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<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.7.90973" ID-Pensoft-Pub="2535-0730-1-51" ID-Pensoft-UUID="ABE1527AFB225DE3B1CDE6FE4E7BB38C" ID-ZooBank="15FBAE73903B4E14BDE3CCC2D65AAD81" ModsDocID="2535-0730-7-1-51" checkinTime="1672842668324" checkinUser="pensoft" docAuthor="Brahma, Shubhranil, Chatterjee, Somnath &amp; Hazra, Niladri" docDate="2023" docId="2E3FA2B05E125EE4AFD91CD4D1CA5C8B" docLanguage="en" docName="EvolutSyst 7(1): 51-66" docOrigin="Evolutionary Systematics 7 (1)" docPubDate="2023-01-03" docSource="http://dx.doi.org/10.3897/evolsyst.7.90973" docTitle="Dasyhelea (Dasyhelea) trigona Brahma &amp; Chatterjee &amp; Hazra 2023, sp. nov." docType="treatment" docUuid="F1CB60B1-616A-4803-AF13-4BDFCF50B374" docUuidSource="ZooBank" docVersion="1" id="ABE1527AFB225DE3B1CDE6FE4E7BB38C" lastPageNumber="51" masterDocId="ABE1527AFB225DE3B1CDE6FE4E7BB38C" masterDocTitle="Five new Indian species of the genus Dasyhelea Kieffer (Diptera, Ceratopogonidae) with a key to the adult males" masterLastPageNumber="66" masterPageNumber="51" pageNumber="51" updateTime="1672842668324" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five new Indian species of the genus Dasyhelea Kieffer (Diptera, Ceratopogonidae) with a key to the adult males</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brahma, Shubhranil</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5007-9704</mods:nameIdentifier>
<mods:affiliation>Iswar Chandra College, Department of Zoology, Belonia, Tripura, 799155, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chatterjee, Somnath</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8572-1779</mods:nameIdentifier>
<mods:affiliation>Entomology Research Unit, Department of Zoology, The University of Burdwan, Burdwan, 713104, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hazra, Niladri</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8149-9445</mods:nameIdentifier>
<mods:affiliation>Entomology Research Unit, Department of Zoology, The University of Burdwan, Burdwan, 713104, India</mods:affiliation>
<mods:nameIdentifier type="email">nhazra@zoo.buruniv.ac.in</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>7</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>51</mods:start>
<mods:end>66</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/evolsyst.7.90973</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.7.90973</mods:identifier>
<mods:identifier type="Pensoft-Pub">2535-0730-1-51</mods:identifier>
<mods:identifier type="ZooBank">15FBAE73903B4E14BDE3CCC2D65AAD81</mods:identifier>
<mods:identifier type="Pensoft-UUID">ABE1527AFB225DE3B1CDE6FE4E7BB38C</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:F1CB60B1-616A-4803-AF13-4BDFCF50B374" httpUri="http://treatment.plazi.org/id/2E3FA2B05E125EE4AFD91CD4D1CA5C8B" lastPageNumber="51" pageId="0" pageNumber="51">
<subSubSection pageId="0" pageNumber="51" type="nomenclature">
<paragraph pageId="0" pageNumber="51">
<taxonomicName LSID="https://zoobank.org/F1CB60B1-616A-4803-AF13-4BDFCF50B374" authority="Brahma &amp; Chatterjee &amp; Hazra, 2023" authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Dasyhelea) trigona" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="trigona" status="sp. nov." subGenus="Dasyhelea">Dasyhelea (Dasyhelea) trigona</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="51">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="type material">
<paragraph pageId="0" pageNumber="51">Type material.</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Holotype</emphasis>
♂, India, West Bengal, Birbhum, Boidhora [
<geoCoordinate degrees="24" direction="north" minutes="14" orientation="latitude" precision="1" seconds="59.4" value="24.249834">24°14'59.4&quot;N</geoCoordinate>
,
<geoCoordinate degrees="87" direction="east" minutes="44" orientation="longitude" precision="1" seconds="20.2" value="87.73894">87°44'20.2&quot;E</geoCoordinate>
], 25.XII.2016, Coll. S. Chatterjee.
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Paratype</emphasis>
: 1♂, India, West Bengal, South 24 Parganas, Bali Island [
<geoCoordinate degrees="22" direction="north" minutes="05" orientation="latitude" precision="1" seconds="19.6" value="22.088778">22°05'19.6&quot;N</geoCoordinate>
,
<geoCoordinate degrees="88" direction="east" minutes="45" orientation="longitude" precision="1" seconds="25.7" value="88.75714">88°45'25.7&quot;E</geoCoordinate>
], 15.V.2017, Coll. S. Brahma.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="diagnosis">
<paragraph pageId="0" pageNumber="51">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="51">
The only species in the subgenus
<taxonomicName authority="Dasyhelea" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Dasyhelea)" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="subGenus" subGenus="Dasyhelea">Dasyhelea Dasyhelea</taxonomicName>
having tubercles on the frontal sclerite, femora with infuscation beyond the mid region and an aedeagus with ventral triangular basal arms.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="description">
<paragraph pageId="0" pageNumber="51">Description.</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Male adult</emphasis>
(n = 2) (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5A-K</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-E Adult male genitalia structures. A Dasyhelea (Dasyhelea) incisura sp. nov.; B Dasyhelea (Dasyhelea) quasifulcillata sp. nov.; C Dasyhelea (Dasyhelea) trigona sp. nov.; D Dasyhelea (Sebessia) falxa sp. nov.; E Dasyhelea (Sebessia) folia sp. nov. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure8" httpUri="https://binary.pensoft.net/fig/790308" pageId="0" pageNumber="51">8C</figureCitation>
). Total body length 2-2.1 mm.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Head</emphasis>
. Dark brown in colour. Eyes separated by half of diameter of ommatidium. Frontal sclerite (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5A</figureCitation>
) 57.5
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 55
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, with oval base, small dome shaped apical portion ending in a threadlike projection and base containing sclerotised tubercles. Antennal flagellum (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5B</figureCitation>
) brown, 0.9 mm long; flagellomere XIII with a distinct apical projection; length ratio of flagellomeres (I-XIII): 17: 14: 11.5-12: 12: 15-15.5: 14-14.5: 14: 15: 15: 29-29.5: 23.5-24: 20: 38.5-39; AR 1.00; flagellomeres III-XIII (visible) with sensilla basiconica. Clypeus (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5C</figureCitation>
) with 4 pairs of setae. Palpus (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5D</figureCitation>
) pale brown; length ratio of palp segments (I-V): 6: 10-10.5: 13: 11-11.5: 14; PR 2.16; palp segment III with 6 capitate sensilla at proximal half.
</paragraph>
<caption doi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Figure 5.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="51">A-K</emphasis>
Adult male of
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Dasyhelea) trigona" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="trigona" subGenus="Dasyhelea">Dasyhelea (Dasyhelea) trigona</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="51">A</emphasis>
Frontal sclerite;
<emphasis bold="true" pageId="0" pageNumber="51">B</emphasis>
Flagellum;
<emphasis bold="true" pageId="0" pageNumber="51">C</emphasis>
Clypeus;
<emphasis bold="true" pageId="0" pageNumber="51">D</emphasis>
Palpus;
<emphasis bold="true" pageId="0" pageNumber="51">E</emphasis>
Thorax;
<emphasis bold="true" pageId="0" pageNumber="51">F</emphasis>
Wing;
<emphasis bold="true" pageId="0" pageNumber="51">G</emphasis>
Femora and tibiae (right to left) of fore, mid and hind legs;
<emphasis bold="true" pageId="0" pageNumber="51">H</emphasis>
Hind tibial comb;
<emphasis bold="true" pageId="0" pageNumber="51">I</emphasis>
Genitalia (left dorsal and right ventral view);
<emphasis bold="true" pageId="0" pageNumber="51">J</emphasis>
Apicolateral process;
<emphasis bold="true" pageId="0" pageNumber="51">K</emphasis>
Aedeagus. Scale bars: 0.05 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Thorax</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5E</figureCitation>
). Brown in colour without any prominent infuscation. Acrostichals 11-12 pairs, biserial; dorsocentrals 10-11 pairs, biserial and 7, uniserial; 13 prealars; scutellum with 6 setae.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Wing</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5F</figureCitation>
). Wing margin with short, prominent macrotrichia; anterior margin of r5 with microtrichia. WL 0.95-1.00 mm, WW 0.35 mm, CR 0.5. Cubital fork lying well before costal extremity.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Legs</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5G</figureCitation>
). Pale brown in colour. Femora infuscated at mid region, mid tibia with diffused infuscation, proximads of tibiae dark brown. Hind tibial comb (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5H</figureCitation>
) with 7 spines, third one from inner longest. TRI 3.14-3.18, TRII 3.14 and TRIII 2.94.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Abdomen</emphasis>
. Brown in colour.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Genitalia</emphasis>
(Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5I</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-E Adult male genitalia structures. A Dasyhelea (Dasyhelea) incisura sp. nov.; B Dasyhelea (Dasyhelea) quasifulcillata sp. nov.; C Dasyhelea (Dasyhelea) trigona sp. nov.; D Dasyhelea (Sebessia) falxa sp. nov.; E Dasyhelea (Sebessia) folia sp. nov. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure8" httpUri="https://binary.pensoft.net/fig/790308" pageId="0" pageNumber="51">8C</figureCitation>
). Tergite 9 broad apically, 100.5-101
<normalizedToken originalValue="μm">μm</normalizedToken>
long and 172-173
<normalizedToken originalValue="μm">μm</normalizedToken>
wide. Apicolateral processes (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5J</figureCitation>
) short with basal notch, 11-12
<normalizedToken originalValue="μm">μm</normalizedToken>
long, with one prominent apical seta, cercus with three setae. Sternite 9 2
<normalizedToken originalValue="×">x</normalizedToken>
wider than length, with convex margin. Gonocoxite stout, elongated, 78
<normalizedToken originalValue="μm">μm</normalizedToken>
long, greatest width 46-57.5
<normalizedToken originalValue="μm">μm</normalizedToken>
, having 3-4 setae dorsally. Gonostylus stout, long, gradually tapering, 92-94
<normalizedToken originalValue="μm">μm</normalizedToken>
in length, 23-30
<normalizedToken originalValue="μm">μm</normalizedToken>
wide basally, 9
<normalizedToken originalValue="μm">μm</normalizedToken>
wide subapically, 7-8
<normalizedToken originalValue="μm">μm</normalizedToken>
wide apically; 2 long setae present at about proximal one fourth and one third length dorsally and 1 long seta at about proximal one third, all directed outwardly; one prominent, seta from about half distance from base, directed inward, 2 prominent setae subapically and 2 at apex. Left basal arm of parameres (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5I</figureCitation>
) 103.5
<normalizedToken originalValue="μm">μm</normalizedToken>
long, and right basal arm 90-92
<normalizedToken originalValue="μm">μm</normalizedToken>
long, both thin, highly sclerotised, joined broadly with 92
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 16-18.5
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, moderately sclerotised, and pointed tipped posteromedian projection of parameres terminating at level of gonocoxite-gonostylus junction. Aedeagus (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-K Adult male of Dasyhelea (Dasyhelea) trigona sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Wing; G Femora and tibiae (right to left) of fore, mid and hind legs; H Hind tibial comb; I Genitalia (left dorsal and right ventral view); J Apicolateral process; K Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure5" httpUri="https://binary.pensoft.net/fig/790305" pageId="0" pageNumber="51">5K</figureCitation>
) broad, 50.5-57.5
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 94-99
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, basal arm 18.5-19
<normalizedToken originalValue="μm">μm</normalizedToken>
long, highly sclerotised, directed slightly posterolaterad, ending in triangular projection ventrally, basal arch 32-44
<normalizedToken originalValue="μm">μm</normalizedToken>
long, posterolateral arm 34.5-39
<normalizedToken originalValue="μm">μm</normalizedToken>
long, stout, ending in point; bifurcated, tubular ventromedian projection.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Female adult.</emphasis>
Unknown.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Pupa and Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="etymology">
<paragraph pageId="0" pageNumber="51">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="51">
The name &quot;
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea trigona" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="trigona">
<emphasis italics="true" pageId="0" pageNumber="51">Dasyhelea trigona</emphasis>
</taxonomicName>
&quot; derived from Latinised version of
<emphasis italics="true" pageId="0" pageNumber="51">triangle</emphasis>
, referring to triangular ventral region of the basal arm of the aedeagus of male genitalia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="remarks">
<paragraph pageId="0" pageNumber="51">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="51">
The new species
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (D.) trigona" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="trigona" subGenus="D.">Dasyhelea (D.) trigona</taxonomicName>
bears similarities with
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. (D.) fulcillata" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="fulcillata" subGenus="D.">D. (D.) fulcillata</taxonomicName>
Yu in Yu et al. 2005 (China) in the shape of the gonocoxite and basal arms of the parameres, but differs in the apicolateral process, absence of setae at the basal half of gonostylus, and the overall structure of the parameres and the aedeagus. It is also close to
<taxonomicName authorityName="Yu" authorityYear="2005" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (D.) abdita" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="abdita" subGenus="D.">Dasyhelea (D.) abdita</taxonomicName>
Yu in Yu et al. 2005 (China) in the structures of parameres and aedeagus, but the basal arm of aedeagus, the apicolateral process and the apex of gonostylus are different. The present species owes resemblances with
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. (D.) actita" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="actita" subGenus="D.">D. (D.) actita</taxonomicName>
Yu in Yu et al. 2005 (Taiwan) in the structure of the posterolateral arms of the aedeagus and the basal arms of the parameres, but the overall structure of parameres and apicolateral processes are dissimilar. The new species and
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. (D.) ampullariae" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="ampullariae" subGenus="D.">D. (D.) ampullariae</taxonomicName>
Macfie, 1934 (Malaysia, and China) look alike in parameres but the structure of the gonostylus, the apicolateral process and the aedeagus disagree. The combination of characters provided in the diagnosis validates
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. (D.) trigona" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="trigona" subGenus="D.">D. (D.) trigona</taxonomicName>
as a new member of this subgenus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="distribution">
<paragraph pageId="0" pageNumber="51">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="51">The materials were collected from the Deltaic Proper of Gangetic of West Bengal at an altitude of 6-60 meters above sea level - new species from India.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="ecological notes">
<paragraph pageId="0" pageNumber="51">Ecological notes.</paragraph>
<paragraph pageId="0" pageNumber="51">
Adults of
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (D.) trigona" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="trigona" subGenus="D.">Dasyhelea (D.) trigona</taxonomicName>
sp. nov. were trapped around a paddy field adjoining a freshwater river (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-E Photographs of collection sites where light traps were operated. A Collection site of Dasyhelea (Dasyhelea) incisura sp. nov.; B Collection site of Dasyhelea (D.) quasifulcillata sp. nov.; C Collection site of Dasyhelea (D.) trigona sp. nov. (adjacent fresh water river shown as different image at upper right corner); D Collection site of Dasyhelea (Sebessia) falxa sp. nov.; E. Collection site of Dasyhelea (S.) folia sp. nov. (adjacent brackish water river is same for both D and E, shown as different image at upper middle)." figureDoi="10.3897/evolsyst.7.90973.figure2" httpUri="https://binary.pensoft.net/fig/790302" pageId="0" pageNumber="51">2C</figureCitation>
); the air temperature and relative humidity were 15-17 °C and 70-75% respectively.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">
Subgenus
<taxonomicName class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Sebessia)" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="subGenus" subGenus="Sebessia">Dasyhelea Sebessia</taxonomicName>
Remm, 1979
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>