307 lines
41 KiB
XML
307 lines
41 KiB
XML
<document id="F419E0250963F675E610E47F91CD83E7" ID-DOI="10.15298/rusentj.32.1.03" ID-ISSN="0132-8069" ID-Zenodo-Dep="10949096" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712198792598" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2023" docId="2F4187C3FFCF166B7A5CFE4920F8740E" docLanguage="en" docName="RussEntomolJ.32.1.16-39.pdf" docOrigin="Russian Entomological Journal 32 (1)" docSource="http://dx.doi.org/10.15298/rusentj.32.1.03" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.1:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="1" docTitle="Pterostichus (Circinatus) tonkinensis Straneo 1980" docType="treatment" docVersion="3" lastPageNumber="22" masterDocId="D378FFBBFFCB166D7966FF9A2163720F" masterDocTitle="New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam" masterLastPageNumber="39" masterPageNumber="16" pageNumber="20" updateTime="1712665435938" updateUser="ExternalLinkService">
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<mods:title id="F55ADF1B7BD44743CB5A8A5A521BB85D">New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam</mods:title>
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<mods:namePart id="BCF62326136D03D735D400AE5BA5B206">Fedorenko, D. N.</mods:namePart>
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<mods:title id="159C7EDBB3DAC58B207DD19CC22FA680">Russian Entomological Journal</mods:title>
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<mods:part id="3F14872C50829D3EB3AA18359B5241FA">
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<mods:date id="9A35D2DE740CF8F26DC28CC721012A8B">2023</mods:date>
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<mods:number id="873EC6C959ED55BF9679D0FEA17FA61B">2023-03-31</mods:number>
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<mods:number id="33973EAEBEA69434B4284FAF3DA9E365">32</mods:number>
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<mods:url id="3C3D2BE5E4A59AAA68FF4383BA7ADA31">http://dx.doi.org/10.15298/rusentj.32.1.03</mods:url>
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<mods:identifier id="0DB05162C4BB6A11B29E8E9D1F4544EB" type="DOI">10.15298/rusentj.32.1.03</mods:identifier>
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<treatment id="2F4187C3FFCF166B7A5CFE4920F8740E" LSID="urn:lsid:plazi:treatment:2F4187C3FFCF166B7A5CFE4920F8740E" httpUri="http://treatment.plazi.org/id/2F4187C3FFCF166B7A5CFE4920F8740E" lastPageId="6" lastPageNumber="22" pageId="4" pageNumber="20">
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<subSubSection id="EFF2655EFFCF16697A5CFE49240B73E3" box="[826,1384,467,492]" pageId="4" pageNumber="20" type="nomenclature">
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<paragraph id="A75736D5FFCF16697A5CFE49240B73E3" blockId="4.[812,1399,467,2058]" box="[826,1384,467,492]" pageId="4" pageNumber="20">
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<heading id="FC1F81B9FFCF16697A5CFE49240B73E3" box="[826,1384,467,492]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="20" reason="8">
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<taxonomicName id="60E84D56FFCF16697A5CFE49240B73E3" ID-CoL="6WLZ2" authority="Straneo, 1980" authorityName="Straneo" authorityYear="1980" box="[826,1384,467,492]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="tonkinensis" subGenus="Circinatus">
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<emphasis id="959CEAC7FFCF16697A5CFE4922A373E4" box="[826,960,467,491]" italics="true" pageId="4" pageNumber="20">Pterostichus</emphasis>
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(
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<emphasis id="959CEAC7FFCF16697AB7FE4E252173E4" box="[977,1090,468,491]" italics="true" pageId="4" pageNumber="20">Circinatus</emphasis>
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)
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<emphasis id="959CEAC7FFCF16697D34FE4925A973E4" box="[1106,1226,467,491]" italics="true" pageId="4" pageNumber="20">tonkinensis</emphasis>
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Straneo, 1980
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCF16697AF6FE6B24707006" box="[912,1299,497,521]" pageId="4" pageNumber="20" type="description">
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<paragraph id="A75736D5FFCF16697AF6FE6B24707006" blockId="4.[812,1399,467,2058]" box="[912,1299,497,521]" pageId="4" pageNumber="20">
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<figureCitation id="3FD32A50FFCF16697AF6FE6B22937006" box="[912,1008,497,521]" captionStart="Figs 6–11" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[202,1388,618,1932]" captionTargetId="figure-261@3.[190,1398,609,1945]" captionTargetPageId="3" captionText="Figs 6–11. Pterostichus: 6, 9 — P. tonkinensis, ♂ from Phia Oac Mt; 7, 10 — P. austrinus sp.n., holotype; 8, 11 — P. laevibasis sp.n., paratype ♂; 6–8 — dorsal habitus; 9–11 — apices of elytra and abdomen, left aspect. Рис. 6–11. Pterostichus: 6, 9 — P. tonkinensis, ♂ с горы Phia Oac; 7, 10 — P. austrinus sp.n., голотип; 8, 11 — P. laevibasis sp.n., паратип ♂; 6–8 — габитус дорЗально; 9–11 — вершина надкрылий и брюшка слева." figureDoi="http://doi.org/10.5281/zenodo.10949100" httpUri="https://zenodo.org/record/10949100/files/figure.png" pageId="4" pageNumber="20">Figs 6, 9</figureCitation>
