400 lines
35 KiB
XML
400 lines
35 KiB
XML
<document ID-CLB-Dataset="68946" ID-DOI="http://dx.doi.org/10.3897/phytokeys.199.83107" ID-GBIF-Dataset="63a02e85-2397-4793-9475-39cb98d81425" ID-PMC="PMC9849001" ID-Pensoft-Pub="1314-2003-199-141" ID-Pensoft-UUID="18229689671F5CE29AE9AE631A94EF55" ID-PubMed="36761878" ModsDocID="1314-2003-199-141" checkinTime="1654860091157" checkinUser="pensoft" docAuthor="Hori, Kiyotaka, Khine, Phyo Kay, Fujiwara, Tao, Shin, Thant & Schneider, Harald" docDate="2022" docId="2DE698DBE2785D06907CB7CA6C9AF0EA" docLanguage="en" docName="PhytoKeys 199: 141-153" docOrigin="PhytoKeys 199" docPubDate="2022-06-09" docSource="http://dx.doi.org/10.3897/phytokeys.199.83107" docTitle="Thylacopteris minuta K. Hori & Khine 2022, sp. nov." docType="treatment" docVersion="5" id="18229689671F5CE29AE9AE631A94EF55" lastPageNumber="141" masterDocId="18229689671F5CE29AE9AE631A94EF55" masterDocTitle="Thylacopteris minuta (Polypodiaceae), a new fern species from Myanmar" masterLastPageNumber="153" masterPageNumber="141" pageNumber="141" updateTime="1732856514493" updateUser="ExternalLinkService">
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<mods:mods id="66D98EC5A66A6149466E276366C8CA82" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5E953068D5D4957CAEE55A2507243310">
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<mods:title id="C1411D5B6185E594B4B4E2B6E39BEE70">Thylacopteris minuta (Polypodiaceae), a new fern species from Myanmar</mods:title>
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</mods:titleInfo>
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<mods:name id="C1846D0EB4E73CA7962D583524B0C9B3" type="personal">
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<mods:role id="8DD64359D34A3DD44173FC8C66D85370">
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<mods:roleTerm id="D20322F351239BF583B6D0D56AD4EC79">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9C4E064C7F6F971CBA2F8355F715445E">Hori, Kiyotaka</mods:namePart>
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<mods:nameIdentifier id="126F3C071DAD6744C2177F01C4DAE4CF" type="ORCID">https://orcid.org/0000-0002-6815-7595</mods:nameIdentifier>
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<mods:affiliation id="F6D4A3B535302CC97D3BB7B945E31405">The Kochi Prefectural Makino Botanical Garden 4200 - 6 Godaisan, Kochi 781 - 8125, Japan</mods:affiliation>
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<mods:nameIdentifier id="DAD52E8796DBB887A0F9499C09D663B5" type="email">khori@makino.or.jp</mods:nameIdentifier>
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</mods:name>
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<mods:name id="FEDB770DD26F1EB0ADB2A51E8CC2B748" type="personal">
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<mods:role id="4D9BDA47650690BFDD10CCD34261694D">
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<mods:roleTerm id="93E6BB8E4A726C1B8DAB1D14C3EE7D62">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="12B4842D137AC44232D530E620C4790A">Khine, Phyo Kay</mods:namePart>
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<mods:affiliation id="CD637EA2214CF09F17BABAC974199335">Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, China</mods:affiliation>
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</mods:name>
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<mods:name id="8259E4DFAC90CF5B5502936E0C7F6764" type="personal">
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<mods:role id="107D580CCCE97FC930ADD35DA4073EB1">
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<mods:roleTerm id="283B7FA0CBEA5041BFD8881562334E2D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8AFABCC6ACB77C930A957839A309FF4F">Fujiwara, Tao</mods:namePart>
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<mods:affiliation id="C848B670F7D52357222175C1C8F87473">Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, China & Makino Herbarium, Tokyo Metropolitan University, 1 - 1 Minami-osawa, Hachioji, Tokyo 192 - 0397, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="5C1B019088FA89BF9207F052B166009C" type="personal">
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<mods:role id="4DE9504E52C85ECC5501D530BE1036F8">
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<mods:roleTerm id="F47E6235CAC62FF0FEB24D63C79D8BDD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4EC766B9F612AA871B809610B7FF3E3D">Shin, Thant</mods:namePart>
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<mods:affiliation id="7606A282DD9396AEE028021AFD9165C4">Forest Research Institute, Yezin, Nay Pyi Taw, Myanmar</mods:affiliation>
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</mods:name>
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<mods:name id="0A17EF6486A2B16B22DC2F8482088F24" type="personal">
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<mods:role id="1CC71DF46B7FAD2A3FF69E39529B453C">
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<mods:roleTerm id="0D63571AD4F9A8EFFCA22121B1104323">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F9F65B18652E5FE4B1FC15AED77BD042">Schneider, Harald</mods:namePart>
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<mods:affiliation id="E7E4E2DB993D2C060B89E95852E4F508">Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, China</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="89DC779E0652B81F6C0A37987B2DCBDB">text</mods:typeOfResource>
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<mods:titleInfo id="EDCC022247D30229BDBF3A45FBA96C19">
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<mods:title id="892C4D7FDFD2CCFAA1D7568F7EF1694B">PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="D32316AB70A0577781B0812ABB7CF134">
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<mods:date id="83C714259358E10AD2DD2D1830255F31">2022</mods:date>
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<mods:detail id="5042D2D5B61D299860CE01BD7D51FEB6" type="pubDate">
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<mods:number id="9979CBFAEC56BA3619523057295ED3A7">2022-06-09</mods:number>
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</mods:detail>
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<mods:detail id="409576BC1D3CAAF60114EC58465559C3" type="volume">
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<mods:number id="159DBAA42D475521955236839B9E0637">199</mods:number>
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</mods:detail>
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<mods:extent id="E89BC7900D36327678777E02E881FB68" unit="page">
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<mods:start id="21F49EDE6FB6E353A75515D0CE9F6C99">141</mods:start>
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<mods:end id="EAEE88E544EAF6DB9D1D2031F5CBA407">153</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="5B9248FE0152996DBC665D0F04D47B87">
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<mods:url id="D75821AE23DEBF100BD03B7E1D395CD7">http://dx.doi.org/10.3897/phytokeys.199.83107</mods:url>
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</mods:location>
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<mods:classification id="BE55A1B0651FB8B1D51CA9B8A1D08C24">journal article</mods:classification>
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<mods:identifier id="DCE0C1F677E66438FDC1625132D89475" type="DOI">http://dx.doi.org/10.3897/phytokeys.199.83107</mods:identifier>
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<mods:identifier id="8FD313E449CE50E8005CE076F40B2AFB" type="Pensoft-Pub">1314-2003-199-141</mods:identifier>
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<mods:identifier id="67E89E48406D42964684B87C6395C385" type="Pensoft-UUID">18229689671F5CE29AE9AE631A94EF55</mods:identifier>
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</mods:mods>
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<treatment id="2DE698DBE2785D06907CB7CA6C9AF0EA" ID-GBIF-Taxon="195832330" LSID="urn:lsid:plazi:treatment:2DE698DBE2785D06907CB7CA6C9AF0EA" httpUri="http://treatment.plazi.org/id/2DE698DBE2785D06907CB7CA6C9AF0EA" lastPageNumber="141" pageId="0" pageNumber="141" scope_family="Polypodiaceae">
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<subSubSection id="76AC44F2E89877E3E33FF2EC37715FB9" pageId="0" pageNumber="141" type="nomenclature">
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<paragraph id="3A8083BDE94F11C3268B833FEB1C32B8" pageId="0" pageNumber="141">
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<taxonomicName id="5663194DB17C87930D1F5F1CF7D53EC9" ID-CoL="9FT6B" LSID="2DE698DB-E278-5D06-907C-B7CA6C9AF0EA" authority="K. Hori & Khine" authorityName="K. Hori & Khine" authorityYear="2022" class="Polypodiopsida" family="Polypodiaceae" genus="Thylacopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thylacopteris minuta" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="minuta" status="sp. nov.">Thylacopteris minuta K.Hori & Khine</taxonomicName>
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<taxonomicNameLabel id="230C84123BEFAC45A614EF8A3D033CA1" pageId="0" pageNumber="141">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="512E1C1E65D7384834EBBC9DC3AC6033" pageId="0" pageNumber="141" type="diagnosis">
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<paragraph id="8158B06380B1549B0F63C860076E539B" pageId="0" pageNumber="141">Diagnosis.</paragraph>
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<paragraph id="BE573DDED4EA9047BED404FE347F2C75" pageId="0" pageNumber="141">
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<taxonomicName id="3673A573424F56E15435214E5EADBFF0" authorityName="K. Hori & Khine" authorityYear="2022" class="Polypodiopsida" family="Polypodiaceae" genus="Thylacopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thylacopteris minuta" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="minuta">
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<emphasis id="C5050D0424841AE1ABB1233372B5DEBE" italics="true" pageId="0" pageNumber="141">Thylacopteris minuta</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName id="C3A94A8AA35D2C17D0A7A3E24C4DF2E4" class="Polypodiopsida" family="Dryopteridaceae" kingdom="Plantae" lsidName="T. papillosa" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="papillosa">
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<emphasis id="EDC152930CBF67D6068C6CFA590E28BE" italics="true" pageId="0" pageNumber="141">T. papillosa</emphasis>
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</taxonomicName>
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with 20-40 sclerenchyma strands per rhizome in cross-section. However,
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<taxonomicName id="5415BEB79241D7BFBB68F98769AC111D" class="Polypodiopsida" family="Dryopteridaceae" kingdom="Plantae" lsidName="T. minuta" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="minuta">
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<emphasis id="23269DC6F9D5C6F8C7A242295F72C517" italics="true" pageId="0" pageNumber="141">T. minuta</emphasis>
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</taxonomicName>
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is distinct from
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<taxonomicName id="06A53C38665A95A6DB6F8965D1759ACD" class="Polypodiopsida" family="Dryopteridaceae" kingdom="Plantae" lsidName="T. papillosa" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="papillosa">
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<emphasis id="BD6F45BEE19DABEB9588E86B88292033" italics="true" pageId="0" pageNumber="141">T. papillosa</emphasis>
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</taxonomicName>
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with sori shallowly sunken vs.
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<taxonomicName id="F268782D0F2F3FD66F9F6DB87C859CAB" class="Polypodiopsida" family="Dryopteridaceae" kingdom="Plantae" lsidName="T. papillosa" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="papillosa">
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<emphasis id="2E611F63255E8E24ED3CEE9F85CE2A91" italics="true" pageId="0" pageNumber="141">T. papillosa</emphasis>
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</taxonomicName>
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sori deeply sunken. In addition, the lamina of
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<taxonomicName id="200777862B7719D405DE65D97F95AD54" class="Polypodiopsida" family="Dryopteridaceae" kingdom="Plantae" lsidName="T. minuta" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="minuta">
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<emphasis id="6D86AE1E3994D5EFD80476A7D5B6D3E5" italics="true" pageId="0" pageNumber="141">T. minuta</emphasis>
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</taxonomicName>
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has a maximal length of 15 cm vs. a maximal length of 59 cm in
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<taxonomicName id="C844C4F76D2864E5E733395865E743B0" class="Polypodiopsida" family="Dryopteridaceae" kingdom="Plantae" lsidName="T. papillosa" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="papillosa">
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<emphasis id="09679584FABF726B9645D03D575BBF3D" italics="true" pageId="0" pageNumber="141">T. papillosa</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="576E87E8CA7C3D4F98DD0C0828EFE650" authorityName="K. Hori & Khine" authorityYear="2022" class="Polypodiopsida" family="Polypodiaceae" genus="Thylacopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thylacopteris minuta" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="minuta">
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<emphasis id="5DC8CC0D2F279892BF62E4C5E0C5B021" italics="true" pageId="0" pageNumber="141">Thylacopteris minuta</emphasis>
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</taxonomicName>
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is distinct from the New Guinea endemic
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<taxonomicName id="B5E0FC69FBFE409AF270231644F8B3D4" class="Polypodiopsida" family="Dryopteridaceae" kingdom="Plantae" lsidName="T. diaphana" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="diaphana">
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<emphasis id="6EDD0EC1B5AD2B38568FF41D4FF530B6" italics="true" pageId="0" pageNumber="141">T. diaphana</emphasis>
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</taxonomicName>
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, which lacks sclerenchyma strands in the rhizome, has superficial sori, and lamina with a maximal length of 45 cm.
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</paragraph>
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</subSubSection>
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<subSubSection id="E98C15EFE1904C0C54B0226F994FF0DF" pageId="0" pageNumber="141" type="materials_examined">
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<paragraph id="5577CC23C7917CD448EB9B176AD8C44B" pageId="0" pageNumber="141">
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<typeStatus id="8AE3594AF168E84CE23247969E7E336D">Type</typeStatus>
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.
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</paragraph>
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<paragraph id="E11A79936667023EDA72DBCE02F968CA" pageId="0" pageNumber="141">
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<materialsCitation id="A5BC6E761BA2216C5B2E5E891E584E86" collectingDate="2019-09-30" collectorName="K. Hori, P. K. Khine, T. Fujiwara, M. Nagashima, P. P. Shwe, A. K. Moe" country="Myanmar" elevation="1325" latitude="20.995777" location="Ywangan Township" longLatPrecision="1" longitude="96.57411" municipality="Ah Lel Chaung reserve" specimenCount="1" stateProvince="Shan State" typeStatus="holotype">
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<collectingCountry id="853DFA0256142FEB94524C2EA48A589B" name="Myanmar">Myanmar</collectingCountry>
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:
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<collectingRegion id="027265BE0EDFD1404E3CA2ADCF85B2BA" country="Myanmar" name="Shan">Shan State</collectingRegion>
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;
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<collectingMunicipality id="821D4A56B0B1DB9A40CA62CE4915CE18">Ah Lel Chaung reserve</collectingMunicipality>
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forest,
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<location id="E63DB54B42D117A2AABA84A83408E884" LSID="urn:lsid:plazi:treatment:2DE698DBE2785D06907CB7CA6C9AF0EA:E63DB54B42D117A2AABA84A83408E884" country="Myanmar" latitude="20.995777" longLatPrecision="1" longitude="96.57411" municipality="Ah Lel Chaung reserve" name="Ywangan Township" stateProvince="Shan State">Ywangan Township</location>
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.
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<geoCoordinate id="ACD8181F5E5C3B6967A3EC48E25946C7" degrees="20" direction="north" minutes="59" orientation="latitude" precision="1" seconds="44.8" value="20.995777">20°59'44.8"N</geoCoordinate>
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,
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<geoCoordinate id="A8F5F474CAA95CD4666ADDEDF601141A" degrees="96" direction="east" minutes="34" orientation="longitude" precision="1" seconds="26.81" value="96.57411">96°34'26.81"E</geoCoordinate>
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, ca.
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<quantity id="B20CEBBC63C0C44DD0BF5D6A068F9DBB" metricMagnitude="3" metricUnit="m" metricValue="1.325" unit="m" value="1325.0">
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<elevation id="21B8391D2F41A20B8D18CEDAE085D0C5" metricMagnitude="3" metricUnit="m" metricValue="1.325" unit="m" value="1325.0">1325 m</elevation>
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</quantity>
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,
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<collectingDate id="7E8E84F05A3BCB0A1D373E080D40E04C" value="2019-09-30">30 Sep. 2019</collectingDate>
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,
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<collectorName id="620C6C99D17CA464D3E41F52E02FD01D">K. Hori</collectorName>
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,
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<collectorName id="B25214CB76C999C68B344B97ACC9C03A">P.K. Khine</collectorName>
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[
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<normalizedToken id="5364BF2C910F52E30DF31BEC8A0475C2" originalValue="“Kine”">"Kine"</normalizedToken>
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],
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<collectorName id="7025D316A9F06DF73A1797893AB06ED4">T. Fujiwara</collectorName>
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,
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<collectorName id="48DE7CC3B708457CE6580F0A61AC2CC5">M. Nagashima</collectorName>
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,
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<collectorName id="12EFDF36B716E624D4807DD6833362FC">P.P. Shwe</collectorName>
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&
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<collectorName id="E6330DDF1FA7A88A30F6AA1AC7F83772">A.K. Moe</collectorName>
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108601 (
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<typeStatus id="E9284C8A5919F3718E50A1FBFF85471B">holotype</typeStatus>
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: MBK 0328421 (herbarium barcode), Figs
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<figureCitation id="C89E0ADC8337959E2732570753045E61" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Thylacopteris minuta K. Hori & P. K. Khine (holotype, Hori et al. 108601 = MBK 0328421, illustration by K. Hori) A habit B abaxial view of a middle segment C adaxial view of a middle segment D cross-section of rhizome E spore F rhizome scales G sporangium. Scale bars: 5 cm (A 1 - A 3); 1 cm (B-C); 0.5 mm (D); 10 µm (E); 20 µm (F); 200 µm (G)." figureDoi="10.3897/phytokeys.199.83107.figure2" httpUri="https://binary.pensoft.net/fig/693967" pageId="0" pageNumber="141">2</figureCitation>
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-
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<figureCitation id="840FC84C987D28D12ED58F1A4E8FE393" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Thylacopteris minuta in its natural habitat (MBK 0328421). Scale bar: 5 cm. (Photo by P. K. Khine)." figureDoi="10.3897/phytokeys.199.83107.figure5" httpUri="https://binary.pensoft.net/fig/693970" pageId="0" pageNumber="141">5</figureCitation>
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isotype: HITBC, RAF)
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</materialsCitation>
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.
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</paragraph>
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<caption id="E7E5DF6E160FCF8298E452146BB23D69" doi="10.3897/phytokeys.199.83107.figure2" httpUri="https://binary.pensoft.net/fig/693967" pageId="0" pageNumber="141" start="Figure 2" startId="F2">
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<paragraph id="CBEBB6345C8F2C3DB3F3E8F935FFC3F8" pageId="0" pageNumber="141">
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<emphasis id="1F00BE761DFF7E5BA3E3EC1E3D7AF602" bold="true" pageId="0" pageNumber="141">Figure 2.</emphasis>
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<taxonomicName id="815F4E36473E0EB9D3623874391D28B9" authorityName="K. Hori & Khine" authorityYear="2022" class="Polypodiopsida" family="Polypodiaceae" genus="Thylacopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thylacopteris minuta" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="minuta">
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<emphasis id="82FC8494E47AA56E495530BF12F924C0" italics="true" pageId="0" pageNumber="141">Thylacopteris minuta</emphasis>
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</taxonomicName>
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K. Hori & P.K.Khine (
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<typeStatus id="9E313079498CCA3EC48534EB281AF8E4">holotype</typeStatus>
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,
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<emphasis id="E383531DDDB647D4BA4F5144D276CE79" italics="true" pageId="0" pageNumber="141">Hori et al. 108601</emphasis>
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=
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<emphasis id="33B04DC0D488F847034B76BECC45B22A" italics="true" pageId="0" pageNumber="141">MBK0328421</emphasis>
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, illustration by K.Hori)
|
|
<emphasis id="6B4545290E017049A1355C6C44C36089" bold="true" pageId="0" pageNumber="141">A</emphasis>
|
|
habit
|
|
<emphasis id="B6667ADF48DEC382E71DB55A96D5D794" bold="true" pageId="0" pageNumber="141">B</emphasis>
|
|
abaxial view of a middle segment
|
|
<emphasis id="A856216AF1DF6CB69F7BDB96C490205D" bold="true" pageId="0" pageNumber="141">C</emphasis>
|
|
adaxial view of a middle segment
|
|
<emphasis id="50AB05C358CDA63EC01EF5310985469A" bold="true" pageId="0" pageNumber="141">D</emphasis>
|
|
cross-section of rhizome
|
|
<emphasis id="56B49CA013EDA3A5FA9D1D57D66ECF6C" bold="true" pageId="0" pageNumber="141">E</emphasis>
|
|
spore
|
|
<emphasis id="B7F4E617B887113D5BD9790DFB555E52" bold="true" pageId="0" pageNumber="141">F</emphasis>
|
|
rhizome scales
|
|
<emphasis id="AC0AB64F9634549A267FBF6A80AB7165" bold="true" pageId="0" pageNumber="141">G</emphasis>
|
|
sporangium. Scale bars:
|
|
<quantity id="FA75445BBD46339FC351666B13BDDC94" metricMagnitude="-2" metricUnit="m" metricValue="5.0" unit="cm" value="5.0">5 cm</quantity>
|
|
(
|
|
<emphasis id="7D6B56245203A73EF588255C92F86037" bold="true" pageId="0" pageNumber="141">A1-A3</emphasis>
|
|
);
|
|
<quantity id="3B7CABA04C0347B83D3873895CD970E9" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="cm" value="1.0">1 cm</quantity>
|
|
(
|
|
<emphasis id="DBAB1AFA15C64AFCFD30CD41EFFF1304" bold="true" pageId="0" pageNumber="141">B-C</emphasis>
|
|
);
|
|
<quantity id="99FAFAD1312EEFC2D98DBDFE37EF8B51" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
|
|
(
|
|
<emphasis id="49578933870BC5CBF1E4D2FE779D3A0D" bold="true" pageId="0" pageNumber="141">D</emphasis>
|
|
); 10
|
|
<normalizedToken id="43981842E087AD365AFE33B270E4A819" originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis id="CC98D0E7E0854914B60BCD48DDAB2744" bold="true" pageId="0" pageNumber="141">E</emphasis>
|
|
); 20
|
|
<normalizedToken id="D0DD778794019FA1238471F58C46F367" originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis id="F70FF8F2580226B170CD254CFDA9CEA8" bold="true" pageId="0" pageNumber="141">F</emphasis>
|
|
); 200
|
|
<normalizedToken id="4BCD4DF39DA9F07FAC3463F7CA98C5B9" originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis id="AEAED862721F7C35191FF5F5FC0BDBF8" bold="true" pageId="0" pageNumber="141">G</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="DE8912DA045C0B1BD1BF3C6C345E3E96" pageId="0" pageNumber="141" type="epilithic">
|
|
<paragraph id="5BD0A8866D458B0BABF4039EEEA006C9" pageId="0" pageNumber="141">Epilithic.</paragraph>
|
|
<paragraph id="D8774E9AF9CBA14C8823DCE4F984F1B1" pageId="0" pageNumber="141">
|
|
<emphasis id="B239B18E860D0F08D4A6680D92878B9A" bold="true" italics="true" pageId="0" pageNumber="141">Rhizome</emphasis>
|
|
long-creeping, weakly branched, 1.0-2.0 mm in diam. (without scales), light brown, densely clothed with scales, phyllopodia sometimes prominent, these 1.0-2.0 mm high, 5.0-10.0 mm apart; 20-40 black sclerenchyma strands in rhizome, longitudinal, scattered in the ground tissue.
|
|
<emphasis id="54D64DA5128F89FC3049CCD89C6E5D46" bold="true" italics="true" pageId="0" pageNumber="141">Rhizome scales</emphasis>
|
|
evenly inserted, dull brown, fragile, adpressed or apically spreading, quite densely set, deciduous, deltoid or ovate, 1.0-1.5 mm long, 0.5-1.0 mm wide, gradually narrowed from base to apex, sometimes with wavy margins, apex acute or rounded.
|
|
<emphasis id="7EB9C93B36B428C699AE2184D0F179CF" bold="true" italics="true" pageId="0" pageNumber="141">Cell walls of rhizome scales</emphasis>
|
|
dark brown, jigsaw-puzzle-shaped and wavy at basal and central part of scales, thickened, densely warty, in a single layer or double layers in basal scales.
|
|
<emphasis id="D69D6BDDE034E43A9D72AE685C7982E9" bold="true" italics="true" pageId="0" pageNumber="141">Fronds</emphasis>
|
|
monomorphic, articulate to rhizome, petiolate.
|
|
<emphasis id="6F84AA9607D2D1175865E54E0F7DD3EC" bold="true" italics="true" pageId="0" pageNumber="141">Stipes</emphasis>
|
|
glabrous, 3.0-5.0 cm long, 0.7-1.0 mm diam, yellowish green.
|
|
<emphasis id="635FECD0ECC2B032FD47E8E3844E3628" bold="true" italics="true" pageId="0" pageNumber="141">Blades</emphasis>
|
|
membranous, lanceolate, 7.0-15.0 cm long, 2.0-4.0 cm wide, equally wide all along or rather wider above the basal part, pinnatisect, yellowish green.
|
|
<emphasis id="B4D68CB63F480FBC7ED503E1733D949D" bold="true" italics="true" pageId="0" pageNumber="141">Segments</emphasis>
|
|
glabrous, 20-30 pairs, lanceolate, ascending at an angle of 90°, 0.8-2.3 cm long, 0.3-0.5 cm wide, apically obtuse, entire at basal margin, crenate at apical margin, lower segments sometimes reduced, apical segments continuously reduced, terminal segments adnate or caudate.
|
|
<emphasis id="6518FC226F8D8EAB1B2C8909BA4D501A" bold="true" italics="true" pageId="0" pageNumber="141">Veins</emphasis>
|
|
free, once-forked, excurrent with terminal hydathodes.
|
|
<emphasis id="EAC9370795A31351C581D4CFDD4914C0" italics="true" pageId="0" pageNumber="141">Sori</emphasis>
|
|
exindusiate, uniserial on each side of costa, placed medially between costa and margin, shallowly sunken, 0.5-1.0 mm in diam., depth of papillae 0.2-0.5 mm, paraphyses absent.
|
|
<emphasis id="A393934DCF6285B8CC5121C7451E76FD" bold="true" italics="true" pageId="0" pageNumber="141">Sporangium</emphasis>
|
|
globe-shaped, long stalked, 200-250
|
|
<normalizedToken id="46A01F4A4D3FBF0DAF1F79E6C0C4AEA6" originalValue="µm">µm</normalizedToken>
|
|
in diam., annulus vertical, indurated cells 10-13.
|
|
<emphasis id="CEC97412BACFFEEF359479FD7B28D395" bold="true" italics="true" pageId="0" pageNumber="141">Spores</emphasis>
|
|
bilateral, oblong, light yellow, 40-60
|
|
<normalizedToken id="E30D01B9B7848894C9ADBE3D15A4E995" originalValue="µm">µm</normalizedToken>
|
|
long, 25-35
|
|
<normalizedToken id="62DB1065C952AF8A5317F9CC6F8B3D94" originalValue="µm">µm</normalizedToken>
|
|
wide in lateral view, laesura 20-25
|
|
<normalizedToken id="AEFF0E739BF82C8154EAA70207FF54B5" originalValue="µm">µm</normalizedToken>
|
|
long, exospore smooth, perispore thin, surface shallowly wrinkled, globules absent.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B44A7D2E1A1288684D05B00341192F14" pageId="0" pageNumber="141" type="distribution">
|
|
<paragraph id="F4037D17105AA6900607A7B9ED684B05" pageId="0" pageNumber="141">Distribution.</paragraph>
|
|
<paragraph id="92CEC459062C93C286E5D5D6DDED8B92" pageId="0" pageNumber="141">Myanmar.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E136955BBCDC0D9EA88BBFDF1AD3223C" pageId="0" pageNumber="141" type="habitat">
|
|
<paragraph id="4D03E065789619ACDEA1428C470E46C1" pageId="0" pageNumber="141">Habitat.</paragraph>
|
|
<paragraph id="4A40017F614977A2FA650456575E98DB" pageId="0" pageNumber="141">Epilithic, growing on shady surfaces of limestones (1-3 m high) in semi-evergreen or evergreen forest; altitude 940-1450 m.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="76C96EDEE54ACD952DB7E4A0B450D8F1" pageId="0" pageNumber="141" type="etymology">
|
|
<paragraph id="8A097FF4270FFEAD01B2A0C0DB28AE56" pageId="0" pageNumber="141">Etymology.</paragraph>
|
|
<paragraph id="5FF889B79E75EB0D4572008064DBA402" pageId="0" pageNumber="141">
|
|
The name refers to the relatively small size of this species compared to other species of
|
|
<taxonomicName id="A9C635F2D6E74DBF2873F466BA10932C" class="Polypodiopsida" family="Polypodiaceae" genus="Thylacopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thylacopteris" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="937FE8B4DD642E0732D8BBE95588FEE5" italics="true" pageId="0" pageNumber="141">Thylacopteris</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="33B314505834C5120115DA6C0D5E8CB6" pageId="0" pageNumber="141" type="materials_examined">
|
|
<paragraph id="F2AA26826D81AB2AB1014B30B35B9832" pageId="0" pageNumber="141">Additional specimens examined.</paragraph>
|
|
<paragraph id="6D19DD054185D475B1A021E9E607F68C" pageId="0" pageNumber="141">
|
|
<materialsCitation id="236DF9BDC1D03BC3F53EA4426C96AC26" collectingDate="2015-09-13" country="Myanmar" county="Lein Le village" elevation="947" latitude="19.994722" location="Pinlaung Township" longLatPrecision="1" longitude="96.65083" municipality="Paunglang Reserve Forest" specimenCount="1" stateProvince="Shan State">
|
|
<collectingCountry id="06667809BB72DFE3FCB524A415CB8760" name="Myanmar">Myanmar</collectingCountry>
|
|
:
|
|
<collectingRegion id="EB70EBB5DA442CC6AD9CBE3C17E6F38B" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
|
; Phaya Taung,
|
|
<collectingCounty id="36C856948327373EA3711804040D562D">Lein Le village</collectingCounty>
|
|
,
|
|
<collectingMunicipality id="78BB7147754DE2D5393D1B9014307945">Paunglang Reserve Forest</collectingMunicipality>
|
|
,
|
|
<location id="28CD0ECF4AB5A78A83E23FB05CDD7944" LSID="urn:lsid:plazi:treatment:2DE698DBE2785D06907CB7CA6C9AF0EA:28CD0ECF4AB5A78A83E23FB05CDD7944" country="Myanmar" county="Lein Le village" latitude="19.994722" longLatPrecision="1" longitude="96.65083" municipality="Paunglang Reserve Forest" name="Pinlaung Township" stateProvince="Shan State">Pinlaung Township</location>
|
|
;
|
|
<geoCoordinate id="D1CA60865646D9DEC08D53BBDC2A6B4A" degrees="19" direction="north" minutes="59" orientation="latitude" precision="1" seconds="41.0" value="19.994722">19°59'41.0"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="8C1DF402D5E44956E46955A9FD45FAA8" degrees="96" direction="east" minutes="39" orientation="longitude" precision="1" seconds="3.0" value="96.65083">96°39'3.0"E</geoCoordinate>
|
|
, ca.
|
|
<quantity id="8CEB4DE36251743AA7F4445AFF6004E0" metricMagnitude="2" metricUnit="m" metricValue="9.47" unit="m" value="947.0">
|
|
<elevation id="A08E0F1B91A048F741665F4D52C7DE6B" metricMagnitude="2" metricUnit="m" metricValue="9.47" unit="m" value="947.0">947 m</elevation>
|
|
</quantity>
|
|
alt.,
|
|
<collectingDate id="37CC8D732685BABC5A2D421B058BCC7E" value="2015-09-13">13 Sep. 2015</collectingDate>
|
|
, Y
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="AA56DBFE5130D8C73EC71661A79C8081" collectingDate="2015-09-16" collectorName="K. Kertsawang, C. Kilgour, C. Puglisi, M. Rodda, P. Srisan-ga, T. Shin, P. P. Hnin" country="Myanmar" county="Lein Le village" elevation="1448" latitude="20.048916" location="Nyaung Phyu village" longLatPrecision="1" longitude="96.7667" municipality="Paunglang Reserve Forest" specimenCount="1" stateProvince="Baba">
|
|
<collectingRegion id="AE052029319B5253C8BF4161B34D34A9" country="Myanmar" name="Chin">Baba</collectingRegion>
|
|
,
|
|
<collectorName id="5D71C07428C88661152A99CA98EB90E4">K. Kertsawang</collectorName>
|
|
,
|
|
<collectorName id="9EF11412AF3F1E55C1BA8CBE25F3F79F">C. Kilgour</collectorName>
|
|
,
|
|
<collectorName id="CFC673423B428662E068E4AAC51E3589">C. Puglisi</collectorName>
|
|
,
|
|
<collectorName id="5E5BDAEFF81B0D14A6F3CC35CFDB18AF">M. Rodda</collectorName>
|
|
,
|
|
<collectorName id="D853D6B21C7ED5EA69741F05FCC5AE6A">P. Srisan-ga</collectorName>
|
|
,
|
|
<collectorName id="532EE5B6B685EB1816E39C675A644111">T. Shin</collectorName>
|
|
&
|
|
<collectorName id="CB424165CB5F262B1AF3D809B22433AE">P.P. Hnin</collectorName>
|
|
103191 (MBK0306471, duplicates on RAF, QBG). ibid., road between
|
|
<location id="3142928B9E85988767054EA959E3B2F8" LSID="urn:lsid:plazi:treatment:2DE698DBE2785D06907CB7CA6C9AF0EA:3142928B9E85988767054EA959E3B2F8" country="Myanmar" county="Lein Le village" latitude="20.048916" longLatPrecision="1" longitude="96.7667" name="Nyaung Phyu village" stateProvince="Baba">Nyaung Phyu village</location>
|
|
and Pinglaung village, Paunglang Reserve Forest, Pinlaung Township;
|
|
<geoCoordinate id="B0061B457FECD3DA05EA418F91D21D6B" degrees="20" direction="north" minutes="02" orientation="latitude" precision="1" seconds="56.1" value="20.048916">20°02'56.1"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="EA2B6664569AB46FB1A7946093B242DC" degrees="96" direction="east" minutes="46" orientation="longitude" precision="1" seconds="00.1" value="96.7667">96°46'00.1"E</geoCoordinate>
|
|
, ca.
|
|
<quantity id="8A5062C314C34A34D43AD694C3F33B75" metricMagnitude="3" metricUnit="m" metricValue="1.448" unit="m" value="1448.0">
|
|
<elevation id="C88167E5B517474811995224E17514F7" metricMagnitude="3" metricUnit="m" metricValue="1.448" unit="m" value="1448.0">1448 m</elevation>
|
|
</quantity>
|
|
alt.,
|
|
<collectingDate id="2BD707487F91A1526E29B24571E0B72C" value="2015-09-16">16 Sep. 2015</collectingDate>
|
|
, ibid., 103361 (MBK0313746, duplicates in RAF, QBG)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="364BFD630875455EDDA7A684921E21C5" pageId="0" pageNumber="141" type="notes">
|
|
<paragraph id="CC1D7ED612B38F4B793A9FF2740D18CC" pageId="0" pageNumber="141">Notes.</paragraph>
|
|
<paragraph id="CD5BBAA83282083FE3C01B1EEADB5633" pageId="0" pageNumber="141">
|
|
The genus
|
|
<taxonomicName id="4001A96DE827591EADD461EAA48E1DF3" class="Polypodiopsida" family="Polypodiaceae" genus="Thylacopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thylacopteris" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="C17A252278260D1B6BEEFD61E48CA110" italics="true" pageId="0" pageNumber="141">Thylacopteris</emphasis>
|
|
</taxonomicName>
|
|
is sometimes confused with
|
|
<taxonomicName id="8625E1EAF47CE62BD191110599BDBDFD" class="Polypodiopsida" family="Polypodiaceae" genus="Goniophlebium" kingdom="Plantae" lsidName="Goniophlebium" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="EED7FECC789150F0E618DC1BD8F4AC2C" italics="true" pageId="0" pageNumber="141">Goniophlebium</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="444F326844F40F59BFD631D744FDEC27" class="Polypodiopsida" family="Polypodiaceae" genus="Polypodium" kingdom="Plantae" lsidName="Polypodium" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="0E0E5E6C7907B0969A0B69CC65353AE1" italics="true" pageId="0" pageNumber="141">Polypodium</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation id="5B867A1085DCE48A3BC9EEDBB7CE155D" author="Roedl-Linder, G" journalOrPublisher="Blumea" pageId="0" pageNumber="141" pagination="351 - 364" refId="B23" refString="Roedl-Linder, G, 1994. A monograph of the fern genus Thylacopteris (Polypodiaceae). Blumea 39: 351 - 364" title="A monograph of the fern genus Thylacopteris (Polypodiaceae)." volume="39" year="1994">
|
|
<normalizedToken id="4E9BF42B079686D538751747A51B470B" originalValue="Rödl-Linder">Roedl-Linder</normalizedToken>
|
|
1994
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="F54D9C001C2F15A288B0F938484D5D17" author="Fraser-Jenkins, CR" journalOrPublisher="Bishen Singh Mahendra Pal Singh, Uttarakhand" pageId="0" pageNumber="141" refId="B11" refString="Fraser-Jenkins, CR, 2020. An Annotated Checklist of Indian Pteridophytes, Part 3: Lomariopsidaceae to Salviniaceae. Bishen Singh Mahendra Pal Singh, Uttarakhand" title="An Annotated Checklist of Indian Pteridophytes, Part 3: Lomariopsidaceae to Salviniaceae." year="2020">Fraser-Jenkins 2020</bibRefCitation>
|
|
). Warty cell walls of rhizome scales (Fig.
|
|
<figureCitation id="F7A79D0FC18FECED87C7A76A27BD53A6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Warty, thickened cell walls of rhizome scales (MBK 0328421; photo taken using DP 20 microscope camera, OLYMPUS, Japan). Scale bar: 2.5 µm." figureDoi="10.3897/phytokeys.199.83107.figure3" httpUri="https://binary.pensoft.net/fig/693968" pageId="0" pageNumber="141">3</figureCitation>
|
|
) can be used to conclusively identify the genus
|
|
<taxonomicName id="F958476B1A4A4D4B14D39D76A4720C56" class="Polypodiopsida" family="Polypodiaceae" genus="Thylacopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thylacopteris" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="2A4CC462E6B61D1E80E7D5B2CE027CD7" italics="true" pageId="0" pageNumber="141">Thylacopteris</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation id="17629A1DBB572C8F603595B688A50634" author="Roedl-Linder, G" journalOrPublisher="Blumea" pageId="0" pageNumber="141" pagination="351 - 364" refId="B23" refString="Roedl-Linder, G, 1994. A monograph of the fern genus Thylacopteris (Polypodiaceae). Blumea 39: 351 - 364" title="A monograph of the fern genus Thylacopteris (Polypodiaceae)." volume="39" year="1994">
|
|
<normalizedToken id="744AE42C7F8B316D7D44F57C7B42B7A0" originalValue="Rödl-Linder">Roedl-Linder</normalizedToken>
|
|
1994
|
|
</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="A739B852A55E3347A51A4D6E5EC3A325" doi="10.3897/phytokeys.199.83107.figure3" httpUri="https://binary.pensoft.net/fig/693968" pageId="0" pageNumber="141" start="Figure 3" startId="F3">
|
|
<paragraph id="A55542C33007402F0FDE8C0ABFDA2CD8" pageId="0" pageNumber="141">
|
|
<emphasis id="D87D1FA964292CC8E97CD1EFF1850AA8" bold="true" pageId="0" pageNumber="141">Figure 3.</emphasis>
|
|
Warty, thickened cell walls of rhizome scales (
|
|
<emphasis id="E4A2683061DD34526A7F37FF6BFE314A" italics="true" pageId="0" pageNumber="141">MBK 0328421</emphasis>
|
|
; photo taken using DP20 microscope camera, OLYMPUS, Japan). Scale bar: 2.5
|
|
<normalizedToken id="5D7612EAFC45EDFB75F1E0EA65038C1A" originalValue="µm">µm</normalizedToken>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="A568B01ED2DB8FA4AA74FD3A66D4F47C" doi="10.3897/phytokeys.199.83107.figure4" httpUri="https://binary.pensoft.net/fig/693969" pageId="0" pageNumber="141" start="Figure 4" startId="F4">
|
|
<paragraph id="B45D5001A91DE1D2631334FCF19506DE" pageId="0" pageNumber="141">
|
|
<emphasis id="D67955B25723CA6DAF5005ACBA558324" bold="true" pageId="0" pageNumber="141">Figure 4.</emphasis>
|
|
The distribution of
|
|
<taxonomicName id="E0811B7626E15EB94F423605DD10A882" authorityName="K. Hori & Khine" authorityYear="2022" class="Polypodiopsida" family="Polypodiaceae" genus="Thylacopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thylacopteris minuta" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="minuta">
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<emphasis id="A621999493D4203E8FB471771E0EC92D" italics="true" pageId="0" pageNumber="141">Thylacopteris minuta</emphasis>
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</taxonomicName>
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sp.nov.
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</paragraph>
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</caption>
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<caption id="DA327A1F8A5708A31E1D876E2BE57C71" doi="10.3897/phytokeys.199.83107.figure5" httpUri="https://binary.pensoft.net/fig/693970" pageId="0" pageNumber="141" start="Figure 5" startId="F5">
|
|
<paragraph id="8028A5B938A2DE129E30C3D25B485ED4" pageId="0" pageNumber="141">
|
|
<emphasis id="AAE2C51A2F9035E2771CBEC12172450C" bold="true" pageId="0" pageNumber="141">Figure 5.</emphasis>
|
|
<taxonomicName id="D18A1BA126E252033C30F6DAD5748EC7" authorityName="K. Hori & Khine" authorityYear="2022" class="Polypodiopsida" family="Polypodiaceae" genus="Thylacopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thylacopteris minuta" order="Polypodiales" pageId="0" pageNumber="141" phylum="Tracheophyta" rank="species" species="minuta">
|
|
<emphasis id="D000455CED9CCB661764B11740B867F0" italics="true" pageId="0" pageNumber="141">Thylacopteris minuta</emphasis>
|
|
</taxonomicName>
|
|
in its natural habitat (
|
|
<emphasis id="DF62639A239C895A8CE482D017268C85" italics="true" pageId="0" pageNumber="141">MBK 0328421</emphasis>
|
|
). Scale bar: 5 cm. (Photo by P.K. Khine).
|
|
</paragraph>
|
|
</caption>
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|
</subSubSection>
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</treatment>
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</document> |