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<document id="74B1B935411641DB824A168A174AC87B" ID-CLB-Dataset="5025" ID-DOI="10.1649/0010-065X-72.3.503" ID-GBIF-Dataset="a3f528b0-9dd1-4059-9e1f-0a88c23f1e46" ID-ISSN="1938-4394" ID-Zenodo-Dep="4789344" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621946641600" checkinUser="carolina" docAuthor="Etzler, Frank E. &amp; Johnson, Paul J." docDate="2018" docId="2E10044760105409FF7C01F48FEBFA70" docLanguage="en" docName="TheColeopBull.72.3.503-521.pdf" docOrigin="The Coleopterists Bulletin 72 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-72.3.503" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Athoplastus lochsa Etzler and Johnson 2018, new species" docType="treatment" docUuid="53CAEDD4-4FC0-4042-ABEF-8D7369F90FCC" docUuidSource="ZooBank" docVersion="9" lastPageNumber="516" masterDocId="D2297C3F601D5404FFBE057D8C77FFA2" masterDocTitle="Athoplastus Johnson and Etzler (Coleoptera: Elateridae: Dendrometrinae), a New Genus of Click Beetle from the Northwestern Continental USA" masterLastPageNumber="521" masterPageNumber="503" pageNumber="516" updateTime="1698990595291" updateUser="ExternalLinkService">
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<mods:title id="1991AD3581B08969B88F5EDD681581BB">Athoplastus Johnson and Etzler (Coleoptera: Elateridae: Dendrometrinae), a New Genus of Click Beetle from the Northwestern Continental USA</mods:title>
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<mods:namePart id="CA63BC0DF33C403B7F3599896E0EB28F">Etzler, Frank E.</mods:namePart>
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<mods:namePart id="B1116D4BCF867B558101B8B0A0A6BA53">Johnson, Paul J.</mods:namePart>
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<mods:title id="F65C099175859B2E13BFF1AC313D76DA">The Coleopterists Bulletin</mods:title>
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<mods:date id="8B7B7107AC5E85318F099C9EE5DEB9BD">2018</mods:date>
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<paragraph id="A606B55160105409FF7C01F48DB2FB1A" blockId="13.[160,629,1161,1288]" pageId="13" pageNumber="516">
<heading id="FD4E023D60105409FF7C01F48DB2FB1A" centered="true" fontSize="9" level="2" pageId="13" pageNumber="516" reason="2">
<emphasis id="94CD694360105409FF7C01F48DB2FB1A" bold="true" pageId="13" pageNumber="516">
<taxonomicName id="61B9CED260105409FF7C01F48E38FB3C" authority="Etzler and Johnson" authorityName="Etzler and Johnson" authorityYear="2018" box="[194,591,1161,1182]" class="Insecta" family="Elateridae" genus="Athoplastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="516" phylum="Arthropoda" rank="species" species="lochsa" status="sp. nov.">
<emphasis id="94CD694360105409FF7C01F48D09FB3C" bold="true" box="[194,382,1161,1182]" italics="true" pageId="13" pageNumber="516">Athoplastus lochsa</emphasis>
Etzler and Johnson
</taxonomicName>
,
<taxonomicNameLabel id="8FFED43860105409FEF101DE8DB2FB1A" box="[335,453,1187,1208]" pageId="13" pageNumber="516" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EEA3E6DA60105409FF4801C38E02FAAA" pageId="13" pageNumber="516" type="description">
<paragraph id="A606B55160105409FF4801C38E68FB71" blockId="13.[160,629,1161,1288]" box="[246,543,1213,1235]" pageId="13" pageNumber="516">
(Adult:
<figureCitation id="3E82A9D460105409FEFC01C38DF2FB71" box="[322,389,1214,1235]" captionStart="Figs" captionStartId="4.[197,236,1615,1634]" captionTargetBox="[245,1120,196,1597]" captionTargetId="figure-0@4.[243,1122,194,1599]" captionTargetPageId="4" captionText="Figs. 14. Athoplastus species, dorsal habitus. 1) A. acutus; 2) A. palouse; 3) A. lochsa; 4) A. rainieri." figureDoi="http://doi.org/10.5281/zenodo.4789354" httpUri="https://zenodo.org/record/4789354/files/figure.png" pageId="13" pageNumber="516">Figs. 3</figureCitation>
,
<figureCitation id="3E82A9D460105409FE2C01C38DA7FB71" box="[402,464,1214,1235]" captionStart="Figs" captionStartId="6.[197,236,1734,1753]" captionTargetBox="[177,1188,196,1716]" captionTargetId="figure-0@6.[175,1190,194,1718]" captionTargetPageId="6" captionText="Figs. 1015. Athoplastus species. 10) A. acutus, prothorax, dorsal view; 11) A. lochsa, prothorax, dorsal view; 12) A. acutus, left antenna; 13) A. lochsa, left antenna; 14) A. palouse, left antenna; 15) A. acutus, female, left wing. Scale bars for antennae = 1 mm; for wing = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4789388" httpUri="https://zenodo.org/record/4789388/files/figure.png" pageId="13" pageNumber="516">11, 13</figureCitation>
,
<figureCitation id="3E82A9D460105409FE6201C38E6FFB71" box="[476,536,1213,1235]" captionStart="Figs" captionStartId="7.[175,214,1615,1634]" captionTargetBox="[246,1077,201,1597]" captionTargetId="figure-1@7.[244,1080,194,1599]" captionTargetPageId="7" captionText="Figs. 1627. Athoplastus species, adeagi, dorsal view, ventral view, and close up of paramere apex, respectively." figureDoi="http://doi.org/10.5281/zenodo.4789394" httpUri="https://zenodo.org/record/4789394/files/figure.png" pageId="13" pageNumber="516">1921</figureCitation>
)
</paragraph>
<paragraph id="A606B55160105409FF6A01A58E02FAAA" blockId="13.[160,629,1161,1288]" pageId="13" pageNumber="516">
Zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="D21F8F8460105409FF1E018E8E02FAAA" box="[160,629,1267,1288]" pageId="13" pageNumber="516">53CAEDD4-4FC0-4042-ABEF-8D7369F90FCC</uuid>
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</paragraph>
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<subSubSection id="EEA3E6DA60105409FF1100608E24FA4E" pageId="13" pageNumber="516" type="diagnosis">
<paragraph id="A606B55160105409FF1100608E24FA4E" blockId="13.[151,639,1309,1729]" pageId="13" pageNumber="516">
<emphasis id="94CD694360105409FF1100608D60FA90" bold="true" box="[175,279,1309,1330]" pageId="13" pageNumber="516">Diagnosis.</emphasis>
This species can be separated from all others in the genus by the combination of relatively sparse pronotal punctation (separated by greater than the diameter of a single puncture) (
<figureCitation id="3E82A9D460105409FD9700108E06FA20" box="[553,625,1389,1410]" captionStart="Figs" captionStartId="6.[197,236,1734,1753]" captionTargetBox="[177,1188,196,1716]" captionTargetId="figure-0@6.[175,1190,194,1718]" captionTargetPageId="6" captionText="Figs. 1015. Athoplastus species. 10) A. acutus, prothorax, dorsal view; 11) A. lochsa, prothorax, dorsal view; 12) A. acutus, left antenna; 13) A. lochsa, left antenna; 14) A. palouse, left antenna; 15) A. acutus, female, left wing. Scale bars for antennae = 1 mm; for wing = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4789388" httpUri="https://zenodo.org/record/4789388/files/figure.png" pageId="13" pageNumber="516">Fig. 11</figureCitation>
), sloped, non-depressed prosternum (
<figureCitation id="3E82A9D460105409FDBE00F58E36FA3F" box="[512,577,1416,1437]" captionStart="Figs" captionStartId="5.[175,214,1615,1634]" captionTargetBox="[206,1118,195,1597]" captionTargetId="figure-0@5.[204,1120,194,1599]" captionTargetPageId="5" captionText="Figs. 59. Athoplastus species. 5) A. acutus, female dorsal habitus; 6) A. acutus, male dorsal habitus, darker coloration; 7) A. acutus, male ventral habitus; 8) A. rainier, prothorax, lateral view (arrow indicates start of area between procoxae); 9) A. acutus, prothorax, lateral view (arrow indicates start of expanded area between procoxae)." figureDoi="http://doi.org/10.5281/zenodo.4789370" httpUri="https://zenodo.org/record/4789370/files/figure.png" pageId="13" pageNumber="516">Fig. 8</figureCitation>
), and antennomeres without secondary expansions (
<figureCitation id="3E82A9D460105409FDE900DF8CC6FA70" captionStart="Figs" captionStartId="6.[197,236,1734,1753]" captionTargetBox="[177,1188,196,1716]" captionTargetId="figure-0@6.[175,1190,194,1718]" captionTargetPageId="6" captionText="Figs. 1015. Athoplastus species. 10) A. acutus, prothorax, dorsal view; 11) A. lochsa, prothorax, dorsal view; 12) A. acutus, left antenna; 13) A. lochsa, left antenna; 14) A. palouse, left antenna; 15) A. acutus, female, left wing. Scale bars for antennae = 1 mm; for wing = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4789388" httpUri="https://zenodo.org/record/4789388/files/figure.png" pageId="13" pageNumber="516">Fig. 13</figureCitation>
). The male genitalia are also distinctive, with the parameres broadly rounded apically (
<figureCitation id="3E82A9D460105409FE4100AA8E30FA4E" box="[511,583,1495,1516]" captionStart="Figs" captionStartId="7.[175,214,1615,1634]" captionTargetBox="[246,1077,201,1597]" captionTargetId="figure-1@7.[244,1080,194,1599]" captionTargetPageId="7" captionText="Figs. 1627. Athoplastus species, adeagi, dorsal view, ventral view, and close up of paramere apex, respectively." figureDoi="http://doi.org/10.5281/zenodo.4789394" httpUri="https://zenodo.org/record/4789394/files/figure.png" pageId="13" pageNumber="516">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EEA3E6DA60105409FF11008F8FF5FBAC" pageId="13" pageNumber="516" type="description">
<paragraph id="A606B55160105409FF11008F8FDCFC56" blockId="13.[151,639,1309,1729]" lastBlockId="13.[685,1173,194,1490]" pageId="13" pageNumber="516">
<emphasis id="94CD694360105409FF11008F8D1BF9A5" bold="true" box="[175,364,1522,1543]" pageId="13" pageNumber="516">Description. Male.</emphasis>
Length 16.0 mm in length, width
<quantity id="614118B460105409FF6903718D53F983" box="[215,292,1548,1569]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="516" unit="mm" value="3.5">3.5 mm</quantity>
(measured across bases of elytra). Red-brown, carina, tips of mandibles, and palps darker; edges of hypomeron and abdomen lighter, almost tawny yellow (
<figureCitation id="3E82A9D460105409FEC703218DCFF9D3" box="[377,440,1628,1649]" captionStart="Figs" captionStartId="4.[197,236,1615,1634]" captionTargetBox="[245,1120,196,1597]" captionTargetId="figure-0@4.[243,1122,194,1599]" captionTargetPageId="4" captionText="Figs. 14. Athoplastus species, dorsal habitus. 1) A. acutus; 2) A. palouse; 3) A. lochsa; 4) A. rainieri." figureDoi="http://doi.org/10.5281/zenodo.4789354" httpUri="https://zenodo.org/record/4789354/files/figure.png" pageId="13" pageNumber="516">Fig. 3</figureCitation>
).
<emphasis id="94CD694360105409FE7103218E7AF9D3" bold="true" box="[463,525,1628,1649]" pageId="13" pageNumber="516">Head:</emphasis>
Ocular index = 63.1.
<emphasis id="94CD694360105409FEB2030A8D1DF92E" bold="true" box="[268,362,1655,1676]" pageId="13" pageNumber="516">Antenna:</emphasis>
Ratio of antennomeres 211: 1: 1.25: 2.9: 3: 3: 2.9: 2.9: 2.9: 2.9: 3.25 (
<figureCitation id="3E82A9D460105409FD9503EC8E05F904" box="[555,626,1681,1702]" captionStart="Figs" captionStartId="6.[197,236,1734,1753]" captionTargetBox="[177,1188,196,1716]" captionTargetId="figure-0@6.[175,1190,194,1718]" captionTargetPageId="6" captionText="Figs. 1015. Athoplastus species. 10) A. acutus, prothorax, dorsal view; 11) A. lochsa, prothorax, dorsal view; 12) A. acutus, left antenna; 13) A. lochsa, left antenna; 14) A. palouse, left antenna; 15) A. acutus, female, left wing. Scale bars for antennae = 1 mm; for wing = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4789388" httpUri="https://zenodo.org/record/4789388/files/figure.png" pageId="13" pageNumber="516">Fig. 13</figureCitation>
).
<emphasis id="94CD694360105409FF2903D18D7FF963" bold="true" box="[151,264,1708,1729]" pageId="13" pageNumber="516">Pronotum:</emphasis>
1.5X wider than long. Lateral margin completely carinate, sinuate anterad divergent hind angles, widening at anterior third; dorsoventrally flattened along lateral edges on anterior half. Punctures distinct, simple and moderately dense on disc, separated by at least diameter of a puncture, with 2 “dimples” on either side of midline just anterad midlength that are nearly impunctate; punctures remain uniform along lateral edges, becoming denser on hind angles.
<emphasis id="94CD694360105409FD1304CC8F46FE64" bold="true" box="[685,817,433,454]" pageId="13" pageNumber="516">Prosternum:</emphasis>
Slightly convex with dense, shallow, subumbilicate punctures; anterior lobe short, broadly rounded, shallowly deflexed. Posterior process short and arcuate, gradually raised ventrally between procoxae, then gradually deflexed posteriorly before becoming parallel to body plane.
<emphasis id="94CD694360105409FD4B072D8F59FDC7" bold="true" box="[757,814,592,613]" pageId="13" pageNumber="516">Legs:</emphasis>
Tarsomere 5 longest on pro- and mesotarsi, tarsomere 5 equal in length to tarsomere 1 on metatarsi.
<emphasis id="94CD694360105409FC8007F88FFFFD38" bold="true" box="[830,904,645,666]" pageId="13" pageNumber="516">Elytra:</emphasis>
Elongate, subparallel, 4.8X length of pronotum; striae shallowly impressed, deeper apically; intervals flat, punctures rough, shallow, and irregularly distributed.
<emphasis id="94CD694360105409FB8E07A888E3FD48" bold="true" box="[1072,1172,725,746]" pageId="13" pageNumber="516">Aedeagus</emphasis>
(
<figureCitation id="3E82A9D460105409FD0B078D8F59FCA7" box="[693,814,751,773]" captionStart="Figs" captionStartId="7.[175,214,1615,1634]" captionTargetBox="[246,1077,201,1597]" captionTargetId="figure-1@7.[244,1080,194,1599]" captionTargetPageId="7" captionText="Figs. 1627. Athoplastus species, adeagi, dorsal view, ventral view, and close up of paramere apex, respectively." figureDoi="http://doi.org/10.5281/zenodo.4789394" httpUri="https://zenodo.org/record/4789394/files/figure.png" pageId="13" pageNumber="516">Figs. 1921</figureCitation>
): Basal piece 1/3 total length, with roughly trapezoidal membranous section on ventral surface (
<figureCitation id="3E82A9D460105409FC8F06588F08FC98" box="[817,895,805,826]" captionStart="Figs" captionStartId="7.[175,214,1615,1634]" captionTargetBox="[246,1077,201,1597]" captionTargetId="figure-1@7.[244,1080,194,1599]" captionTargetPageId="7" captionText="Figs. 1627. Athoplastus species, adeagi, dorsal view, ventral view, and close up of paramere apex, respectively." figureDoi="http://doi.org/10.5281/zenodo.4789394" httpUri="https://zenodo.org/record/4789394/files/figure.png" pageId="13" pageNumber="516">Fig. 20</figureCitation>
); median lobe just slightly longer than parameres when measured from basal struts, slightly deflexed ventrally at tip, median carina on ventral surface, basal struts coming off at an obtuse angle; parameres subparallel, gradually narrowing toward tip, tip with expanded lateral expansion bearing 4 setae (
<figureCitation id="3E82A9D460105409FC6C06B98856FC7B" box="[978,1057,964,985]" captionStart="Figs" captionStartId="7.[175,214,1615,1634]" captionTargetBox="[246,1077,201,1597]" captionTargetId="figure-1@7.[244,1080,194,1599]" captionTargetPageId="7" captionText="Figs. 1627. Athoplastus species, adeagi, dorsal view, ventral view, and close up of paramere apex, respectively." figureDoi="http://doi.org/10.5281/zenodo.4789394" httpUri="https://zenodo.org/record/4789394/files/figure.png" pageId="13" pageNumber="516">Fig. 21</figureCitation>
), which is broadly rounded apically.
</paragraph>
<paragraph id="A606B55160105409FD7B06848FF5FBAC" blockId="13.[685,1173,194,1490]" box="[709,898,1017,1038]" pageId="13" pageNumber="516">
<emphasis id="94CD694360105409FD7B06848F62FBAC" bold="true" box="[709,789,1017,1038]" pageId="13" pageNumber="516">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="EEA3E6DA60105409FD7B016988E6FBFC" pageId="13" pageNumber="516" type="materials_examined">
<paragraph id="A606B55160105409FD7B016988E6FBFC" blockId="13.[685,1173,194,1490]" pageId="13" pageNumber="516">
<emphasis id="94CD694360105409FD7B01698FAEFB8B" bold="true" box="[709,985,1044,1065]" pageId="13" pageNumber="516">Specimens Examined (1).</emphasis>
<materialsCitation id="16D1BF0C60105409FC54016988E6FBFC" ID-GBIF-Occurrence="3333036308" collectingDate="1985-04-29" collectionCode="PJJC, USNM" collectorName="P. J. Johnson" country="United States of America" county="Idaho Co." location="Lochsa" pageId="13" pageNumber="516" specimenCount="1" stateProvince="Idaho" typeStatus="holotype">
<emphasis id="94CD694360105409FC5401698F74FBE1" bold="true" pageId="13" pageNumber="516">
<typeStatus id="79020BF360105409FC540169883FFB8B" box="[1002,1096,1044,1065]" pageId="13" pageNumber="516" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="DEAEF5C160105409FBE2016988F8FB8B" box="[1116,1167,1044,1065]" name="United States of America" pageId="13" pageNumber="516">USA</collectingCountry>
:
<collectingRegion id="647D7BB360105409FD1301538E89FBE1" box="[685,766,1070,1091]" country="United States of America" name="Idaho" pageId="13" pageNumber="516">IDAHO</collectingRegion>
:
</emphasis>
<collectingCounty id="4F67CDDD60105409FCB701538F10FBE1" box="[777,871,1070,1091]" pageId="13" pageNumber="516">Idaho Co.</collectingCounty>
;
<location id="A366E38A60105409FCCE01538FC5FBE1" LSID="urn:lsid:plazi:treatment:2E10044760105409FF7C01F48FEBFA70:A366E38A60105409FCCE01538FC5FBE1" box="[880,946,1070,1091]" country="United States of America" county="Idaho Co." name="Lochsa" pageId="13" pageNumber="516" stateProvince="Idaho">Lochsa</location>
RNA [Research Natural Area];
<date id="D207939160105409FD5001348F27FBFC" box="[750,848,1097,1118]" pageId="13" pageNumber="516" value="1985-04-29">
<collectingDate id="C2436A7960105409FD5001348F27FBFC" box="[750,848,1097,1118]" pageId="13" pageNumber="516" value="1985-04-29">29.iv.1985</collectingDate>
</date>
;
<collectorName id="0B4CD08760105409FCE501348FBCFBFC" box="[859,971,1097,1118]" pageId="13" pageNumber="516">P.J. Johnson</collectorName>
(1,
<collectionCode id="C0A82D9460105409FC4C01348857FBFC" box="[1010,1056,1097,1118]" pageId="13" pageNumber="516">PJJC</collectionCode>
to
<collectionCode id="C0A82D9460105409FBFE013488F0FBFC" box="[1088,1159,1097,1118]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="516" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EEA3E6DA60105409FD7B011E8F7EFB0C" pageId="13" pageNumber="516" type="distribution">
<paragraph id="A606B55160105409FD7B011E8F7EFB0C" blockId="13.[685,1173,194,1490]" pageId="13" pageNumber="516">
<emphasis id="94CD694360105409FD7B011E8F31FBDA" bold="true" box="[709,838,1123,1144]" pageId="13" pageNumber="516">Distribution.</emphasis>
This species is only known from the Lochsa River Valley in
<collectingRegion id="647D7BB360105409FC7601038F89FB31" box="[968,1022,1150,1171]" country="United States of America" name="Idaho" pageId="13" pageNumber="516">Idaho</collectingRegion>
County,
<collectingRegion id="647D7BB360105409FBE0010388E3FB31" box="[1118,1172,1150,1171]" country="United States of America" name="Idaho" pageId="13" pageNumber="516">Idaho</collectingRegion>
(
<figureCitation id="3E82A9D460105409FD0A01E48E8AFB0C" box="[692,765,1177,1198]" captionStart="Figs" captionStartId="9.[175,214,1615,1634]" captionTargetBox="[219,1104,195,1597]" captionTargetId="figure-1@9.[217,1106,194,1599]" captionTargetPageId="9" captionText="Figs. 2830. Athoplastus species. 28) A. rainier, female, internal genitalia, v = vagina, u = uterus, acgld = accessory gland, bc = bursa copulatrix (after Han et al. 2012); 29) Distribution map of examined specimens, with rivers; 30) Neighbor-joining tree of Montana populations of A. acutus, COI mtDNA, 500 runs, scale bar = 1% difference." figureDoi="http://doi.org/10.5281/zenodo.4789442" httpUri="https://zenodo.org/record/4789442/files/figure.png" pageId="13" pageNumber="516">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EEA3E6DA60105409FD7B01CE8FE0FABA" pageId="13" pageNumber="516" type="etymology">
<paragraph id="A606B55160105409FD7B01CE8FE0FABA" blockId="13.[685,1173,194,1490]" pageId="13" pageNumber="516">
<emphasis id="94CD694360105409FD7B01CE8F4FFB6A" bold="true" box="[709,824,1203,1224]" pageId="13" pageNumber="516">Etymology.</emphasis>
The specific name “
<taxonomicName id="61B9CED260105409FC4A01CF8845FB6A" authorityName="Etzler and Johnson" authorityYear="2018" box="[1012,1074,1202,1224]" class="Insecta" family="Elateridae" genus="Athoplastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="516" phylum="Arthropoda" rank="species" species="lochsa">
<emphasis id="94CD694360105409FC4A01CF8845FB6A" box="[1012,1074,1202,1224]" italics="true" pageId="13" pageNumber="516">lochsa</emphasis>
</taxonomicName>
” is a noun in apposition representing the Nez Perce name for the Lochsa River, along which the only known specimen was collected.
</paragraph>
</subSubSection>
<subSubSection id="EEA3E6DA60105409FD7B00608FEBFA70" pageId="13" pageNumber="516" type="discussion">
<paragraph id="A606B55160105409FD7B00608FEBFA70" blockId="13.[685,1173,194,1490]" pageId="13" pageNumber="516">
<emphasis id="94CD694360105409FD7B00608FDEFA90" bold="true" box="[709,937,1309,1330]" pageId="13" pageNumber="516">Taxonomic Remarks.</emphasis>
While it is best to describe a species based on more than
<specimenCount id="B0BF7ED860105409FB8E00458EAFFAC5" pageId="13" pageNumber="516" type="generic">one specimen</specimenCount>
, the unique pronotal punctation and male genitalia support recognition as a new species. We also feel that this description will promote collecting trips to collect more specimens of this species and expand its known range.
</paragraph>
</subSubSection>
</treatment>
</document>