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,
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<figureCitation id="3FD32A50FFCF16697A9BFE6B25217006" box="[1021,1090,497,521]" captionStart="Figs 12–21" captionStartId="5.[226,263,1945,1964]" captionTargetBox="[189,1399,319,1919]" captionTargetId="figure-122@5.[189,1399,314,1923]" captionTargetPageId="5" captionText="Figs 12–21. Aedeagus median lobe: 12–13 — Pterostichus glabellus sp.n.; 14–15 — P. unisetosus sp.n.; 16–17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20–21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — left lateral aspect; 13, 15, 17, 19, 21 — right lateral aspect. Scale bars 1 mm. Рис. 12–21. СреднЯЯ долЯ Эдеагуса: 12–13 — Pterostichus glabellus sp.n.; 14–15 — P. unisetosus sp.n.; 16–17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20–21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — слева; 13, 15, 17, 19, 21 — справа. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949104" httpUri="https://zenodo.org/record/10949104/files/figure.png" pageId="4" pageNumber="20">16–17</figureCitation>
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,
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<figureCitation id="3FD32A50FFCF16697D28FE6B25F77006" box="[1102,1172,497,521]" captionStart="Figs 22–34" captionStartId="7.[226,263,1921,1940]" captionTargetBox="[190,1399,697,1904]" captionTargetId="figure-163@7.[189,1399,694,1904]" captionText="Figs 22–34. Aedeagus: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28–29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22–26 — median lobe; 28–34 — right paramere, scale as for respective median lobe; 22–25, 27 — ventrobasal aspect; 29–34 — left lateral aspect; 26, 28 — dorsal aspect. Scale bars 1 mm. Рис. 22–34. Эдеагус: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28–29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22–26 — среднЯЯ долЯ; 28–34 — праваЯ парамера в масштабе соответствуюЩей средней доли; 22–25, 27 — вентробаЗально; 29–34 — слева; 26, 28 — сверху. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949106" httpUri="https://zenodo.org/record/10949106/files/figure.png" pageId="4" pageNumber="20">24, 32</figureCitation>
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, 35–36, 47.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCF16697A34FD80251370DA" pageId="4" pageNumber="20" type="materials_examined">
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<paragraph id="A75736D5FFCF16697A34FD80251370DA" blockId="4.[812,1399,467,2058]" pageId="4" pageNumber="20">
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MATERIAL.
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<typeStatus id="78538877FFCF16697AAAFD8025747022" box="[972,1047,538,557]" pageId="4" pageNumber="20" type="paratype">Paratype</typeStatus>
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<specimenCount id="B1EEFD5CFFCF16697D78FD8D25497022" box="[1054,1066,535,557]" count="1" pageId="4" pageNumber="20" type="female">♀</specimenCount>
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(
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<collectionCode id="C1F9AE10FFCF16697D50FD80251F7022" box="[1078,1148,538,557]" pageId="4" pageNumber="20">MSNM</collectionCode>
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; digital images), with labels: ‘
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<collectingCountry id="DFFF7645FFCF16697A56FDA8221B704A" box="[816,888,562,581]" name="Vietnam" pageId="4" pageNumber="20">Tonkin</collectingCountry>
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/ Montes Mauson [Mau Son]/ April,Mai 2-3000´/ H.Fruhstorfer’, red printed, combined with handwritten text ‘
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<taxonomicName id="60E84D56FFCF16697C23FDD02239707A" authorityName="Förster" authorityYear="1862" class="Insecta" family="Braconidae" genus="Allotypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="genus">
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<typeStatus id="78538877FFCF16697C23FDD02239707A" pageId="4" pageNumber="20" type="allotype">Allotypus</typeStatus>
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</taxonomicName>
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/
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<emphasis id="959CEAC7FFCF16697A0EFDF822D1707A" box="[872,946,610,629]" italics="true" pageId="4" pageNumber="20">tonkinen</emphasis>
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/
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<emphasis id="959CEAC7FFCF16697AD8FDF822B6707B" box="[958,981,610,628]" italics="true" pageId="4" pageNumber="20">sis</emphasis>
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Stran.’, ‘
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<taxonomicName id="60E84D56FFCF16697D4FFDF825F7707A" authorityName="Bonelli" authorityYear="1810" box="[1065,1172,610,629]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="genus">
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<emphasis id="959CEAC7FFCF16697D4FFDF825F7707A" box="[1065,1172,610,629]" italics="true" pageId="4" pageNumber="20">Pterostichus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="60E84D56FFCF16697DC4FDF82460707A" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[1186,1283,610,629]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="tonkinensis" status="sp. nov.">
