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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.95.97156" ID-Pensoft-Pub="1314-4049-95-61" ID-Pensoft-UUID="72171DE948FB58008EB2E623130744DB" ModsDocID="1314-4049-95-61" checkinTime="1675973707820" checkinUser="pensoft" docAuthor="Tang, Song-Ming, Vadthanarat, Santhiti, He, Jun, Raghoonundon, Bhavesh, Yu, Feng-Ming, Karunarathna, Samantha C., Li, Shu-Hong &amp; Raspe ́, Olivier" docDate="2023" docId="2E6B44B8597F5D7ABB7D7B319222AE0D" docLanguage="en" docName="MycoKeys 95: 61-82" docOrigin="MycoKeys 95" docPubDate="2023-02-08" docSource="http://dx.doi.org/10.3897/mycokeys.95.97156" docTitle="Termitomyces tigrinus S. M. Tang &amp; Raspe ́ 2023, sp. nov." docType="treatment" docVersion="1" id="72171DE948FB58008EB2E623130744DB" lastPageNumber="61" masterDocId="72171DE948FB58008EB2E623130744DB" masterDocTitle="Morphological and molecular analyses reveal two new species of Termitomyces (Agaricales, Lyophyllaceae) and morphological variability of T. intermedius" masterLastPageNumber="82" masterPageNumber="61" pageNumber="61" updateTime="1675973707820" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Morphological and molecular analyses reveal two new species of Termitomyces (Agaricales, Lyophyllaceae) and morphological variability of T. intermedius</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tang, Song-Ming</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6174-7314</mods:nameIdentifier>
<mods:affiliation>Biotechnology and Germplasm Resources Institute, Yunnan Academy of Agricultural Sciences, Kunming 650205, China &amp; School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; College of Agriculture &amp; Biological Sciences, Dali University, Dali 671003, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vadthanarat, Santhiti</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9035-0375</mods:nameIdentifier>
<mods:affiliation>School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>He, Jun</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7027-7206</mods:nameIdentifier>
<mods:affiliation>Biotechnology and Germplasm Resources Institute, Yunnan Academy of Agricultural Sciences, Kunming 650205, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Raghoonundon, Bhavesh</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6671-2404</mods:nameIdentifier>
<mods:affiliation>School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yu, Feng-Ming</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9133-8645</mods:nameIdentifier>
<mods:affiliation>School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Karunarathna, Samantha C.</mods:namePart>
<mods:affiliation>Qujing Normal University, Qujing, Yunnan 655011, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shu-Hong</mods:namePart>
<mods:affiliation>Biotechnology and Germplasm Resources Institute, Yunnan Academy of Agricultural Sciences, Kunming 650205, China</mods:affiliation>
<mods:nameIdentifier type="email">shuhongfungi@126.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Raspe ́, Olivier</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8426-2133</mods:nameIdentifier>
<mods:affiliation>School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">Olivier.Ras@mfu.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>95</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>61</mods:start>
<mods:end>82</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.95.97156</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.95.97156</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-95-61</mods:identifier>
<mods:identifier type="Pensoft-UUID">72171DE948FB58008EB2E623130744DB</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:2E6B44B8597F5D7ABB7D7B319222AE0D" httpUri="http://treatment.plazi.org/id/2E6B44B8597F5D7ABB7D7B319222AE0D" lastPageNumber="61" pageId="0" pageNumber="61">
<subSubSection pageId="0" pageNumber="61" type="nomenclature">
<paragraph pageId="0" pageNumber="61">
<taxonomicName LSID="2E6B44B8-597F-5D7A-BB7D-7B319222AE0D" authority="S. M. Tang &amp; Raspe ́" authorityName="S. M. Tang &amp; Raspe ́" authorityYear="2023" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Termitomyces tigrinus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="tigrinus" status="sp. nov.">Termitomyces tigrinus S.M. Tang &amp; Raspé</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="61">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Termitomyces tigrinus a-c basidiomata (a HKAS 107560, b, c HKAS 107561) d cheilocystidia e basidia with acute sterigmata apex f basidia with obtuse sterigmata apex g pileipellis h basidiospores. Scale bars: 1 cm (a-c); 10 μm (d-f); 20 μm (g); 5 μm (h). Photographs by Song-Ming Tang." figureDoi="10.3897/mycokeys.95.97156.figure4" httpUri="https://binary.pensoft.net/fig/807373" pageId="0" pageNumber="61">Fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="etymology">
<paragraph pageId="0" pageNumber="61">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="61">
The epithet
<normalizedToken originalValue="“tigrinus”">&quot;tigrinus&quot;</normalizedToken>
refers to the alternating greyish white and dark grey zones on the pileus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="type material">
<paragraph pageId="0" pageNumber="61">Type material.</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">
<emphasis italics="true" pageId="0" pageNumber="61">Holotype</emphasis>
. China
</emphasis>
. Yunnan Province: Chuxiong County, Fuming, elev. 1,800 m, 16 July 2019, Song-Ming Tang (
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Holotype</emphasis>
: HKAS 107560,
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">isotype</emphasis>
: MFLU 22-0143).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="diagnosis">
<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="61">
Differs from other
<taxonomicName authorityName="R.Heim" authorityYear="1942" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Termitomyces" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Termitomyces</emphasis>
</taxonomicName>
species in having a regular alternating greyish white and dark grey zones on the pileus, and subclavate stipe.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">Description.</paragraph>
<paragraph pageId="0" pageNumber="61">
Basidiomata medium-sized. Pileus 7-9 cm in diam., convex to plano-convex, circular when seen from above, dark grey (1F1-2) at the centre, greyish white (1B1) to grey (1C1-1D1) towards margin, with regular alternating dark grey and greyish white zones, densely tomentose to tomentose-squamulose, hairs grey to drab, in dry conditions, often cracked into large or small scales; margin exceeding lamellae, undate; perforatorium small, as an acute papilla, dark grey (1C1). context 1-2 mm thick half-way to the margin, tough, white (1A1). Lamellae close, ventricose, 3-5 mm wide, adnexed, crowded, white (1A1) at first, then cream to greyish pink when mature; lamellar edge eroded; lamellulae in 1-2 tiers. Stipe 5-7
<normalizedToken originalValue="×">x</normalizedToken>
1-3.5 cm, central, subclavate, white (1A1) at the apex, greyish white (1B1) to grey (1C1-1D1) toward the base, smooth; context white (1A1), solid, fibrous. Annulus absent. Pseudorhiza terete, strongly tapering; surface grey (1D1-2) to dark grey (1F1-1F2), smooth; context solid, fibrous. Odour slightly fragrant. Taste not distinctive.
</paragraph>
<paragraph pageId="0" pageNumber="61">
Basidia of two types conspicuously different by the apex of sterigmata being either acute or obtuse, first type rather abundant, clavate, sterigmata apex acute, mostly 4-spored sometimes 1-spored or 2-spored, 25-32
<normalizedToken originalValue="×">x</normalizedToken>
7-12
<normalizedToken originalValue="μm">μm</normalizedToken>
, av. 28
<normalizedToken originalValue="±">+/-</normalizedToken>
2.4
<normalizedToken originalValue="×">x</normalizedToken>
11
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5
<normalizedToken originalValue="μm">μm</normalizedToken>
, sterigmata 1-3
<normalizedToken originalValue="μm">μm</normalizedToken>
long; the second type fewer in number, clavate, sterigmata apex obtuse, mostly 1-spored, 2-spored, sometimes 4-spored, 24-30
<normalizedToken originalValue="×">x</normalizedToken>
9-13
<normalizedToken originalValue="μm">μm</normalizedToken>
, av. 26
<normalizedToken originalValue="±">+/-</normalizedToken>
2.2
<normalizedToken originalValue="×">x</normalizedToken>
12
<normalizedToken originalValue="±">+/-</normalizedToken>
0.7
<normalizedToken originalValue="μm">μm</normalizedToken>
, sterigmata 2-4
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Basidiospores [90/5/2] (6.1-) 7.2-9.6 (-10.1)
<normalizedToken originalValue="×">x</normalizedToken>
(3.3-) 5.2-7.3 (-7.9)
<normalizedToken originalValue="μm">μm</normalizedToken>
, Lm
<normalizedToken originalValue="×">x</normalizedToken>
Wm = 8.1
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
<normalizedToken originalValue="×">x</normalizedToken>
6.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8
<normalizedToken originalValue="μm">μm</normalizedToken>
, Q = (1.01-) 1.20-2.03 (-2.30), Qm = 1.53
<normalizedToken originalValue="±">+/-</normalizedToken>
0.20, broadly ellipsoid to ellipsoid, colourless, thin-walled, smooth. Hymenophoral trama regular, element parallel, 51-100
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, made up of thin-walled, ellipsoid to clavate inflated hyphae 16-18
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, filamentous hyphae abundant, 5-10
<normalizedToken originalValue="μm">μm</normalizedToken>
wide. Subhymenium 8-21
<normalizedToken originalValue="μm">μm</normalizedToken>
thick, with 1-2 layers of ovoid, subglobose, fusiform, ellipsoid or irregular cells, 8-11
<normalizedToken originalValue="×">x</normalizedToken>
3-6
<normalizedToken originalValue="μm">μm</normalizedToken>
. Pleurocystidia absent. Lamellar edge composed mostly of undifferentiated, basidiole-like cells. Cheilocystidia few, broadly clavate, 17-36
<normalizedToken originalValue="×">x</normalizedToken>
9-16
<normalizedToken originalValue="μm">μm</normalizedToken>
, av. 28
<normalizedToken originalValue="±">+/-</normalizedToken>
2.0
<normalizedToken originalValue="×">x</normalizedToken>
14
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
<normalizedToken originalValue="μm">μm</normalizedToken>
. Pileipellis 2-layered, suprapellis an ixocutis 22-51
<normalizedToken originalValue="×">x</normalizedToken>
5-7
<normalizedToken originalValue="μm">μm</normalizedToken>
av. 30
<normalizedToken originalValue="±">+/-</normalizedToken>
5.7
<normalizedToken originalValue="×">x</normalizedToken>
6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5
<normalizedToken originalValue="μm">μm</normalizedToken>
, cylindrical hyphae with obtuse apex, thin-walled, hyaline in KOH; subpellis made up of inflated elements, 31-81
<normalizedToken originalValue="×">x</normalizedToken>
7-14
<normalizedToken originalValue="μm">μm</normalizedToken>
, av. 58
<normalizedToken originalValue="±">+/-</normalizedToken>
7.6
<normalizedToken originalValue="×">x</normalizedToken>
10
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8
<normalizedToken originalValue="μm">μm</normalizedToken>
. Clamp connections not seen in any tissues.
</paragraph>
<caption doi="10.3897/mycokeys.95.97156.figure4" httpUri="https://binary.pensoft.net/fig/807373" pageId="0" pageNumber="61" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 4.</emphasis>
<taxonomicName authorityName="S. M. Tang &amp; Raspe ́" authorityYear="2023" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Termitomyces tigrinus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="tigrinus">
<emphasis italics="true" pageId="0" pageNumber="61">Termitomyces tigrinus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="61">a-c</emphasis>
basidiomata (
<emphasis bold="true" pageId="0" pageNumber="61">a</emphasis>
HKAS 107560,
<emphasis bold="true" pageId="0" pageNumber="61">b, c</emphasis>
HKAS 107561)
<emphasis bold="true" pageId="0" pageNumber="61">d</emphasis>
cheilocystidia
<emphasis bold="true" pageId="0" pageNumber="61">e</emphasis>
basidia with acute sterigmata apex
<emphasis bold="true" pageId="0" pageNumber="61">f</emphasis>
basidia with obtuse sterigmata apex
<emphasis bold="true" pageId="0" pageNumber="61">g</emphasis>
pileipellis
<emphasis bold="true" pageId="0" pageNumber="61">h</emphasis>
basidiospores. Scale bars: 1 cm (
<emphasis bold="true" pageId="0" pageNumber="61">a-c</emphasis>
); 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="61">d-f</emphasis>
); 20
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="61">g</emphasis>
); 5
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="61">h</emphasis>
). Photographs by Song-Ming Tang.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="habitat">
<paragraph pageId="0" pageNumber="61">Habitat and distribution.</paragraph>
<paragraph pageId="0" pageNumber="61">Basidiomata scattered on soil with decaying litter under which termites have built their nest. Occurring in summer. So far only known from southwestern China.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="additional species examined">
<paragraph pageId="0" pageNumber="61">Additional species examined.</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">China</emphasis>
. Yunnan Province, Chuxiong city, 20 July 2019, Jun He (HKAS 107561).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="notes">
<paragraph pageId="0" pageNumber="61">Notes.</paragraph>
<paragraph pageId="0" pageNumber="61">
<taxonomicName authorityName="S. M. Tang &amp; Raspe ́" authorityYear="2023" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Termitomyces tigrinus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="tigrinus">
<emphasis italics="true" pageId="0" pageNumber="61">Termitomyces tigrinus</emphasis>
</taxonomicName>
is distinguished from other
<taxonomicName authorityName="R.Heim" authorityYear="1942" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Termitomyces" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Termitomyces</emphasis>
</taxonomicName>
by densely tomentose to tomentose-squamulose pileus with regularly alternating greyish white and dark grey zones, and a small, dark grey perforatorium as an acute papilla, two conspicuously different types of basidia, broadly ellipsoid to ellipsoid basidiospores, clavate, thin-walled cheilocystidia that are rare (HKAS 107560), or absent (in HKAS 107561).
</paragraph>
<paragraph pageId="0" pageNumber="61">
Morphologically,
<taxonomicName class="Agaricomycetes" kingdom="Fungi" lsidName="T. tigrinus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="tigrinus">
<emphasis italics="true" pageId="0" pageNumber="61">T. tigrinus</emphasis>
</taxonomicName>
is similar to
<taxonomicName class="Agaricomycetes" kingdom="Fungi" lsidName="T. robustus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="robustus">
<emphasis italics="true" pageId="0" pageNumber="61">T. robustus</emphasis>
</taxonomicName>
(Beeli) R. Heim in having grey to black stipe. However,
<taxonomicName class="Agaricomycetes" kingdom="Fungi" lsidName="T. robustus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="robustus">
<emphasis italics="true" pageId="0" pageNumber="61">T. robustus</emphasis>
</taxonomicName>
has a blunt perforatorium (
<bibRefCitation DOI="https://doi.org/10.2307/3666672" author="Heim, R" journalOrPublisher="Archives du Museum National d'Histoire Naturelle" pageId="0" pageNumber="61" refId="B12" refString="Heim, R, 1951. Les Termitomyces du Congo belge recueillis par Madame Goossens-Fontana. Bulletin du Jardin Botanique de l'Etat [Bruxelles] 21(3/4): 205-222. https://doi.org/10.2307/3666672" title="Les Termitomyces du Congo belge recueillis par Madame Goossens-Fontana. Bulletin du Jardin Botanique de l'Etat [Bruxelles] 21 (3 / 4): 205 - 222." url="https://doi.org/10.2307/3666672" year="1951">Heim 1951</bibRefCitation>
), smaller basidia (20-25
<normalizedToken originalValue="×">x</normalizedToken>
7-8
<normalizedToken originalValue="μm">μm</normalizedToken>
) and basidiospores (7.0-8.0
<normalizedToken originalValue="×">x</normalizedToken>
5.0-5.5
<normalizedToken originalValue="μm">μm</normalizedToken>
) (
<bibRefCitation author="Otieno, NC" journalOrPublisher="Sydowia" pageId="0" pageNumber="61" pagination="160 - 165" refId="B31" refString="Otieno, NC, 1969. Further contributions to a knowledge of termite fungi in East Africa: The genus Termitomyces Heim. Sydowia 22: 160 - 165" title="Further contributions to a knowledge of termite fungi in East Africa: The genus Termitomyces Heim." volume="22" year="1969">Otieno 1969</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="61">
<taxonomicName class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Termitomyces entolomoides" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="entolomoides">
<emphasis italics="true" pageId="0" pageNumber="61">Termitomyces entolomoides</emphasis>
</taxonomicName>
R. Heim with
<taxonomicName class="Agaricomycetes" kingdom="Fungi" lsidName="T. tigrinus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="tigrinus">
<emphasis italics="true" pageId="0" pageNumber="61">T. tigrinus</emphasis>
</taxonomicName>
in having a tapering upwards stipe, dark grey pileus and greyish white to grey stipe. However,
<taxonomicName class="Agaricomycetes" kingdom="Fungi" lsidName="T. entolomoides" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="entolomoides">
<emphasis italics="true" pageId="0" pageNumber="61">T. entolomoides</emphasis>
</taxonomicName>
has been originally described from Congo by
<bibRefCitation DOI="https://doi.org/10.2307/3666672" author="Heim, R" journalOrPublisher="Archives du Museum National d'Histoire Naturelle" pageId="0" pageNumber="61" refId="B12" refString="Heim, R, 1951. Les Termitomyces du Congo belge recueillis par Madame Goossens-Fontana. Bulletin du Jardin Botanique de l'Etat [Bruxelles] 21(3/4): 205-222. https://doi.org/10.2307/3666672" title="Les Termitomyces du Congo belge recueillis par Madame Goossens-Fontana. Bulletin du Jardin Botanique de l'Etat [Bruxelles] 21 (3 / 4): 205 - 222." url="https://doi.org/10.2307/3666672" year="1951">Heim (1951)</bibRefCitation>
, has a small basidioma (pileus 3-4 cm diam.), a grey pseudorhiza and relatively smaller basidiospores (6.2-6.6
<normalizedToken originalValue="×">x</normalizedToken>
4-4.2
<normalizedToken originalValue="μm">μm</normalizedToken>
;
<bibRefCitation DOI="https://doi.org/10.2307/3666672" author="Heim, R" journalOrPublisher="Archives du Museum National d'Histoire Naturelle" pageId="0" pageNumber="61" refId="B12" refString="Heim, R, 1951. Les Termitomyces du Congo belge recueillis par Madame Goossens-Fontana. Bulletin du Jardin Botanique de l'Etat [Bruxelles] 21(3/4): 205-222. https://doi.org/10.2307/3666672" title="Les Termitomyces du Congo belge recueillis par Madame Goossens-Fontana. Bulletin du Jardin Botanique de l'Etat [Bruxelles] 21 (3 / 4): 205 - 222." url="https://doi.org/10.2307/3666672" year="1951">Heim 1951</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="61">
In our multi-locus phylogeny,
<taxonomicName class="Agaricomycetes" kingdom="Fungi" lsidName="T. tigrinus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="tigrinus">
<emphasis italics="true" pageId="0" pageNumber="61">T. tigrinus</emphasis>
</taxonomicName>
is closely related to
<taxonomicName class="Agaricomycetes" kingdom="Fungi" lsidName="T. intermedius" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="intermedius">
<emphasis italics="true" pageId="0" pageNumber="61">T. intermedius</emphasis>
</taxonomicName>
,
<taxonomicName class="Agaricomycetes" kingdom="Fungi" lsidName="T. radicatus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="radicatus">
<emphasis italics="true" pageId="0" pageNumber="61">T. radicatus</emphasis>
</taxonomicName>
Natarajan and
<taxonomicName class="Agaricomycetes" kingdom="Fungi" lsidName="T. striatus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="striatus">
<emphasis italics="true" pageId="0" pageNumber="61">T. striatus</emphasis>
</taxonomicName>
. However,
<taxonomicName class="Agaricomycetes" kingdom="Fungi" lsidName="T. intermedius" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="intermedius">
<emphasis italics="true" pageId="0" pageNumber="61">T. intermedius</emphasis>
</taxonomicName>
has a cylindrical stipe, and cheilocystidia clavate to pyriform, narrowly lageniform, lageniform or broadly lageniform (this study).
<taxonomicName class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Termitomyces radicatus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="radicatus">
<emphasis italics="true" pageId="0" pageNumber="61">Termitomyces radicatus</emphasis>
</taxonomicName>
has a pale orange pileus, dark brown perforatorium, relatively smaller pileus (1.5-3.5 cm), and cylindrical stipe (
<bibRefCitation DOI="https://doi.org/10.2307/4118066" author="Pegler, DN" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="61" pagination="717 - 736" refId="B33" refString="Pegler, DN, Vanhaecke, M, 1994. Termitomyces of Southeast Asia. Kew Bulletin 49 (4): 717 - 736, DOI: https://doi.org/10.2307/4118066" title="Termitomyces of Southeast Asia." url="https://doi.org/10.2307/4118066" volume="49" year="1994">Pegler and Vanhaecke 1994</bibRefCitation>
).
<taxonomicName authorityName="R.Heim" authorityYear="1942" baseAuthorityName="Beeli" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Termitomyces striatus" order="Agaricales" pageId="0" pageNumber="61" phylum="Basidiomycota" rank="species" species="striatus">
<emphasis italics="true" pageId="0" pageNumber="61">Termitomyces striatus</emphasis>
</taxonomicName>
has a white to ocher pileus, irregularity fibrous striate on the stipe surface, long pseudorhiza (30-100 cm), with squama of various sizes and shapes on the surface cheilocystidia and pleurocystidia pyriform,broadly clavate, cylindrical or ovoid, (20-45
<normalizedToken originalValue="×">x</normalizedToken>
11-22
<normalizedToken originalValue="μm">μm</normalizedToken>
;
<bibRefCitation author="Heim, R" journalOrPublisher="Boubee, Paris" pageId="0" pageNumber="61" refId="B13" refString="Heim, R, 1977. Termites et champignons. Les champignons termitophiles d'Afrique noire et d'Asie meridionale. Ed. Boubee, Paris" title="Termites et champignons. Les champignons termitophiles d'Afrique noire et d'Asie meridionale. Ed." year="1977">Heim 1977</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>