223 lines
34 KiB
XML
223 lines
34 KiB
XML
<document id="4829EC6431EDE17F3165E72C5167975C" ID-CLB-Dataset="4951" ID-DOI="10.11646/zootaxa.4984.1.27" ID-GBIF-Dataset="e01cc40a-5281-4611-8775-13c9f91b57d7" ID-ISSN="1175-5326" ID-Zenodo-Dep="4928138" ID-ZooBank="7D854C7C-E117-4C20-90FA-5E83896609B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623432413404" checkinUser="plazi" docAuthor="Golovatch, Sergei I." docDate="2021" docId="2F5287828C6EFFACFF7186ACFEE1FBE1" docLanguage="en" docName="zootaxa.4984.1.27.pdf" docOrigin="Zootaxa 4984 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hedinomorpha jaegeri Golovatch 2021, spec. nov." docType="treatment" docVersion="8" lastPageNumber="376" masterDocId="D36BFFFA8C6BFFA4FFE68262FFE0F844" masterDocTitle="Review of the millipede genus Hedinomorpha Verhoeff, 1934, with descriptions of three new species collected in China by Professor Jochen Martens (Diplopoda Polydesmida, Paradoxosomatidae)" masterLastPageNumber="384" masterPageNumber="368" pageNumber="373" updateTime="1699010600760" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="80678A012B91DEC36C2058B848242E08">Review of the millipede genus Hedinomorpha Verhoeff, 1934, with descriptions of three new species collected in China by Professor Jochen Martens (Diplopoda Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:name id="808AA2DC8E2EDB2BFF82916D0F4EF90E" type="personal">
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<mods:namePart id="0CB4F7CCA003EF0F308F1F8D59909170">Golovatch, Sergei I.</mods:namePart>
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<mods:title id="68A17C9D95F93E588A554FF83072F299">Zootaxa</mods:title>
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<mods:part id="B469DD662304018AE6191EFBB20B7393">
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<mods:date id="FA22E5FB269472519A280832A5B40825">2021</mods:date>
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<mods:number id="B3CABF1E3028F3607CB03F499B72D16E">2021-06-10</mods:number>
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<mods:number id="042538F805CAD725C8BC43B5960081E0">4984</mods:number>
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<mods:identifier id="57ECF3850B535EE85492335F32155359" type="DOI">10.11646/zootaxa.4984.1.27</mods:identifier>
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<treatment id="2F5287828C6EFFACFF7186ACFEE1FBE1" ID-DOI="http://doi.org/10.5281/zenodo.4927894" ID-GBIF-Taxon="184413793" ID-Zenodo-Dep="4927894" LSID="urn:lsid:plazi:treatment:2F5287828C6EFFACFF7186ACFEE1FBE1" httpUri="http://treatment.plazi.org/id/2F5287828C6EFFACFF7186ACFEE1FBE1" lastPageId="8" lastPageNumber="376" pageId="5" pageNumber="373">
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<subSubSection id="EFE1651F8C6EFFA1FF7186ACFDC9FCAD" box="[151,553,1230,1257]" pageId="5" pageNumber="373" type="nomenclature">
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<paragraph id="A74436948C6EFFA1FF7186ACFDC9FCAD" blockId="5.[151,553,1230,1293]" box="[151,553,1230,1257]" pageId="5" pageNumber="373">
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<heading id="FC0C81F88C6EFFA1FF7186ACFDC9FCAD" bold="true" box="[151,553,1230,1257]" fontSize="11" level="1" pageId="5" pageNumber="373" reason="1">
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<emphasis id="958FEA868C6EFFA1FF7186ACFDC9FCAD" bold="true" box="[151,553,1230,1257]" pageId="5" pageNumber="373">
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<taxonomicName id="60FB4D178C6EFFA1FF7186ACFE4AFCAD" authority="Golovatch, 2021" authorityName="Golovatch" authorityYear="2021" box="[151,426,1230,1257]" class="Diplopoda" family="Paradoxosomatidae" genus="Hedinomorpha" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
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<emphasis id="958FEA868C6EFFA1FF7186ACFE4AFCAD" bold="true" box="[151,426,1230,1257]" italics="true" pageId="5" pageNumber="373">Hedinomorpha jaegeri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8EBC57FD8C6EFFA1FE5786ADFDC9FCAD" box="[433,553,1231,1257]" pageId="5" pageNumber="373" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EFE1651F8C6EFFA1FF718691FEF4FD49" box="[151,276,1267,1293]" pageId="5" pageNumber="373" type="description">
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<paragraph id="A74436948C6EFFA1FF718691FEF4FD49" blockId="5.[151,553,1230,1293]" box="[151,276,1267,1293]" pageId="5" pageNumber="373">
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<figureCitation id="3FC02A118C6EFFA1FF718691FEF4FD49" box="[151,276,1267,1293]" captionStart-0="FIGURES 16–19" captionStart-1="FIGURES 20–24" captionStart-2="FIGURES 25–28" captionStartId-0="6.[151,264,1943,1967]" captionStartId-1="7.[151,264,1620,1644]" captionStartId-2="8.[151,264,1703,1727]" captionTargetBox-0="[197,1391,181,1919]" captionTargetBox-1="[159,1423,189,1585]" captionTargetBox-2="[151,1436,957,1680]" captionTargetId-0="figure-16@6.[197,1391,181,1919]" captionTargetId-1="figure-26@7.[151,1436,181,1596]" captionTargetId-2="figure-416@8.[151,1436,957,1680]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 16–19. Hedinomorpha jaegeri spec. nov., holotype male. 16 Habitus, dorsal view. 17 Anterior half of body, ventral view. 18 Midbody segments, dorsal view. 19 Caudal part of body, dorsal view. Pictures by K. Makarov, taken not to scale." captionText-1="FIGURES 20–24. Hedinomorpha jaegeri spec. nov., holotype male. 20 Tip of epiproct, dorsal view. 21 Hypoproct, ventral view. 22 Sternal lobe between coxae 4, caudal view. 23–24. Right gonopod, submesal (23) and ventrolateral (24) views, respectively. Abbreviations: d—subapical tooth; fe—femorite; g—gutter; k—midway spine; l—apicolateral postfemoral lobe; lo—lobule; q—lamina lateralis; sg—seminal groove; sl—solenomere; sph—solenophore. Scale bar: 1.0 mm (20–22) and 0.5 mm (23–24)." captionText-2="FIGURES 25–28. Hedinomorpha jaegeri spec. nov., holotype male. Right gonopod, lateral (25), submesal (26), mesal (27) and ventral (28) views, respectively. Pictures by K. Makarov, taken not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.4928150" figureDoi-1="http://doi.org/10.5281/zenodo.4928152" figureDoi-2="http://doi.org/10.5281/zenodo.4928156" httpUri-0="https://zenodo.org/record/4928150/files/figure.png" httpUri-1="https://zenodo.org/record/4928152/files/figure.png" httpUri-2="https://zenodo.org/record/4928156/files/figure.png" pageId="5" pageNumber="373">Figs 16–28</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="EFE1651F8C6EFFA1FF718758FC4DFDD9" pageId="5" pageNumber="373" type="materials_examined">
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<paragraph id="A74436948C6EFFA1FF718758FB17FD3D" blockId="5.[151,1437,1338,2013]" pageId="5" pageNumber="373">
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<emphasis id="958FEA868C6EFFA1FF718758FE61FD10" bold="true" box="[151,385,1338,1364]" pageId="5" pageNumber="373">Material examined.</emphasis>
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<materialsCitation id="17933CC98C6EFFA1FE6E8759FB17FD3D" ID-GBIF-Occurrence="3334677303" collectingDate="1997-07-20" collectionCode="SMF" collectorName="P. Jager & J. Martens" country="China" elevation="1700" location="Houshenzi" municipality="Taibai Shan Mts" pageId="5" pageNumber="373" specimenCount="2" specimenCount-male="1" stateProvince="Shaanxi" typeStatus="holotype">
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<typeStatus id="784088368C6EFFA1FE6E8759FE10FD11" box="[392,496,1339,1365]" pageId="5" pageNumber="373" type="holotype">Holotype</typeStatus>
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male and
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<specimenCount id="B1FDFD1D8C6EFFA1FD838759FD51FD11" box="[613,689,1339,1365]" pageId="5" pageNumber="373" type="male">1 male</specimenCount>
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<typeStatus id="784088368C6EFFA1FD5E8759FCF8FD11" box="[696,792,1339,1365]" pageId="5" pageNumber="373" type="paratype">paratype</typeStatus>
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(
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<collectionCode id="C1EAAE518C6EFFA1FCCE8759FC85FD11" box="[808,869,1339,1365]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="373">SMF</collectionCode>
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),
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<collectingCountry id="DFEC76048C6EFFA1FC9F8759FC5FFD11" box="[889,959,1339,1365]" name="China" pageId="5" pageNumber="373">China</collectingCountry>
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,
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<collectingRegion id="653FF8768C6EFFA1FC2C8759FB89FD11" box="[970,1129,1339,1365]" country="China" name="Shaanxi" pageId="5" pageNumber="373">Shaanxi Prov.</collectingRegion>
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,
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<collectorName id="0A0E53428C6EFFA1FB928759FAAAFD11" box="[1140,1354,1339,1365]" pageId="5" pageNumber="373">
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<collectingMunicipality id="4720ACEE8C6EFFA1FB928759FACEFD11" box="[1140,1326,1339,1365]" pageId="5" pageNumber="373">Taibai Shan Mts</collectingMunicipality>
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, S
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</collectorName>
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flanks, above
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<location id="A224604F8C6EFFA1FF07873DFEBCFD3D" LSID="urn:lsid:plazi:treatment:2F5287828C6EFFACFF7186ACFEE1FBE1:A224604F8C6EFFA1FF07873DFEBCFD3D" box="[225,348,1375,1401]" country="China" municipality="Taibai Shan Mts" name="Houshenzi" pageId="5" pageNumber="373" stateProvince="Shaanxi">Houshenzi</location>
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,
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<elevation id="2CD6D1A78C6EFFA1FE81873DFE18FD3D" box="[359,504,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="373" unit="m" value="1700.0">
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<quantity id="60039B718C6EFFA1FE81873DFE5CFD3D" box="[359,444,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="373" unit="m" value="1700.0">1700 m</quantity>
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a.s.l.
