187 lines
20 KiB
XML
187 lines
20 KiB
XML
<document id="70EAEF6CE31A1817ED33A02B2DFF611C" ID-DOI="10.15298/rusentj.32.1.03" ID-ISSN="0132-8069" ID-Zenodo-Dep="10949096" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712198792598" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2023" docId="2F4187C3FFCA166C79AAFDB02435718E" docLanguage="en" docName="RussEntomolJ.32.1.16-39.pdf" docOrigin="Russian Entomological Journal 32 (1)" docSource="http://dx.doi.org/10.15298/rusentj.32.1.03" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.1:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="1" docTitle="(Morphohaptoderus) Tschitscherine 1898" docType="treatment" docVersion="5" lastPageNumber="17" masterDocId="D378FFBBFFCB166D7966FF9A2163720F" masterDocTitle="New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam" masterLastPageNumber="39" masterPageNumber="16" pageNumber="17" updateTime="1712734223118" updateUser="ExternalLinkService">
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<mods:titleInfo id="068339A00DEBF4AAEBF40EAC291439D2">
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<mods:title id="75068079A634F0404F95DA78678E46EE">New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam</mods:title>
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<mods:name id="866423F90B1B2DAE4C007455B109A541" type="personal">
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<mods:namePart id="6412F24BF616958953728582E27C8054">Fedorenko, D. N.</mods:namePart>
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<mods:title id="00AE99A4CB30BDDE540BB2F481E5379C">Russian Entomological Journal</mods:title>
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<mods:part id="713BD2ED34F401802414025D81217375">
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<mods:date id="E822B2FA93F11C6905A408F6B511F726">2023</mods:date>
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<mods:detail id="64241EEB3A76D8855D623A696DF89C23" type="pubDate">
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<mods:number id="230179044BAD9817FF94B7761DD68F84">2023-03-31</mods:number>
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<mods:number id="9B37204C76C1ECD4AD0110325E5C76D9">32</mods:number>
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<mods:url id="61B42C6629BFA9878F93FE01605C7669">http://dx.doi.org/10.15298/rusentj.32.1.03</mods:url>
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<mods:identifier id="9287C774C421737F4EAA12CA08A0DDB9" type="DOI">10.15298/rusentj.32.1.03</mods:identifier>
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<mods:identifier id="CED8D533EC1EC03DD7771AF2D2F42F3A" type="ISSN">0132-8069</mods:identifier>
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<treatment id="2F4187C3FFCA166C79AAFDB02435718E" ID-DOI="http://doi.org/10.5281/zenodo.10952380" ID-Zenodo-Dep="10952380" LSID="urn:lsid:plazi:treatment:2F4187C3FFCA166C79AAFDB02435718E" httpUri="http://treatment.plazi.org/id/2F4187C3FFCA166C79AAFDB02435718E" lastPageNumber="17" pageId="1" pageNumber="17">
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<subSubSection id="EFF2655EFFCA166C79AAFDB02394704D" box="[204,759,554,578]" pageId="1" pageNumber="17" type="nomenclature">
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<paragraph id="A75736D5FFCA166C79AAFDB02394704D" blockId="1.[189,776,554,2069]" box="[204,759,554,578]" pageId="1" pageNumber="17">
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<heading id="FC1F81B9FFCA166C79AAFDB02394704D" box="[204,759,554,578]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="17" reason="3">
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Subgenus
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<taxonomicName id="60E84D56FFCA166C785DFDB02394704D" ID-CoL="8MBT6" authority="Tschitscherine, 1898" authorityName="Tschitscherine" authorityYear="1898" box="[315,759,554,578]" kingdom="Animalia" pageId="1" pageNumber="17" rank="subGenus" subGenus="Morphohaptoderus">
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<emphasis id="959CEAC7FFCA166C785DFDB0236C704D" bold="true" box="[315,527,554,578]" italics="true" pageId="1" pageNumber="17">Morphohaptoderus</emphasis>
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Tschitschérine, 1898
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCA166C7984FDD520157075" pageId="1" pageNumber="17" type="reference_group">
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<paragraph id="A75736D5FFCA166C7984FDD520157075" blockId="1.[189,776,554,2069]" pageId="1" pageNumber="17">
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<bibRefCitation id="C3794B24FFCA166C7984FDD520B2706D" author="Tschitscherine T." box="[226,465,591,610]" pageId="1" pageNumber="17" pagination="1 - 224" refId="ref20688" refString="Tschitscherine T. 1898 a. Materiaux pour servir a l'etude des feroniens. IV // Horae Soc. Ent. Ross. T. 32. P. 1 - 224." type="journal article" year="1898">Tschitschérine, 1898a: 190</bibRefCitation>
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(
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<taxonomicName id="60E84D56FFCA166C7887FDCA2345706D" authority="Leach, 1817" box="[481,550,592,610]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Feronia">
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<emphasis id="959CEAC7FFCA166C7887FDCA2345706D" box="[481,550,592,610]" italics="true" pageId="1" pageNumber="17">Feronia</emphasis>
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</taxonomicName>
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subg.);
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<bibRefCitation id="C3794B24FFCA166C7B15FDD52262706D" author="Sciaky R." box="[627,769,591,610]" pageId="1" pageNumber="17" pagination="1 - 19" refId="ref20291" refString="Sciaky R. 1994. Revision of Pterostichus subg. Morphohaptoderus Tschitscherine, 1898 with description of ten new species from China (Coleoptera: Carabidae) // Koleopt. Rundsch. Bd. 64. P. 1 - 19." type="book chapter" year="1994">Sciaky, 1994: 1</bibRefCitation>
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;
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<bibRefCitation id="C3794B24FFCA166C79DBFDFD20107075" author="Shi H. & Sciaky R. & Liang H. & Zhou H." box="[189,371,615,635]" pageId="1" pageNumber="17" pagination="101 - 135" refId="ref20376" refString="Shi H., Sciaky R., Liang H., Zhou H. 2013. A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions // Zootaxa. Vol. 3664. No. 2. P. 101 - 135." type="book chapter" year="2013">Shi et al., 2013: 105</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCA166C7984FD1A237C70A5" pageId="1" pageNumber="17" type="materials_examined">
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<paragraph id="A75736D5FFCA166C7984FD1A237C70A5" blockId="1.[189,776,554,2069]" pageId="1" pageNumber="17">
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<typeStatus id="78538877FFCA166C7984FD1A206D709C" box="[226,270,640,659]" pageId="1" pageNumber="17">Type</typeStatus>
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species:
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<taxonomicName id="60E84D56FFCA166C7807FDE523B4709D" authority="Tschitscherine, 1889" authorityName="Tschitscherine" authorityYear="1889" box="[353,727,639,658]" class="Insecta" family="Carabidae" genus="Haptoderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="maximus">
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<emphasis id="959CEAC7FFCA166C7807FDE5237A709D" box="[353,537,639,658]" italics="true" pageId="1" pageNumber="17">Haptoderus maximus</emphasis>
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<bibRefCitation id="C3794B24FFCA166C7B46FDE523B4709D" author="Tschitscherine T." box="[544,727,639,658]" pageId="1" pageNumber="17" pagination="185 - 198" refId="ref20594" refString="Tschitscherine T. 1889. Insecta, a Cl. G. N. Potanin in China et in Mongolia novissime lecta. Insectes rapportes par Mr. Potanin de son voyage fait en 1884 - 85 - 86. VI. Genre Pterostichus // Horae Soc. Ent. Ross. T. 23. P. 185 - 198." type="journal article" year="1889">Tschitschérine, 1889</bibRefCitation>
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</taxonomicName>
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, subsequently designated by Jeannel [1937].
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCA166C7984FD352196711A" pageId="1" pageNumber="17" type="diagnosis">
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<paragraph id="A75736D5FFCA166C7984FD352196711A" blockId="1.[189,776,554,2069]" pageId="1" pageNumber="17">
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DIAGNOSIS. A polytypic subgenus of
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<taxonomicName id="60E84D56FFCA166C7B3DFD3523B770CB" authorityName="Bonelli" authorityYear="1810" box="[603,724,687,708]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="959CEAC7FFCA166C7B3DFD3523B770CB" box="[603,724,687,708]" italics="true" pageId="1" pageNumber="17">Pterostichus</emphasis>
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</taxonomicName>
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from southern
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<collectingCountry id="DFFF7645FFCA166C7871FD50203270EF" box="[279,337,714,736]" name="China" pageId="1" pageNumber="17">China</collectingCountry>
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to northern
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<collectingCountry id="DFFF7645FFCA166C78ADFD50234270EF" box="[459,545,714,736]" name="Vietnam" pageId="1" pageNumber="17">Vietnam</collectingCountry>
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, defined chiefly by
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<emphasis id="959CEAC7FFCA166C7B83FD50239970D0" box="[741,762,714,735]" italics="true" pageId="1" pageNumber="17">cx</emphasis>
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3 trisetose,
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<emphasis id="959CEAC7FFCA166C787FFD7F204A70F5" box="[281,297,741,762]" italics="true" pageId="1" pageNumber="17">tr</emphasis>
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3 setose and the body apterous, with metepisterna short.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCA166C7984FC80240F7028" pageId="1" pageNumber="17" type="description">
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<paragraph id="A75736D5FFCA166C7984FC80230671E5" blockId="1.[189,776,554,2069]" pageId="1" pageNumber="17">
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REDESCRIPTION (For recognition only). Body smallto medium-sized, BL 4.8–
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<quantity id="60109B30FFCA166C78DDFCAE209C7145" box="[443,511,820,842]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="17" unit="mm" value="13.0">13 mm</quantity>
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. Dorsum shiny black, often iridescent, sometimes dull in female. Head rather small, eyes convex to flat, neck constriction indistinct. Antennae filiform, moderately long, scape with preapical seta, pedicel with ventral seta and often with a few short setae toward apex; antennomeres 4–11 and often extreme apex of antennomere 3 pubescent. Submentum bi- or quadrisetose.
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</paragraph>
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<paragraph id="A75736D5FFCA166C7984FC75239B762F" blockId="1.[189,776,554,2069]" pageId="1" pageNumber="17">Pronotum flattened, each side mostly finely beaded, with basal angle distinct and two basolateral sulci on each side.</paragraph>
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<paragraph id="A75736D5FFCA166C7984FBBE23DD7700" blockId="1.[189,776,554,2069]" pageId="1" pageNumber="17">
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Elytra rather wide at bases, with striae 1–7 inside humeral angle; humeri and humeral tooth mostly distinct, parascutellar seta present and parascutellar striole short or missing. Interval 3 with discal setae varying from species to species in number as well as in position, sometimes multiple or missing. USS consisting of 16–17 (mostly 16) or 14–15
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<collectingCountry id="DFFF7645FFCA166C7B1DFB3023F976B0" box="[635,666,1194,1215]" name="United States of America" pageId="1" pageNumber="17">US</collectingCountry>
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in larger or smaller species, respectively;
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<collectingCountry id="DFFF7645FFCA166C78BDFB5E209976D6" box="[475,506,1220,1241]" name="United States of America" pageId="1" pageNumber="17">US</collectingCountry>
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arranged into a row 6–1–1– … (
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<collectingCountry id="DFFF7645FFCA166C7985FB45206776FA" box="[227,260,1247,1269]" name="United States of America" pageId="1" pageNumber="17">US</collectingCountry>
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1–6 —
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<collectingCountry id="DFFF7645FFCA166C782BFB45200F76FB" box="[333,364,1247,1268]" name="United States of America" pageId="1" pageNumber="17">US</collectingCountry>
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7 —
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<collectingCountry id="DFFF7645FFCA166C78F9FB4520A276FA" box="[415,449,1247,1269]" name="United States of America" pageId="1" pageNumber="17">US</collectingCountry>
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8–…),
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<emphasis id="959CEAC7FFCA166C7B65FB7A234276FA" box="[515,545,1248,1269]" italics="true" pageId="1" pageNumber="17">i.e.</emphasis>
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, sparser at middle, and adjoining stria 8 while
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<collectingCountry id="DFFF7645FFCA166C78F9FB6020DC7700" box="[415,447,1274,1295]" name="United States of America" pageId="1" pageNumber="17">US</collectingCountry>
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3 adjoining striae 8 and 9.
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</paragraph>
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<paragraph id="A75736D5FFCA166C7984FA8E233177BF" blockId="1.[189,776,554,2069]" pageId="1" pageNumber="17">
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Body setation otherwise complete, including clypeus bisetose, two supraocular setae, two elytral apical setae in stria 7,
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<emphasis id="959CEAC7FFCA166C79B1FAD0218B7750" box="[215,232,1354,1375]" italics="true" pageId="1" pageNumber="17">fe</emphasis>
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1 with three posterior setae,
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<emphasis id="959CEAC7FFCA166C7B65FAD023777750" box="[515,532,1354,1375]" italics="true" pageId="1" pageNumber="17">fe</emphasis>
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2 with two anteroventral setae in basal three fifths and 2–4 anterodorsal setae in apical two fifths,
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<emphasis id="959CEAC7FFCA166C7854FAE52020779B" box="[306,323,1407,1428]" italics="true" pageId="1" pageNumber="17">fe</emphasis>
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3 bisetose;
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<emphasis id="959CEAC7FFCA166C78DBFA1A20A9779A" box="[445,458,1408,1429]" italics="true" pageId="1" pageNumber="17">ti</emphasis>
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3 mostly without lateral setae. Tarsomere 5 glabrous or setose ventrally.
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</paragraph>
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<paragraph id="A75736D5FFCA166C7984FA2E20F37465" blockId="1.[189,776,554,2069]" pageId="1" pageNumber="17">
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Prosternal process flat or with indistinct median groove, apex truncate and indistinctly bifid in anteroventral view due to the median groove more impressed at extreme apex, angles rounded or blunt; inclination flat, with lateral beads converging ventrad to enclose triangular dorsal part or nearly so. Abdominal sternite VII bi- (
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<specimenCount id="B1EEFD5CFFCA166C78B5F9AD208A7440" box="[467,489,1591,1615]" count="1" pageId="1" pageNumber="17" type="male">♂</specimenCount>
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) or quadrisetose (
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<specimenCount id="B1EEFD5CFFCA166C7BFBF9AD23C97440" box="[669,682,1591,1615]" count="1" pageId="1" pageNumber="17" type="female">♀</specimenCount>
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) at apex, not modified in male.
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</paragraph>
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<paragraph id="A75736D5FFCA166C7984F9F523C574FF" blockId="1.[189,776,554,2069]" pageId="1" pageNumber="17">
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Legs rather slender;
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<emphasis id="959CEAC7FFCA166C78D0F9EA20A0748A" box="[438,451,1648,1669]" italics="true" pageId="1" pageNumber="17">ti</emphasis>
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1 anterior face smooth and glabrous;
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<emphasis id="959CEAC7FFCA166C7860F910207A7490" box="[262,281,1674,1695]" italics="true" pageId="1" pageNumber="17">ta</emphasis>
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2 and
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<emphasis id="959CEAC7FFCA166C7807F91020177490" box="[353,372,1674,1695]" italics="true" pageId="1" pageNumber="17">ta</emphasis>
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3 mostly with tarsomeres 1–3 carinate laterally and bisulcate dorsolaterally; posterior (inner) sulcus being more shallow than anterior (outer) one and varying from distinct to totally obliterate between species.
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</paragraph>
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<paragraph id="A75736D5FFCA166C7984F96E2303759F" blockId="1.[189,776,554,2069]" pageId="1" pageNumber="17">Aedeagus rather typical for the genus, median lobe in lateral view strongly curved to geniculate, with apical orifice left dorsolateral to nearly dorsal; left paramere quadrate, with a deep dorsobasal fissure; right paramere short. Everted and inflated internal sac curved first leftward and then ventrad and slightly basad, sometimes apicad only.</paragraph>
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<paragraph id="A75736D5FFCA166C7984F80E240F7028" blockId="1.[189,776,554,2069]" lastBlockId="1.[812,1399,315,897]" pageId="1" pageNumber="17">
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Female tergite VIII moderate in length (moderately wide), with basolateral apodemes slightly less sclerotized toward tips; sternite VIII mostly well sclerotized laterobasally, densely setulose along apical margin to glabrous or nearly so. Tergite IX: laterotergite with apical margin rounded and densely setose, setae varying between species in length; gonosubcoxite with a short row of fairly short and dense setae along latero-apical margin; gonocoxite triangular, more or less crescent, with preapical nematiform seta, 2–3 outer (ventral) and one, sometimes two, inner (dorsal) ensiform setae. Reproductive tract rather characteristic of the genus:
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<emphasis id="959CEAC7FFCA166C7DFAFE5B245A73D9" box="[1180,1337,449,470]" italics="true" pageId="1" pageNumber="17">bursa copulatrix</emphasis>
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trapezoidal, with apex invaginated and sclerotized; spermatheca long, subdivided into long and narrow seminal canal and shorter subclavate receptacle; spermathecal gland duct long.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCA166C7A34FDB12435718E" pageId="1" pageNumber="17" type="discussion">
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<paragraph id="A75736D5FFCA166C7A34FDB12435718E" blockId="1.[812,1399,315,897]" pageId="1" pageNumber="17">
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COMMENTS. This subgenus includes over 30 described and a number of undescribed species from
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<collectingCountry id="DFFF7645FFCA166C7D95FDDC24537053" box="[1267,1328,582,604]" name="China" pageId="1" pageNumber="17">China</collectingCountry>
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. Three species described below are very similar to one another in body size and appearance and share similar male and female genitalia, but their some, derived, characters are unusual for the subgenus. These polythetic characters include some elytral setae missing, parascutellar or discal, or both, and the inclination of the prosternal process not apically beaded. Furthermore, the
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<emphasis id="959CEAC7FFCA166C7AB1FC9B228F7119" box="[983,1004,769,790]" italics="true" pageId="1" pageNumber="17">cx</emphasis>
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3 setae tend to be reduced from three to two in number. The posterior two setae are short or vestigial yet constantly present in
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<taxonomicName id="60E84D56FFCA166C7D4FFCAD25FD7144" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[1065,1182,822,844]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="glabellus" status="sp.n.">
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<emphasis id="959CEAC7FFCA166C7D4FFCAD25FD7144" box="[1065,1182,822,844]" italics="true" pageId="1" pageNumber="17">P. glabellus</emphasis>
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</taxonomicName>
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<emphasis id="959CEAC7FFCA166C7DC1FCAC25BB7144" bold="true" box="[1191,1240,822,843]" pageId="1" pageNumber="17">
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<taxonomicNameLabel id="8EAF57BCFFCA166C7DC1FCAC25BB7144" box="[1191,1240,822,843]" pageId="1" pageNumber="17" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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while the inner seta is absent from one (
|
||
<taxonomicName id="60E84D56FFCA166C7D54FCCB25D57169" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[1074,1206,849,870]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="unisetosus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFCA166C7D54FCCB25D57169" box="[1074,1206,849,870]" italics="true" pageId="1" pageNumber="17">P. unisetosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFCA166C7DA4FCCB25907169" bold="true" box="[1218,1267,849,870]" pageId="1" pageNumber="17">
|
||
<taxonomicNameLabel id="8EAF57BCFFCA166C7DA4FCCB25907169" box="[1218,1267,849,870]" pageId="1" pageNumber="17" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, one of
|
||
<specimenCount id="B1EEFD5CFFCA166C7C34FCCB22F3718E" count="2" pageId="1" pageNumber="17" type="generic">two specimens</specimenCount>
|
||
examined) or both (
|
||
<taxonomicName id="60E84D56FFCA166C7D3CFCF625BB718E" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[1114,1240,876,897]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="primitivus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFCA166C7D3CFCF625BB718E" box="[1114,1240,876,897]" italics="true" pageId="1" pageNumber="17">P. primitivus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFCA166C7DB9FCF12473718F" bold="true" box="[1247,1296,875,896]" pageId="1" pageNumber="17">
|
||
<taxonomicNameLabel id="8EAF57BCFFCA166C7DB9FCF12473718F" box="[1247,1296,875,896]" pageId="1" pageNumber="17" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) sides.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |