269 lines
37 KiB
XML
269 lines
37 KiB
XML
<document id="2064003351FEAFEE380CDDFCDBBC461A" ID="10.11646/zootaxa.4216.6.3" ID-CLB-Dataset="37326" ID-DOI="10.5281/zenodo.242442" ID-GBIF-Dataset="cd7b958e-d906-4cac-8582-f412619f550d" ID-ISSN="1175-5326" ID-Zenodo-Dep="242442" ID-ZooBank="099626D3-839B-438D-94B8-C89A5227A574" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484320646478" checkinUser="plazi" docAuthor="Ojeda-Escoto, Juan Carlos, Villalobos, José Luis & Álvarez, Fernando" docDate="2017" docId="2F0387BA9B18FFEE86F7FF1EFBE417D0" docLanguage="en" docName="zootaxa.4216.6.3.pdf" docOrigin="Zootaxa 4216 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudothelphusa acahuizotla Ojeda-Escoto, Villalobos & Álvarez, 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="564" masterDocId="D33AFFC29B1AFFEB8660FFE1FF8C1728" masterDocTitle="Three new species of freshwater crabs of the genus Pseudothelphusa (De Saussure, 1857) (Crustacea: Decapoda: Pseudothelphusidae) from Mexico" masterLastPageNumber="571" masterPageNumber="559" pageNumber="561" updateTime="1698435521958" updateUser="ExternalLinkService">
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<mods:title id="98386FCA0D70A397B72C7B081B40C0AB">Three new species of freshwater crabs of the genus Pseudothelphusa (De Saussure, 1857) (Crustacea: Decapoda: Pseudothelphusidae) from Mexico</mods:title>
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<mods:namePart id="CDB501B44723E32BD4631F0E18CB507E">Ojeda-Escoto, Juan Carlos</mods:namePart>
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<mods:namePart id="0C0FED33A7B10B7F140AD8C7EAC36022">Villalobos, José Luis</mods:namePart>
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<mods:namePart id="839EFA8099F8FDE4EFC5456DB2192911">Álvarez, Fernando</mods:namePart>
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<mods:title id="278F7E7427B8FE62886D45579C9CF08B">Zootaxa</mods:title>
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<mods:date id="3B2312F4D698C6BADF09AB8E2C22369B">2017</mods:date>
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<treatment id="2F0387BA9B18FFEE86F7FF1EFBE417D0" ID-DOI="http://doi.org/10.5281/zenodo.6040723" ID-GBIF-Taxon="127378521" ID-Zenodo-Dep="6040723" LSID="urn:lsid:plazi:treatment:2F0387BA9B18FFEE86F7FF1EFBE417D0" httpUri="http://treatment.plazi.org/id/2F0387BA9B18FFEE86F7FF1EFBE417D0" lastPageId="5" lastPageNumber="564" pageId="2" pageNumber="561">
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<paragraph id="A71536AC9B18FFE986F7FF1EFDCE1632" blockId="2.[151,578,255,315]" box="[151,578,255,282]" pageId="2" pageNumber="561">
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<heading id="FC5D81C09B18FFE986F7FF1EFDCE1632" bold="true" box="[151,578,255,282]" fontSize="11" level="1" pageId="2" pageNumber="561" reason="1">
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<emphasis id="95DEEABE9B18FFE986F7FF1EFDCE1632" bold="true" box="[151,578,255,282]" pageId="2" pageNumber="561">
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<taxonomicName id="60AA4D2F9B18FFE986F7FF1EFE751631" ID-CoL="4PD83" box="[151,505,255,281]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="acahuizotla" status="sp. nov.">
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<emphasis id="95DEEABE9B18FFE986F7FF1EFE751631" bold="true" box="[151,505,255,281]" italics="true" pageId="2" pageNumber="561">Pseudothelphusa acahuizotla</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8EED57C59B18FFE98460FEE1FDCE1632" box="[512,578,256,282]" pageId="2" pageNumber="561" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="A71536AC9B18FFE986F7FEC3FE9D1613" blockId="2.[151,578,255,315]" box="[151,273,290,315]" pageId="2" pageNumber="561">
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(
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<figureCitation id="3F912A299B18FFE986FFFEC3FF621613" box="[159,238,290,315]" captionStart="FIGURE 2" captionStartId="3.[151,250,1526,1548]" captionTargetBox="[195,1391,194,1505]" captionTargetId="figure@3.[195,1392,193,1505]" captionTargetPageId="3" captionText="FIGURE 2. Pseudothelphusa acahuizotla n. sp., male holotype, CNCR 31708: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242444/files/figure.png" pageId="2" pageNumber="561">Figs. 2</figureCitation>
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,
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<figureCitation id="3F912A299B18FFE9869AFEC2FE841613" box="[250,264,291,315]" captionStart="FIGURE 3" captionStartId="4.[151,250,1933,1955]" captionTargetBox="[153,1436,373,1911]" captionTargetId="figure@4.[151,1436,369,1912]" captionTargetPageId="4" captionText="FIGURE 3. Pseudothelphusa acahuizotla n. sp., male holotype CNCR 31708, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudomarginal projection; mp, mesial process; ms, marginal suture; tps, terminal pore setae." httpUri="https://zenodo.org/record/242445/files/figure.png" pageId="2" pageNumber="561">3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="EFB065279B18FFE986F7FE8BFE6416E3" pageId="2" pageNumber="561" type="materials_examined">
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<paragraph id="A71536AC9B18FFE986F7FE8BFE6416E3" blockId="2.[151,1437,362,1971]" pageId="2" pageNumber="561">
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<materialsCitation id="17C23CF19B18FFE986F7FE8BFE6416E3" ID-GBIF-Occurrence="1424565296" collectingDate="2014-07-26" collectionCode="CNCR" collectorName="S. M. Martinez-Zam" country="Mexico" county="Bravo" elevation="820" latitude="17.360556" location="Municipality" longLatPrecision="21" longitude="-99.467224" municipality="Chilpancingo" pageId="2" pageNumber="561" specimenCode="CNCR 31708" specimenCount="2" stateProvince="Guerrero" typeStatus="holotype">
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<emphasis id="95DEEABE9B18FFE986F7FE8BFEC916AB" bold="true" box="[151,325,362,387]" pageId="2" pageNumber="561">
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<typeStatus id="7811880E9B18FFE986F7FE8BFF5E16AB" box="[151,210,362,387]" pageId="2" pageNumber="561">Type</typeStatus>
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material.
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</emphasis>
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Male
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<typeStatus id="7811880E9B18FFE987EEFE8BFE7F16AB" box="[398,499,362,387]" pageId="2" pageNumber="561" type="holotype">holotype</typeStatus>
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, cl 22.0 mm, cb
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<quantity id="60529B499B18FFE984CFFE8BFC9B16AA" box="[687,791,362,387]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="2" pageNumber="561" unit="mm" value="35.8">35.8 mm</quantity>
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; small stream in
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<taxonomicName id="60AA4D2F9B18FFE985B7FE8BFBD216AB" box="[983,1118,362,387]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="564" phylum="Arthropoda" rank="species" species="acahuizotla" status="sp. nov.">Acahuizotla</taxonomicName>
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(
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<geoCoordinate id="C29E506B9B18FFE9820EFE8AFB7616AA" box="[1134,1274,363,387]" direction="north" orientation="latitude" pageId="2" pageNumber="561" precision="15" value="17.360556">17°21'38" N</geoCoordinate>
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,
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<geoCoordinate id="C29E506B9B18FFE98366FE8AFA1416AA" box="[1286,1432,363,387]" direction="west" orientation="longitude" pageId="2" pageNumber="561" precision="15" value="-99.467224">99°28'02" W</geoCoordinate>
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;
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<quantity id="60529B499B18FFE986F7FE6EFF6F168E" box="[151,227,399,423]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="561" unit="m" value="820.0">
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<elevation id="2C87D19F9B18FFE986F7FE6EFF6F168E" box="[151,227,399,423]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="561" unit="m" value="820.0">820 m</elevation>
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</quantity>
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asl),
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<location id="A27560779B18FFE9874AFE6FFE37168F" LSID="urn:lsid:plazi:treatment:2F0387BA9B18FFEE86F7FF1EFBE417D0:A27560779B18FFE9874AFE6FFE37168F" box="[298,443,398,423]" country="Mexico" county="Bravo" latitude="17.360556" longLatPrecision="21" longitude="-99.467224" municipality="Chilpancingo" name="Municipality" pageId="2" pageNumber="561" stateProvince="Guerrero">Municipality</location>
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of
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<collectingMunicipality id="4771ACD69B18FFE9878AFE6FFD0D168F" box="[490,641,398,423]" pageId="2" pageNumber="561">Chilpancingo</collectingMunicipality>
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de los
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<collectingCounty id="4E744E209B18FFE98480FE6EFCAB168E" box="[736,807,399,422]" pageId="2" pageNumber="561">Bravo</collectingCounty>
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, state of
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<collectingRegion id="656EF84E9B18FFE985F9FE6EFB8D168F" box="[921,1025,399,423]" country="Mexico" name="Guerrero" pageId="2" pageNumber="561">Guerrero</collectingRegion>
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,
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<collectingCountry id="DFBD763C9B18FFE98271FE6FFBE6168F" box="[1041,1130,398,423]" name="Mexico" pageId="2" pageNumber="561">Mexico</collectingCountry>
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;
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<date id="D314106C9B18FFE9821AFE6FFA94168F" box="[1146,1304,398,423]" pageId="2" pageNumber="561" value="2014-07-26">
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<collectingDate id="C350E9849B18FFE9821AFE6FFA94168F" box="[1146,1304,398,423]" pageId="2" pageNumber="561" value="2014-07-26">26 July 2014</collectingDate>
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</date>
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; coll.
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<collectorName id="0A5F537A9B18FFE98306FE6EFEB116E3" pageId="2" pageNumber="561">S.M. Martínez-Zam</collectorName>
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||
;
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<specimenCode id="F70C9ED79B18FFE98727FE52FE6916E3" box="[327,485,435,459]" collectionCode="CNCR" pageId="2" pageNumber="561">
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<collectionCode id="C1BBAE699B18FFE98727FE52FE1F16E3" box="[327,403,435,459]" pageId="2" pageNumber="561">CNCR</collectionCode>
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31708
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</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="EFB065279B18FFE886A7FE37FC191158" lastPageId="3" lastPageNumber="562" pageId="2" pageNumber="561" type="description">
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<paragraph id="A71536AC9B18FFE986A7FE37FA1D1453" blockId="2.[151,1437,362,1971]" pageId="2" pageNumber="561">
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<emphasis id="95DEEABE9B18FFE986A7FE37FEDD16C7" bold="true" box="[199,337,470,495]" pageId="2" pageNumber="561">Description</emphasis>
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. Dorsal surface of carapace flat, punctate, with low tubercles on area behind orbits, adjacent to anterolateral margin at branchial region (
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<figureCitation id="3F912A299B18FFE98400FE1BFD2F153B" box="[608,675,506,531]" captionStart="FIGURE 2" captionStartId="3.[151,250,1526,1548]" captionTargetBox="[195,1391,194,1505]" captionTargetId="figure@3.[195,1392,193,1505]" captionTargetPageId="3" captionText="FIGURE 2. Pseudothelphusa acahuizotla n. sp., male holotype, CNCR 31708: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242444/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
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A). Gastric region inflated, anterior portion slightly curving to reach superior frontal border, marked by acute frontal bend of carapace, inclined towards median groove, with low, rounded tubercles. Front vertically deflexed, smooth, bilobed (
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<figureCitation id="3F912A299B18FFE98508FDA3FC231573" box="[872,943,578,603]" captionStart="FIGURE 2" captionStartId="3.[151,250,1526,1548]" captionTargetBox="[195,1391,194,1505]" captionTargetId="figure@3.[195,1392,193,1505]" captionTargetPageId="3" captionText="FIGURE 2. Pseudothelphusa acahuizotla n. sp., male holotype, CNCR 31708: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242444/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
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D); inferior frontal border visible in dorsal view; ventrally slightly projected, sinuous, with evident middle concavity, slight convexity at antennular fossae level. Postfrontal lobes well formed, without anterior depression (
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<figureCitation id="3F912A299B18FFE985C3FD6BFC60158B" box="[931,1004,650,675]" captionStart="FIGURE 2" captionStartId="3.[151,250,1526,1548]" captionTargetBox="[195,1391,194,1505]" captionTargetId="figure@3.[195,1392,193,1505]" captionTargetPageId="3" captionText="FIGURE 2. Pseudothelphusa acahuizotla n. sp., male holotype, CNCR 31708: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242444/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
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D). Frontal portion between superior frontal border and postfrontal lobes flat. Median frontal groove narrow, deep, anteriorly finished in V-shaped notch, dividing superior frontal border; posteriorly separating postfrontal lobes, becoming obsolete little after. Dorsal area behind orbits concave. Cervical grooves well marked, wide, curved, ending before reaching anterolateral margin. Regions moderately indicated, gastric, branchial swollen, cardiac, intestinal depressed. Anterolateral margin prominent in lateral view, serrated, with subacute granules from cervical groove to middle portion of branchial region; area between orbit and cervical grove with 8–10 granules. Posterior margin of carapace straight (
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<figureCitation id="3F912A299B18FFE9834AFC83FAE11453" box="[1322,1389,866,891]" captionStart="FIGURE 2" captionStartId="3.[151,250,1526,1548]" captionTargetBox="[195,1391,194,1505]" captionTargetId="figure@3.[195,1392,193,1505]" captionTargetPageId="3" captionText="FIGURE 2. Pseudothelphusa acahuizotla n. sp., male holotype, CNCR 31708: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242444/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
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A).
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</paragraph>
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<paragraph id="A71536AC9B18FFE986A7FC67FDFC13CB" blockId="2.[151,1437,362,1971]" pageId="2" pageNumber="561">
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Orbits with external angle faintly developed, granulated, with rounded basal notch; internal angle with high border. Basal article of antenna separated from anterior frontal margin by narrow hiatus. In frontal view, antennules, antennular fossae partially visible, fossae wider in middle; interantennular septum partially covered by inferior surface of frontal border (
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<figureCitation id="3F912A299B18FFE9845FFC13FD071323" box="[575,651,1010,1035]" captionStart="FIGURE 2" captionStartId="3.[151,250,1526,1548]" captionTargetBox="[195,1391,194,1505]" captionTargetId="figure@3.[195,1392,193,1505]" captionTargetPageId="3" captionText="FIGURE 2. Pseudothelphusa acahuizotla n. sp., male holotype, CNCR 31708: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242444/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
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D). Opercular plate of antennal gland ovoid, with small middle constriction, tuft of setae on external third. Epistome, portions surrounding buccal cavity densely setose; laterally with rounded granules; epistomal tooth triangular, apex directed downwards at same level of interantennular septum, between palps of third maxillipeds when closing buccal cavity (
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<figureCitation id="3F912A299B18FFE985B7FBBFFB91135F" box="[983,1053,1118,1143]" captionStart="FIGURE 2" captionStartId="3.[151,250,1526,1548]" captionTargetBox="[195,1391,194,1505]" captionTargetId="figure@3.[195,1392,193,1505]" captionTargetPageId="3" captionText="FIGURE 2. Pseudothelphusa acahuizotla n. sp., male holotype, CNCR 31708: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242444/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
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D). Opening of efferent branchial channel subquadrate, ratio width/length 1.26. Third maxilliped with ischium trapezoidal, slightly longer than wide; merus narrower than ischium, anterior margin widely rounded without shallow, rounded notch before insertion of palp; ratio exopod/ischium 0.75 (
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<figureCitation id="3F912A299B18FFE9846AFB2BFDC213CB" box="[522,590,1226,1251]" captionStart="FIGURE 2" captionStartId="3.[151,250,1526,1548]" captionTargetBox="[195,1391,194,1505]" captionTargetId="figure@3.[195,1392,193,1505]" captionTargetPageId="3" captionText="FIGURE 2. Pseudothelphusa acahuizotla n. sp., male holotype, CNCR 31708: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242444/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
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C).
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</paragraph>
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<paragraph id="A71536AC9B18FFE986A7FB0FFE79125B" blockId="2.[151,1437,362,1971]" pageId="2" pageNumber="561">
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Chelipeds asymmetrical. Merus of larger cheliped with row of blunt tubercles along internal margin, carpus with strong triangular spine on internal margin. Chela with internal surface smooth, globose; fingers gaping, curved inwards distally, with low, triangular teeth on cutting edges (
|
||
<figureCitation id="3F912A299B18FFE98525FAD7FC071267" box="[837,907,1334,1359]" captionStart="FIGURE 2" captionStartId="3.[151,250,1526,1548]" captionTargetBox="[195,1391,194,1505]" captionTargetId="figure@3.[195,1392,193,1505]" captionTargetPageId="3" captionText="FIGURE 2. Pseudothelphusa acahuizotla n. sp., male holotype, CNCR 31708: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242444/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
|
||
B). Dactylus rather narrow, dorsal surface with longitudinal rows of small pits.
|
||
</paragraph>
|
||
<paragraph id="A71536AC9B18FFE986A7FA9FFC89109B" blockId="2.[151,1437,362,1971]" pageId="2" pageNumber="561">
|
||
Male gonopod robust. In mesial view, distal crest of caudomarginal projection moderately curved at same level of lateral crest (
|
||
<figureCitation id="3F912A299B18FFE9872FFA43FE181293" box="[335,404,1442,1467]" captionStart="FIGURE 3" captionStartId="4.[151,250,1933,1955]" captionTargetBox="[153,1436,373,1911]" captionTargetId="figure@4.[151,1436,369,1912]" captionTargetPageId="4" captionText="FIGURE 3. Pseudothelphusa acahuizotla n. sp., male holotype CNCR 31708, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudomarginal projection; mp, mesial process; ms, marginal suture; tps, terminal pore setae." httpUri="https://zenodo.org/record/242445/files/figure.png" pageId="2" pageNumber="561">Fig. 3</figureCitation>
|
||
A). Distal lobe of caudomarginal projection ending as acute, triangular tooth, rising from superior angle of proximal lobe, separated by narrow V-shaped notch; apex not reaching cephalic margin. Proximal lobe of caudomarginal projection longer than wide, subtriangular, wider distally, cephalo-proximally directed; cephalic margin curved. In cephalic view (
|
||
<figureCitation id="3F912A299B18FFE98411F9EFFD20110F" box="[625,684,1550,1575]" captionStart="FIGURE 3" captionStartId="4.[151,250,1933,1955]" captionTargetBox="[153,1436,373,1911]" captionTargetId="figure@4.[151,1436,369,1912]" captionTargetPageId="4" captionText="FIGURE 3. Pseudothelphusa acahuizotla n. sp., male holotype CNCR 31708, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudomarginal projection; mp, mesial process; ms, marginal suture; tps, terminal pore setae." httpUri="https://zenodo.org/record/242445/files/figure.png" pageId="2" pageNumber="561">Fig 3</figureCitation>
|
||
B), distal lobe of caudomarginal projection oriented cephalically, at same level as proximal lobe; internal surface flat. Proximal lobe of caudomarginal projection parallel to principal axis of gonopod. Mesial process approximately trapezoidal, wider section perpendicular to longitudinal axis of gonopod; distolateral angle with acute spine, caudo-laterally directed. Superior margin rounded, reaching internal angle forming subtriangular expansion closing apical cavity cephalically, in contact with internal surface of caudomarginal projection; same margin developing into central crest inside apical cavity, delimitng field of spines. Lateral border slightly concave, inferior angle rounded. In lateral view (
|
||
<figureCitation id="3F912A299B18FFE98583F907FBA611D7" box="[995,1066,1766,1791]" captionStart="FIGURE 3" captionStartId="4.[151,250,1933,1955]" captionTargetBox="[153,1436,373,1911]" captionTargetId="figure@4.[151,1436,369,1912]" captionTargetPageId="4" captionText="FIGURE 3. Pseudothelphusa acahuizotla n. sp., male holotype CNCR 31708, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudomarginal projection; mp, mesial process; ms, marginal suture; tps, terminal pore setae." httpUri="https://zenodo.org/record/242445/files/figure.png" pageId="2" pageNumber="561">Fig. 3</figureCitation>
|
||
C), lateral crest rounded, lateral spine, spine on caudal surface of mesial process visible, middle constriction on lateral surface of principal axis, strongly marked. In caudal view (
|
||
<figureCitation id="3F912A299B18FFE9847EF8CFFDEE106F" box="[542,610,1838,1863]" captionStart="FIGURE 3" captionStartId="4.[151,250,1933,1955]" captionTargetBox="[153,1436,373,1911]" captionTargetId="figure@4.[151,1436,369,1912]" captionTargetPageId="4" captionText="FIGURE 3. Pseudothelphusa acahuizotla n. sp., male holotype CNCR 31708, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudomarginal projection; mp, mesial process; ms, marginal suture; tps, terminal pore setae." httpUri="https://zenodo.org/record/242445/files/figure.png" pageId="2" pageNumber="561">Fig. 3</figureCitation>
|
||
D), distal third of gonopod tilted mesially, caudal crest concave, caudal surface of mesial process armed with well-formed central conical spine. In apical view (
|
||
<figureCitation id="3F912A299B18FFE982E0F8B3FB301043" box="[1152,1212,1874,1899]" captionStart="FIGURE 3" captionStartId="4.[151,250,1933,1955]" captionTargetBox="[153,1436,373,1911]" captionTargetId="figure@4.[151,1436,369,1912]" captionTargetPageId="4" captionText="FIGURE 3. Pseudothelphusa acahuizotla n. sp., male holotype CNCR 31708, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudomarginal projection; mp, mesial process; ms, marginal suture; tps, terminal pore setae." httpUri="https://zenodo.org/record/242445/files/figure.png" pageId="2" pageNumber="561">Fig 3</figureCitation>
|
||
E), apical cavity Ushaped, with opening of sperm channel in caudal position; field of spines wide, close to lateral crest, armed with 20–24 terminal pore setae, caudo-cephalically oriented.
|
||
</paragraph>
|
||
<caption id="F3D566249B19FFE886F7FA17FDBA1103" httpUri="https://zenodo.org/record/242444/files/figure.png" pageId="3" pageNumber="562" targetBox="[195,1391,194,1505]" targetPageId="3">
|
||
<paragraph id="A71536AC9B19FFE886F7FA17FDBA1103" blockId="3.[151,1436,1526,1579]" pageId="3" pageNumber="562">
|
||
<emphasis id="95DEEABE9B19FFE886F7FA17FE9B1124" bold="true" box="[151,279,1526,1548]" pageId="3" pageNumber="562">FIGURE 2.</emphasis>
|
||
<taxonomicName id="60AA4D2F9B19FFE88741FA17FDC31124" box="[289,591,1526,1548]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="acahuizotla" status="sp. nov.">
|
||
<emphasis id="95DEEABE9B19FFE88741FA17FDC31124" box="[289,591,1526,1548]" italics="true" pageId="3" pageNumber="562">Pseudothelphusa acahuizotla</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95DEEABE9B19FFE8843AFA16FD1B1124" bold="true" box="[602,663,1527,1548]" pageId="3" pageNumber="562">
|
||
<taxonomicNameLabel id="8EED57C59B19FFE8843AFA16FD1B1124" box="[602,663,1527,1548]" pageId="3" pageNumber="562" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, CNCR 31708: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A71536AC9B19FFE886A7F9B6FC191158" blockId="3.[151,1436,1623,2036]" box="[199,917,1623,1648]" pageId="3" pageNumber="562">
|
||
<emphasis id="95DEEABE9B19FFE886A7F9B6FD881158" bold="true" box="[199,516,1623,1648]" pageId="3" pageNumber="562">Geographical distribution.</emphasis>
|
||
Only known from the type locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFB065279B19FFE886A7F99AFE57119F" pageId="3" pageNumber="562" type="etymology">
|
||
<paragraph id="A71536AC9B19FFE886A7F99AFE57119F" blockId="3.[151,1436,1623,2036]" pageId="3" pageNumber="562">
|
||
<emphasis id="95DEEABE9B19FFE886A7F99AFEC211BC" bold="true" box="[199,334,1659,1684]" pageId="3" pageNumber="562">Etymology.</emphasis>
|
||
The specific epithet is the name of the
|
||
<typeStatus id="7811880E9B19FFE88564F99DFCB811BC" box="[772,820,1660,1684]" pageId="3" pageNumber="562">type</typeStatus>
|
||
locality,
|
||
<taxonomicName id="60AA4D2F9B19FFE885FAF99AFBA811BC" box="[922,1060,1659,1684]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="564" phylum="Arthropoda" rank="species" species="acahuizotla" status="sp. nov.">Acahuizotla</taxonomicName>
|
||
, in the state of
|
||
<collectingRegion id="656EF84E9B19FFE882ADF99AFAB911BB" box="[1229,1333,1659,1683]" country="Mexico" name="Guerrero" pageId="3" pageNumber="562">Guerrero</collectingRegion>
|
||
,
|
||
<collectingCountry id="DFBD763C9B19FFE88320F99AFA1411BC" box="[1344,1432,1659,1684]" name="Mexico" pageId="3" pageNumber="562">
|
||
<collectingRegion id="656EF84E9B19FFE88320F99AFA1411BC" box="[1344,1432,1659,1684]" country="Mexico" name="Mexico" pageId="3" pageNumber="562">Mexico</collectingRegion>
|
||
</collectingCountry>
|
||
, used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFB065279B19FFEE86A7F920FBE417D0" lastPageId="5" lastPageNumber="564" pageId="3" pageNumber="562" type="discussion">
|
||
<paragraph id="A71536AC9B19FFEF86A7F920FC3E1615" blockId="3.[151,1436,1623,2036]" lastBlockId="4.[151,1436,151,317]" lastPageId="4" lastPageNumber="563" pageId="3" pageNumber="562">
|
||
<emphasis id="95DEEABE9B19FFE886A7F920FEB711F2" bold="true" box="[199,315,1729,1754]" pageId="3" pageNumber="562">Remarks.</emphasis>
|
||
Regarding the external morphology,
|
||
<taxonomicName id="60AA4D2F9B19FFE884BCF920FBAA11F1" box="[732,1062,1729,1753]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="acahuizotla">
|
||
<emphasis id="95DEEABE9B19FFE884BCF920FBAA11F1" box="[732,1062,1729,1753]" italics="true" pageId="3" pageNumber="562">Pseudothelphusa acahuizotla</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="60AA4D2F9B19FFE882D8F923FF5E11D5" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="belliana">
|
||
<emphasis id="95DEEABE9B19FFE882D8F923FAA311F1" box="[1208,1327,1729,1753]" italics="true" pageId="3" pageNumber="562">P. belliana</emphasis>
|
||
<bibRefCitation id="C33B4B5D9B19FFE88356F920FF5E11D5" author="Rathbun" pageId="3" pageNumber="562" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537." type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
, described from Chautipan,
|
||
<collectingRegion id="656EF84E9B19FFE88471F904FDF611D5" box="[529,634,1765,1789]" country="Mexico" name="Guerrero" pageId="3" pageNumber="562">Guerrero</collectingRegion>
|
||
; both species have a flat and punctate carapace with round granules on the frontal and branchial surfaces; the front is vertical with a well-marked superior frontal border bearing low, rounded tubercles.
|
||
<taxonomicName id="60AA4D2F9B19FFE8870FF8CAFD35106B" box="[367,697,1835,1859]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="acahuizotla" status="sp. nov.">
|
||
<emphasis id="95DEEABE9B19FFE8870FF8CAFD35106B" box="[367,697,1835,1859]" italics="true" pageId="3" pageNumber="562">Pseudothelphusa acahuizotla</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95DEEABE9B19FFE884A1F8CAFC8C106C" bold="true" box="[705,768,1835,1860]" pageId="3" pageNumber="562">
|
||
<taxonomicNameLabel id="8EED57C59B19FFE884A1F8CAFC8C106C" box="[705,768,1835,1860]" pageId="3" pageNumber="562" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="60AA4D2F9B19FFE885F8F8CDFB9D106B" box="[920,1041,1835,1859]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="belliana">
|
||
<emphasis id="95DEEABE9B19FFE885F8F8CDFB9D106B" box="[920,1041,1835,1859]" italics="true" pageId="3" pageNumber="562">P. belliana</emphasis>
|
||
</taxonomicName>
|
||
in having the dorsal surface of the carapace smooth, without black and short bristles on the posterolateral regions, and in the following characters of the male gonopod: the size and form of the distal lobe of the caudomarginal projection which is small and triangular, separated from the proximal lobe by a V-shaped narrow notch; and by the shape of the mesial process which is approximately trapezoidal with the lateral spine at the level of the superior margin and in the form of the inferior margin; caudal surface of mesial process with an elongated and rounded spine.
|
||
<taxonomicName id="60AA4D2F9B19FFE88219F83AFA1010DB" box="[1145,1436,2011,2035]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="belliana">
|
||
<emphasis id="95DEEABE9B19FFE88219F83AFA1010DB" box="[1145,1436,2011,2035]" italics="true" pageId="3" pageNumber="562">Pseudothelphusa belliana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="C33B4B5D9B1EFFEF86FEFF76FE5D1798" author="Rathbun" box="[158,465,151,176]" pageId="4" pageNumber="563" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537." type="journal article" year="1898">Rathbun, 1898: 515, fig. 4</bibRefCitation>
|
||
), has a male gonopod with a distal lobe of the caudomarginal projection slender and separated from the mesial process by a narrow incision; the mesial process is subtriangular, and the superior margin is widely curved; the lateral spine is located on the distal half of the lateral margin; below the spine the lateral margin is straight, and continues proximally to end in the principal axis of the gonopod without forming an inferior margin; the caudal surface of the mesial process has a conical tubercle.
|
||
</paragraph>
|
||
<caption id="F3D566249B1EFFEF86F7F86CFBB110C9" httpUri="https://zenodo.org/record/242445/files/figure.png" pageId="4" pageNumber="563" targetBox="[153,1436,373,1911]" targetPageId="4">
|
||
<paragraph id="A71536AC9B1EFFEF86F7F86CFBB110C9" blockId="4.[151,1436,1933,2017]" pageId="4" pageNumber="563">
|
||
<emphasis id="95DEEABE9B1EFFEF86F7F86CFE9F108B" bold="true" box="[151,275,1933,1955]" pageId="4" pageNumber="563">FIGURE 3.</emphasis>
|
||
<taxonomicName id="60AA4D2F9B1EFFEF877AF86CFDC8108B" box="[282,580,1933,1955]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="563" phylum="Arthropoda" rank="species" species="acahuizotla" status="sp. nov.">
|
||
<emphasis id="95DEEABE9B1EFFEF877AF86CFDC8108B" box="[282,580,1933,1955]" italics="true" pageId="4" pageNumber="563">Pseudothelphusa acahuizotla</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95DEEABE9B1EFFEF842BF86FFD09108B" bold="true" box="[587,645,1934,1955]" pageId="4" pageNumber="563">
|
||
<taxonomicNameLabel id="8EED57C59B1EFFEF842BF86FFD09108B" box="[587,645,1934,1955]" pageId="4" pageNumber="563" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype CNCR 31708, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudomarginal projection; mp, mesial process; ms, marginal suture; tps, terminal pore setae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A71536AC9B1FFFEE86A7FF76FBE417D0" blockId="5.[151,1436,151,248]" pageId="5" pageNumber="564">
|
||
The new species is distributed in southeastern
|
||
<collectingRegion id="656EF84E9B1FFFEE84ABFF79FCBF1798" box="[715,819,152,176]" country="Mexico" name="Guerrero" pageId="5" pageNumber="564">Guerrero</collectingRegion>
|
||
, close to
|
||
<taxonomicName id="60AA4D2F9B1FFFEE85FAFF78FB9C1798" box="[922,1040,152,176]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="belliana">
|
||
<emphasis id="95DEEABE9B1FFFEE85FAFF78FB9C1798" box="[922,1040,152,176]" italics="true" pageId="5" pageNumber="564">P. belliana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60AA4D2F9B1FFFEE8225FF78FB681798" box="[1093,1252,152,176]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="hoffmannae">
|
||
<emphasis id="95DEEABE9B1FFFEE8225FF78FB681798" box="[1093,1252,152,176]" italics="true" pageId="5" pageNumber="564">P. hoffmannae</emphasis>
|
||
</taxonomicName>
|
||
, all occurring in the Papagayo River basin, where
|
||
<taxonomicName id="60AA4D2F9B1FFFEE846DFF5CFDE517FC" box="[525,617,188,212]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="galloi">
|
||
<emphasis id="95DEEABE9B1FFFEE846DFF5CFDE517FC" box="[525,617,188,212]" italics="true" pageId="5" pageNumber="564">P. galloi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60AA4D2F9B1FFFEE8418FF5CFD6817FC" box="[632,740,188,212]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="nelsoni">
|
||
<emphasis id="95DEEABE9B1FFFEE8418FF5CFD6817FC" box="[632,740,188,212]" italics="true" pageId="5" pageNumber="564">P. nelsoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60AA4D2F9B1FFFEE857AFF5CFC4317FC" box="[794,975,189,212]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="guerreroensis">
|
||
<emphasis id="95DEEABE9B1FFFEE857AFF5CFC4317FC" box="[794,975,189,212]" italics="true" pageId="5" pageNumber="564">P. guerreroensis</emphasis>
|
||
</taxonomicName>
|
||
are also distributed. With the description of
|
||
<taxonomicName id="60AA4D2F9B1FFFEE86D5FF00FED817D0" box="[181,340,224,248]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="acahuizotla" status="sp. nov.">
|
||
<emphasis id="95DEEABE9B1FFFEE86D5FF00FED817D0" box="[181,340,224,248]" italics="true" pageId="5" pageNumber="564">P. acahuizotla</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95DEEABE9B1FFFEE873AFF3EFE1517D0" bold="true" box="[346,409,223,248]" pageId="5" pageNumber="564">
|
||
<taxonomicNameLabel id="8EED57C59B1FFFEE873AFF3EFE1517D0" box="[346,409,223,248]" pageId="5" pageNumber="564" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the number of recognized species in the state increases to eight.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |