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<document id="318074863503761FB7284008BABA8B5D" ID-CLB-Dataset="25956" ID-DOI="10.11646/zootaxa.4658.1.7" ID-GBIF-Dataset="fc545a57-4242-4f8d-916d-b99fba5c6bca" ID-ISSN="1175-5326" ID-Zenodo-Dep="3373177" ID-ZooBank="1D952E2C-9DCD-4E8D-9F03-405B5415A0E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566365256020" checkinUser="plazi" docAuthor="Volynkin, Anton V., Černý, Karel, Saldaitis, Aidas &amp; Ivanova, Maria S." docDate="2019" docId="2F32EF65FFECFFC1AD972580FBA3FCF9" docLanguage="en" docName="zootaxa.4658.1.7.pdf" docOrigin="Zootaxa 4658 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyana titovi Volynkin &amp; Černý &amp; Saldaitis &amp; Ivanova 2019, sp. nov." docType="treatment" docUuid="4D9B45FE-C743-483A-99AF-EDC56939F7B6" docUuidSource="ZooBank" docVersion="9" lastPageNumber="160" masterDocId="D30B971DFFE9FFC4AD002519FFA0FFA0" masterDocTitle="On the correct identification of ‘ Cyana dohertyi (Elwes, 1890) ’ from northern Indochina and China, with descriptions of two new species and a new subspecies (Lepidoptera, Erebidae, Arctiinae)" masterLastPageNumber="167" masterPageNumber="155" pageNumber="160" updateTime="1698751969439" updateUser="ExternalLinkService">
<mods:mods id="FC910858C190BFB2AC04DDB29122D644" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F943003FE123416A7A118927A6BDD61">
<mods:title id="B06665745A8DC953ACEA727D22EE36FE">On the correct identification of ‘ Cyana dohertyi (Elwes, 1890) ’ from northern Indochina and China, with descriptions of two new species and a new subspecies (Lepidoptera, Erebidae, Arctiinae)</mods:title>
</mods:titleInfo>
<mods:name id="7F6C2478017AA1FE65C04C1A6A916052" type="personal">
<mods:role id="EB9F86FFA04B6778D3A6E2091A59AB6D">
<mods:roleTerm id="825B09FFF0F398ECAA8F81AAA4625330">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="68FFE808A1033F543B8AE1B46B8089EA">Volynkin, Anton V.</mods:namePart>
</mods:name>
<mods:name id="F7B946E25E4B613212391F18AF13A550" type="personal">
<mods:role id="BE240DBE243A82E60D64BAD87EDE60CD">
<mods:roleTerm id="C38299D060497AB5C80819CB09E48D7C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75B3AF66273EE1EA9DAE737B32976D3D">Černý, Karel</mods:namePart>
</mods:name>
<mods:name id="434CF5CE800B76B771FDD19091E564B4" type="personal">
<mods:role id="4E550494BC9BD4ABA8C4BD80F9EEA629">
<mods:roleTerm id="FE9788F8F3D303F468F47E6A1866F596">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B9152A5F38781B75A3F555D0FF6AC5C">Saldaitis, Aidas</mods:namePart>
</mods:name>
<mods:name id="3F6726FC6C4C8E328A64FA6DBF21B165" type="personal">
<mods:role id="05721285AC3FBF7F0C52B955F27430B8">
<mods:roleTerm id="503A905EF944122730A9F095EF157BB8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A40647AE7E54F2D3EC4A623499F2A19B">Ivanova, Maria S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5C4C4F7CCAD006D11EEFB9ABFF1AFDCF">text</mods:typeOfResource>
<mods:relatedItem id="B3ABE17215F7CB45F118E6932410305E" type="host">
<mods:titleInfo id="2F35245DD3F37D266DA47023DF20B284">
<mods:title id="1F373F89AD2B1E72D0F6B199E4FA8002">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="50D2383A93AED72FC48504095D2A1BEB">
<mods:date id="5CF132E285FB632BE31B364A15238DF5">2019</mods:date>
<mods:detail id="3A9191D88984B7876F4057E7D8FCD389" type="pubDate">
<mods:number id="DB5C13B48A928D861A84C8519022A60C">2019-08-21</mods:number>
</mods:detail>
<mods:detail id="A2FF83951A7C0A3619A7AFBA7B79663C" type="volume">
<mods:number id="7AD6C5DC78574CD086D5835459D6728B">4658</mods:number>
</mods:detail>
<mods:detail id="1CB80C4BEB8B47FE4075FAE246B82CC0" type="issue">
<mods:number id="F114C4743AF4EC630845E9C8DEAC796A">1</mods:number>
</mods:detail>