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<emphasis id="959CEAC7FFCF16697DC4FDF82460707A" box="[1186,1283,610,629]" italics="true" pageId="4" pageNumber="20">tonkinensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8EAF57BCFFCF16697C6FFDF82452707A" box="[1289,1329,610,629]" pageId="4" pageNumber="20" rank="species">n.sp.</taxonomicNameLabel>
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/ det. S. L. Straneo 1977/
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<taxonomicName id="60E84D56FFCF16697AA0FDE025747082" authorityName="Förster" authorityYear="1862" box="[966,1047,634,653]" class="Insecta" family="Braconidae" genus="Allotypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="genus">
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<emphasis id="959CEAC7FFCF16697AA0FDE025747082" box="[966,1047,634,653]" italics="true" pageId="4" pageNumber="20">
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<typeStatus id="78538877FFCF16697AA0FDE025747082" box="[966,1047,634,653]" pageId="4" pageNumber="20" type="allotype">Allotypus</typeStatus>
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</emphasis>
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</taxonomicName>
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’, ‘From/
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<emphasis id="959CEAC7FFCF16697D0BFDE025C07082" box="[1133,1187,634,653]" italics="true" pageId="4" pageNumber="20">Berlin</emphasis>
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/ Mus.’, ‘Gift for/
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<emphasis id="959CEAC7FFCF16697C23FDE024137082" box="[1349,1392,634,653]" italics="true" pageId="4" pageNumber="20">1977</emphasis>
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/ determ.’;
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<specimenCount id="B1EEFD5CFFCF16697AE4FD0822D670AA" box="[898,949,655,677]" count="2" pageId="4" pageNumber="20" type="male">2♂♂</specimenCount>
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(
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<collectionCode id="C1F9AE10FFCF16697AA5FD08256870AA" box="[963,1035,658,677]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="20" type="Museum">ZMMU</collectionCode>
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):
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<collectingCountry id="DFFF7645FFCF16697D7AFD08250B70AA" box="[1052,1128,658,677]" name="Vietnam" pageId="4" pageNumber="20">Vietnam</collectingCountry>
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,
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<quantity id="60109B30FFCF16697D12FD0825CF70AA" box="[1140,1196,658,677]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="20" unit="km" value="40.0">40 km</quantity>
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W of
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<collectingRegion id="652CF837FFCF16697D8DFD08242770AA" box="[1259,1348,658,677]" country="Vietnam" name="Cao Bang" pageId="4" pageNumber="20">Cao Bang</collectingRegion>
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, Phia Oac Mt.,
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<geoCoordinate id="C2DC5012FFCF16697AE7FD33228C70B3" box="[897,1007,681,700]" degrees="22" direction="north" minutes="36" orientation="latitude" pageId="4" pageNumber="20" precision="15" seconds="50" value="22.61389">22°36´50´´N</geoCoordinate>
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<geoCoordinate id="C2DC5012FFCF16697A90FD33250070B3" box="[1014,1123,681,700]" degrees="105" direction="east" minutes="52" orientation="longitude" pageId="4" pageNumber="20" precision="15" seconds="0" value="105.86667">105°52´0´´E</geoCoordinate>
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, h~
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<quantity id="60109B30FFCF16697DE5FD3025AA70B2" box="[1155,1225,682,701]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="20" unit="m" value="1800.0">1800 m</quantity>
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, deciduous bamboo forest,
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<date id="D3561015FFCF16697A0CFD5822B870DA" box="[874,987,706,725]" pageId="4" pageNumber="20" value="2018-10-03" valueMax="2018-10-11" valueMin="2018-10-03">3–11.X.2018</date>
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(D.
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<taxonomicName id="60E84D56FFCF16697D60FD58250570DA" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[1030,1126,706,725]" pageId="4" pageNumber="26" rank="subGenus" status="subgen. nov." subGenus="Fedorenko">Fedorenko</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCF16697A34FD40246E71A5" pageId="4" pageNumber="20" type="diagnosis">
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<paragraph id="A75736D5FFCF16697A34FD40246E71A5" blockId="4.[812,1399,467,2058]" pageId="4" pageNumber="20">
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DIAGNOSIS. This new species is recognizable among all the consubgeners by a very slight constriction between pro- and mesothorax so that the pronotum and the elytra are subequally wide (
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<emphasis id="959CEAC7FFCF16697A0BFCB022E27130" box="[877,897,810,831]" italics="true" pageId="4" pageNumber="20">vs</emphasis>
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. narrow) at their bases, the pronotum thus being subquadrate (
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<emphasis id="959CEAC7FFCF16697ACFFCDF22DF7155" box="[937,956,837,858]" italics="true" pageId="4" pageNumber="20">vs</emphasis>
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. circular to oval), and the elytra being subparallel-sided. The most similar species,
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<taxonomicName id="60E84D56FFCF16697DF4FCFA2461717B" authorityName="Fedorenko" authorityYear="2019" box="[1170,1282,863,885]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="caobang">
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<emphasis id="959CEAC7FFCF16697DF4FCFA2461717B" box="[1170,1282,863,885]" italics="true" pageId="4" pageNumber="20">P. caobang</emphasis>
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</taxonomicName>
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<bibRefCitation id="C3794B24FFCF16697C6BFCC5223E7180" author="Fedorenko D. N." pageId="4" pageNumber="20" pagination="358 - 369" refId="ref19561" refString="Fedorenko D. N. 2019. New species of Pterostichus subgenus Circinatus (Coleoptera: Carabidae: Pterostichini) from Vietnam // Russian Entomol. J. Vol. 28. No. 4. P. 358 - 369. doi: 10.15298 / rusentj. 28.4.03." type="journal article" year="2019">Fedorenko, 2019</bibRefCitation>
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, is distinctive in having
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<emphasis id="959CEAC7FFCF16697D5DFCE0252F7180" box="[1083,1100,890,911]" italics="true" pageId="4" pageNumber="20">fe</emphasis>
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3 bisetose,
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<emphasis id="959CEAC7FFCF16697DD4FCE025A27180" box="[1202,1217,890,911]" italics="true" pageId="4" pageNumber="20">tr</emphasis>
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3 seta vestigial, and the body appearance characteristic of the subgenus.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCF166B7A34FC3520A67683" lastPageId="6" lastPageNumber="22" pageId="4" pageNumber="20" type="description">
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<paragraph id="A75736D5FFCF16697A34FC3524147695" blockId="4.[812,1399,467,2058]" pageId="4" pageNumber="20">
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REDESCRIPTION. With characters of the subgenus [
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<bibRefCitation id="C3794B24FFCF16697C32FC35229671D0" author="Allegro G. & Sciaky R." pageId="4" pageNumber="20" pagination="3 - 18" refId="ref19009" refString="Allegro G., Sciaky R. 2010. Review of the Chinese subgenus Circinatus of Pterostichus, with description of six new species (Coleoptera, Carabidae) // Boll. Mus. Stor. Nat. Verona. Vol. 34. Bot. & Zool. P. 3 - 18." type="book chapter" year="2010">Allegro, Sciaky, 2010</bibRefCitation>
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;
|
||
<bibRefCitation id="C3794B24FFCF16697D65FC5325D271D0" author="Shi H. & Liang H." box="[1027,1201,969,991]" pageId="4" pageNumber="20" pagination="1 - 92" refId="ref20430" refString="Shi H., Liang H. 2015. The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini) and new species descriptions // Zookeys. Vol. 536. P. 1 - 92." type="journal article" year="2015">Shi, Liang, 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C3794B24FFCF16697DD9FC53240D71D0" author="Fedorenko D. N." box="[1215,1390,969,991]" pageId="4" pageNumber="20" pagination="358 - 369" refId="ref19561" refString="Fedorenko D. N. 2019. New species of Pterostichus subgenus Circinatus (Coleoptera: Carabidae: Pterostichini) from Vietnam // Russian Entomol. J. Vol. 28. No. 4. P. 358 - 369. doi: 10.15298 / rusentj. 28.4.03." type="journal article" year="2019">
|
||
<taxonomicName id="60E84D56FFCF16697DD9FC53244A71D0" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[1215,1321,969,991]" pageId="4" pageNumber="26" rank="subGenus" status="subgen. nov." subGenus="Fedorenko">Fedorenko</taxonomicName>
|
||
, 2019
|
||
</bibRefCitation>
|
||
]. Body (
|
||
<figureCitation id="3FD32A50FFCF16697A09FC7E22C971F5" box="[879,938,996,1018]" captionStart="Figs 6–11" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[202,1388,618,1932]" captionTargetId="figure-261@3.[190,1398,609,1945]" captionTargetPageId="3" captionText="Figs 6–11. Pterostichus: 6, 9 — P. tonkinensis, ♂ from Phia Oac Mt; 7, 10 — P. austrinus sp.n., holotype; 8, 11 — P. laevibasis sp.n., paratype ♂; 6–8 — dorsal habitus; 9–11 — apices of elytra and abdomen, left aspect. Рис. 6–11. Pterostichus: 6, 9 — P. tonkinensis, ♂ с горы Phia Oac; 7, 10 — P. austrinus sp.n., голотип; 8, 11 — P. laevibasis sp.n., паратип ♂; 6–8 — габитус дорЗально; 9–11 — вершина надкрылий и брюшка слева." figureDoi="http://doi.org/10.5281/zenodo.10949100" httpUri="https://zenodo.org/record/10949100/files/figure.png" pageId="4" pageNumber="20">Fig. 6</figureCitation>
|
||
) apterous and rather small, BL 10.5–10.6 mm, shiny black and iridescent. Extreme lateral margin of pronotum reddish, more widely so at basal angles. Antennae, palps, extreme femoral apices, and tarsi red. Dorsal microsculpture isodiametric and distinct on head, hardly traceable on pronotum and elytra, consisting of very small, moderately transverse meshes or very dense transverse lines, respectively.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFCF16697A34FB0525BF77FA" blockId="4.[812,1399,467,2058]" pageId="4" pageNumber="20">Head very small, without neck constriction. Eyes convex, genae about a third as long and meeting neck at obtuse angle. Clypeus indistinctly sinuate at apical margin, frons smooth, frontal sulci short, finely and densely punctate at bottom only, parallel and very deep behind straight frontoclypeal suture, then curved toward while not reaching the level of anterior supra-ocular setae, each extended onto clypeus into a fine impressed line, with convexity outward, to adjoin clypeal seta. Labrum rectangular, subsinuate apically, with a shallow round pit at middle of apical half. Mandibular scrobe with longitudinal sulcus. Labial palpomere 3 obliquely truncate and very slightly narrower at apex than preapically. Submentum quadrisetose. Antennae just reaching pronotal base.</paragraph>
|
||
<paragraph id="A75736D5FFCF16697A34FA60259D75DB" blockId="4.[812,1399,467,2058]" pageId="4" pageNumber="20">Pronotum quadrate, broadest two fifths from apex; sides from subequally rounded all along to nearly straight in basal half, slightly more rounded at basal angles, these slightly obtuse and blunt. Base barely oblique on sides, two fifths wider than apex, moderately punctate at bottom of and a little inside inner basolateral sulci, more sparsely so toward basal angles. Apex nearly truncate, apical angles very slightly acute, a little projecting and slightly blunted. Marginal beads fine and obliterate in middle third of base only. Lateral groove in form of a fine and well impressed line inside lateral bead, finely and densely punctate at bottom and slightly inside in apical four fifths. Inner basolateral sulci in form of deep impressed lines running parallel to each other in basal two fifths, almost reaching base; outer sulci as short and oblique lines hardly traceable between basal and lateral margins. Median line fine, deep, indistinctly crenulate. Basal and apical transverse impressions indistinct. Disc smooth, sublateral line inside lateral groove missing or vague.</paragraph>
|
||
<paragraph id="A75736D5FFCF166B7A34F8432390709E" blockId="4.[812,1399,467,2058]" lastBlockId="6.[189,776,315,1537]" lastPageId="6" lastPageNumber="22" pageId="4" pageNumber="20">
|
||
Elytra very convex, oval, broadest at about middle, rounded toward humeri and in apical half, nearly straight in between; apices rounded combined; preapical sinuation rather shallow, plica distinct in lateral view. Base slightly oblique, humeri with a distinct tooth; basal ridge transverse and nearly straight, humeral angle obtuse. Striae deep, minutely punctate at bottom, stria 9 finely punctate, those 1–6 adjoining basal ridge, stria 2 adjoining parascutellar seta; parascutellar striole missing (
|
||
<emphasis id="959CEAC7FFCD166B7B83FE5B226273D9" box="[741,769,449,470]" italics="true" pageId="6" pageNumber="22">i.e.</emphasis>
|
||
, substituted for obliterate primary base of stria 1). Intervals convex, very so toward apex, intervals 7, 5 and 3 confluent apicad in succession separate from intervals 2 and 8. This latter narrow and costate at base and at apex. Reflexed lateral margin narrow, without additional carinae. Interval 3 with single discal seta, d2, adjoining stria 2 just behind middle, stria 7 with two preapical setae. USS: 6–1–9 or 6–1–8,
|
||
<collectingCountry id="DFFF7645FFCD166B7B4CFDE12329709F" box="[554,586,635,656]" name="United States of America" pageId="6" pageNumber="22">US</collectingCountry>
|
||
8 at about middle.
|
||
</paragraph>
|
||
<caption id="F397665DFFCE16687984F80324587A1E" ID-DOI="http://doi.org/10.5281/zenodo.10949104" ID-Zenodo-Dep="10949104" httpUri="https://zenodo.org/record/10949104/files/figure.png" pageId="5" pageNumber="21" startId="5.[226,263,1945,1964]" targetBox="[189,1399,319,1919]" targetPageId="5" targetType="figure">
|
||
<paragraph id="A75736D5FFCE16687984F803204375D2" blockId="5.[189,1399,1945,2065]" pageId="5" pageNumber="21">
|
||
Figs 12–21. Aedeagus median lobe: 12–13 —
|
||
<taxonomicName id="60E84D56FFCE16687B13F803225575A3" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[629,822,1945,1964]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="glabellus" status="sp. nov.">
|
||
<emphasis id="959CEAC7FFCE16687B13F803225575A3" box="[629,822,1945,1964]" italics="true" pageId="5" pageNumber="21">Pterostichus glabellus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFCE16687A5AF803220475A3" bold="true" box="[828,871,1945,1964]" pageId="5" pageNumber="21">
|
||
<taxonomicNameLabel id="8EAF57BCFFCE16687A5AF803220475A3" box="[828,871,1945,1964]" pageId="5" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 14–15 —
|
||
<taxonomicName id="60E84D56FFCE16687AACF800255E75A3" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[970,1085,1946,1964]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="unisetosus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFCE16687AACF800255E75A3" box="[970,1085,1946,1964]" italics="true" pageId="5" pageNumber="21">P. unisetosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFCE16687D25F803250E75A3" bold="true" box="[1091,1133,1945,1964]" pageId="5" pageNumber="21">
