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<mods:title id="3EE4D37463E79206054AD0ED27317980">A taxonomic revision of the Meranoplus F. Smith of Madagascar (Hymenoptera: Formicidae: Myrmicinae) with keys to species and diagnosis of the males.</mods:title>
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<mods:namePart id="B4ADC900A4F66D27ACD0F7AAB56E39DB">Boudinot, B. E.</mods:namePart>
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<mods:namePart id="0C78CF0044412A2B767C978E01868156">B. L. Fisher</mods:namePart>
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<paragraph id="264F9769A797E3524B240AD74C325CB6" box="[118,567,118,138]" pageId="32" pageNumber="333" type="mainText">
<taxonomicName id="BE4DAE921DCE65D2AFCE7B7AA8572EA1" authority="Boudinot &amp; Fisher" authorityName="Boudinot &amp; Fisher" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="sylvarius">
<pageBreakToken id="B40BC216991279A823838DC21B720F5E" pageId="32" pageNumber="333" start="start">Meranoplus</pageBreakToken>
sylvarius Boudinot &amp; Fisher
</taxonomicName>
<taxonomicNameLabel id="7003BF03E499E1D72B78FCFA30BECC5C" pageId="32" pageNumber="333" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C7D5B25906EEF67B93B010430A9B6412" pageId="32" pageNumber="333" type="description">
<paragraph id="20FA8B3BCC0296C12B8D8A2A71A453E5" box="[118,399,146,165]" pageId="32" pageNumber="333" type="mainText">(Figs. 10, 23, 28, 29, 57-62, 66)</paragraph>
</subSubSection>
<subSubSection id="9AAA65F02CFB42DBEF1206A3A2E48E48" pageId="32" pageNumber="333" type="materials_examined">
<paragraph id="EF64A3343A4A1960E23524992F2E33F8" box="[118,1125,198,273]" pageId="32" pageNumber="333" type="mainText">Holotype worker, MADAGASCAR: Fianarantsoa, Vevembe, 22°47.46' S, 47°10.91' E, 600 m, rainforest transition to montane forest, collection code BLF14131, 23-24Apr2006 (B.L. Fisher et al.) (CASC: CASENT017529). Paratype gyne with same data as holotype worker (CASC: CASENT0317567).</paragraph>
<paragraph id="EEC57BFA3E807724834C9175A9F0C516" box="[118,1121,282,413]" pageId="32" pageNumber="333" type="mainText">Paratype workers with same data as holotype worker (BMNH: CASENT0317527; CASC: CASENT0317536, CASENT0317566, CASENT0070611, CASENT0317528, CASENT0317540-CASENT0317542, CASENT0317557-CASENT0317559, CASENT0317544, CASENT0317547, CASENT0317548, CASENT0317548-CASENT0317552, CASENT0317554-CASENT0317556; MHNG: CASENT0317538; MCZ: CASENT0317543; USNM: CASENT0317549; NHMB: CASENT0317553).</paragraph>
</subSubSection>
<subSubSection id="1E8D91004DE54599AA91D94E97F62630" lastPageId="36" lastPageNumber="337" pageId="32" pageNumber="333" type="description">
<paragraph id="31A545BBF552FC7FC0EE7A145C07FEF4" box="[118,226,446,465]" pageId="32" pageNumber="333" type="mainText">Description</paragraph>
<paragraph id="305D43603825214E93602B9072857D75" box="[118,1121,474,662]" pageId="32" pageNumber="333" type="mainText">
Worker (holotype in parentheses). HL 1.03-1.13 (1.06), HW 0.90-1.02 (0.96), HLA 0.37-0.41 (0.39), CW 0.36-0.38 (0.38), CDD 0.09-0.11 (0.11), SL 0.73-0.80 (0.78), EL 0.22-0.23 (0.23), EW 0.15-0.16 (0.16), PML 0.72-0.80 (0.76), PWA 0.88-0.98 (0.94), PWP 0.47-0.53 (0.48), SPL 0.19-0.22 (0.21), WL 1.04-1.17 (1.12), PTL 0.23-0.25 (0.25), PTH 0.43-0.49 (0.46), PPL 0.23-0.24 (0.24), PPH 0.40-0.45 (0.42), ATW 1.23-1.34 (1.32), ATL 1.27-1.48 (1.42), CI 87.4-90.6 (90.1), SI 77.7-81.3 (81.1), OMI 58.3-59.6 (59.4), CDI 23.9
<normalizedToken id="22D0B6F7C89FEB28E94C3BF62743DBBB" originalValue="-" pageId="32" pageNumber="333">--</normalizedToken>
31.2 (27.3), SEI 324-355 (336), PMI 122.7-124.4 (122.8), PWI 51.5-55.3 (51.5), CS 1.0-1.1 (1.0), EYE 37.7-40.4 (38.5) (7 measured).
</paragraph>
<paragraph id="DF5FAF1C07354B67BBA1766DE5F59D6D" box="[155,450,672,691]" pageId="32" pageNumber="333" type="mainText">Chocolate brown to almost black.</paragraph>
<paragraph id="2441255EFAB999D3392EFD1CF660928D" box="[118,1121,700,916]" pageId="32" pageNumber="333" type="mainText">
Head longer than broad (CI 77-82). Mandibles with nearly parallel striation. Face with dilute
<taxonomicName id="6F27AE1A07A9902C40299D74D30CFE4D" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
which veer away from posterior margin of head; area anterior to nuchal carina without rugae or areolation. Area around eyes costate;
<taxonomicName id="9F430CE0DB7C375823188B5569346F21" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
in malar area somewhat rugose. Frontal carinae comparatively narrow, not concealing margin of eyes in full-face view. Scrobal carina weakly developed; anterior half bulging. Anterior margin of clypeus with weakly bilobed lamina; lobes close-set (CDI 24-31). Middle portion of clypeus shining and weakly costate; lateral thirds of middle portion with about two
<taxonomicName id="C762C8B6AB1EE71E1B17A633F5B59A2E" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
; middle third without
<taxonomicName id="42E7FC507EA3E62E09238E321AA31B1C" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
. Eyes large (EYE 35-41). Maximum length of eye distinctly less than
<normalizedToken id="EACCC5406D58D8766F5BD796255FF42B" originalValue="¾" pageId="32" pageNumber="333">3/4</normalizedToken>
length of malar area (OMI 55-60). Face with erect to subdecumbent setae of various lengths. Setae ventrally and anterior to the eye mostly subdecumbent.
</paragraph>
<paragraph id="E28DF730840B29A80C1A7269F3D6076A" box="[118,1122,925,1169]" pageId="32" pageNumber="333" type="mainText">
Promesonotal shield longer than broad (PMI 122-125); posterolateral corners close-set (PWI 51-55). Dorsum of promesonotum with dilute
<taxonomicName id="9AD306BB0D6DE11E0D7917097C7B8EE7" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
, mostly concentrated in anterior half; interstices with setae raised on tiny nodules. Promesonotal shield setae decumbent to suberect, of mixed lengths. Lateral face of pronotum with weak rugae; setae raised on nodules. Lateral face of mesonotum smooth and shining. Katepi-anepisternal suture present; mesopleural-mesonotal suture present; mesopleural-metapleural suture absent to present. Mesopleuron smooth and strongly shining, with very dilute (1 to 4), very weak indications of raised sculpture other than in posteroventral area. Metapleural lobes well-developed. Propodeal spines thornlike; dorsomedian carina absent; dorsolateral carina extends dorsolaterally to promesonotum. Posterior face of propodeum smooth and shining; very weak
<taxonomicName id="B2FD02F9BA023EA2AEB88652D9A214E3" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
may be present in top third, which otherwise has setae raised on nodules.
</paragraph>
<paragraph id="194E04EF07D18583DC5E88C77C7C18B1" box="[118,1121,1178,1394]" pageId="32" pageNumber="333" type="mainText">
Petiole cuneate in profile, without pedicel. Anteroventral process of petiole a simple denticle. Postpetiole nodiform, about as tall as long (PTI 50-55). Sub-post-petiolar process well-developed, with a strong inflection between posterior half of sternum. Dorsal face of postpetiole with weak to indistinct rugae; lateral face with weak to indistinct rugae; posterior face smooth and shining except for nodulose setae and transverse
<taxonomicName id="4A158E4FFDD3CD3C950211D3C781D1AA" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
near extreme base. Fourth abdominal tergum (ATIV) comparatively small (ATW/WL 1.15-1.23, ATL/HL 1.20-1.33). ATIV shining, with fine areolate sculpture between stellate setiferous punctures; smooth near base of tergum. Setae of ATIV of mixed lengths; shortest subdecumbent and longer than width of metafemora; longest setae about twice as long as shortest.
</paragraph>
<paragraph id="CA6C02701A2F30B15F7293879920BC77" box="[118,1121,1403,1478]" pageId="32" pageNumber="333" type="mainText">Queen (paratype). HL 1.22, HW 1.17, HLA 0.41, CW 0.44, CDD 0.13, SL 0.82, EL 0.29, EW 0.22, SPL 0.20, WL 1.61, PTL 0.31, PTH 0.56, PPL 0.29, PPH 0.56, ATW 1.82, ATL 1.89, CI 95.2, SI 70.6, OMI 71.4, CDI 29.1, SEI 285, CS 1.2, EYE 42.4.</paragraph>
<paragraph id="A4C4458165FC7EEC952E1AED6149E6B6" box="[155,367,1487,1506]" pageId="32" pageNumber="333" type="mainText">Color: chocolate brown.</paragraph>
<paragraph id="AA0A7FD607F529CE34E2E42B0FD6FBE5" box="[118,1121,1515,1591][][][][118,1120,118,194]" lastPageId="36" lastPageNumber="337" pageId="32" pageNumber="333" type="mainText">
Large (WL 1.61). Head longer than broad (CI 95). Mandibles with more-or-less parallel striae. Face with weakly rugose
<taxonomicName id="E9E2465C554391AAEE5CB4EB11576446" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
which continue to the posterior margin of the head. Area above eye areolate; behind and beneath eye costate. Scrobal carina weakly bulging in anterior half. Anterior margin of clypeus with two well-developed
<pageBreakToken id="77EB4C5B1CED37194D834CF07CBD0694" pageId="36" pageNumber="337" start="start">,</pageBreakToken>
close-set laminar lobes (CDI 29). Middle portion of clypeus with strong
<taxonomicName id="D3FE36B51B461B70BB97F3E7FF61231A" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
in lateral thirds; middle third without sculpture. Eyes large (EYE 42). Maximum eye length about
<normalizedToken id="2FC9FA4365B1EEDFD18BBC29DD10F10F" originalValue="¾" pageId="36" pageNumber="337">3/4</normalizedToken>
length of malar area (OMI 71). Scapes long (SI 71). Face with erect to subdecumbent setae; longest setae about 1.5 length of shortest setae.
</paragraph>
<caption id="2F3D4982BB36B837B86559400CB42019" box="[118,1122,1539,1581]" pageId="33" pageNumber="334" type="caption">
<paragraph id="4964055166FD54D9E3C42E3EB11962BA" box="[118,1122,1539,1581]" pageId="33" pageNumber="334" type="caption">
<pageBreakToken id="B036EBDD6D569BEFB07F5453D9024D2C" pageId="33" pageNumber="334" start="start">FIGURES</pageBreakToken>
57-59.
<taxonomicName id="53F6E9BEDAC454D3876F893B94CDAF29" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="sylvarius">M. sylvarius</taxonomicName>
<taxonomicNameLabel id="E751A4AE36AA689C9527FA1AD84D8F01" pageId="33" pageNumber="334" rank="species">sp. n.</taxonomicNameLabel>
, holotype worker (CASENT0317539). 57. Body in lateral view. 58. Body in dorsal view. 59. Head in full-face view.
</paragraph>
</caption>
<caption id="D857109C9336AA99C34B7607729E1A4E" box="[118,1122,1539,1581]" pageId="34" pageNumber="335" type="caption">
<paragraph id="A7DA7FBBD259F12C0D9224B8D3784610" box="[118,1122,1539,1581]" pageId="34" pageNumber="335" type="caption">
<pageBreakToken id="3683CAC652612876B0C664DD5255A534" pageId="34" pageNumber="335" start="start">FIGURES</pageBreakToken>
60-62.
<taxonomicName id="13C956F25AF330D70197EA02694C4C24" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="sylvarius">M. sylvarius</taxonomicName>
<taxonomicNameLabel id="E83B6FF98E5D9756B100648562960E9C" pageId="34" pageNumber="335" rank="species">sp. n.</taxonomicNameLabel>
, paratype gyne (CASENT0317567; S. Hartman 2012). 60. Body in lateral view. 61. Body in dorsal view. 62. Head in full-face view.
</paragraph>
</caption>
<caption id="8C19A276E6AEF656BE8D391EA8B49D9D" box="[118,1122,1499,1588]" pageId="35" pageNumber="336" type="caption">
<paragraph id="0F7A30D06EF6B1D77882C95EE396D7E8" box="[118,1122,1499,1588]" pageId="35" pageNumber="336" type="caption">
<pageBreakToken id="66521A9C1F518C1BE14AC425B44067FE" pageId="35" pageNumber="336" start="start">FIGURES</pageBreakToken>
63-66. Distribution maps of Malagasy
<taxonomicName id="4ED63A4F66018086905F17CAB45FB39D" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">Meranoplus</taxonomicName>
species; type localities are indicated by red markers. Elevation is indicated by color: green 0-100 m, peach 101-500 m, pink 501-1000 m, grey 1001-2000+ m (map provided by NASA). 63.
<taxonomicName id="74B7D4EC467E700E5FE85CD860DFCA00" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="cryptomys">M. cryptomys</taxonomicName>
<taxonomicNameLabel id="2BC7B7C097D5699A0315EB24F7C04027" pageId="35" pageNumber="336" rank="species">sp. n.</taxonomicNameLabel>
64.
<taxonomicName id="1E2DAC7ECDAF87D821C6AA6B7A596B48" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="mayri">M. mayri</taxonomicName>
. 65.
<taxonomicName id="0B47B5FCB289078FFB9491CB86280C74" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="radamae">M. radamae</taxonomicName>
; the exact type locality is unknown, but is in &quot;Central Madagascar&quot;. 66.
<taxonomicName id="44DD79C41F9BDAA0D02931DCE6FBDA4D" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="sylvarius">M. sylvarius</taxonomicName>
<taxonomicNameLabel id="8682D4F341F85B631F22536404F62770" pageId="35" pageNumber="336" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="BE4723C03CB001DAED6FD5B91CF6A69D" box="[118,1122,203,447]" pageId="36" pageNumber="337" type="mainText">
<pageBreakToken id="3A649B59501BF297EFE5EB342FDEA35D" pageId="36" pageNumber="337">Mesosoma</pageBreakToken>
longer than tall. Anterolateral corners of pronotum angled; dorsolateral margin of pronotum ecarinate. Mesosomal dorsum with little sculpture: pronotum smooth medially, rugose laterally; scutum smooth and shining in anterior half, with dilute rugae in posterior half; scutellum with transverse rugae branching more frequently in lateral portions. Mesosomal dorsum with subdecumbent to erect setae of mixed lengths. Lateral face of pronotum rugose anteriorly; with setae raised on strong nodules. Katepisternum mostly smooth and shining; without
<taxonomicName id="E6D5C98A84570A3C543BE5556FA643C9" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
extending longitudinally margin to margin. Anteroventral portion of anepisternum smooth and shining; dorsal portion with longitudinal rugae, interstices roughened. Metapleuron roughened with rugae; area anteroventral to propodeal spiracle smooth and shining. Propodeal spines thorn-like, long (SPL 0.20). Meso and metatibial spurs absent. Wings unknown.
</paragraph>
<paragraph id="8DF0EF043C9EACBA8D5355833830D531" box="[118,1121,456,587]" pageId="36" pageNumber="337" type="mainText">
Petiole cuneate in profile; subpetiolar process dentiform. Postpetiole nodiform; sternal process large. Dorsal and lateral faces of postpetiole rugose; posterior face roughened near base and with dilute longitudinal
<taxonomicName id="9686B5386C4D061CA104C613E7934322" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
. Forth abdominal tergum (ATIV) with sides convex; nearly as broad as long (ATW/GL 0.96). ATIV with fine, weak areolate sculpture between stellate setiferous punctures; weak, dilute
<taxonomicName id="A7BDA5079BD1DD95F7C562428CF64E90" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="costae">costae</taxonomicName>
present in basal region. ATIV with subdecumbent short setae; suberect setae about 1.5-2 times as long as shorter setae.
</paragraph>
</subSubSection>
<subSubSection id="FF691492533672AD2BD011DFAEDEC840" pageId="36" pageNumber="337" type="diagnosis">
<paragraph id="2D43E7A8015F2ACBACFAFAFD3F9F47C1" box="[118,208,624,643]" pageId="36" pageNumber="337" type="mainText">Diagnosis</paragraph>
<paragraph id="E465E6EA3271C21550026E1F3394151A" box="[118,1121,653,728]" pageId="36" pageNumber="337" type="mainText">
Worker. The absence of meso- and metatibial spurs is unique in the Malagasy fauna. This
<taxonomicNameLabel id="5AD5D6743D14E76C00190922726E60A0" pageId="36" pageNumber="337" rank="species">new species</taxonomicNameLabel>
may be separated from
<taxonomicName id="B2C562356137AA7B5586EF8E554B1306" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="radamae">M. radamae</taxonomicName>
by the absence of a dorsomedial costa extending from the base of the propodeal spine to the promesonotum.
</paragraph>
<paragraph id="1CD90343CB87ECDB9C1B17BD6716CE7C" box="[118,1121,737,812]" pageId="36" pageNumber="337" type="mainText">
Gyne. The combination of stellate setiferous punctures, comparatively large size (WL 1.61), dark chocolatebrown color, malar area about
<normalizedToken id="9FC958E286EA921A101F5706BFAE22DE" originalValue="¾" pageId="36" pageNumber="337">3/4</normalizedToken>
the length of the eye (OMI 71.4), long scapes (SI 70.6), uniquely identify gynes of this species in the Malagasy fauna.
</paragraph>
</subSubSection>
<subSubSection id="0B87D98786EEF20CCF569C09D0282088" pageId="36" pageNumber="337" type="discussion">
<paragraph id="263ADDD720E4B6C9DD8A297CBE1A6E73" box="[118,219,849,868]" pageId="36" pageNumber="337" type="mainText">Comments</paragraph>
<paragraph id="F0C46546F1BC01CAF5F3E3BC3646FFE1" box="[118,1121,878,953]" pageId="36" pageNumber="337" type="mainText">
The type series of
<taxonomicName id="607243F0F91CCF4BEB119D9A43D0BE90" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="sylvarius">M. sylvarius</taxonomicName>
is designated from a large nest collection from the rainforest/montane forest of Vevembe, the only known locale of this species. Restriction to moist forest habitat is unique for Malagasy
<taxonomicName id="9CD8B55574B21DB772F4EDDAD37446FD" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">Meranoplus</taxonomicName>
.
</paragraph>
<paragraph id="B6A32EDD1195E37B4E665A5D63384E21" box="[118,1123,962,1122]" pageId="36" pageNumber="337" type="mainText">
Although not collected in sympatry with
<taxonomicName id="A91ACBBD594E882E348B333D23EA1209" authority=", M. sylvarius" authorityName=", M. sylvarius" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="radamae">M. radamae, M. sylvarius</taxonomicName>
is morphologically distinct enough to support the hypothesis of reproductive isolation. The lack of meso- and metatibial spurs is unique in the Malagasy
<taxonomicName id="E65A1CBEECC383BBFE86469469B71442" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="fauna">Meranoplus fauna</taxonomicName>
. Although
<taxonomicName id="35131DC7FCAA0896389A5C4DB53F6C3C" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="radamae">M. radamae</taxonomicName>
shows much geographic variation in its sculpturation, the trends of sculpture reduction across body parts never move in synchrony to such extremes, suggesting that the almost complete reduction of sculpture on specimens of
<taxonomicName id="6FE12B51D7CD9870CD1A8D742F0B4244" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="sylvarius">M. sylvarius</taxonomicName>
would be highly unlikely under circumstances of gene flow.
</paragraph>
</subSubSection>
<subSubSection id="CD7074D9A674E92B4D2E27718FB2F8E8" pageId="36" pageNumber="337" type="etymology">
<paragraph id="CCFCC73571B28C46A5AF396AC74E2F57" box="[118,219,1159,1177]" pageId="36" pageNumber="337" type="mainText">Etymology</paragraph>
<paragraph id="15C09B2B937D691F53E84AA731F439DC" box="[118,1122,1187,1234]" pageId="36" pageNumber="337" type="mainText">
The
<taxonomicNameLabel id="7C115D4CB2899034A4FCBF7AAEFB64EB" pageId="36" pageNumber="337" rank="species">new species</taxonomicNameLabel>
name is Latin for
<normalizedToken id="2D53B2282A42E55137906B35D9B41AAB" originalValue="“forester”" pageId="36" pageNumber="337">&quot;forester&quot;</normalizedToken>
, as this species is unique in the Malagasy
<taxonomicName id="01A7DEA90DD3BF5690BA5A59D16F3C97" class="Insecta" family="Formicidae" genus="Meranoplus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="fauna">Meranoplus fauna</taxonomicName>
for its restriction to rainforest habitat. The specific epithet is a noun in apposition and thus invariant.
</paragraph>
</subSubSection>
<subSubSection id="9187A742D990722ABDC8ED6A9505856E" pageId="36" pageNumber="337" type="materials_examined">
<paragraph id="CB485A43ACF570E7DC57BD068C77F5D7" box="[118,397,1271,1290]" pageId="36" pageNumber="337" type="mainText">Additional material examined</paragraph>
<paragraph id="9EA8F36E0824CE74E6866EFCBE57DBF1" box="[118,1122,1300,1347]" pageId="36" pageNumber="337" type="mainText">MADAGASCAR: Fianarantsoa, Vevembe, 22°47.46' S, 47°10.91' E, 600 m, montane forest, collection code BLF14314, 23-24Apr2006 (B.L. Fisher et al.).</paragraph>
</subSubSection>
</treatment>
</document>