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<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.97.115682" ID-Pensoft-Pub="2747-8106-97-53" ID-Pensoft-UUID="4A3318772B185D35B1E7EC9578EF4A47" ID-ZooBank="933EC356F21C45AF9CFA563E64D27953" ModsDocID="2747-8106-97-53" checkinTime="1702462880200" checkinUser="pensoft" docAuthor="Neubauer, Thomas A. &amp; Wesselingh, Frank P." docDate="2023" docId="2E19A21579EA5C9786B5B8F84E408DA4" docLanguage="en" docName="Zitteliana 97: 53-88" docOrigin="Zitteliana 97" docPubDate="2023-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.97.115682" docTitle="Graecoanatolica ? sp." docType="treatment" docVersion="1" id="4A3318772B185D35B1E7EC9578EF4A47" lastPageNumber="53" masterDocId="4A3318772B185D35B1E7EC9578EF4A47" masterDocTitle="The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms" masterLastPageNumber="88" masterPageNumber="53" pageNumber="53" updateTime="1702462880200" updateUser="pensoft">
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<mods:title id="16E24221C882F0167BB7873AE060068E">The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms</mods:title>
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<mods:namePart id="7E38C99974300E8E3FEB1A58EF3763C4">Neubauer, Thomas A.</mods:namePart>
<mods:nameIdentifier id="0F7A3BFFB4F8A3FEFA577460B5C3387E" type="ORCID">https://orcid.org/0000-0002-1398-9941</mods:nameIdentifier>
<mods:affiliation id="C7CD33DF7ECADCC8B1FFF9B3664BCFEE">SNSB - Bavarian State Collection for Palaeontology and Geology, Richard-Wagner-Strasse 10, 80333 Munich, Germany &amp; Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
<mods:nameIdentifier id="81AF2BAD775850E2F6D75E0E8C16117E" type="email">neubauer@snsb.de</mods:nameIdentifier>
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<mods:roleTerm id="4E0B37FEA7B2C52A1026619C87A8D1F3">Author</mods:roleTerm>
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<mods:namePart id="040556D60570C88336ECB426D5DA20DD">Wesselingh, Frank P.</mods:namePart>
<mods:nameIdentifier id="4FC752ED138A67D2EE06D0CB527AA940" type="ORCID">https://orcid.org/0000-0003-3655-0701</mods:nameIdentifier>
<mods:affiliation id="7F85D4F75D60673EBB08CD95872DABD6">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands &amp; Department of Earth Sciences, Utrecht University, Princetonlaan 8 a, 3584 CB Utrecht, Netherlands</mods:affiliation>
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<mods:title id="126105BED94F87DDB163487A15B5E1DF">Zitteliana</mods:title>
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<mods:date id="5FF276B9AE30C049275188BB59A95BCB">2023</mods:date>
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<mods:number id="DDF9CBA74A0FDA507930281434FFDC11">2023-12-12</mods:number>
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<mods:number id="2F58CCE0F0EF61F0592B0BC3BA83AD49">97</mods:number>
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<mods:url id="3CEC0173BACE82978A8289094B6FCCFF">http://dx.doi.org/10.3897/zitteliana.97.115682</mods:url>
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<mods:classification id="18CCF05969744817EFCDDAB2CE0E371B">journal article</mods:classification>
<mods:identifier id="60555CE7FBFA37255FC4AB735FA9826C" type="DOI">http://dx.doi.org/10.3897/zitteliana.97.115682</mods:identifier>
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<treatment id="2E19A21579EA5C9786B5B8F84E408DA4" LSID="urn:lsid:plazi:treatment:2E19A21579EA5C9786B5B8F84E408DA4" httpUri="http://treatment.plazi.org/id/2E19A21579EA5C9786B5B8F84E408DA4" lastPageNumber="53" pageId="0" pageNumber="53">
<subSubSection id="22EC1320C5FF288A29A479C8AB250950" pageId="0" pageNumber="53" type="nomenclature">
<paragraph id="3FEC1EBC3290D6868607F383F2A7AD7E" pageId="0" pageNumber="53">
<taxonomicName id="18ABE5F575119344EACDCBAE9CA83792" ID-CoL="7P4J5" LSID="2E19A215-79EA-5C97-86B5-B8F84E408DA4" authority="? sp." authorityName="? sp." class="Gastropoda" family="Hydrobiidae" genus="Graecoanatolica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graecoanatolica" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">Graecoanatolica? sp.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="96DB93D05EFA685527F3E1CADA532AB9" pageId="0" pageNumber="53" type="description">
<paragraph id="F88D3AEF9C860F14DD227927901E8FE7" pageId="0" pageNumber="53">
<figureCitation id="934C54B45E40E374B656478FB766EC0F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caspiinae (Hydrobiidae) of the Kolankaya I fauna. A-E. Graecoanatolica alcicekorum sp. nov., holotype, RGM 962617, sample 2. F-I. Graecoanatolica? sp., RGM 1365349, sample 1. Scale bars: 1 mm (1 - D, F, G); 100 μm (E, H, I)." figureDoi="10.3897/zitteliana.97.115682.figure6" httpUri="https://binary.pensoft.net/fig/950743" pageId="0" pageNumber="53">Fig. 6F-I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="02FF490A8D35C9FF8B97723F24C9C9A5" pageId="0" pageNumber="53" type="materials_examined">
<paragraph id="3F5CE7AE17E701D4FF4F205472ACCC17" pageId="0" pageNumber="53">Material.</paragraph>
<paragraph id="3FEE027C7859C6A592391A0E490C35FC" pageId="0" pageNumber="53">
<specimenCount id="C18E6330DB694BE013872E2D14D4193B" type="generic">1 specimen</specimenCount>
(RGM 1365349) from sample 1.