|
||
<taxonomicNameLabel id="8EAF57BCFFCE16687D25F803250E75A3" box="[1091,1133,1945,1964]" pageId="5" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 16–17 —
|
||
<taxonomicName id="60E84D56FFCE16687DB7F800242975A3" authorityName="Fedorenko" authorityYear="2023" box="[1233,1354,1945,1964]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="tonkinensis">
|
||
<emphasis id="959CEAC7FFCE16687DB7F800242975A3" box="[1233,1354,1945,1964]" italics="true" pageId="5" pageNumber="21">P. tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
; 18– 19 —
|
||
<taxonomicName id="60E84D56FFCE16687997F828203A75CB" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[241,345,1970,1988]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="austrinus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFCE16687997F828203A75CB" box="[241,345,1970,1988]" italics="true" pageId="5" pageNumber="21">P. austrinus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFCE16687838F82B20EA75CB" bold="true" box="[350,393,1969,1988]" pageId="5" pageNumber="21">
|
||
<taxonomicNameLabel id="8EAF57BCFFCE16687838F82B20EA75CB" box="[350,393,1969,1988]" pageId="5" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 20–21 —
|
||
<taxonomicName id="60E84D56FFCE1668788CF828233875CB" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[490,603,1969,1988]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="subirideus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFCE1668788CF828233875CB" box="[490,603,1969,1988]" italics="true" pageId="5" pageNumber="21">P. subirideus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFCE16687B06F82B23E875CB" bold="true" box="[608,651,1969,1988]" pageId="5" pageNumber="21">
|
||
<taxonomicNameLabel id="8EAF57BCFFCE16687B06F82B23E875CB" box="[608,651,1969,1988]" pageId="5" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 12, 14, 16, 18, 20 — left lateral aspect; 13, 15, 17, 19, 21 — right lateral aspect. Scale bars 1 mm.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFCE16687984F87C24587A1E" blockId="5.[189,1399,1945,2065]" pageId="5" pageNumber="21">
|
||
Рис. 12–21. СреднЯЯ долЯ Эдеагуса: 12–13 —
|
||
<taxonomicName id="60E84D56FFCE16687B18F87F225D75F7" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[638,830,2021,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="glabellus" status="sp. nov.">
|
||
<emphasis id="959CEAC7FFCE16687B18F87F225D75F7" box="[638,830,2021,2040]" italics="true" pageId="5" pageNumber="21">Pterostichus glabellus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFCE16687A22F87F220D75F7" bold="true" box="[836,878,2021,2040]" pageId="5" pageNumber="21">
|
||
<taxonomicNameLabel id="8EAF57BCFFCE16687A22F87F220D75F7" box="[836,878,2021,2040]" pageId="5" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 14–15 —
|
||
<taxonomicName id="60E84D56FFCE16687AA9F87C252275F7" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[975,1089,2022,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="unisetosus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFCE16687AA9F87C252275F7" box="[975,1089,2022,2040]" italics="true" pageId="5" pageNumber="21">P. unisetosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFCE16687D20F87F251275F7" bold="true" box="[1094,1137,2021,2040]" pageId="5" pageNumber="21">
|
||
<taxonomicNameLabel id="8EAF57BCFFCE16687D20F87F251275F7" box="[1094,1137,2021,2040]" pageId="5" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 16–17 —
|
||
<taxonomicName id="60E84D56FFCE16687DB4F87C242875F7" authorityName="Fedorenko" authorityYear="2023" box="[1234,1355,2021,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="tonkinensis">
|
||
<emphasis id="959CEAC7FFCE16687DB4F87C242875F7" box="[1234,1355,2021,2040]" italics="true" pageId="5" pageNumber="21">P. tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
; 18– 19 —
|
||
<taxonomicName id="60E84D56FFCE16687993F864203D7A1F" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[245,350,2046,2064]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="austrinus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFCE16687993F864203D7A1F" box="[245,350,2046,2064]" italics="true" pageId="5" pageNumber="21">P. austrinus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFCE16687803F86720F37A1F" bold="true" box="[357,400,2045,2064]" pageId="5" pageNumber="21">
|
||
<taxonomicNameLabel id="8EAF57BCFFCE16687803F86720F37A1F" box="[357,400,2045,2064]" pageId="5" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 20–21 —
|
||
<taxonomicName id="60E84D56FFCE16687890F864230A7A1F" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[502,617,2045,2064]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="subirideus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFCE16687890F864230A7A1F" box="[502,617,2045,2064]" italics="true" pageId="5" pageNumber="21">P. subirideus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFCE16687B09F86723F97A1F" bold="true" box="[623,666,2045,2064]" pageId="5" pageNumber="21">
|
||
<taxonomicNameLabel id="8EAF57BCFFCE16687B09F86723F97A1F" box="[623,666,2045,2064]" pageId="5" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 12, 14, 16, 18, 20 — слева; 13, 15, 17, 19, 21 — справа. Масштаб 1 мм.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A75736D5FFCD166B7984FD0C23D87194" blockId="6.[189,776,315,1537]" pageId="6" pageNumber="22">