<mods:extent id="85A563EBF67B54FEA3AFC21A93CB91D7" unit="page">
<mods:start id="33638DB75C53586223AB2449242897A5">155</mods:start>
<mods:end id="038A2CDFDE409BC2478EC274D33D7716">167</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="44C90824AE91075A23920A7C5E954BC4">journal article</mods:classification>
<mods:identifier id="8AF7033247C245B3D1077E5AA543A95D" type="CLB-Dataset">25956</mods:identifier>
<mods:identifier id="04F4B9E894E0C9C2F36C5E6BA0F5E07F" type="DOI">10.11646/zootaxa.4658.1.7</mods:identifier>
<mods:identifier id="2D77627A3C21B1EC4AE34D909E9F95E6" type="GBIF-Dataset">fc545a57-4242-4f8d-916d-b99fba5c6bca</mods:identifier>
<mods:identifier id="DD00EB899063C06361CDE3709C1BC87F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AC291B85D9648B74A068B53A4F37B105" type="Zenodo-Dep">3373177</mods:identifier>
<mods:identifier id="67110ED9F5FBFAA6B6C8399CA201BC52" type="ZooBank">1D952E2C-9DCD-4E8D-9F03-405B5415A0E9</mods:identifier>
</mods:mods>
<treatment id="2F32EF65FFECFFC1AD972580FBA3FCF9" ID-GBIF-Taxon="159217307" LSID="urn:lsid:zoobank.org:act:4D9B45FE-C743-483A-99AF-EDC56939F7B6" httpUri="http://treatment.plazi.org/id/2F32EF65FFECFFC1AD972580FBA3FCF9" lastPageNumber="160" pageId="5" pageNumber="160">
<subSubSection id="EF810DF8FFECFFC1AD972580FD56FF14" box="[151,758,152,180]" pageId="5" pageNumber="160" type="nomenclature">
<paragraph id="A7245E73FFECFFC1AD972580FD56FF14" blockId="5.[151,971,152,245]" box="[151,758,152,180]" pageId="5" pageNumber="160">
<heading id="FC6CE91FFFECFFC1AD972580FD56FF14" bold="true" box="[151,758,152,180]" fontSize="11" level="1" pageId="5" pageNumber="160" reason="1">
<emphasis id="95EF8261FFECFFC1AD972580FD56FF14" bold="true" box="[151,758,152,180]" pageId="5" pageNumber="160">
<taxonomicName id="609B25F0FFECFFC1AD972580FD2CFF14" ID-CoL="BFNW5" authority="Volynkin, Cerny &amp; Ivanova" authorityName="Volynkin &amp; Černý &amp; Saldaitis &amp; Ivanova" authorityYear="2019" box="[151,652,152,180]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="titovi" status="sp. nov.">
<emphasis id="95EF8261FFECFFC1AD972580FE89FF13" bold="true" box="[151,297,153,179]" italics="true" pageId="5" pageNumber="160">Cyana titovi</emphasis>
Volynkin, Černý &amp; Ivanova
</taxonomicName>
,
<taxonomicNameLabel id="8EDC3F1AFFECFFC1AF992583FD56FF14" box="[665,758,154,180]" pageId="5" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EF810DF8FFECFFC1AD9725A4FC6BFF55" pageId="5" pageNumber="160" type="description">
<paragraph id="A7245E73FFECFFC1AD9725A4FE29FF77" blockId="5.[151,971,152,245]" box="[151,393,189,216]" pageId="5" pageNumber="160">
(
<figureCitation id="3FA042F6FFECFFC1AD9F25A4FEA4FF77" box="[159,260,189,216]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1911,1936]" captionTargetBox="[187,1404,181,1878]" captionTargetId="figure@3.[182,1404,181,1885]" captionTargetPageId="3" captionText="FIGURES 1–8. Cyana spp.: adults. 1, C. lada, holotype ♂, Vietnam (Coll. MWM/ZSM); 2, C. lada, paratype ♂, Vietnam (Coll. MWM/ZSM); 3, C. lada, paratype ♂, Vietnam (Coll. MWM/ZSM); 4, C. lada, paratype ♀, Vietnam (Coll. MWM/ZSM); 5, C. titovi, holotype ♂, Vietnam (©NHMUK); 6, C. titovi, paratype ♀, Vietnam (CAV); 7, C. aurantifascia, holotype ♂, Vietnam, (©NHMUK); 8, C. aurantifascia, paratype ♀, Vietnam (CAV)." httpUri="https://zenodo.org/record/3373179/files/figure.png" pageId="5" pageNumber="160">Figs 5, 6</figureCitation>
,