</paragraph>
</subSubSection>
<subSubSection id="6F07E99C8A46B7D19023A9FCC2393591" pageId="0" pageNumber="53" type="remarks">
<paragraph id="AC12444E34E88C77892D445522128247" pageId="0" pageNumber="53">Remarks.</paragraph>
<paragraph id="A43444D155F80A987A9789C31DBB2F26" pageId="0" pageNumber="53">
At first glance, this shell reminds of
<taxonomicName id="9EA5C31F6383C18BC007FB9944CA4065" authorityName="Neubauer &amp; Wesselingh" authorityYear="2023" class="Gastropoda" family="Hydrobiidae" genus="Iraklimelania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iraklimelania minutissima" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="minutissima">
<emphasis id="630A3D351441D8778C9BF69596E1595B" italics="true" pageId="0" pageNumber="53">Iraklimelania minutissima</emphasis>
</taxonomicName>
sp. nov. (see below) in terms of the very small, slender shell with convex whorls. However, the protoconch of
<taxonomicName id="C666795F7D8BAF5FA6DFA873C68C72A1" class="Gastropoda" family="Hydrobiidae" genus="Graecoanatolica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graecoanatolica" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="AE268D5F19B33D0408AF07ADFA6F00F3" italics="true" pageId="0" pageNumber="53">Graecoanatolica</emphasis>
</taxonomicName>
? sp. is higher, whorls are more regularly convex, the aperture is oblique in lateral view, and the umbilicus is clearly visible. In all these characteristics it closely resembles extant
<taxonomicName id="896A3E6523C40A213D98FDDA2C297D65" class="Gastropoda" family="Hydrobiidae" genus="Graecoanatolica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graecoanatolica dinarica" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="dinarica">
<emphasis id="421CBA467B132F6F801736B710C84320" italics="true" pageId="0" pageNumber="53">Graecoanatolica dinarica</emphasis>
</taxonomicName>
<normalizedToken id="1513885A5452707776F3E3E903EC6337" originalValue="Kebapçı">Kebapci</normalizedToken>
, Koca &amp;
<normalizedToken id="AD59D919C7EA5C5D835BF12F6185313A" originalValue="Yıldırım">Yildirim</normalizedToken>
, 2012, for which we have studied topotypical material from
<normalizedToken id="A0B5C9C1A55B38B2EE496A6D6691ABCB" originalValue="İncirlipınar">Incirlipinar</normalizedToken>
in Afyonkarahisar region. That species differs from
<taxonomicName id="604719C5814F639A28BD950C76A46F5D" class="Gastropoda" family="Hydrobiidae" genus="Graecoanatolica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graecoanatolica" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="313E66D12A943597000D140BE6EE45CA" italics="true" pageId="0" pageNumber="53">Graecoanatolica</emphasis>
</taxonomicName>
? sp. in the larger size, the comparatively larger apex, the faster whorl expansion rate, and the often irregular growth (
<bibRefCitation id="FAD2767FE113BD80C915AD0A3B558EB9" DOI="https://doi.org/10.3906/zoo-1011-10" author="Kebapci, Ue" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="53" pagination="399 - 411" refId="B77" refString="Kebapci, Ue, Bahadir Koca, S, Yildirim, MZ, 2012. Revision of Graecoanatolica (Gastropoda: Hydrobiidae) species in Turkey. Turkish Journal of Zoology 36: 399 - 411, DOI: https://doi.org/10.3906/zoo-1011-10" title="Revision of Graecoanatolica (Gastropoda: Hydrobiidae) species in Turkey." url="https://doi.org/10.3906/zoo-1011-10" volume="36" year="2012">
<normalizedToken id="E9E73D22BAB81A9FF81E2C42A698EC04" originalValue="Kebapçı">Kebapci</normalizedToken>
et al. 2012
</bibRefCitation>
). Yet, the growth lines of
<taxonomicName id="7F91542651986B9AF38700CBB3DC1C13" class="Gastropoda" family="Hydrobiidae" genus="Graecoanatolica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graecoanatolica" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="CACA995522A2B06877D73EF33BB154D9" italics="true" pageId="0" pageNumber="53">Graecoanatolica</emphasis>
</taxonomicName>
? sp. are nearly orthocline, which has not been observed in any
<taxonomicName id="537387137204344E00D168FA7F5A7FBC" class="Gastropoda" family="Hydrobiidae" genus="Graecoanatolica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graecoanatolica" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="6857751C0BEE5893A885A9FAEF722880" italics="true" pageId="0" pageNumber="53">Graecoanatolica</emphasis>
</taxonomicName>
species, which is why we only tentatively associate the species with this genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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