|
||
Ventral side: propleura, as well as meso- and metathorax except medially, with coarse and dense punctation, abdominal sternites II–VI more finely and rather densely punctate along sides. Prosternum without median groove; prosternal process apically truncate, not beaded, in lateral view subrectangular and blunt; inclination wide and concave, with sides edged. Metepisternum short, est3L/W 0.87. Abdominal lateral bead entire. Male sternite VII (
|
||
<figureCitation id="3FD32A50FFCD166B78CCFCCB20827168" box="[426,481,849,871]" captionStart="Figs 6–11" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[202,1388,618,1932]" captionTargetId="figure-261@3.[190,1398,609,1945]" captionTargetPageId="3" captionText="Figs 6–11. Pterostichus: 6, 9 — P. tonkinensis, ♂ from Phia Oac Mt; 7, 10 — P. austrinus sp.n., holotype; 8, 11 — P. laevibasis sp.n., paratype ♂; 6–8 — dorsal habitus; 9–11 — apices of elytra and abdomen, left aspect. Рис. 6–11. Pterostichus: 6, 9 — P. tonkinensis, ♂ с горы Phia Oac; 7, 10 — P. austrinus sp.n., голотип; 8, 11 — P. laevibasis sp.n., паратип ♂; 6–8 — габитус дорЗально; 9–11 — вершина надкрылий и брюшка слева." figureDoi="http://doi.org/10.5281/zenodo.10949100" httpUri="https://zenodo.org/record/10949100/files/figure.png" pageId="6" pageNumber="22">Fig. 9</figureCitation>
|
||
) with a pair of small paramedian tubercles, a shallow transverse impression in between and two apical setae 1/3 sternite length distant from apex.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFCD166B7984FC3B20E471DE" blockId="6.[189,776,315,1537]" pageId="6" pageNumber="22">
|
||
Legs:
|
||
<emphasis id="959CEAC7FFCD166B7879FC3B205371B9" box="[287,304,929,950]" italics="true" pageId="6" pageNumber="22">fe</emphasis>
|
||
3 unisetose due to distal seta missing; tarsomere 5 glabrous ventrally.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFCD166B7984FC4C20A67683" blockId="6.[189,776,315,1537]" pageId="6" pageNumber="22">
|
||
Aedeagus (
|
||
<figureCitation id="3FD32A50FFCD166B7829FC4C20A371E3" box="[335,448,982,1004]" captionStart="Figs 12–21" captionStartId="5.[226,263,1945,1964]" captionTargetBox="[189,1399,319,1919]" captionTargetId="figure-122@5.[189,1399,314,1923]" captionTargetPageId="5" captionText="Figs 12–21. Aedeagus median lobe: 12–13 — Pterostichus glabellus sp.n.; 14–15 — P. unisetosus sp.n.; 16–17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20–21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — left lateral aspect; 13, 15, 17, 19, 21 — right lateral aspect. Scale bars 1 mm. Рис. 12–21. СреднЯЯ долЯ Эдеагуса: 12–13 — Pterostichus glabellus sp.n.; 14–15 — P. unisetosus sp.n.; 16–17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20–21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — слева; 13, 15, 17, 19, 21 — справа. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949104" httpUri="https://zenodo.org/record/10949104/files/figure.png" pageId="6" pageNumber="22">Figs 16–17</figureCitation>
|
||
,
|
||
<figureCitation id="3FD32A50FFCD166B78ADFC4C236971E4" box="[459,522,982,1003]" captionStart="Figs 22–34" captionStartId="7.[226,263,1921,1940]" captionTargetBox="[190,1399,697,1904]" captionTargetId="figure-163@7.[189,1399,694,1904]" captionText="Figs 22–34. Aedeagus: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28–29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22–26 — median lobe; 28–34 — right paramere, scale as for respective median lobe; 22–25, 27 — ventrobasal aspect; 29–34 — left lateral aspect; 26, 28 — dorsal aspect. Scale bars 1 mm. Рис. 22–34. Эдеагус: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28–29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22–26 — среднЯЯ долЯ; 28–34 — праваЯ парамера в масштабе соответствуюЩей средней доли; 22–25, 27 — вентробаЗально; 29–34 — слева; 26, 28 — сверху. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949106" httpUri="https://zenodo.org/record/10949106/files/figure.png" pageId="6" pageNumber="22">24, 32</figureCitation>
|
||
, 35–36, 47): Median lobe in lateral view strongly curved, nearly geniculate, with an oblong and shallow concavity at middle of right side; apex in dorsal view narrow and slightly curved to the left. Right paramere short triangular. Everted and inflated internal sac curved leftward and ventrad, with two large preapical bulbs, left being larger than right.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFF2655EFFCD166B7984FB0B23BE76F8" pageId="6" pageNumber="22" type="distribution">
|
||
<paragraph id="A75736D5FFCD166B7984FB0B239176A8" blockId="6.[189,776,315,1537]" box="[226,754,1169,1191]" pageId="6" pageNumber="22">
|
||
DISTRIBUTION. Known from the
|
||
<typeStatus id="78538877FFCD166B7B26FB0B230976A9" box="[576,618,1169,1190]" pageId="6" pageNumber="22">type</typeStatus>
|
||
locality only.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFCD166B7984FB3123BE76F8" blockId="6.[189,776,315,1537]" pageId="6" pageNumber="22">
|
||
HABITATS AND HABITS. A forest-dwelling species. The
|
||
<typeStatus id="78538877FFCD166B798EFB5C207176D4" box="[232,274,1222,1243]" pageId="6" pageNumber="22">type</typeStatus>
|
||
specimens were collected at ca.
|
||
<quantity id="60109B30FFCD166B7B2BFB5C23D876D4" box="[589,699,1222,1244]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="6" pageNumber="22" unit="m" value="750.0" valueMax="900.0" valueMin="600.0">600–900 m</quantity>
|
||
and the specimens from the Phia Oac Mt at
|
||
<quantity id="60109B30FFCD166B7B7BFB7B230E76F9" box="[541,621,1249,1270]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="22" unit="m" value="1800.0">1.800 m</quantity>
|
||
elevations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFF2655EFFCD166B7984FB6120F8740E" pageId="6" pageNumber="22" type="discussion">
|
||
<paragraph id="A75736D5FFCD166B7984FB6120F8740E" blockId="6.[189,776,315,1537]" pageId="6" pageNumber="22">
|
||
COMMENTS. This species was described based on
|
||
<specimenCount id="B1EEFD5CFFCD166B7BB0FB6120477723" count="3" pageId="6" pageNumber="22" type="generic">three specimens</specimenCount>
|
||
, male
|
||
<typeStatus id="78538877FFCD166B780EFA8C20DF7723" box="[360,444,1302,1324]" pageId="6" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="B1EEFD5CFFCD166B7896FA8C231F7724" box="[496,636,1302,1323]" count="2" pageId="6" pageNumber="22" type="generic" typeStatus="paratype">two paratypes</specimenCount>
|
||
, male and female, from the
|
||
<location id="A237600EFFCD166B7829FAAB20B27749" LSID="urn:lsid:plazi:treatment:2F4187C3FFCF166B7A5CFE4920F8740E:A237600EFFCD166B7829FAAB20B27749" box="[335,465,1329,1350]" country="Vietnam" county="Phia Oac Mt." municipality="This" name="Mau Son Mts" pageId="6" pageNumber="22" stateProvince="Lang Son">Mau Son Mts</location>
|
||
in
|
||
<collectingRegion id="652CF837FFCD166B7896FAAB23C87748" box="[496,683,1329,1351]" country="Vietnam" name="Lang Son" pageId="6" pageNumber="22">Lang Son Province</collectingRegion>
|
||
, northern
|
||
<collectingCountry id="DFFF7645FFCD166B79DBFAD12070776E" box="[189,275,1355,1377]" name="Vietnam" pageId="6" pageNumber="22">Vietnam</collectingCountry>
|
||
.
|
||
<location id="A237600EFFCD166B787CFAD120E3776E" LSID="urn:lsid:plazi:treatment:2F4187C3FFCF166B7A5CFE4920F8740E:A237600EFFCD166B787CFAD120E3776E" box="[282,384,1355,1377]" country="Vietnam" county="Phia Oac Mt." municipality="This" name="According" pageId="6" pageNumber="22" stateProvince="Lang Son">According</location>
|
||
to the description, the
|
||
<typeStatus id="78538877FFCD166B7B30FAD123C9776E" box="[598,682,1355,1377]" pageId="6" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
is
|
||
<quantity id="60109B30FFCD166B7BA2FAD12265776E" box="[708,774,1355,1377]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="22" unit="mm" value="11.0">11 mm</quantity>
|
||
in length (
|
||
<emphasis id="959CEAC7FFCD166B7878FAFD205A7773" box="[286,313,1383,1404]" italics="true" pageId="6" pageNumber="22">vs.</emphasis>
|
||
9.2 mm in the female
|
||
<typeStatus id="78538877FFCD166B7B6AFAFC233C7774" box="[524,607,1382,1403]" pageId="6" pageNumber="22" type="paratype">paratype</typeStatus>
|
||
I measured using scale bar).
|
||
<collectingMunicipality id="4733ACAFFFCD166B7842FA1B202C7798" box="[292,335,1409,1431]" pageId="6" pageNumber="22">This</collectingMunicipality>
|
||
latter specimen is very slightly different from the examined specimens from the
|
||
<collectingCounty id="4E364E59FFCD166B7B63FA0123E677BE" box="[517,645,1435,1457]" pageId="6" pageNumber="22">Phia Oac Mt.</collectingCounty>
|
||
in having the pronotum barely wider, with base slightly wider (
|
||
<emphasis id="959CEAC7FFCD166B7BFCFA2D23D677C3" box="[666,693,1463,1484]" italics="true" pageId="6" pageNumber="22">vs.</emphasis>
|
||
narrower) than the elytral bases combined, and the elytral humeral tooth nearly indistinct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |