees="22" direction="south" minutes="14" orientation="latitude" precision="925" value="-22.233334">22°14'S</geoCoordinate>
, 
<geoCoordinate id="13738F1D3597111D493BEEBEACE631C3" degrees="166" direction="east" minutes="50" orientation="longitude" precision="925" value="166.83333">166°50'E</geoCoordinate>
; 
<quantity id="8D59CCAB8F4DB491B9224B63FE5AC294" metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">
<elevation id="27142EF15F9E23234984180FF5048447" metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">280 m</elevation>
</quantity>
; 
<collectingDate id="C791D7907D822CE775C9AB8D66CD9EA9" value="2004-11-25" valueMax="2004-11-26" valueMin="2004-11-25">25-26.xi.2004</collectingDate>
; day hand collecting, rainforest; QM party; [QM]
</materialsCitation>
.
</paragraph>
<caption id="1428AC2433E5DDDD6F34BB7B3AB290A4" doi="10.3897/zookeys.1044.63775.figure45" httpUri="https://binary.pensoft.net/fig/554058" pageId="0" pageNumber="375" start="Figure 45" startId="F11">
<paragraph id="44B0132C63475A3D2A5C0EAC3A7B95FD" pageId="0" pageNumber="375">
<emphasis id="369E56E93D344C67E42E62D10EDEDF29" bold="true" pageId="0" pageNumber="375">Figure 45.</emphasis>
A diagram of the variation in aedeagus tip form and spermatheca 
<typeStatus id="8A96C1BDA4F07BA6B928054B6D245004">types</typeStatus>
included in the 
<taxonomicName id="6259776BB471F30B875D41B68B13368F" authorityName="Heller" authorityYear="1916" class="Insecta" family="Carabidae" genus="Adelopomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adelopomorpha glabra" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="CA04B765F978516559506BD6DC421C12" italics="true" pageId="0" pageNumber="375">Adelopomorpha glabra</emphasis>
</taxonomicName>
species group. Outlines of the aedeagi are based on dissected males from the listed locality. Spermatheca and gland outlines are somewhat stylized, showing generalized 
<typeStatus id="0784CCBF94A4DCB05C12B9777468CE13">type</typeStatus>
as in characters 34-37.
</paragraph>
</caption>
<caption id="A892629FBE77BF23699650D27E5176AA" doi="10.3897/zookeys.1044.63775.figures46-47" httpUri="https://binary.pensoft.net/fig/554059" pageId="0" pageNumber="375" start="Figures 46, 47" startId="F12">
<paragraph id="32C88DAA17AFF8F943A77C3F8CBE4242" pageId="0" pageNumber="375">
<emphasis id="CBC22BEB67317654D19764C7CAEB0E94" bold="true" pageId="0" pageNumber="375">Figures 46, 47.</emphasis>
Humeral region of left elytron of 
<emphasis id="378502266A5AB6A01E40172CBEDD323E" bold="true" pageId="0" pageNumber="375">46</emphasis>
<taxonomicName id="718B7C116CC6384B65824A8014BBC6F6" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus erwini" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="erwini">
<emphasis id="85A16232BC78D5E224682892F71DE2A0" italics="true" pageId="0" pageNumber="375">Coptocarpus erwini</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="208D0E809B8C0A96DBB736F1D2F1A977" bold="true" pageId="0" pageNumber="375">47</emphasis>
<taxonomicName id="AEA40ACB346195821606FAE10BD42337" lsidName="C. cyllodinus" pageId="0" pageNumber="375" rank="species" species="cyllodinus">
<emphasis id="58FF65966F3FA954C5620A1905AFAF56" italics="true" pageId="0" pageNumber="375">C. cyllodinus</emphasis>
</taxonomicName>
(Fauvel). Arrow indicates the submarginal carina.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DAF1A02AEBCFF54A121B8C0FB81A9EB7" pageId="0" pageNumber="375" type="diagnosis">
<paragraph id="3C51D0B94F58D9034FC47FEAB3ED802F" pageId="0" pageNumber="375">Diagnosis.</paragraph>
<paragraph id="9B59D04C9F2F1FFFCEF66A7E8AD501F6" pageId="0" pageNumber="375">
A very distinctive, large species for the New Caledonian fauna with a deep notch below the eye that may receive the antenna (Fig. 
<figureCitation id="BBECE012540B23A8719E32172C1C0C9D" captionStart="Figures 49–51" captionStartId="F14" captionText="Figures 49 - 51. Head in left lateral view of 49 Coptocarpus microps sp. nov. 50 C. cyllodinus (Fauvel) 51 C. magnus sp. nov. Arrow indicates location of notch beneath the eye." figureDoi="10.3897/zookeys.1044.63775.figures49-51" httpUri="https://binary.pensoft.net/fig/554061" pageId="0" pageNumber="375">50</figureCitation>
); eye and orbit very prominent, almost shelf-like; male metafemur with large, ventral tooth (Fig. 
<figureCitation id="7B75C58C60FA3B9AB51DE7207C2867D5" captionStart="Figure 48" captionStartId="F13" captionText="Figure 48. Left hind leg, anterior view of Coptocarpus cyllodinus (Fauvel) male." figureDoi="10.3897/zookeys.1044.63775.figure48" httpUri="https://binary.pensoft.net/fig/554060" pageId="0" pageNumber="375">48</figureCitation>
), clypeal setae absent, elytral stria 6 and 7 impressed throughout, 5 only impressed in apical 1/2, 1-4 not evident or only shallowly impressed near the apex. Aedeagus with distinct, broad apical blade (Fig. 
<figureCitation id="B766DE48E0986F7D3D312F304C95C871" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">24</figureCitation>
).
</paragraph>
<caption id="233651F7A16CBB25DB706BD160C87FFC" doi="10.3897/zookeys.1044.63775.figure48" httpUri="https://binary.pensoft.net/fig/554060" pageId="0" pageNumber="375" start="Figure 48" startId="F13">
<paragraph id="E2A8C486D3AE1DCA441A5AFD2A4D643E" pageId="0" pageNumber="375">
<emphasis id="F76E96D93974D93E333E6A9485F3EB09" bold="true" pageId="0" pageNumber="375">Figure 48.</emphasis>
Left hind leg, anterior view of 
<taxonomicName id="485EE848EF68F9C5ECCFE7C86DA9ACBF" authorityName="Will &amp; Guéorguiev" authorityYear="2021" baseAuthorityName="Fauvel" baseAuthorityYear="1882" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus cyllodinus" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="cyllodinus">
<emphasis id="10D80D18B05BD7516F51F79F1E5D837A" italics="true" pageId="0" pageNumber="375">Coptocarpus cyllodinus</emphasis>
</taxonomicName>
(Fauvel) male.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0F04DAEFB766CFA84E54B178F18DF0E7" pageId="0" pageNumber="375" type="redescription">
<paragraph id="28B2340A3A3591DAAF25ED91CE2B2806" pageId="0" pageNumber="375">Redescription.</paragraph>
<paragraph id="4FF91A3A90F049DC5ED7239F9D9EFBC8" pageId="0" pageNumber="375">
<emphasis id="F4E83108A865541EEDA3FAD2D6141ADB" bold="true" pageId="0" pageNumber="375">
<emphasis id="E6B45B8FF4DC0EDE00BB13D42F197B7F" italics="true" pageId="0" pageNumber="375">Habitus</emphasis>
.
</emphasis>
Large sized for New Caledonia oodines (BL: 7.35-8.45 mm, BW: 3.60-4.40 mm), with elliptic, moderately convex body (Fig. 
<figureCitation id="E1EF77FD2764234A8529370670250595" captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Dorsal habitus of 18 C. erwini sp. nov. 19 holotype of Coptocarpus cyllodinus, with labels 20 C. amieuensis sp. nov. 21 C. lescheni sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.63775.figures18-21" httpUri="https://binary.pensoft.net/fig/554053" pageId="0" pageNumber="375">19</figureCitation>
). 
<emphasis id="6D8697D6090BB0A94B72C482C6291FC6" bold="true" pageId="0" pageNumber="375">
<emphasis id="41106A578F413F842C9CD6E656963A4E" italics="true" pageId="0" pageNumber="375">Color and luster</emphasis>
.
</emphasis>
Head dark reddish black to black; clypeus and mandibles reddish brown; palpi, antennae and tarsi yellow to yellow brown; pronotum, elytra and ventral surface of body black; legs (excl. tarsi) deep reddish brown to reddish black. Integument moderately shiny, mostly without spectral iridescence, excluding proepisterna and abdominal ventrites on sides where it is present. 
<emphasis id="7D221F96950D75ED8B75EF19628E793E" bold="true" pageId="0" pageNumber="375">
<emphasis id="94B9842DC4D2C152BF4CCC8931264F2B" italics="true" pageId="0" pageNumber="375">Microsculpture and punctation</emphasis>
.
</emphasis>
Dorsal surface of head (excluding mandibles) and pronotum, with isodiametric meshes well impressed and visible; elytra with isodiametric meshes weakly visible; prosternum with scarcely-visible isodiametric sculpticells, ventrites 4-6 medially with scarcely-visible slightly transverse meshes, rest of ventral surface with sculpticells mostly not evident. Dorsal surface lacking macropunctation; head, pronotum and elytra with micropunctation, smaller on head and most of pronotum, larger on basal 1/4 of pronotum and elytra; micropunctation regular on metacoxae, very scattered on last abdominal ventrites, absent on rest of ventral surface, abdominal ventrites 1-3 macropunctate on sides, 2-5 finely coarsely wrinkled on sides. 
<emphasis id="E4ED769926C48D85B7644AA223FF30AB" bold="true" pageId="0" pageNumber="375">
<emphasis id="9E80209F785B2BE5F0E4B965CD968813" italics="true" pageId="0" pageNumber="375">Chaetotaxy</emphasis>
.
</emphasis>
Labrum with six setae, each in its own socket, four lateral setae larger than medial two. Clypeal setae absent. Supraorbital setae absent. Elytron without discal setiferous punctures. Abdominal ventrites 1-5 without ambulatory setae. 
<emphasis id="D2F59053E94D40D5AB062C729D07B85B" bold="true" pageId="0" pageNumber="375">
<emphasis id="2D4965951AF868DB1FC0A8EAB7304D10" italics="true" pageId="0" pageNumber="375">Head</emphasis>
.
</emphasis>
Approximately 2 1/2 
<normalizedToken id="8AF883FA5BA80D45F3B36BEBC9F004E5" originalValue="×">x</normalizedToken>
as wide as pronotum (Suppl. material 2: Table S1). Eye size large, eye prominently protruded, EyW/HW: 1.38-1.48. Labrum anterior margin somewhat concave. Frontoclypeal suture evident, but slightly impressed. Antenna relatively short, with pubescence starting from segment 4. Last labial palpomere swollen, blunt at apex, longer and larger than penultimate one. Mentum tooth with paramedial border distinct. 
<emphasis id="1C804A2D472DCA2B6935EFECE16D6CC3" bold="true" pageId="0" pageNumber="375">
<emphasis id="FC89D3CD43CF726EA6A5E0ABF6504264" italics="true" pageId="0" pageNumber="375">Thorax</emphasis>
.
</emphasis>
Pronotum ~ 1 2/3 wider than long (PW/PL: 1.64-1.85); width at apex 2-2 2/3 
<normalizedToken id="E7A995EE1499663CC5D5F62646A31680" originalValue="×">x</normalizedToken>
less than at widest point (PW/PA: 1.95-2.38). Disc with midline fine, shallowly impressed, and ended well before apical and basal margins, without apical transverse impression; anterior angles weakly projected, rounded at tips, anterior margin concave, submarginal sulcus laterally present, lacking in medial 3/10. Prosternal process rhomboid, elongate, broadly pointed at apex, bordered at apex, not bordered at sides. Mesosternum anteriorly without medial tubercle. Metepisternum ~ 1 1/2 
<normalizedToken id="4E0A01C819189A0A15E4702A52FA8038" originalValue="×">x</normalizedToken>
wider than long, with lateral margin convex, coadunation with epipleuron long, located medially and posteriorly; suture between metepisternum and metepimeron barely distinct. 
<emphasis id="1849409FB0C1B406E5A51250391E5957" bold="true" pageId="0" pageNumber="375">
<emphasis id="35B7383CFDFE31204EDE532100EAD75D" italics="true" pageId="0" pageNumber="375">Elytra</emphasis>
.
</emphasis>
Broad, slightly longer than wide (EL/EW: 1.05-1.17). Basal margin forming small, sharp tooth at shoulder, medially nearing parascutellar puncture to which it is joined by impressed basal section of stria 2. Humeral submarginal carina present. Apical sinuation shallow, barely distinct. Elytral striae 6 and 7 entirely impressed, 5 impressed only at posterior 1/3, 3 and 4 only at apical 1/7 or 1/6, anterior continuations of striae 3-5, whole of striae 1 and 2 and parascutellar striole not impressed. Granulation in marginal furrow discontinuous, interrupted briefly along second elytral quarter. 
<emphasis id="BB625C171FF2E65B17952FAFA0981688" bold="true" pageId="0" pageNumber="375">
<emphasis id="64D8FBC7B85651D3C3BD81FC252060CF" italics="true" pageId="0" pageNumber="375">Legs</emphasis>
.
</emphasis>
Mesotibia evenly dilated apically. Male protarsomeres 1-3 expanded, 1 subelongate, nearly as long as wide, 2 and 3 wider than long; ventral adhesive setae present only at apical 1/3-1/4 surface of protarsomere 1 and entire surfaces of 2 and 3. Male basomesotarsus slightly flattened, with apical 2/3 provided with some short, stout, tooth-like setae ventrally; female basomesotarsus with finer setae ventrally. 
<emphasis id="E9AEB2BE3232C6CB56604DDE7FDE7606" bold="true" pageId="0" pageNumber="375">
<emphasis id="F1350E4DCFD0195A2884DD0AF6CF144B" italics="true" pageId="0" pageNumber="375">Female genitalia</emphasis>
.
</emphasis>
Gonocoxite 2 subtriangular. Spermatheca narrow and long, nearly straight in basal 1/2, coiled in distal 1/2, undifferentiated; spermathecal gland connected near basal 1/6. 
<emphasis id="2B0F9C0CF1B010C740F8229140EEF229" bold="true" pageId="0" pageNumber="375">
<emphasis id="6B35EA05EDD17A7B31A5EAB8892E7905" italics="true" pageId="0" pageNumber="375">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view long, considerably bent ventrally, tapering distally, slightly broadened at apex (Fig. 
<figureCitation id="362A1331A6E24AB4B13E417BAEB71596" captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">34</figureCitation>
), with relatively large basal bulb; lobe in dorsal view with apical lamella strongly bent to right, with right side quite concave and left side slightly and evenly convex (Fig. 
<figureCitation id="40566C33B32BF368627AC71C7D622375" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">24</figureCitation>
); basal bulb forming edge dorsally with evident crest; sclerotized portion of endophallus visible in repose with three sclerites.
</paragraph>
</subSubSection>
<subSubSection id="01D9C5524B42FCF87C405F8B459B5016" pageId="0" pageNumber="375" type="notes">
<paragraph id="A64D802B68E0F7A9504EC69B2F24D0A3" pageId="0" pageNumber="375">Notes.</paragraph>
<paragraph id="4E3B6897765C100100601838DE6CED9C" pageId="0" pageNumber="375">Fauvel stated that this species has only four setae on the anterior edge of the labrum. However, the type specimen has a slightly malformed labrum with three setae on the right and two on the left. The median seta on the right is minute.</paragraph>
</subSubSection>
</treatment>
</document>