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</elevation>
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, primary broadleaved forest,
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<date id="D34510548C6EFFA1FCA4873DFC2DFD3D" box="[834,973,1375,1401]" pageId="5" pageNumber="373" value="1997-07-20">
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||
<collectingDate id="C301E9BC8C6EFFA1FCA4873DFC2DFD3D" box="[834,973,1375,1401]" pageId="5" pageNumber="373" value="1997-07-20">20.VII.1997</collectingDate>
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</date>
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,
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<collectorName id="0A0E53428C6EFFA1FC3F873DFBCEFD3D" box="[985,1070,1375,1401]" pageId="5" pageNumber="373">P. Jäger</collectorName>
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&
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<collectorName id="0A0E53428C6EFFA1FBB4873DFB27FD3D" box="[1106,1223,1375,1401]" pageId="5" pageNumber="373">J. Martens</collectorName>
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leg.
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</materialsCitation>
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</paragraph>
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<paragraph id="A74436948C6EFFA1FF2187E1FC4DFDD9" blockId="5.[151,1437,1338,2013]" box="[199,941,1411,1437]" pageId="5" pageNumber="373">
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<emphasis id="958FEA868C6EFFA1FF2187E1FEF4FDD9" bold="true" box="[199,276,1411,1437]" pageId="5" pageNumber="373">Name.</emphasis>
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This species is dedicated to Peter Jäger, the main collector.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFE1651F8C6EFFA1FF2187C4FB7EFE16" pageId="5" pageNumber="373" type="diagnosis">
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<paragraph id="A74436948C6EFFA1FF2187C4FB7EFE16" blockId="5.[151,1437,1338,2013]" pageId="5" pageNumber="373">
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<emphasis id="958FEA868C6EFFA1FF2187C4FEDAFD84" bold="true" box="[199,314,1446,1472]" pageId="5" pageNumber="373">Diagnosis</emphasis>
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. Differs from other species of the genus by the following combination of characters: colouration brown to nearly pallid, devoid of a vivid colour pattern; paraterga poorly-developed, their pore-bearing and poreless calluses with two or three lateral incisions, respectively, mostly sharp caudally, delimited by a distinct sulcus only dorsally and a very narrow sulcus ventrally, both sulci being complete; pleurosternal carinae present only on segment 2, and the peculiar shapes of the solenophore and its outgrowths (see also Key below).
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</paragraph>
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</subSubSection>
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<subSubSection id="EFE1651F8C6EFFACFF218438FEE1FBE1" lastPageId="8" lastPageNumber="376" pageId="5" pageNumber="373" type="description">
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<paragraph id="A74436948C6EFFA1FF218438FB75FE31" blockId="5.[151,1437,1338,2013]" box="[199,1173,1626,1653]" pageId="5" pageNumber="373">
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<emphasis id="958FEA868C6EFFA1FF218438FE4CFE30" bold="true" box="[199,428,1626,1652]" pageId="5" pageNumber="373">Description (male).</emphasis>
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All characters as in
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<taxonomicName id="60FB4D178C6EFFA1FD768439FCA7FE30" authority="Golovatch, 2021" authorityName="Golovatch" authorityYear="2021" box="[656,839,1627,1652]" class="Diplopoda" family="Paradoxosomatidae" genus="Hedinomorpha" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="martensorum" status="sp. nov.">
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<emphasis id="958FEA868C6EFFA1FD768439FCA7FE30" box="[656,839,1627,1652]" italics="true" pageId="5" pageNumber="373">H. martensorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8EBC57FD8C6EFFA1FCA88439FC20FE31" box="[846,960,1627,1653]" pageId="5" pageNumber="373" rank="species">spec. nov.</taxonomicNameLabel>
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, except as follows.
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</paragraph>
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<paragraph id="A74436948C6EFFA1FF21841DFDA7FF41" blockId="5.[151,1437,1338,2013]" pageId="5" pageNumber="373">
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Length of holo- and
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<typeStatus id="784088368C6EFFA1FE48841DFDEEFEDD" box="[430,526,1663,1689]" pageId="5" pageNumber="373" type="paratype">paratype</typeStatus>
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ca 14 or
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<quantity id="60039B718C6EFFA1FD9E841DFD29FEDD" box="[632,713,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="373" unit="mm" value="10.0">10 mm</quantity>
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, width of midbody pro- and metazonae 0.95 and 1.1 or 0.85 and 1.0 mm, respectively. Colouration in alcohol mainly brown (
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<typeStatus id="784088368C6EFFA1FCB784C1FC56FEFA" box="[849,950,1699,1726]" pageId="5" pageNumber="373" type="holotype">holotype</typeStatus>
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) (
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<figureCitation id="3FC02A118C6EFFA1FC3684C1FBB2FEFA" box="[976,1106,1699,1726]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[197,1391,181,1919]" captionTargetId="figure-16@6.[197,1391,181,1919]" captionTargetPageId="6" captionText="FIGURES 16–19. Hedinomorpha jaegeri spec. nov., holotype male. 16 Habitus, dorsal view. 17 Anterior half of body, ventral view. 18 Midbody segments, dorsal view. 19 Caudal part of body, dorsal view. Pictures by K. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.4928150" httpUri="https://zenodo.org/record/4928150/files/figure.png" pageId="5" pageNumber="373">Figs 16–19</figureCitation>
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) or pallid, devoid of a vivid colour pattern; in
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<typeStatus id="784088368C6EFFA1FE8484A5FE27FEA5" box="[354,455,1735,1761]" pageId="5" pageNumber="373" type="holotype">holotype</typeStatus>
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, legs and tip of epiproct lighter, nearly pallid; antennomeres 6 and 7 slightly infuscate, light brown, but tip of antennae pallid.
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</paragraph>
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<paragraph id="A74436948C6EFFACFF21856DFAE4FA79" blockId="5.[151,1437,1338,2013]" lastBlockId="8.[151,1437,151,933]" lastPageId="8" lastPageNumber="376" pageId="5" pageNumber="373">
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Clypeolabral region rather densely setose. In width, collum <segment 3=4 <2 = 5–16 <head, body gently and gradually tapering on segments 17–20. Tegument generally shining and finely shagreened, most metaterga slightly tuberculate, in places rugulose to rugose above and below paraterga; surface below paraterga very delicately microgranulate; strictures between pro- and metazonae densely ribbed (
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<figureCitation id="3FC02A118C6EFFA1FC988519FBCCFFD1" box="[894,1068,1915,1941]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[197,1391,181,1919]" captionTargetId="figure-16@6.[197,1391,181,1919]" captionTargetPageId="6" captionText="FIGURES 16–19. Hedinomorpha jaegeri spec. nov., holotype male. 16 Habitus, dorsal view. 17 Anterior half of body, ventral view. 18 Midbody segments, dorsal view. 19 Caudal part of body, dorsal view. Pictures by K. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.4928150" httpUri="https://zenodo.org/record/4928150/files/figure.png" pageId="5" pageNumber="373">Figs 16, 18, 19</figureCitation>
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). Tergal setae often abraded, but remaining ones thin and moderately long, mostly ca 1/3 as long as metaterga; setation pattern traceable as 2+
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<quantity id="60039B718C6EFFA1FAB885FDFA69FFFD" box="[1374,1417,1951,1977]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="373" unit="in" value="2.0">2 in</quantity>
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a transverse pre-sulcus row, setae being borne on evident, but small bosses/tubercles; about 3+3 similar tuberculations arranged in two transverse rows, one each in pre-sulcus (with 2+2 setigerous) and post-sulcus halves of metaterga, but rear row always devoid of setae or their insertion points. Dorsal surface of collum and segment 2 smooth, that of metaterga 3–18 uneven, slightly tuberculate, with a mostly unusually deep, microgranulate and wide transverse sulcus, and an axial line. Paraterga poorly developed, largely set at upper 1/3 of metatergal height, clearly drawn only anteriad and somewhat projecting beyond fore tergal margin only on segment 2; caudal corner drawn into a small and sharp tooth, but never extended behind rear tergal margin thereafter (
|
||
<figureCitation id="3FC02A118C6CFFA3FBFD85BCFB7FFFBD" box="[1051,1183,2014,2041]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[197,1391,181,1919]" captionTargetId="figure-16@6.[197,1391,181,1919]" captionTargetPageId="6" captionText="FIGURES 16–19. Hedinomorpha jaegeri spec. nov., holotype male. 16 Habitus, dorsal view. 17 Anterior half of body, ventral view. 18 Midbody segments, dorsal view. 19 Caudal part of body, dorsal view. Pictures by K. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.4928150" httpUri="https://zenodo.org/record/4928150/files/figure.png" pageId="7" pageNumber="375">Figs 16–19</figureCitation>
|
||
). Lateral calluses subhorizontal, narrow, with two (pore-bearing segments) or three (poreless ones) small setigerous incisions at lateral margin, delimited by complete sulci both dorsally and ventrally, but ventral sulcus very narrow and indistinct. Ozopores lateral, almost invisible from above, each lying inside an ovoid pit at ca 1/4–1/5 off caudal margin. Transverse metatergal sulci exceptionally evident, deep and wide, microgranulate at bottom, slightly sinuate medially, reaching the bases of paraterga, present on segments 3–19, but less distinct on segments 3, 4 and 19. Axial line sulciform, present on both halves of metazonae. Pleurosternal carina a small squarish lappet only on segment 2, carinae totally missing thereafter (
|
||
<figureCitation id="3FC02A118C63FFACFE88830DFE21F9CD" box="[366,449,367,393]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[197,1391,181,1919]" captionTargetId="figure-16@6.[197,1391,181,1919]" captionTargetPageId="6" captionText="FIGURES 16–19. Hedinomorpha jaegeri spec. nov., holotype male. 16 Habitus, dorsal view. 17 Anterior half of body, ventral view. 18 Midbody segments, dorsal view. 19 Caudal part of body, dorsal view. Pictures by K. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.4928150" httpUri="https://zenodo.org/record/4928150/files/figure.png" pageId="8" pageNumber="376">Fig. 17</figureCitation>
|
||
). Epiproct (
|
||
<figureCitation id="3FC02A118C63FFACFDA5830DFD79F9CD" box="[579,665,367,393]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[197,1391,181,1919]" captionTargetId="figure-16@6.[197,1391,181,1919]" captionTargetPageId="6" captionText="FIGURES 16–19. Hedinomorpha jaegeri spec. nov., holotype male. 16 Habitus, dorsal view. 17 Anterior half of body, ventral view. 18 Midbody segments, dorsal view. 19 Caudal part of body, dorsal view. Pictures by K. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.4928150" httpUri="https://zenodo.org/record/4928150/files/figure.png" pageId="8" pageNumber="376">Figs 19</figureCitation>
|
||
,
|
||
<figureCitation id="3FC02A118C63FFACFD42830DFD21F9CD" box="[676,705,367,393]" captionStart="FIGURES 20–24" captionStartId="7.[151,264,1620,1644]" captionTargetBox="[159,1423,189,1585]" captionTargetId="figure-26@7.[151,1436,181,1596]" captionTargetPageId="7" captionText="FIGURES 20–24. Hedinomorpha jaegeri spec. nov., holotype male. 20 Tip of epiproct, dorsal view. 21 Hypoproct, ventral view. 22 Sternal lobe between coxae 4, caudal view. 23–24. Right gonopod, submesal (23) and ventrolateral (24) views, respectively. Abbreviations: d—subapical tooth; fe—femorite; g—gutter; k—midway spine; l—apicolateral postfemoral lobe; lo—lobule; q—lamina lateralis; sg—seminal groove; sl—solenomere; sph—solenophore. Scale bar: 1.0 mm (20–22) and 0.5 mm (23–24)." figureDoi="http://doi.org/10.5281/zenodo.4928152" httpUri="https://zenodo.org/record/4928152/files/figure.png" pageId="8" pageNumber="376">20</figureCitation>
|
||
) as usual, rather long, faintly concave to subtruncate at tip, lateral pre-apical papillae small. Hypoproct (
|
||
<figureCitation id="3FC02A118C63FFACFDAE83F1FD7BF9E9" box="[584,667,403,429]" captionStart="FIGURES 20–24" captionStartId="7.[151,264,1620,1644]" captionTargetBox="[159,1423,189,1585]" captionTargetId="figure-26@7.[151,1436,181,1596]" captionTargetPageId="7" captionText="FIGURES 20–24. Hedinomorpha jaegeri spec. nov., holotype male. 20 Tip of epiproct, dorsal view. 21 Hypoproct, ventral view. 22 Sternal lobe between coxae 4, caudal view. 23–24. Right gonopod, submesal (23) and ventrolateral (24) views, respectively. Abbreviations: d—subapical tooth; fe—femorite; g—gutter; k—midway spine; l—apicolateral postfemoral lobe; lo—lobule; q—lamina lateralis; sg—seminal groove; sl—solenomere; sph—solenophore. Scale bar: 1.0 mm (20–22) and 0.5 mm (23–24)." figureDoi="http://doi.org/10.5281/zenodo.4928152" httpUri="https://zenodo.org/record/4928152/files/figure.png" pageId="8" pageNumber="376">Fig. 21</figureCitation>
|
||
) roundly subtrapeziform, caudal margin with 1+1 long setae borne on flat rounded knobs. Sterna without modifications except for a lower, roundly subtrapeziform, slightly concave and setose lobe between coxae 4 (
|
||
<figureCitation id="3FC02A118C63FFACFDC583B9FD9CF9B1" box="[547,636,475,501]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[197,1391,181,1919]" captionTargetId="figure-16@6.[197,1391,181,1919]" captionTargetPageId="6" captionText="FIGURES 16–19. Hedinomorpha jaegeri spec. nov., holotype male. 16 Habitus, dorsal view. 17 Anterior half of body, ventral view. 18 Midbody segments, dorsal view. 19 Caudal part of body, dorsal view. Pictures by K. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.4928150" httpUri="https://zenodo.org/record/4928150/files/figure.png" pageId="8" pageNumber="376">Figs 17</figureCitation>
|
||
,
|
||
<figureCitation id="3FC02A118C63FFACFD6D83B9FD4BF9B1" box="[651,683,475,501]" captionStart="FIGURES 20–24" captionStartId="7.[151,264,1620,1644]" captionTargetBox="[159,1423,189,1585]" captionTargetId="figure-26@7.[151,1436,181,1596]" captionTargetPageId="7" captionText="FIGURES 20–24. Hedinomorpha jaegeri spec. nov., holotype male. 20 Tip of epiproct, dorsal view. 21 Hypoproct, ventral view. 22 Sternal lobe between coxae 4, caudal view. 23–24. Right gonopod, submesal (23) and ventrolateral (24) views, respectively. Abbreviations: d—subapical tooth; fe—femorite; g—gutter; k—midway spine; l—apicolateral postfemoral lobe; lo—lobule; q—lamina lateralis; sg—seminal groove; sl—solenomere; sph—solenophore. Scale bar: 1.0 mm (20–22) and 0.5 mm (23–24)." figureDoi="http://doi.org/10.5281/zenodo.4928152" httpUri="https://zenodo.org/record/4928152/files/figure.png" pageId="8" pageNumber="376">22</figureCitation>
|
||
). Legs moderately long and slender, obviously incrassate in male compared to a still unknown female, midbody ones ca 1.3–1.4x as long as body height, mostly less densely setose, ventral brushes on tarsi present only until leg-pair 9 or 10, gradually thinning out thereafter (
|
||
<figureCitation id="3FC02A118C63FFACFB428041FB17FA79" box="[1188,1271,547,573]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[197,1391,181,1919]" captionTargetId="figure-16@6.[197,1391,181,1919]" captionTargetPageId="6" captionText="FIGURES 16–19. Hedinomorpha jaegeri spec. nov., holotype male. 16 Habitus, dorsal view. 17 Anterior half of body, ventral view. 18 Midbody segments, dorsal view. 19 Caudal part of body, dorsal view. Pictures by K. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.4928150" httpUri="https://zenodo.org/record/4928150/files/figure.png" pageId="8" pageNumber="376">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="F384661C8C6DFFA2FF7185F5FA8CFF90" ID-DOI="http://doi.org/10.5281/zenodo.4928150" ID-Zenodo-Dep="4928150" httpUri="https://zenodo.org/record/4928150/files/figure.png" pageId="6" pageNumber="374" startId="6.[151,264,1943,1967]" targetBox="[197,1391,181,1919]" targetPageId="6">
|
||
<paragraph id="A74436948C6DFFA2FF7185F5FA8CFF90" blockId="6.[151,1436,1943,2004]" pageId="6" pageNumber="374">
|
||
<emphasis id="958FEA868C6DFFA2FF7185F5FEAEFFEB" bold="true" box="[151,334,1943,1967]" pageId="6" pageNumber="374">FIGURES 16–19</emphasis>
|
||
.
|
||
<taxonomicName id="60FB4D178C6DFFA2FEBC85FAFDA2FFEB" authority="Golovatch, 2021" authorityName="Golovatch" authorityYear="2021" box="[346,578,1944,1968]" class="Diplopoda" family="Paradoxosomatidae" genus="Hedinomorpha" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="374" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
|
||
<emphasis id="958FEA868C6DFFA2FEBC85FAFDA2FFEB" box="[346,578,1944,1968]" italics="true" pageId="6" pageNumber="374">Hedinomorpha jaegeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="958FEA868C6DFFA2FDAE85FAFD4FFFF4" bold="true" box="[584,687,1944,1968]" pageId="6" pageNumber="374">
|
||
<taxonomicNameLabel id="8EBC57FD8C6DFFA2FDAE85FAFD4FFFF4" box="[584,687,1944,1968]" pageId="6" pageNumber="374" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male. 16 Habitus, dorsal view. 17 Anterior half of body, ventral view. 18 Midbody segments, dorsal view. 19 Caudal part of body, dorsal view. Pictures by K. Makarov, taken not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F384661C8C6CFFA3FF718436FEFCFEB8" ID-DOI="http://doi.org/10.5281/zenodo.4928152" ID-Zenodo-Dep="4928152" httpUri="https://zenodo.org/record/4928152/files/figure.png" pageId="7" pageNumber="375" startId="7.[151,264,1620,1644]" targetBox="[159,1423,189,1585]" targetPageId="7">
|
||
<paragraph id="A74436948C6CFFA3FF718436FEFCFEB8" blockId="7.[151,1436,1620,1788]" pageId="7" pageNumber="375">
|
||
<emphasis id="958FEA868C6CFFA3FF718436FEAFFE28" bold="true" box="[151,335,1620,1644]" pageId="7" pageNumber="375">FIGURES 20–24</emphasis>
|
||
.
|
||
<taxonomicName id="60FB4D178C6CFFA3FEB88436FDA8FE28" authority="Golovatch, 2021" authorityName="Golovatch" authorityYear="2021" box="[350,584,1620,1644]" class="Diplopoda" family="Paradoxosomatidae" genus="Hedinomorpha" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="375" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
|
||
<emphasis id="958FEA868C6CFFA3FEB88436FDA8FE28" box="[350,584,1620,1644]" italics="true" pageId="7" pageNumber="375">Hedinomorpha jaegeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="958FEA868C6CFFA3FDB68436FD59FE28" bold="true" box="[592,697,1620,1644]" pageId="7" pageNumber="375">
|
||
<taxonomicNameLabel id="8EBC57FD8C6CFFA3FDB68436FD59FE28" box="[592,697,1620,1644]" pageId="7" pageNumber="375" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male. 20 Tip of epiproct, dorsal view. 21 Hypoproct, ventral view. 22 Sternal lobe between coxae 4, caudal view. 23–24. Right gonopod, submesal (23) and ventrolateral (24) views, respectively. Abbreviations: d—subapical tooth; fe—femorite; g—gutter; k—midway spine; l—apicolateral postfemoral lobe; lo—lobule; q—lamina lateralis; sg—seminal groove; sl—solenomere; sph—solenophore. Scale bar: 1.0 mm (20–22) and 0.5 mm (23–24).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A74436948C63FFACFF218025FEE1FBE1" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="376">
|
||
Gonopods (
|
||
<figureCitation id="3FC02A118C63FFACFEAF8025FE7FFA25" box="[329,415,583,609]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[197,1391,181,1919]" captionTargetId="figure-16@6.[197,1391,181,1919]" captionTargetPageId="6" captionText="FIGURES 16–19. Hedinomorpha jaegeri spec. nov., holotype male. 16 Habitus, dorsal view. 17 Anterior half of body, ventral view. 18 Midbody segments, dorsal view. 19 Caudal part of body, dorsal view. Pictures by K. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.4928150" httpUri="https://zenodo.org/record/4928150/files/figure.png" pageId="8" pageNumber="376">Figs 17</figureCitation>
|
||
,
|
||
<figureCitation id="3FC02A118C63FFACFE4C8025FE11FA25" box="[426,497,583,609]" captionStart-0="FIGURES 20–24" captionStart-1="FIGURES 25–28" captionStartId-0="7.[151,264,1620,1644]" captionStartId-1="8.[151,264,1703,1727]" captionTargetBox-0="[159,1423,189,1585]" captionTargetBox-1="[151,1436,957,1680]" captionTargetId-0="figure-26@7.[151,1436,181,1596]" captionTargetId-1="figure-416@8.[151,1436,957,1680]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 20–24. Hedinomorpha jaegeri spec. nov., holotype male. 20 Tip of epiproct, dorsal view. 21 Hypoproct, ventral view. 22 Sternal lobe between coxae 4, caudal view. 23–24. Right gonopod, submesal (23) and ventrolateral (24) views, respectively. Abbreviations: d—subapical tooth; fe—femorite; g—gutter; k—midway spine; l—apicolateral postfemoral lobe; lo—lobule; q—lamina lateralis; sg—seminal groove; sl—solenomere; sph—solenophore. Scale bar: 1.0 mm (20–22) and 0.5 mm (23–24)." captionText-1="FIGURES 25–28. Hedinomorpha jaegeri spec. nov., holotype male. Right gonopod, lateral (25), submesal (26), mesal (27) and ventral (28) views, respectively. Pictures by K. Makarov, taken not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.4928152" figureDoi-1="http://doi.org/10.5281/zenodo.4928156" httpUri-0="https://zenodo.org/record/4928152/files/figure.png" httpUri-1="https://zenodo.org/record/4928156/files/figure.png" pageId="8" pageNumber="376">23–28</figureCitation>
|
||
) largely suberect,
|
||
<emphasis id="958FEA868C63FFACFD588025FCE3FA24" box="[702,771,583,608]" italics="true" pageId="8" pageNumber="376">in situ</emphasis>
|
||
their distal parts coiled and in touch with each other; coxite subcylindrical, setose distoventrally, ca 2/3 as long as femorite (
|
||
<emphasis id="958FEA868C63FFACFCBD8008FC91FAC0" bold="true" box="[859,881,618,644]" pageId="8" pageNumber="376">fe</emphasis>
|
||
); prefemoral (= densely setose) part nearly half as long as
|
||
<emphasis id="958FEA868C63FFACFF0B80ECFEE3FAEC" bold="true" box="[237,259,654,680]" pageId="8" pageNumber="376">fe</emphasis>
|
||
; seminal groove (
|
||
<emphasis id="958FEA868C63FFACFE2B80EDFE06FAED" bold="true" box="[461,486,655,681]" pageId="8" pageNumber="376">sg</emphasis>
|
||
) running along
|
||
<emphasis id="958FEA868C63FFACFD7E80ECFD4EFAEC" bold="true" box="[664,686,654,680]" pageId="8" pageNumber="376">fe</emphasis>
|
||
at bottom of a shallow mesal groove/gutter (
|
||
<emphasis id="958FEA868C63FFACFB4480EDFB50FAED" bold="true" box="[1186,1200,655,681]" pageId="8" pageNumber="376">g</emphasis>
|
||
) before moving onto a distinct flagelliform solenomere (
|
||
<emphasis id="958FEA868C63FFACFDF980D1FDD2FA89" bold="true" box="[543,562,691,717]" pageId="8" pageNumber="376">sl</emphasis>
|
||
) squeezed between a particularly strongly developed lamina lateralis (
|
||
<emphasis id="958FEA868C63FFACFAA480D1FAB2FA89" bold="true" box="[1346,1362,691,717]" pageId="8" pageNumber="376">q</emphasis>
|
||
) and a much smaller lamina medialis, and supported by a more complex postfemoral part; the latter forming a subtriangular, apical, postfemoral, lateral lobe (
|
||
<emphasis id="958FEA868C63FFACFDD38099FDDDFB51" bold="true" box="[565,573,763,789]" pageId="8" pageNumber="376">l</emphasis>
|
||
) demarcated at base by a transverse sulcus, and being largely represented by a long, ribbon-shaped, loosely coiled, distally barbed and apically subacuminate solenophore branch (
|
||
<emphasis id="958FEA868C63FFACFAE2817DFACEFB7D" bold="true" box="[1284,1326,799,825]" pageId="8" pageNumber="376">sph</emphasis>
|
||
) supplied with a distinct spine (
|
||
<emphasis id="958FEA868C63FFACFE628121FE74FB19" bold="true" box="[388,404,835,861]" pageId="8" pageNumber="376">k</emphasis>
|
||
) near midway and a small rounded lobule (
|
||
<emphasis id="958FEA868C63FFACFC958121FC9BFB19" bold="true" box="[883,891,835,861]" pageId="8" pageNumber="376">l</emphasis>
|
||
) in distal 2/3. A large, ribbon-shaped, curved and apically acuminate process (
|
||
<emphasis id="958FEA868C63FFACFE308105FE06FBC5" bold="true" box="[470,486,871,897]" pageId="8" pageNumber="376">p</emphasis>
|
||
) set at base of
|
||
<emphasis id="958FEA868C63FFACFD698105FD77FBC5" bold="true" box="[655,663,871,897]" pageId="8" pageNumber="376">l</emphasis>
|
||
in addition to both main branches (
|
||
<emphasis id="958FEA868C63FFACFBC18105FBDAFBC5" bold="true" box="[1063,1082,871,897]" pageId="8" pageNumber="376">sl</emphasis>
|
||
and
|
||
<emphasis id="958FEA868C63FFACFB978105FB7BFBC5" bold="true" box="[1137,1179,871,897]" pageId="8" pageNumber="376">sph</emphasis>
|
||
) and a small subapical tooth (
|
||
<emphasis id="958FEA868C63FFACFF0781E9FF11FBE1" bold="true" box="[225,241,907,933]" pageId="8" pageNumber="376">d</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |