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<document id="67B5246725F038D4C6A6885E72B99CFC" ID-DOI="10.11646/zootaxa.2062.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5320706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629618014742" checkinUser="felipe" docAuthor="García-López, Alejandra, Morón, Miguel Ángel, Micó, Estefanía &amp; Galante, Eduardo" docDate="2009" docId="2D3DDF298F355D29FF4591D3FABA0099" docLanguage="en" docName="zt02062p045.pdf" docOrigin="Zootaxa 2062 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2062.1.3" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Phyllophaga (Phyllophaga) larai Garcia-Lopez &amp; Moron 2009, sp. nov." docType="treatment" docVersion="3" lastPageNumber="43" masterDocId="D104A7518F315D2FFFD2932CFFD50256" masterDocTitle="Two new species of Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) from Costa Rica" masterLastPageNumber="45" masterPageNumber="37" pageNumber="41" updateTime="1699084035901" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="EDA056CEC856D8365FCD6A26A5AD9526" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E2191D7E2D54F2195911AB7533CD9C9F">
<mods:title id="E80A36043138B68D8B5EE0AFEBD80CFC">Two new species of Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="ACAD81969C862A746ED49278ACBE2AAF" type="personal">
<mods:role id="0F01AF5AD8A3EFCB542AAF04A34E332B">
<mods:roleTerm id="ADA738C76A54AF9A17AA4C8193C25ADD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D85C0499F96FE2F795B69733D218BB70">García-López, Alejandra</mods:namePart>
<mods:affiliation id="81B2532F7E5550917C05B82531800F04">Centro Iberoamericano de la Biodiversidad (CIBIO), Universidad de Alicante, 03080 Alicante, Spain.</mods:affiliation>
</mods:name>
<mods:name id="14C46D3AE70BF4AF8DF17B90404E394F" type="personal">
<mods:role id="01F3EC13271731A5159480629B340BDE">
<mods:roleTerm id="6CF759352E6F17180691C477FFAF66E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A5CB5099B51E46A04AB473780AF4D432">Morón, Miguel Ángel</mods:namePart>
<mods:affiliation id="FD4447F9226D7C9CCBDFF8733FC5FEA0">Departamento de Entomología, Instituto de Ecología, A. C. Apdo. Postal 63, Xalapa, Veracruz 91000, México.</mods:affiliation>
</mods:name>
<mods:name id="15509B3917BB0C4711DA749B02FAF387" type="personal">
<mods:role id="043A8481504BAE67B600E7C5BBA3F14D">
<mods:roleTerm id="816E34BE98B3DA1C81F089902912A163">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D98A579399B77FACC9A60C514E3600BF">Micó, Estefanía</mods:namePart>
<mods:affiliation id="96CAEBF913BB655F74ABC52C070EE605">Centro Iberoamericano de la Biodiversidad (CIBIO), Universidad de Alicante, 03080 Alicante, Spain.</mods:affiliation>
</mods:name>
<mods:name id="A21A811AF11196A93542FEF27A2408C9" type="personal">
<mods:role id="204D951246CE4E599FC80FB61D54A9A8">
<mods:roleTerm id="765409B3F50084732F9C2329F24B7F1B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AE6C617C184BD0EACA4D0E2FF4865747">Galante, Eduardo</mods:namePart>
<mods:affiliation id="EFA2BD4DE865A482FCECEA3E6FC5F32A">Centro Iberoamericano de la Biodiversidad (CIBIO), Universidad de Alicante, 03080 Alicante, Spain.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D81F6193EFF1F15702B2A97E1B2EFBE8">text</mods:typeOfResource>
<mods:relatedItem id="5220DFF9C39F3D079F604EB7628F971D" type="host">
<mods:titleInfo id="EA8CE0D57FAEBE5FFB04F67D3C900B6A">
<mods:title id="C784EDDA5C1D9903F814693949B61AB9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1C6B8DCB993A8193EFF97FEDBD8433D4">
<mods:date id="60377738DF4D151D5D863FD4C026615A">2009</mods:date>
<mods:detail id="9A149E86ED93F6F6BFFA334D8C6AE82E" type="pubDate">
<mods:number id="CFA269B91DD85844EC7830A28AFFDBE6">2009-04-03</mods:number>
</mods:detail>
<mods:detail id="FF5062D2D341E4D817227197112D8BD8" type="volume">
<mods:number id="18B457BB64363112184498DD368A8FCA">2062</mods:number>
</mods:detail>
<mods:detail id="13CEF4FF7072AD00D52C04DA6A5F6768" type="issue">
<mods:number id="9F958ED8DD97D0F03FAB994A0EB913FA">1</mods:number>
</mods:detail>
<mods:extent id="DF54FA53E5EFB012125BFBC26465D00C" unit="page">
<mods:start id="55920E86BE1A5E024B15191280420CF5">37</mods:start>
<mods:end id="6AA9A42257DE9C3B6851ABFFD7BB2A72">45</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="F43BAFE1B15D14D7994167F62F48F0CA">
<mods:url id="C2266288F15B002263E15C8C5B49B22A">https://biotaxa.org/Zootaxa/article/view/zootaxa.2062.1.3</mods:url>
</mods:location>
<mods:classification id="7CB30F974A6272D8BF6C20F5509E1C27">journal article</mods:classification>
<mods:identifier id="0F0AF9C63FE7BB1D44C958E077CC2E9E" type="DOI">10.11646/zootaxa.2062.1.3</mods:identifier>
<mods:identifier id="703C0AF771C32282E1F12E4312968166" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DC70EB4283A323AB706D6E05C954FE7B" type="Zenodo-Dep">5320706</mods:identifier>
</mods:mods>
<treatment id="2D3DDF298F355D29FF4591D3FABA0099" LSID="urn:lsid:plazi:treatment:2D3DDF298F355D29FF4591D3FABA0099" httpUri="http://treatment.plazi.org/id/2D3DDF298F355D29FF4591D3FABA0099" lastPageId="6" lastPageNumber="43" pageId="4" pageNumber="41">
<subSubSection id="ED8E3DB48F355D2BFF4591D3FC6C014F" box="[151,953,767,793]" pageId="4" pageNumber="41" type="nomenclature">
<paragraph id="A52B6E3F8F355D2BFF4591D3FC6C014F" blockId="4.[151,953,767,832]" box="[151,953,767,793]" pageId="4" pageNumber="41">
<heading id="FE63D9538F355D2BFF4591D3FC6C014F" bold="true" box="[151,953,767,793]" fontSize="11" level="1" pageId="4" pageNumber="41" reason="1">
<emphasis id="97E0B22D8F355D2BFF4591D3FC6C014F" bold="true" box="[151,953,767,793]" pageId="4" pageNumber="41">
<taxonomicName id="629415BC8F355D2BFF4591D3FC81014F" authority="Garcia-Lopez &amp; Moron" authorityName="Garcia-Lopez &amp; Moron" authorityYear="2009" box="[151,852,767,793]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="larai" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="97E0B22D8F355D2BFF4591D3FDF1014F" bold="true" box="[151,548,767,793]" italics="true" pageId="4" pageNumber="41">Phyllophaga (Phyllophaga) larai</emphasis>
García-López &amp; Morón
</taxonomicName>
<taxonomicNameLabel id="8CD30F568F355D2BFC8E91D3FC6C014F" box="[860,953,767,793]" pageId="4" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ED8E3DB48F355D2BFF45900AFC3E042E" pageId="4" pageNumber="41" type="description">
<paragraph id="A52B6E3F8F355D2BFF45900AFEE10116" blockId="4.[151,953,767,832]" box="[151,308,806,832]" pageId="4" pageNumber="41">
(
<figureCitation id="3DAF72BA8F355D2BFF4D900AFF270116" box="[159,242,806,832]" captionStart="FIGURE 5" captionStartId="5.[151,255,1154,1178]" captionText="FIGURE 5. Phyllophaga larai sp. nov. dorsal view of holotype." pageId="4" pageNumber="41">Figs. 5</figureCitation>
,
<figureCitation id="3DAF72BA8F355D2BFF2D900AFEF90116" box="[255,300,806,832]" captionStart="FIGURES 6–9" captionStartId="5.[151,271,1978,2002]" captionTargetBox="[163,1398,1247,1936]" captionTargetId="figure-57@5.[151,1436,1232,1955]" captionTargetPageId="5" captionText="FIGURES 6–9. Phyllophaga larai, male genital capsule. 6—parameres, dorsal view. 7—Lateral view with aedeagus extended. 8—Parameres, ventroapical view. 9—Aedeagus, distal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320712" httpUri="https://zenodo.org/record/5320712/files/figure.png" pageId="4" pageNumber="41">6–9</figureCitation>
)
</paragraph>
<paragraph id="A52B6E3F8F355D2BFF45905FFC3E042E" blockId="4.[151,1437,883,1880]" pageId="4" pageNumber="41">
<emphasis id="97E0B22D8F355D2BFF45905FFE1001DB" bold="true" box="[151,453,883,909]" pageId="4" pageNumber="41">
Description.
<typeStatus id="7A2FD09D8F355D2BFE96905FFE1501DB" box="[324,448,883,909]" pageId="4" pageNumber="41" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male: Total body length:
<quantity id="626CC3DA8F355D2BFCC49058FC5F01D8" box="[790,906,884,910]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="4" pageNumber="41" unit="mm" value="21.5">21.5 mm</quantity>
. Humeral width: 9.0 mm. Clypeus, frons, pronotum and elytra reddish brown; head shiny with erect, slender, long setae; pronotum weakly pruinose, with erect, mixed long and short setae on entire surface; mouth parts, sterna, pygidium and legs shiny reddish brown (
<figureCitation id="3DAF72BA8F355D2BFF2390C8FEED01A8" box="[241,312,996,1022]" captionStart="FIGURE 5" captionStartId="5.[151,255,1154,1178]" captionText="FIGURE 5. Phyllophaga larai sp. nov. dorsal view of holotype." pageId="4" pageNumber="41">Fig. 5</figureCitation>
). Clypeus 3.8 times wider than long, densely punctate, with scattered erect short setae. Frons 5.5 times wider than dorsal diameter of eye. Eye canthus long and wide, with 9–10 setae. Mentum with anterior border briefly sinuate. Pronotum 1.8 times wider than long and 2.4 wider than frons. Pronotal disk with deep, round punctures irregularly separated from one another by 1–4 diameters; lateral borders strongly angled, lateral marginal bead crenulate and progressively dentate toward posterior corner, with regularly located, long, slender setae; basal bead indicated by setiferous punctures; anterior angles nearly acute, prominent; posterior angles obtuse, prominent. Scutellum 1.6 times wider than long. Elytra 2.4 longer than wide, densely setiferous punctuate. Pygidium moderately convex, shiny, densely setiferous punctate, with scattered long, erect setae on distal half of disk; apical margin with 12 long, slender setae; basal margin effaced medially. Anal plate with row of 14 long setae near posterior border. Mesotibia with upper apical spur acute, 1.1 times longer than lower spur. Upper apical spur of the metatibia with apex rounded, 1.2 times longer than basal metatarsomere, and 1.3 times longer than lower spur. Protarsomeres 1–3 each with subapical, small tubercle. Genital capsule with one weak lateroapical rounded prominence at each side, apex straight, progressively narrowed, with apical border slightly notched (
<figureCitation id="3DAF72BA8F355D2BFD1496E5FCFB07B5" box="[710,814,1481,1507]" captionStart="FIGURES 6–9" captionStartId="5.[151,271,1978,2002]" captionTargetBox="[163,1398,1247,1936]" captionTargetId="figure-57@5.[151,1436,1232,1955]" captionTargetPageId="5" captionText="FIGURES 6–9. Phyllophaga larai, male genital capsule. 6—parameres, dorsal view. 7—Lateral view with aedeagus extended. 8—Parameres, ventroapical view. 9—Aedeagus, distal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320712" httpUri="https://zenodo.org/record/5320712/files/figure.png" pageId="4" pageNumber="41">Fig. 6–9</figureCitation>
). Aedeagus with large, wide sclerotized support and dorsoapical tuft of 4 long bristles and ventrobasal patches of minute spines; ventroapical border of the sclerotized support downturned, widely rounded; preapical area of inner sac ventrally with patches of microspines, ending in one triangular sclerotized plate covered with microspines (
<figureCitation id="3DAF72BA8F355D2BFB419515FAD80405" box="[1171,1293,1593,1619]" captionStart="FIGURES 6–9" captionStartId="5.[151,271,1978,2002]" captionTargetBox="[163,1398,1247,1936]" captionTargetId="figure-57@5.[151,1436,1232,1955]" captionTargetPageId="5" captionText="FIGURES 6–9. Phyllophaga larai, male genital capsule. 6—parameres, dorsal view. 7—Lateral view with aedeagus extended. 8—Parameres, ventroapical view. 9—Aedeagus, distal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320712" httpUri="https://zenodo.org/record/5320712/files/figure.png" pageId="4" pageNumber="41">Figs. 6–9</figureCitation>
). Length of genital capsule from apex of parameres to border of phallobase:
<quantity id="626CC3DA8F355D2BFC589572FC32042E" box="[906,999,1630,1656]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="41" unit="mm" value="5.6">5.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="ED8E3DB48F355D29FF1495AFFD38005B" lastPageId="6" lastPageNumber="43" pageId="4" pageNumber="41" type="materials_examined">
<paragraph id="A52B6E3F8F355D2BFF1495AFFC5D050E" blockId="4.[151,1437,883,1880]" pageId="4" pageNumber="41">
<emphasis id="97E0B22D8F355D2BFF1495AFFE6C04CB" bold="true" box="[198,441,1667,1693]" pageId="4" pageNumber="41">Material examined</emphasis>
(
<specimenCount id="B392A5B68F355D2BFE0295A8FDBE04C8" box="[464,619,1668,1694]" pageId="4" pageNumber="41" type="generic">5 specimens</specimenCount>
).
<materialsCitation id="15FC64628F355D2BFD5795A8FAC20495" collectingDate="2007-07-12" collectionCode="INBio" collectorName="W. Arana." country="Costa Rica" county="Parque Nacional" elevation="1000" latitude="9.640278" location="Cerro Bitarkara" longLatPrecision="21" longitude="-83.1375" municipality="Cerere" pageId="4" pageNumber="41" specimenCount="1" stateProvince="Limon" typeStatus="holotype">
<typeStatus id="7A2FD09D8F355D2BFD5795A8FD2704C8" box="[645,754,1668,1694]" pageId="4" pageNumber="41" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="DD832EAF8F355D2BFC9D95A8FBD104C8" box="[847,1028,1668,1694]" name="Costa Rica" pageId="4" pageNumber="41">COSTA RICA</collectingCountry>
:
<collectingRegion id="6750A0DD8F355D2BFBC495A8FBB204C8" box="[1046,1127,1668,1694]" country="Costa Rica" name="Limon" pageId="4" pageNumber="41">Limón</collectingRegion>
,
<collectingCounty id="4C4A16B38F355D2BFBAB95A8FA9704C8" box="[1145,1346,1668,1694]" pageId="4" pageNumber="41">Parque Nacional</collectingCounty>
Hitoy-
<collectingMunicipality id="454FF4458F355D2BFF459585FF3D0495" box="[151,232,1705,1731]" pageId="4" pageNumber="41">Cerere</collectingMunicipality>
,
<location id="A04B38E48F355D2BFF219585FE790495" LSID="urn:lsid:plazi:treatment:2D3DDF298F355D29FF4591D3FABA0099:A04B38E48F355D2BFF219585FE790495" box="[243,428,1705,1731]" country="Costa Rica" county="Parque Nacional" latitude="9.640278" longLatPrecision="21" longitude="-83.1375" municipality="Cerere" name="Cerro Bitarkara" pageId="4" pageNumber="41" stateProvince="Limon">Cerro Bitárkara</location>
,
<quantity id="626CC3DA8F355D2BFE659585FDC70495" box="[439,530,1705,1731]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="41" unit="m" value="1000.0">
<elevation id="2EB9890C8F355D2BFE659585FDC70495" box="[439,530,1705,1731]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="41" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, (
<geoCoordinate id="C0A008F88F355D2BFDF49585FD140495" box="[550,705,1705,1731]" degrees="9" direction="north" minutes="38" orientation="latitude" pageId="4" pageNumber="41" precision="15" seconds="25" value="9.640278">9º 38’ 25’’ N</geoCoordinate>
,
<geoCoordinate id="C0A008F88F355D2BFD1E9585FCAE0495" box="[716,891,1705,1731]" degrees="83" direction="west" minutes="08" orientation="longitude" pageId="4" pageNumber="41" precision="15" seconds="15" value="-83.1375">83º 08’ 15’’ W</geoCoordinate>
),
<date id="D12A48FF8F355D2BFC5D9585FBE50495" box="[911,1072,1705,1731]" pageId="4" pageNumber="41" value="2007-07-12">
<collectingDate id="C16EB1178F355D2BFC5D9585FBE50495" box="[911,1072,1705,1731]" pageId="4" pageNumber="41" value="2007-07-12">July 12, 2007</collectingDate>
</date>
.
<collectorName id="08610BE98F355D2BFBE89585FB640495" box="[1082,1201,1705,1731]" pageId="4" pageNumber="41">W. Arana.</collectorName>
(
<collectionCode id="C385F6FA8F355D2BFB6D9585FADA0495" box="[1215,1295,1705,1731]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="4" pageNumber="41">INBio</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="15FC64628F355D2BFAF09585FC8704BE" collectingDate="2007-06-14" collectionCode="R" collectorName="Arana" country="Costa Rica" county="Parque Nacional" elevation="1000" latitude="9.640278" location="Cerro Bitarkara" longLatPrecision="21" longitude="-83.1375" municipality="Cerere" pageId="4" pageNumber="41" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
<typeStatus id="7A2FD09D8F355D2BFAF09585FA4D0495" box="[1314,1432,1705,1731]" pageId="4" pageNumber="41" type="paratype">Paratypes</typeStatus>
: same data as holotype except:
<date id="D12A48FF8F355D2BFE2E95E2FD7604BE" box="[508,675,1742,1768]" pageId="4" pageNumber="41" value="2007-06-14">
<collectingDate id="C16EB1178F355D2BFE2E95E2FD7604BE" box="[508,675,1742,1768]" pageId="4" pageNumber="41" value="2007-06-14">June 14, 2007</collectingDate>
</date>
. W.
<collectorName id="08610BE98F355D2BFD0495E2FCC804BE" box="[726,797,1742,1768]" pageId="4" pageNumber="41">Arana</collectorName>
y
<collectionCode id="C385F6FA8F355D2BFCE995E2FC8704BE" box="[827,850,1742,1768]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="41">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="15FC64628F355D2BFC8C95E2FEB40565" collectingDate="2007-06-16" collectionCode="V" collectorName="A. Garcia &amp; M. Moraga &amp; Guatuso, P. N." country="Costa Rica" county="Gonzalez" elevation="1000" latitude="9.786112" location="Pejibaye" longLatPrecision="21" longitude="-83.75166" municipality="Jimenez" pageId="4" pageNumber="41" specimenCount="1" stateProvince="Cartago" typeStatus="paratype">
<collectingCounty id="4C4A16B38F355D2BFC8C95E2FC1804BE" box="[862,973,1742,1768]" pageId="4" pageNumber="41">González</collectingCounty>
(2);
<collectingRegion id="6750A0DD8F355D2BFBD795E2FBB004BE" box="[1029,1125,1742,1768]" country="Costa Rica" name="Cartago" pageId="4" pageNumber="41">Cartago</collectingRegion>
,
<collectingMunicipality id="454FF4458F355D2BFBA395E2FB0104BE" box="[1137,1236,1742,1768]" pageId="4" pageNumber="41">Jiménez</collectingMunicipality>
,
<location id="A04B38E48F355D2BFB0D95E2FA9204BE" LSID="urn:lsid:plazi:treatment:2D3DDF298F355D29FF4591D3FABA0099:A04B38E48F355D2BFB0D95E2FA9204BE" box="[1247,1351,1742,1768]" country="Costa Rica" county="Gonzalez" latitude="9.786112" longLatPrecision="21" longitude="-83.75166" municipality="Jimenez" name="Pejibaye" pageId="4" pageNumber="41" stateProvince="Cartago">Pejibaye</location>
,
<location id="A04B38E48F355D2BFA8195E2FE940558" LSID="urn:lsid:plazi:treatment:2D3DDF298F355D29FF4591D3FABA0099:A04B38E48F355D2BFA8195E2FE940558" country="Costa Rica" county="Gonzalez" latitude="9.786112" longLatPrecision="21" longitude="-83.75166" municipality="Jimenez" name="Selva. Alb. El Copal" pageId="4" pageNumber="41" stateProvince="Cartago">Selva. Alb. El Copal</location>
,
<quantity id="626CC3DA8F355D2BFE9D95D8FE780558" box="[335,429,1780,1806]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="41" unit="m" value="1000.0">
<elevation id="2EB9890C8F355D2BFE9D95D8FE780558" box="[335,429,1780,1806]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="41" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, (
<geoCoordinate id="C0A008F88F355D2BFE1195D8FDB70558" box="[451,610,1780,1806]" degrees="9" direction="north" minutes="47" orientation="latitude" pageId="4" pageNumber="41" precision="15" seconds="10" value="9.786112">9º 47’ 10” N</geoCoordinate>
,
<geoCoordinate id="C0A008F88F355D2BFDBD95D8FCC00558" box="[623,789,1780,1806]" degrees="83" direction="west" minutes="45" orientation="longitude" pageId="4" pageNumber="41" precision="15" seconds="6" value="-83.75166">83º 45’ 6” W</geoCoordinate>
).
<date id="D12A48FF8F355D2BFCF895D8FC020558" box="[810,983,1780,1806]" pageId="4" pageNumber="41" value="2007-06-16">
<collectingDate id="C16EB1178F355D2BFCF895D8FC020558" box="[810,983,1780,1806]" pageId="4" pageNumber="41" value="2007-06-16">June 16, 2007</collectingDate>
</date>
.
<collectorName id="08610BE98F355D2BFC3695D8FB880558" box="[996,1117,1780,1806]" pageId="4" pageNumber="41">A. García</collectorName>
,
<collectorName id="08610BE98F355D2BFBB895D8FB270558" box="[1130,1266,1780,1806]" pageId="4" pageNumber="41">M. Moraga</collectorName>
(1);
<collectingRegion id="6750A0DD8F355D2BFAFD95D8FA4D0558" box="[1327,1432,1780,1806]" country="Costa Rica" name="Alajuela" pageId="4" pageNumber="41">Alajuela</collectingRegion>
,
<collectorName id="08610BE98F355D2BFF459435FE960565" box="[151,323,1817,1843]" pageId="4" pageNumber="41">Guatuso, P. N.</collectorName>
<collectionCode id="C385F6FA8F355D2BFE9E9435FEB40565" box="[332,353,1817,1843]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="41" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="15FC64628F355D2BFEBF9435FE65050E" collectingDate="2008-05-05" collectorName="Falda N &amp; J. A. Azofeifa" country="Costa Rica" elevation="1160" latitude="10.695833" location="Montezuma" longLatPrecision="1" longitude="-85.01617" municipality="Tenorio" pageId="4" pageNumber="41" specimenCount="1" stateProvince="Cartago" typeStatus="paratype">
<collectingMunicipality id="454FF4458F355D2BFEBF9435FE190565" box="[365,460,1817,1843]" pageId="4" pageNumber="41">Tenorio</collectingMunicipality>
,
<collectorName id="08610BE98F355D2BFE0B9435FDEC0565" box="[473,569,1817,1843]" pageId="4" pageNumber="41">Falda N</collectorName>
cerro
<location id="A04B38E48F355D2BFD549435FCC20565" LSID="urn:lsid:plazi:treatment:2D3DDF298F355D29FF4591D3FABA0099:A04B38E48F355D2BFD549435FCC20565" box="[646,791,1817,1843]" country="Costa Rica" latitude="10.695833" longLatPrecision="1" longitude="-85.01617" municipality="Tenorio" name="Montezuma" pageId="4" pageNumber="41" stateProvince="Cartago">Montezuma</location>
,
<quantity id="626CC3DA8F355D2BFCF69435FC550565" box="[804,896,1817,1843]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="4" pageNumber="41" unit="m" value="1160.0">
<elevation id="2EB9890C8F355D2BFCF69435FC550565" box="[804,896,1817,1843]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="4" pageNumber="41" unit="m" value="1160.0">1160 m</elevation>
</quantity>
, (
<geoCoordinate id="C0A008F88F355D2BFC449435FBB40565" box="[918,1121,1817,1843]" degrees="10" direction="north" minutes="41" orientation="latitude" pageId="4" pageNumber="41" precision="1" seconds="45.00" value="10.695833">10º 41' 45.00&quot; N</geoCoordinate>
,
<geoCoordinate id="C0A008F88F355D2BFBBC9435FAE40565" box="[1134,1329,1817,1843]" degrees="85" direction="west" minutes="0" orientation="longitude" pageId="4" pageNumber="41" precision="1" seconds="58.20" value="-85.01617">85º 0' 58.20&quot; W</geoCoordinate>
),
<date id="D12A48FF8F355D2BFA959435FF00050E" pageId="4" pageNumber="41" value="2008-05-05">
<collectingDate id="C16EB1178F355D2BFA959435FF00050E" pageId="4" pageNumber="41" value="2008-05-05">May 5, 2008</collectingDate>
</date>
.
<collectorName id="08610BE98F355D2BFF329412FE51050E" box="[224,388,1854,1880]" pageId="4" pageNumber="41">J. A. Azofeifa</collectorName>
(1)
</materialsCitation>
.
<materialsCitation id="15FC64628F355D2BFE6E9412FC51050E" box="[444,900,1854,1880]" collectionCode="CEUA, MXAL" country="Costa Rica" pageId="4" pageNumber="41" specimenCount="1" stateProvince="Cartago" typeStatus="paratype">
<typeStatus id="7A2FD09D8F355D2BFE6E9412FDFB050E" box="[444,558,1854,1880]" pageId="4" pageNumber="41" type="paratype">Paratypes</typeStatus>
deposited in
<collectionCode id="C385F6FA8F355D2BFD199412FCCB050E" box="[715,798,1854,1880]" pageId="4" pageNumber="41">CEUA</collectionCode>
,
<collectionCode id="C385F6FA8F355D2BFCF89412FC51050E" box="[810,900,1854,1880]" pageId="4" pageNumber="41">MXAL</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="F1EB3EB78F345D2AFF4597AEFC9C06CC" box="[151,841,1154,1178]" pageId="5" pageNumber="42" startId="5.[151,255,1154,1178]">
<paragraph id="A52B6E3F8F345D2AFF4597AEFC9C06CC" blockId="5.[151,841,1154,1178]" box="[151,841,1154,1178]" pageId="5" pageNumber="42">
<emphasis id="97E0B22D8F345D2AFF4597AEFECD06CC" bold="true" box="[151,280,1154,1178]" pageId="5" pageNumber="42">FIGURE 5.</emphasis>
<taxonomicName id="629415BC8F345D2AFEF297AFFE3606CC" authorityName="Garcia-Lopez &amp; Moron" authorityYear="2009" box="[288,483,1155,1178]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="larai" status="sp. nov.">
<emphasis id="97E0B22D8F345D2AFEF297AFFE3606CC" box="[288,483,1155,1178]" italics="true" pageId="5" pageNumber="42">Phyllophaga larai</emphasis>
</taxonomicName>
<emphasis id="97E0B22D8F345D2AFE3B97AEFDEB06CC" bold="true" box="[489,574,1154,1178]" pageId="5" pageNumber="42">
<taxonomicNameLabel id="8CD30F568F345D2AFE3B97AEFDEB06CC" box="[489,574,1154,1178]" pageId="5" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
dorsal view of holotype.
</paragraph>
</caption>
<caption id="F1EB3EB78F345D2AFF459496FBEA05A2" ID-DOI="http://doi.org/10.5281/zenodo.5320712" ID-Zenodo-Dep="5320712" httpUri="https://zenodo.org/record/5320712/files/figure.png" pageId="5" pageNumber="42" startId="5.[151,271,1978,2002]" targetBox="[163,1398,1247,1936]" targetPageId="5">
<paragraph id="A52B6E3F8F345D2AFF459496FBEA05A2" blockId="5.[151,1435,1978,2036]" pageId="5" pageNumber="42">
<emphasis id="97E0B22D8F345D2AFF459496FE930584" bold="true" box="[151,326,1978,2002]" pageId="5" pageNumber="42">FIGURES 6–9.</emphasis>
<taxonomicName id="629415BC8F345D2AFE9C9497FDC30584" authorityName="Garcia-Lopez &amp; Moron" authorityYear="2009" box="[334,534,1979,2002]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="larai">
<emphasis id="97E0B22D8F345D2AFE9C9497FDC30584" box="[334,534,1979,2002]" italics="true" pageId="5" pageNumber="42">Phyllophaga larai</emphasis>
</taxonomicName>
, male genital capsule. 6—parameres, dorsal view. 7—Lateral view with aedeagus extended. 8—Parameres, ventroapical view. 9—Aedeagus, distal view. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="A52B6E3F8F375D29FF1493BBFEEC02A9" blockId="6.[151,1436,151,719]" pageId="6" pageNumber="43">
<emphasis id="97E0B22D8F375D29FF1493BBFE6C02E7" bold="true" box="[198,441,151,177]" pageId="6" pageNumber="43">
<typeStatus id="7A2FD09D8F375D29FF1493BBFEED02E7" box="[198,312,151,177]" pageId="6" pageNumber="43" type="paratype">Paratype</typeStatus>
variation.
</emphasis>
Similar to the
<typeStatus id="7A2FD09D8F375D29FDB593B4FD1802E4" box="[615,717,152,178]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
except in total body length: 20.3–22.0 mm. The extension of the pruinose area of the pronotum is variable. Protibial apical spur straight, sharply pointed, shorter than basal protarsomere.
</paragraph>
<paragraph id="A52B6E3F8F375D29FF149227FB11031A" blockId="6.[151,1436,151,719]" pageId="6" pageNumber="43">
<emphasis id="97E0B22D8F375D29FF149227FEB10373" bold="true" box="[198,356,267,293]" pageId="6" pageNumber="43">Type locality</emphasis>
. Cerro Bitárkara. Área de Conservación La Amistad-Caribe, Parque Nacional Hitoy-Cerere. Province of
<collectingRegion id="6750A0DD8F375D29FEF4921EFEAD031A" box="[294,376,306,332]" country="Costa Rica" name="Limon" pageId="6" pageNumber="43">Limón</collectingRegion>
.
<collectingCountry id="DD832EAF8F375D29FE51921EFDD3031A" box="[387,518,306,332]" name="Costa Rica" pageId="6" pageNumber="43">Costa Rica</collectingCountry>
. (9º 38’ 25’’ latitude N, 83º 08’ 15’’ longitude W) (
<figureCitation id="3DAF72BA8F375D29FB8C921EFB60031A" box="[1118,1205,306,332]" captionStart="FIGURE 10" captionStartId="6.[151,255,1891,1915]" captionTargetBox="[192,1391,778,1844]" captionTargetId="figure-273@6.[151,1436,743,1898]" captionTargetPageId="6" captionText="FIGURE 10. Distribution of Phyllophaga larai, P. copalensis, P. guapiles and P. guapiloides in Costa Rica. Life Zones map provided by GIS Unit of Inbio. Distribution of P. guapiles and P. guapiloides taken from ATTA database (InBio)." figureDoi="http://doi.org/10.5281/zenodo.5320714" httpUri="https://zenodo.org/record/5320714/files/figure.png" pageId="6" pageNumber="43">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="A52B6E3F8F375D29FF149274FD38005B" blockId="6.[151,1436,151,719]" pageId="6" pageNumber="43">
<emphasis id="97E0B22D8F375D29FF149274FE5C0324" bold="true" box="[198,393,344,370]" pageId="6" pageNumber="43">Biological data.</emphasis>
Phenology: May–July. Other species of
<taxonomicName id="629415BC8F375D29FCA29275FBD30324" authorityName="Harris" authorityYear="1826" box="[880,1030,345,370]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="97E0B22D8F375D29FCA29275FBD30324" box="[880,1030,345,370]" italics="true" pageId="6" pageNumber="43">Phyllophaga</emphasis>
</taxonomicName>
flying at the same time and place were
<taxonomicName id="629415BC8F375D29FF0892ACFE3703CC" authority="(Moron)" baseAuthorityName="Moron" box="[218,482,384,410]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="chorotega">
<emphasis id="97E0B22D8F375D29FF0892ACFEA403CF" box="[218,369,384,409]" italics="true" pageId="6" pageNumber="43">P. chorotega</emphasis>
(Morón)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FE2292ACFD0D03CF" authority="(Moser)" baseAuthorityName="Moser" baseAuthorityYear="1918" box="[496,728,384,410]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="densata">
<emphasis id="97E0B22D8F375D29FE2292ACFDB903CF" box="[496,620,384,409]" italics="true" pageId="6" pageNumber="43">P. densata</emphasis>
(Moser)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FD3492ACFC0403CC" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1888" box="[742,977,384,410]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="97E0B22D8F375D29FD3492ACFCBB03CF" box="[742,878,384,409]" italics="true" pageId="6" pageNumber="43">P. gigantea</emphasis>
(Bates)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FC0C92ACFAAD03CC" authority="(Moron &amp; Solis)" baseAuthorityName="Moron &amp; Solis" box="[990,1400,384,410]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="guanacasteca">
<emphasis id="97E0B22D8F375D29FC0C92ACFB7703CF" box="[990,1186,384,409]" italics="true" pageId="6" pageNumber="43">P. guanacasteca</emphasis>
(Morón &amp; Solís)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FA5492ACFEBB0396" authority="(Saylor)" baseAuthorityName="Saylor" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="guapiles">
<emphasis id="97E0B22D8F375D29FA5492ACFF2B0396" italics="true" pageId="6" pageNumber="43">P. guapiles</emphasis>
(Saylor)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FEAE928BFCD70396" authority="(Moron &amp; Solis)" baseAuthorityName="Moron &amp; Solis" box="[380,770,422,448]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="guapiloides">
<emphasis id="97E0B22D8F375D29FEAE928BFDFC0396" box="[380,553,423,448]" italics="true" pageId="6" pageNumber="43">P. guapiloides</emphasis>
(Morón &amp; Solís)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FCC2928BFBD00396" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1888" box="[784,1029,422,448]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="lissopyge">
<emphasis id="97E0B22D8F375D29FCC2928BFC750396" box="[784,928,423,448]" italics="true" pageId="6" pageNumber="43">P. lissopyge</emphasis>
(Bates)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FBC1928BFAAD0396" authority="(Moron &amp; Solis)" baseAuthorityName="Moron &amp; Solis" box="[1043,1400,422,448]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="lorencita">
<emphasis id="97E0B22D8F375D29FBC1928BFB760396" box="[1043,1187,423,448]" italics="true" pageId="6" pageNumber="43">P. lorencita</emphasis>
(Morón &amp; Solís)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FA54928BFE5E03B1" authority="(Saylor)" baseAuthorityName="Saylor" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="nevermanni">
<emphasis id="97E0B22D8F375D29FA54928BFEF403B0" italics="true" pageId="6" pageNumber="43">P. nevermanni</emphasis>
(Saylor)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FE4592E1FDA103B1" authority="(Moser)" baseAuthorityName="Moser" baseAuthorityYear="1918" box="[407,628,461,487]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="orosina">
<emphasis id="97E0B22D8F375D29FE4592E1FDDE03B0" box="[407,523,461,486]" italics="true" pageId="6" pageNumber="43">P. orosina</emphasis>
(Moser)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FD5292E1FC5F03B1" authority="(Chapin)" baseAuthorityName="Chapin" box="[640,906,461,487]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="panamana">
<emphasis id="97E0B22D8F375D29FD5292E1FCCD03B0" box="[640,792,461,486]" italics="true" pageId="6" pageNumber="43">P. panamana</emphasis>
(Chapin)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FC4492E1FBA403B1" authority="(Bates)" baseAuthorityName="Bates" box="[918,1137,461,487]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="97E0B22D8F375D29FC4492E1FBC703B0" box="[918,1042,461,486]" italics="true" pageId="6" pageNumber="43">P. prolixus</emphasis>
(Bates)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FBAE92E1FA4203B1" authority="(Blanchard)" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[1148,1431,461,487]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="pruinosa">
<emphasis id="97E0B22D8F375D29FBAE92E1FAD403B0" box="[1148,1281,461,486]" italics="true" pageId="6" pageNumber="43">P. pruinosa</emphasis>
(Blanchard)
</taxonomicName>
,
<taxonomicName id="629415BC8F375D29FF4592D8FE620058" authority="(Blanchard)" baseAuthorityName="Blanchard" box="[151,439,500,526]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="setidorsis">
<emphasis id="97E0B22D8F375D29FF4592D8FEF1005B" box="[151,292,500,525]" italics="true" pageId="6" pageNumber="43">P. setidorsis</emphasis>
(Blanchard)
</taxonomicName>
and
<taxonomicName id="629415BC8F375D29FE2292D8FD58005B" box="[496,653,500,525]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="copalensis" status="sp. nov.">
<emphasis id="97E0B22D8F375D29FE2292D8FD58005B" box="[496,653,500,525]" italics="true" pageId="6" pageNumber="43">P. copalensis</emphasis>
</taxonomicName>
<emphasis id="97E0B22D8F375D29FD4292DFFD38005B" bold="true" box="[656,749,499,525]" pageId="6" pageNumber="43">
<taxonomicNameLabel id="8CD30F568F375D29FD4292DFFD38005B" box="[656,749,499,525]" pageId="6" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="ED8E3DB48F375D29FF149136FBDC00D4" pageId="6" pageNumber="43" type="discussion">
<paragraph id="A52B6E3F8F375D29FF149136FBDC00D4" blockId="6.[151,1436,151,719]" pageId="6" pageNumber="43">
<emphasis id="97E0B22D8F375D29FF149136FEEA0062" bold="true" box="[198,319,538,564]" pageId="6" pageNumber="43">Remarks.</emphasis>
<taxonomicName id="629415BC8F375D29FE9B9136FDF70065" authorityName="Garcia-Lopez &amp; Moron" authorityYear="2009" box="[329,546,538,563]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="larai">
<emphasis id="97E0B22D8F375D29FE9B9136FDF70065" box="[329,546,538,563]" italics="true" pageId="6" pageNumber="43">Phyllophaga larai</emphasis>
</taxonomicName>
is most closely related to
<taxonomicName id="629415BC8F375D29FCB69136FC2A0065" box="[868,1023,538,563]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="copalensis">
<emphasis id="97E0B22D8F375D29FCB69136FC2A0065" box="[868,1023,538,563]" italics="true" pageId="6" pageNumber="43">P. copalensis</emphasis>
</taxonomicName>
.
<taxonomicName id="629415BC8F375D29FBC29136FB3D0065" authorityName="Garcia-Lopez &amp; Moron" authorityYear="2009" box="[1040,1256,538,563]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="larai">
<emphasis id="97E0B22D8F375D29FBC29136FB3D0065" box="[1040,1256,538,563]" italics="true" pageId="6" pageNumber="43">Phyllophaga larai</emphasis>
</taxonomicName>
is also closely related to
<taxonomicName id="629415BC8F375D29FEC7916DFE23000D" authority="Saylor" authorityName="Saylor. Moron and Solis" authorityYear="2001" box="[277,502,577,603]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="guapiles">
<emphasis id="97E0B22D8F375D29FEC7916DFE48000C" box="[277,413,577,602]" italics="true" pageId="6" pageNumber="43">P. guapiles</emphasis>
Saylor
</taxonomicName>
from the same area of
<collectingCountry id="DD832EAF8F375D29FCCB916DFC71000D" box="[793,932,577,603]" name="Costa Rica" pageId="6" pageNumber="43">Costa Rica</collectingCountry>
. The differences between
<taxonomicName id="629415BC8F375D29FB3E916DFA92000C" authorityName="Garcia-Lopez &amp; Moron" authorityYear="2009" box="[1260,1351,577,602]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="larai">
<emphasis id="97E0B22D8F375D29FB3E916DFA92000C" box="[1260,1351,577,602]" italics="true" pageId="6" pageNumber="43">P. larai</emphasis>
</taxonomicName>
and
<taxonomicName id="629415BC8F375D29FA54916DFF2F00D7" authorityName="Saylor. Moron and Solis" authorityYear="2001" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="guapiles">
<emphasis id="97E0B22D8F375D29FA54916DFF2F00D7" italics="true" pageId="6" pageNumber="43">P. guapiles</emphasis>
</taxonomicName>
are identical to those found between
<taxonomicName id="629415BC8F375D29FD639144FC9C00D7" box="[689,841,616,641]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="copalensis">
<emphasis id="97E0B22D8F375D29FD639144FC9C00D7" box="[689,841,616,641]" italics="true" pageId="6" pageNumber="43">P. copalensis</emphasis>
</taxonomicName>
and
<taxonomicName id="629415BC8F375D29FC509144FBD700D7" authorityName="Saylor. Moron and Solis" authorityYear="2001" box="[898,1026,616,641]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="guapiles">
<emphasis id="97E0B22D8F375D29FC509144FBD700D7" box="[898,1026,616,641]" italics="true" pageId="6" pageNumber="43">P. guapiles</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="ED8E3DB48F375D29FF1491A2FABA0099" pageId="6" pageNumber="43" type="etymology">
<paragraph id="A52B6E3F8F375D29FF1491A2FABA0099" blockId="6.[151,1436,151,719]" pageId="6" pageNumber="43">
<emphasis id="97E0B22D8F375D29FF1491A2FE8600FE" bold="true" box="[198,339,654,680]" pageId="6" pageNumber="43">Etymology.</emphasis>
With a sincere and warm embrace, this species is dedicated to the memory of Juan Ignacio Lara Pozuelo (Nacho), a tireless and eternal friend. His name will now inhabit the tropical forests forever.
</paragraph>
</subSubSection>
</treatment>
</document>
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