<figureCitation id="3FA042F6FFECFFC1AC1025A4FEF8FF77" box="[272,344,189,215]" captionStart="FIGURES 29–32" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[161,1435,181,1907]" captionTargetId="figure@8.[151,1435,181,1920]" captionTargetPageId="8" captionText="FIGURES 29–32. Cyana spp.: male genitalia. 29, C. titovi, holotype, Vietnam, slide AV2302♂ Volynkin; 30, C. titovi, paratype, Vietnam, slide AV1703♂ Volynkin; 31, C. aurantifascia, holotype, Vietnam, slide AV2299♂ Volynkin; 32, C. aurantifascia, paratype, Vietnam, slide AV2300♂ Volynkin." httpUri="https://zenodo.org/record/3373189/files/figure.png" pageId="5" pageNumber="160">29, 30</figureCitation>
,
<figureCitation id="3FA042F6FFECFFC1AC6425A4FE21FF77" box="[356,385,189,215]" captionStart="FIGURES 37–42" captionStartId="10.[151,264,1879,1904]" captionTargetBox="[156,1390,185,1827]" captionTargetId="figure@10.[151,1435,181,1853]" captionTargetPageId="10" captionText="FIGURES 37–42. Cyana spp.: female genitalia. 37, C. lada, paratype, Vietnam, slide ZSM Arct. 103/2017♀ Volynkin; 38, C. gazella, N India, slide MWM 34493 Volynkin; 39, C. titovi, paratype, Vietnam, slide AV2297♀ Volynkin; 40, C. aurantifascia, paratype, Vietnam, slide AV2301♀ Volynkin; 41, C. sikkimensis, NE India, slide MWM 34428 Volynkin; 42, C. abiens, China, slide ZSM Arct. 83/2017♀ Volynkin." httpUri="https://zenodo.org/record/3373193/files/figure.png" pageId="5" pageNumber="160">39</figureCitation>
)
</paragraph>
<paragraph id="A7245E73FFECFFC1AD9725C3FC6BFF55" blockId="5.[151,971,152,245]" box="[151,971,218,245]" pageId="5" pageNumber="160">
<uri id="D30A5271FFECFFC1AD9725C3FC6BFF55" box="[151,971,218,245]" pageId="5" pageNumber="160">
urn:lsid:zoobank.org:act:
<uuid id="D33D64A6FFECFFC1ACA625C3FC6BFF55" box="[422,971,218,245]" pageId="5" pageNumber="160">4D9B45FE-C743-483A-99AF-EDC56939F7B6</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="EF810DF8FFECFFC1AD97243AFBA3FCF9" pageId="5" pageNumber="160" type="materials_examined">
<paragraph id="A7245E73FFECFFC1AD97243AFC8BFEC1" blockId="5.[151,1437,290,857]" pageId="5" pageNumber="160">
<emphasis id="95EF8261FFECFFC1AD97243AFEE8FE9D" bold="true" box="[151,328,290,317]" pageId="5" pageNumber="160">Type material.</emphasis>
<materialsCitation id="17F3542EFFECFFC1AC51243AFC8BFEC1" ID-GBIF-Occurrence="2350486505" collectingDate="2015-08" collectionCode="NHMUK" collectorName="Le Luong Thanh" country="Vietnam" location="Ba Na Mt." pageId="5" pageNumber="160" specimenCount="2" specimenCount-male="2" stateProvince="Da Nang" typeStatus="holotype">
<emphasis id="95EF8261FFECFFC1AC51243AFE1EFE9D" bold="true" box="[337,446,291,317]" pageId="5" pageNumber="160">
<typeStatus id="7820E0D1FFECFFC1AC51243AFE1EFE9D" box="[337,446,291,317]" pageId="5" pageNumber="160" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="3FA042F6FFECFFC1ACCF243AFDBBFE9D" box="[463,539,290,317]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1911,1936]" captionTargetBox="[187,1404,181,1878]" captionTargetId="figure@3.[182,1404,181,1885]" captionTargetPageId="3" captionText="FIGURES 1–8. Cyana spp.: adults. 1, C. lada, holotype ♂, Vietnam (Coll. MWM/ZSM); 2, C. lada, paratype ♂, Vietnam (Coll. MWM/ZSM); 3, C. lada, paratype ♂, Vietnam (Coll. MWM/ZSM); 4, C. lada, paratype ♀, Vietnam (Coll. MWM/ZSM); 5, C. titovi, holotype ♂, Vietnam (©NHMUK); 6, C. titovi, paratype ♀, Vietnam (CAV); 7, C. aurantifascia, holotype ♂, Vietnam, (©NHMUK); 8, C. aurantifascia, paratype ♀, Vietnam (CAV)." httpUri="https://zenodo.org/record/3373179/files/figure.png" pageId="5" pageNumber="160">Figs 5</figureCitation>
,
<figureCitation id="3FA042F6FFECFFC1AF29243AFDE9FE9D" box="[553,585,291,317]" captionStart="FIGURES 29–32" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[161,1435,181,1907]" captionTargetId="figure@8.[151,1435,181,1920]" captionTargetPageId="8" captionText="FIGURES 29–32. Cyana spp.: male genitalia. 29, C. titovi, holotype, Vietnam, slide AV2302♂ Volynkin; 30, C. titovi, paratype, Vietnam, slide AV1703♂ Volynkin; 31, C. aurantifascia, holotype, Vietnam, slide AV2299♂ Volynkin; 32, C. aurantifascia, paratype, Vietnam, slide AV2300♂ Volynkin." httpUri="https://zenodo.org/record/3373189/files/figure.png" pageId="5" pageNumber="160">29</figureCitation>
):
<specimenCount id="B19D95FAFFECFFC1AF60243AFDD7FE9D" box="[608,631,291,317]" pageId="5" pageNumber="160" type="male">♂</specimenCount>
, “
<date id="D32578B3FFECFFC1AF8F243BFCA3FE9D" box="[655,771,290,317]" pageId="5" pageNumber="160" value="2015-08">
<collectingDate id="C361815BFFECFFC1AF8F243BFCA3FE9D" box="[655,771,290,317]" pageId="5" pageNumber="160" value="2015-08">VIII.2015</collectingDate>
</date>
, Central
<collectingCountry id="DF8C1EE3FFECFFC1AE6D243AFC6FFE9D" box="[877,975,291,317]" name="Vietnam" pageId="5" pageNumber="160">Vietnam</collectingCountry>
,
<collectingRegion id="655F9091FFECFFC1AEDC243AFB13FE9D" box="[988,1203,291,317]" country="Vietnam" name="Da Nang" pageId="5" pageNumber="160">Da Nang Province</collectingRegion>
,
<location id="A24408A8FFECFFC1A9C1243AFA9EFE9D" LSID="urn:lsid:plazi:treatment:2F32EF65FFECFFC1AD972580FBA3FCF9:A24408A8FFECFFC1A9C1243AFA9EFE9D" box="[1217,1342,291,317]" country="Vietnam" name="Ba Na Mt." pageId="5" pageNumber="160" stateProvince="Da Nang">Ba Na Mt.</location>
, leg.
<collectorName id="0A6E3BA5FFECFFC1A87F243AFE94FEC1" pageId="5" pageNumber="160">Le Luong Thanh</collectorName>
”, slide AV2302
<specimenCount id="B19D95FAFFECFFC1ACE3245EFE55FEC1" box="[483,501,327,353]" pageId="5" pageNumber="160" type="male">♂</specimenCount>
Volynkin (Coll.
<collectionCode id="C18AC6B6FFECFFC1AFB0245EFCBFFEC1" box="[688,799,327,353]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="5" pageNumber="160" type="Museum">NHMUK</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="A7245E73FFECFFC1ADC72472FC6BFE6D" blockId="5.[151,1437,290,857]" pageId="5" pageNumber="160">
<materialsCitation id="17F3542EFFECFFC1ADC72472FBACFE25" ID-GBIF-Occurrence="2416598479" box="[199,1036,363,389]" collectingDate="2015-09" collectingDateMax="2016-08" collectingDateMin="2015-09" collectionCode="CAV" collectorName="Coll. &amp; Le Luong Thanh" country="Vietnam" county="Central" elevation="1400" location="Coll." municipality="Annamite Mts." pageId="5" pageNumber="160" specimenCount="14" specimenCount-female="9" specimenCount-male="5" stateProvince="Thua Thien-Hue Province" typeStatus="paratype">
<emphasis id="95EF8261FFECFFC1ADC72472FE9FFE25" bold="true" box="[199,319,363,389]" pageId="5" pageNumber="160">
<typeStatus id="7820E0D1FFECFFC1ADC72472FE9FFE25" box="[199,319,363,389]" pageId="5" pageNumber="160" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="B19D95FAFFECFFC1AC4D2472FED9FE25" box="[333,377,363,389]" pageId="5" pageNumber="160" type="male">1 ♂</specimenCount>
, same data as in the holotype, slide AV2297
<specimenCount id="B19D95FAFFECFFC1AE6D2472FCDAFE25" box="[877,890,363,389]" pageId="5" pageNumber="160" type="female">♀</specimenCount>
(
<location id="A24408A8FFECFFC1AE892472FC61FE25" LSID="urn:lsid:plazi:treatment:2F32EF65FFECFFC1AD972580FBA3FCF9:A24408A8FFECFFC1AE892472FC61FE25" box="[905,961,363,389]" country="Vietnam" county="Central" municipality="Annamite Mts." name="Coll." pageId="5" pageNumber="160" stateProvince="Thua Thien-Hue Province">Coll.</location>
<collectionCode id="C18AC6B6FFECFFC1AEC82472FBA5FE25" box="[968,1029,363,389]" pageId="5" pageNumber="160">CAV</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="17F3542EFFECFFC1A9172472FD4CFE09" ID-GBIF-Occurrence="2416598478" collectingDate="2015-09" collectingDateMax="2016-08" collectingDateMin="2015-09" collectionCode="CAV" collectorName="Coll. &amp; Le Luong Thanh" country="Vietnam" county="Central" elevation="1400" location="Coll." municipality="Annamite Mts." pageId="5" pageNumber="160" specimenCount="14" specimenCount-female="9" specimenCount-male="5" stateProvince="Thua Thien-Hue Province" typeStatus="paratype">
<specimenCount id="B19D95FAFFECFFC1A9172472FBE3FE25" box="[1047,1091,363,389]" pageId="5" pageNumber="160" type="male">1 ♂</specimenCount>
,
<specimenCount id="B19D95FAFFECFFC1A94E2472FBD8FE25" box="[1102,1144,363,389]" pageId="5" pageNumber="160" type="female">8 ♀</specimenCount>
, same locality and collec- tor,
<date id="D32578B3FFECFFC1ADC32497FE83FE09" box="[195,291,398,425]" pageId="5" pageNumber="160" value="2015-09">
<collectingDate id="C361815BFFECFFC1ADC32497FE83FE09" box="[195,291,398,425]" pageId="5" pageNumber="160" value="2015-09">IX.2015</collectingDate>
</date>
, slides AV1703
<specimenCount id="B19D95FAFFECFFC1ACD52496FE48FE09" box="[469,488,399,425]" pageId="5" pageNumber="160" type="male">♂</specimenCount>
,
<location id="A24408A8FFECFFC1ACF32496FDF9FE09" LSID="urn:lsid:plazi:treatment:2F32EF65FFECFFC1AD972580FBA3FCF9:A24408A8FFECFFC1ACF32496FDF9FE09" box="[499,601,399,425]" country="Vietnam" county="Central" municipality="Annamite Mts." name="Volynkin" pageId="5" pageNumber="160" stateProvince="Thua Thien-Hue Province">Volynkin</location>
(
<collectorName id="0A6E3BA5FFECFFC1AF682496FD01FE09" box="[616,673,399,425]" pageId="5" pageNumber="160">Coll.</collectorName>
<collectionCode id="C18AC6B6FFECFFC1AFA72496FD44FE09" box="[679,740,399,425]" pageId="5" pageNumber="160">CAV</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="17F3542EFFECFFC1AFF72496FC6BFE6D" ID-GBIF-Occurrence="2350486509" collectingDate="2015-09" collectingDateMax="2016-08" collectingDateMin="2015-09" collectionCode="CAV" collectorName="Coll. &amp; Le Luong Thanh" country="Vietnam" county="Central" elevation="1400" location="Coll." municipality="Annamite Mts." pageId="5" pageNumber="160" specimenCount="14" specimenCount-female="9" specimenCount-male="5" stateProvince="Thua Thien-Hue Province" typeStatus="paratype">
<specimenCount id="B19D95FAFFECFFC1AFF72496FC83FE09" box="[759,803,399,425]" pageId="5" pageNumber="160" type="male">2 ♂</specimenCount>
,
<date id="D32578B3FFECFFC1AE2F2496FC01FE09" box="[815,929,399,425]" pageId="5" pageNumber="160" value="2016-08">
<collectingDate id="C361815BFFECFFC1AE2F2496FC01FE09" box="[815,929,399,425]" pageId="5" pageNumber="160" value="2016-08">VIII.2016</collectingDate>
</date>
,
<collectingCounty id="4E4526FFFFECFFC1AEAC2496FC5EFE09" box="[940,1022,399,425]" pageId="5" pageNumber="160">Central</collectingCounty>
<collectingCountry id="DF8C1EE3FFECFFC1A9052496FBC7FE09" box="[1029,1127,399,425]" name="Vietnam" pageId="5" pageNumber="160">Vietnam</collectingCountry>
,
<collectingRegion id="655F9091FFECFFC1A9712496FA38FE09" box="[1137,1432,399,425]" country="Vietnam" name="Thua Thien-Hue" pageId="5" pageNumber="160">Thua Thien-Hue Province</collectingRegion>
,
<collectingMunicipality id="4740C409FFECFFC1AD9724AAFEE2FE6D" box="[151,322,435,461]" pageId="5" pageNumber="160">Annamite Mts.</collectingMunicipality>
,
<location id="A24408A8FFECFFC1AC4D24AAFE44FE6D" LSID="urn:lsid:plazi:treatment:2F32EF65FFECFFC1AD972580FBA3FCF9:A24408A8FFECFFC1AC4D24AAFE44FE6D" box="[333,484,435,461]" country="Vietnam" county="Central" municipality="Annamite Mts." name="Bach Ma Mt." pageId="5" pageNumber="160" stateProvince="Thua Thien-Hue Province">Bach Ma Mt.</location>
,
<quantity id="6063F396FFECFFC1ACF024AAFDE0FE6D" box="[496,576,435,461]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="160" unit="m" value="1400.0">
<elevation id="2CB6B940FFECFFC1ACF024AAFDE0FE6D" box="[496,576,435,461]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="160" unit="m" value="1400.0">1400m</elevation>
</quantity>
, leg.
<collectorName id="0A6E3BA5FFECFFC1AF7C24AAFC98FE6D" box="[636,824,435,461]" pageId="5" pageNumber="160">Le Luong Thanh</collectorName>
(
<collectorName id="0A6E3BA5FFECFFC1AE4724AAFCDFFE6D" box="[839,895,435,461]" pageId="5" pageNumber="160">Coll.</collectorName>
<collectionCode id="C18AC6B6FFECFFC1AE8624AAFC63FE6D" box="[902,963,435,461]" pageId="5" pageNumber="160">CAV</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph id="A7245E73FFECFFC1AECA24AAFC6EFE6D" blockId="5.[151,1437,290,857]" box="[970,974,435,461]" pageId="5" pageNumber="160">.</paragraph>
<paragraph id="A7245E73FFECFFC1ADC724CFFA2AFD4D" blockId="5.[151,1437,290,857]" pageId="5" pageNumber="160">
<emphasis id="95EF8261FFECFFC1ADC724CFFE98FE51" bold="true" box="[199,312,470,497]" pageId="5" pageNumber="160">Diagnosis</emphasis>
. Forewing length is 14–14.5 mm in males (14.5 mm in the
<typeStatus id="7820E0D1FFECFFC1AECB24CEFB8CFE52" box="[971,1068,471,498]" pageId="5" pageNumber="160" type="holotype">holotype</typeStatus>
) and 17.5 mm in females.
<taxonomicName id="609B25F0FFECFFC1A85424CEFF6FFDB5" authorityName="Volynkin &amp; Černý &amp; Saldaitis &amp; Ivanova" authorityYear="2019" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="titovi">
<emphasis id="95EF8261FFECFFC1A85424CEFF6FFDB5" italics="true" pageId="5" pageNumber="160">Cyana titovi</emphasis>
</taxonomicName>
(
<figureCitation id="3FA042F6FFECFFC1ADDE24E2FE9FFDB6" box="[222,319,506,534]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1911,1936]" captionTargetBox="[187,1404,181,1878]" captionTargetId="figure@3.[182,1404,181,1885]" captionTargetPageId="3" captionText="FIGURES 1–8. Cyana spp.: adults. 1, C. lada, holotype ♂, Vietnam (Coll. MWM/ZSM); 2, C. lada, paratype ♂, Vietnam (Coll. MWM/ZSM); 3, C. lada, paratype ♂, Vietnam (Coll. MWM/ZSM); 4, C. lada, paratype ♀, Vietnam (Coll. MWM/ZSM); 5, C. titovi, holotype ♂, Vietnam (©NHMUK); 6, C. titovi, paratype ♀, Vietnam (CAV); 7, C. aurantifascia, holotype ♂, Vietnam, (©NHMUK); 8, C. aurantifascia, paratype ♀, Vietnam (CAV)." httpUri="https://zenodo.org/record/3373179/files/figure.png" pageId="5" pageNumber="160">Figs 5, 6</figureCitation>
) is similar externally to
<taxonomicName id="609B25F0FFECFFC1AF4D24E2FD5FFDB5" authorityName="Volynkin, Cerny &amp; Dubatolov" authorityYear="2017" box="[589,767,507,533]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="aurantifascia">
<emphasis id="95EF8261FFECFFC1AF4D24E2FD5FFDB5" box="[589,767,507,533]" italics="true" pageId="5" pageNumber="160">C. aurantifascia</emphasis>
</taxonomicName>
(
<figureCitation id="3FA042F6FFECFFC1AE0E24E2FCD0FDB5" box="[782,880,507,533]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1911,1936]" captionTargetBox="[187,1404,181,1878]" captionTargetId="figure@3.[182,1404,181,1885]" captionTargetPageId="3" captionText="FIGURES 1–8. Cyana spp.: adults. 1, C. lada, holotype ♂, Vietnam (Coll. MWM/ZSM); 2, C. lada, paratype ♂, Vietnam (Coll. MWM/ZSM); 3, C. lada, paratype ♂, Vietnam (Coll. MWM/ZSM); 4, C. lada, paratype ♀, Vietnam (Coll. MWM/ZSM); 5, C. titovi, holotype ♂, Vietnam (©NHMUK); 6, C. titovi, paratype ♀, Vietnam (CAV); 7, C. aurantifascia, holotype ♂, Vietnam, (©NHMUK); 8, C. aurantifascia, paratype ♀, Vietnam (CAV)." httpUri="https://zenodo.org/record/3373179/files/figure.png" pageId="5" pageNumber="160">Figs 7, 8</figureCitation>
), but differs clearly by its smaller size and smaller black spots on forewing. The male genital capsule of the new species (
<figureCitation id="3FA042F6FFECFFC1AE912706FBADFD99" box="[913,1037,543,569]" captionStart="FIGURES 29–32" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[161,1435,181,1907]" captionTargetId="figure@8.[151,1435,181,1920]" captionTargetPageId="8" captionText="FIGURES 29–32. Cyana spp.: male genitalia. 29, C. titovi, holotype, Vietnam, slide AV2302♂ Volynkin; 30, C. titovi, paratype, Vietnam, slide AV1703♂ Volynkin; 31, C. aurantifascia, holotype, Vietnam, slide AV2299♂ Volynkin; 32, C. aurantifascia, paratype, Vietnam, slide AV2300♂ Volynkin." httpUri="https://zenodo.org/record/3373189/files/figure.png" pageId="5" pageNumber="160">Figs 29, 30</figureCitation>
) is similar to that of
<taxonomicName id="609B25F0FFECFFC1A9EB2706FA3CFD99" authorityName="Volynkin, Cerny &amp; Dubatolov" authorityYear="2017" box="[1259,1436,543,569]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="aurantifascia">
<emphasis id="95EF8261FFECFFC1A9EB2706FA3CFD99" box="[1259,1436,543,569]" italics="true" pageId="5" pageNumber="160">C. aurantifascia</emphasis>
</taxonomicName>
(
<figureCitation id="3FA042F6FFECFFC1AD9F275AFEBBFDFD" box="[159,283,579,605]" captionStart="FIGURES 29–32" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[161,1435,181,1907]" captionTargetId="figure@8.[151,1435,181,1920]" captionTargetPageId="8" captionText="FIGURES 29–32. Cyana spp.: male genitalia. 29, C. titovi, holotype, Vietnam, slide AV2302♂ Volynkin; 30, C. titovi, paratype, Vietnam, slide AV1703♂ Volynkin; 31, C. aurantifascia, holotype, Vietnam, slide AV2299♂ Volynkin; 32, C. aurantifascia, paratype, Vietnam, slide AV2300♂ Volynkin." httpUri="https://zenodo.org/record/3373189/files/figure.png" pageId="5" pageNumber="160">Figs 31, 32</figureCitation>
), but has slightly more convex costal margin of valva. The vesica of
<taxonomicName id="609B25F0FFECFFC1A902275AFBF9FDFD" authorityName="Volynkin &amp; Černý &amp; Saldaitis &amp; Ivanova" authorityYear="2019" box="[1026,1113,579,605]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="titovi">
<emphasis id="95EF8261FFECFFC1A902275AFBF9FDFD" box="[1026,1113,579,605]" italics="true" pageId="5" pageNumber="160">C. titovi</emphasis>
</taxonomicName>
differs from that of
<taxonomicName id="609B25F0FFECFFC1A832275AFF47FD21" authorityName="Volynkin, Cerny &amp; Dubatolov" authorityYear="2017" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="aurantifascia">
<emphasis id="95EF8261FFECFFC1A832275AFF47FD21" italics="true" pageId="5" pageNumber="160">C. aurantifascia</emphasis>
</taxonomicName>
by the much smaller largest diverticulum, the broader 1
<superScript id="50EEF33BFFECFFC1AE49277FFCF4FDD4" attach="left" box="[841,852,614,628]" fontSize="6" pageId="5" pageNumber="160">st</superScript>
distal diverticulum bearing a broader cluster of much more robust spinules, and the smaller 2
<superScript id="50EEF33BFFECFFC1AF472790FDF7FD37" attach="left" box="[583,599,649,663]" fontSize="6" pageId="5" pageNumber="160">nd</superScript>
distal diverticulum. The female genitalia of
<taxonomicName id="609B25F0FFECFFC1A9412792FB39FD05" authorityName="Volynkin &amp; Černý &amp; Saldaitis &amp; Ivanova" authorityYear="2019" box="[1089,1177,651,677]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="titovi">
<emphasis id="95EF8261FFECFFC1A9412792FB39FD05" box="[1089,1177,651,677]" italics="true" pageId="5" pageNumber="160">C. titovi</emphasis>
</taxonomicName>
(
<figureCitation id="3FA042F6FFECFFC1A9A92792FB59FD05" box="[1193,1273,651,677]" captionStart="FIGURES 37–42" captionStartId="10.[151,264,1879,1904]" captionTargetBox="[156,1390,185,1827]" captionTargetId="figure@10.[151,1435,181,1853]" captionTargetPageId="10" captionText="FIGURES 37–42. Cyana spp.: female genitalia. 37, C. lada, paratype, Vietnam, slide ZSM Arct. 103/2017♀ Volynkin; 38, C. gazella, N India, slide MWM 34493 Volynkin; 39, C. titovi, paratype, Vietnam, slide AV2297♀ Volynkin; 40, C. aurantifascia, paratype, Vietnam, slide AV2301♀ Volynkin; 41, C. sikkimensis, NE India, slide MWM 34428 Volynkin; 42, C. abiens, China, slide ZSM Arct. 83/2017♀ Volynkin." httpUri="https://zenodo.org/record/3373193/files/figure.png" pageId="5" pageNumber="160">Fig. 39</figureCitation>
) can be distinguished from those of
<taxonomicName id="609B25F0FFECFFC1AC8B27B6FD9CFD69" authorityName="Volynkin, Cerny &amp; Dubatolov" authorityYear="2017" box="[395,572,687,713]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="aurantifascia">
<emphasis id="95EF8261FFECFFC1AC8B27B6FD9CFD69" box="[395,572,687,713]" italics="true" pageId="5" pageNumber="160">C. aurantifascia</emphasis>
</taxonomicName>
(
<figureCitation id="3FA042F6FFECFFC1AF4A27B6FD39FD69" box="[586,665,687,713]" captionStart="FIGURES 37–42" captionStartId="10.[151,264,1879,1904]" captionTargetBox="[156,1390,185,1827]" captionTargetId="figure@10.[151,1435,181,1853]" captionTargetPageId="10" captionText="FIGURES 37–42. Cyana spp.: female genitalia. 37, C. lada, paratype, Vietnam, slide ZSM Arct. 103/2017♀ Volynkin; 38, C. gazella, N India, slide MWM 34493 Volynkin; 39, C. titovi, paratype, Vietnam, slide AV2297♀ Volynkin; 40, C. aurantifascia, paratype, Vietnam, slide AV2301♀ Volynkin; 41, C. sikkimensis, NE India, slide MWM 34428 Volynkin; 42, C. abiens, China, slide ZSM Arct. 83/2017♀ Volynkin." httpUri="https://zenodo.org/record/3373193/files/figure.png" pageId="5" pageNumber="160">Fig. 40</figureCitation>
) by the slightly longer ductus bursae, the larger signum bursae, and the narrower and twisted distal membranous section of appendix bursae (that is broad and globular in
<taxonomicName id="609B25F0FFECFFC1A9C827CAFADAFD4D" authorityName="Volynkin, Cerny &amp; Dubatolov" authorityYear="2017" box="[1224,1402,723,749]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="aurantifascia">
<emphasis id="95EF8261FFECFFC1A9C827CAFADAFD4D" box="[1224,1402,723,749]" italics="true" pageId="5" pageNumber="160">C. aurantifascia</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="A7245E73FFECFFC1ADC727EFFCAFFCB2" blockId="5.[151,1437,290,857]" box="[199,783,758,786]" pageId="5" pageNumber="160">
<emphasis id="95EF8261FFECFFC1ADC727EFFEF9FCB1" bold="true" box="[199,345,758,785]" pageId="5" pageNumber="160">Distribution</emphasis>
. Central
<collectingCountry id="DF8C1EE3FFECFFC1ACC027EEFDBFFCB1" box="[448,543,759,785]" name="Vietnam" pageId="5" pageNumber="160">Vietnam</collectingCountry>
(
<collectingRegion id="655F9091FFECFFC1AF2E27EEFCA4FCB2" box="[558,772,759,786]" country="Vietnam" name="Da Nang" pageId="5" pageNumber="160">Da Nang Province</collectingRegion>
).
</paragraph>
<paragraph id="A7245E73FFECFFC1ADC72602FBA3FCF9" blockId="5.[151,1437,290,857]" pageId="5" pageNumber="160">
<emphasis id="95EF8261FFECFFC1ADC72602FEE8FC95" bold="true" box="[199,328,795,821]" pageId="5" pageNumber="160">Etymology</emphasis>
. The species name is dedicated to Dr. Sergey V. Titov (Pavlodar,
<collectingCountry id="DF8C1EE3FFECFFC1A9252602FB0CFC96" box="[1061,1196,795,822]" name="Kazakhstan" pageId="5" pageNumber="160">Kazakhstan</collectingCountry>
), an expert in Central Asian Noctuoidea and friend of senior and fourth authors of the present paper.
</paragraph>
</subSubSection>
</treatment>
</document>
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