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<mods:title id="7119732E9DEF40973039AAF6E49A3940">Revision of Leocrates Kinberg, 1866 and Leocratides Ehlers, 1908 (Annelida, Errantia, Hesionidae)</mods:title>
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<emphasis id="97C4EAE0FF84734BFF23FB20F58D65FA" bold="true" box="[151,495,1237,1264]" italics="true" pageId="103" pageNumber="104">Paraleocrates djangkarensis</emphasis>
(Augener &amp; Pettibone
<emphasis id="97C4EAE0FF84734BFCA4FB20F74A65E5" bold="true" box="[784,808,1237,1263]" italics="true" pageId="103" pageNumber="104">in</emphasis>
<bibRefCitation id="C1214B03FF84734BFC84FB20F79365E5" author="Pettibone" box="[816,1009,1237,1263]" firstAuthor="Pettibone" pageId="103" pageNumber="104" pagination="199 - 270" refId="ref88038" refString="Pettibone, M. H. (1970) Polychaeta Errantia of the Siboga Expedition, 4. Some additional polychaetes of the Polynoidae, Hesionidae, Nereidae, Goniadidae, Eunicidae, and Onuphidae, selected as new species by the late Dr. Hermann Augener with remarks on other related species. Siboga Expeditie Monographie, 24 (1 d), 199 - 270." type="journal article" year="1970">Pettibone, 1970</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="8CF7579BFF84734BFBB5FB23F00965FA" box="[1025,1131,1238,1264]" pageId="103" pageNumber="104" rank="species">n. comb.</taxonomicNameLabel>
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<figureCitation id="3D8B2A77FF84734BFF23FB0CF5616419" box="[151,259,1273,1299]" captionStart="FIGURE 61" captionStartId="104.[151,250,1204,1229]" captionTargetBox="[173,1416,184,1171]" captionTargetId="figure@104.[170,1418,181,1177]" captionTargetPageId="104" captionText="FIGURE 61. Paraleocrates djankarensis (Augener &amp; Pettibone in Pettibone, 1970), new combination, holotype (ZMA V. Pol. 534.3). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 3, left parapodium, anterior view, MGS (inset: lower neurochaetal blade). E. Chaetiger 8, left parapodium, anterior view, MGS (insets: lower neurochaetal blades). Scale bars: A: 1.4 mm, B: 0.6 mm, C: 0.8 mm, D: 0.2 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672674" httpUri="https://zenodo.org/record/3672674/files/figure.png" pageId="103" pageNumber="104">Figure 61</figureCitation>
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<paragraph id="A50F36F2FF84734BFF23FACBF075645C" blockId="103.[151,1047,1341,1397]" box="[151,1047,1341,1366]" pageId="103" pageNumber="104">
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<emphasis id="97C4EAE0FF84734BFF23FACBF5EC645C" box="[151,398,1342,1366]" italics="true" pageId="103" pageNumber="104">Leocrates djangkarensis</emphasis>
Augener &amp; Pettibone
<emphasis id="97C4EAE0FF84734BFDC7FACBF6E5645F" box="[627,647,1342,1365]" italics="true" pageId="103" pageNumber="104">in</emphasis>
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,
<figureCitation id="3D8B2A77FF84734BFC2AFAC8F071645C" box="[926,1043,1341,1366]" captionStart-0="FIGURE 25" captionStart-1="FIGURE 26" captionStartId-0="48.[151,250,1873,1898]" captionStartId-1="50.[151,250,1826,1851]" captionTargetBox-0="[172,1416,815,1848]" captionTargetBox-1="[173,1415,925,1798]" captionTargetId-0="figure@48.[170,1418,813,1849]" captionTargetId-1="figure@50.[170,1418,921,1801]" captionTargetPageId-0="48" captionTargetPageId-1="50" captionText-0="FIGURE 25. Leocrates ahlfeldae n. sp., holotype (USNM 48942).A.Anterior region, dorsal view. B. Anterior end, dorsal view. C. Pharynx, frontal view, MGS. D. Chaetiger 7, left parapodium, anterior view (insets: notacicular and neuracicular lobes). E. Upper neurochaetal blades. F. Lower neurochaetal blades. Scale bars: A: 1.1 mm, B: 0.3 mm, C, D: 0.5 mm, E: 100 μm, F: 150 μm." captionText-1="FIGURE 26. Leocrates chinensis Kinberg, 1866, holotype of L. anonymous Hessle, 1925 (UUZM 672). A. Anterior region, dorsal view. B. Anterior end, pharynx exposed with two lateral vesicles. C. Same, left lateral view. D. Same, frontal view of pharynx after pressing the body to swell lateral vesicles (inset: lateral view of wedge-shaped upper jaw tip). E. Chaetiger 7, left parapodium, anterior view (insets: notacicular lobe, neurochaetal blade). Scale bars:A: 1.9 mm, B: 1 mm, C, D: 0.6 mm, E: 0.4 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3672600" figureDoi-1="http://doi.org/10.5281/zenodo.3672602" httpUri-0="https://zenodo.org/record/3672600/files/figure.png" httpUri-1="https://zenodo.org/record/3672602/files/figure.png" pageId="103" pageNumber="104">Figs 2526</figureCitation>
.
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<treatmentCitationGroup id="85A011DCFF84734BFF23FAA8F7CB647F" box="[151,937,1372,1397]" pageId="103" pageNumber="104">
<treatmentCitation id="241110E3FF84734BFF23FAA8F670647E" author="Horst, R." box="[151,530,1372,1396]" page="193" pageId="103" pageNumber="104" year="1924">
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<emphasis id="97C4EAE0FF84734BFF23FAA8F53D647E" box="[151,351,1372,1396]" italics="true" pageId="103" pageNumber="104">Leocrates chinensis</emphasis>
:
<bibRefCitation id="C1214B03FF84734BFED9FAA8F670647E" author="Horst" box="[365,530,1372,1396]" firstAuthor="Horst" pageId="103" pageNumber="104" pagination="145 - 198" refId="ref85770" refString="Horst, R. (1924) Polychaeta Errantia of the Siboga-Expedition, 3. Nereidae and Hesionidae. Siboga-Expeditie Monographs, 24 (1 c), 145 - 198." type="journal article" year="1924">Horst 1924: 193</bibRefCitation>
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(
<emphasis id="97C4EAE0FF84734BFD95FAA8F609647E" box="[545,619,1373,1396]" italics="true" pageId="103" pageNumber="104">partim,</emphasis>
Sta. 105) (
<emphasis id="97C4EAE0FF84734BFD6FFAA8F763647E" box="[731,769,1373,1396]" italics="true" pageId="103" pageNumber="104">non</emphasis>
<bibRefCitation id="C1214B03FF84734BFCB3FAA9F7FC647F" author="Kinberg" box="[775,926,1372,1397]" firstAuthor="Kinberg" pageId="103" pageNumber="104" pagination="239 - 258" refId="ref86199" refString="Kinberg, J. G. H. (1866) Annulata nova (Nephtydea, Phyllodocea, Alciopea, Hesionida, Glycerea, Goniadea, Syllidea, Ariciea, Spiodea, Aonidea, Cirratulida, Opheliacea). Ofversigt af Kongelige Vetenskaps-Aakademiens Forhandlingar, 22 (Volume Corresponding to 1865), 4, 239 - 258. [http: // www. biodiversitylibrary. org / item / 100715 # page / 253 / mode / 1 up]" type="journal article" year="1866">Kinberg, 1866</bibRefCitation>
).
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<emphasis id="97C4EAE0FF84734BFF23FA56F52264B7" bold="true" box="[151,320,1442,1469]" pageId="103" pageNumber="104">Type material</emphasis>
.
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</collectingCountry>
.
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(
<collectionCode id="C3A1AE37FF84734BFDEAFA56F6FF64B7" box="[606,669,1443,1469]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="103" pageNumber="104">ZMA</collectionCode>
<collectorName id="08455324FF84734BFD11FA56F69064B7" box="[677,754,1443,1469]" pageId="103" pageNumber="104">V. Pol.</collectorName>
534.3), Sulu Islands, off
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NE
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, R
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, Sta. 105 (
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</quantity>
, dredge, coral,
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<collectingDate id="C14AE9DAFF84734BFCBCFA32F7E964EB" box="[776,907,1479,1505]" pageId="103" pageNumber="104" value="1899-07-04">4 Jul. 1899</collectingDate>
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.
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<collectingCountry id="DDA77662FF84734BFC2CFA33F06F64EB" box="[920,1037,1478,1505]" name="Indonesia" pageId="103" pageNumber="104">Indonesia</collectingCountry>
</emphasis>
,
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(
<collectionCode id="C3A1AE37FF84734BFB39FA32F0AE64EB" box="[1165,1228,1479,1505]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="103" pageNumber="104">ZMA</collectionCode>
<collectorName id="08455324FF84734BFB61FA32F14064EB" box="[1237,1314,1479,1505]" pageId="103" pageNumber="104">V. Pol.</collectorName>
2104), off
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,
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<collectorName id="08455324FF84734BFEF2FA1EF5C9670F" box="[326,427,1515,1541]" pageId="103" pageNumber="104">Island, R</collectorName>
/
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,
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5 (
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),
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<elevation id="2E9DD1C1FF84734BFC3EFA1EF7B1670F" box="[906,979,1515,1541]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="103" pageNumber="104" unit="m" value="330.0">330 m</elevation>
</quantity>
, deep-sea trawl, muddy bottom,
<date id="D10E1032FF84734BFAF1FA1EF4AD6723" pageId="103" pageNumber="104" value="1899-03-10">
<collectingDate id="C14AE9DAFF84734BFAF1FA1EF4AD6723" pageId="103" pageNumber="104" value="1899-03-10">10 Mar. 1899</collectingDate>
</date>
(used for variation).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FF847345FF73F9C7F1656014" lastPageId="105" lastPageNumber="106" pageId="103" pageNumber="104" type="description">
<paragraph id="A50F36F2FF84734BFF73F9C7F7EB660B" blockId="103.[151,1437,1442,1973]" pageId="103" pageNumber="104">
<emphasis id="97C4EAE0FF84734BFF73F9C7F5336747" bold="true" box="[199,337,1586,1613]" pageId="103" pageNumber="104">Description</emphasis>
.
<typeStatus id="7A0B8850FF84734BFED4F9C6F5AA6747" box="[352,456,1587,1613]" pageId="103" pageNumber="104" type="holotype">Holotype</typeStatus>
(ZMZ V. Pol. 534.3), complete, slightly distorted, dehydrated, most body cirri missing (
<figureCitation id="3D8B2A77FF84734BFF14F9A2F56A6778" box="[160,264,1623,1650]" captionStart="FIGURE 61" captionStartId="104.[151,250,1204,1229]" captionTargetBox="[173,1416,184,1171]" captionTargetId="figure@104.[170,1418,181,1177]" captionTargetPageId="104" captionText="FIGURE 61. Paraleocrates djankarensis (Augener &amp; Pettibone in Pettibone, 1970), new combination, holotype (ZMA V. Pol. 534.3). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 3, left parapodium, anterior view, MGS (inset: lower neurochaetal blade). E. Chaetiger 8, left parapodium, anterior view, MGS (insets: lower neurochaetal blades). Scale bars: A: 1.4 mm, B: 0.6 mm, C: 0.8 mm, D: 0.2 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672674" httpUri="https://zenodo.org/record/3672674/files/figure.png" pageId="103" pageNumber="104">Fig. 61A</figureCitation>
). Body obconic, wider anteriorly, tapered posteriorly,
<quantity id="62489B17FF84734BFCD7F9A2F7D0677B" box="[867,946,1623,1649]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="103" pageNumber="104" unit="mm" value="12.0">12 mm</quantity>
long,
<quantity id="62489B17FF84734BFC4CF9A2F05B677B" box="[1016,1081,1623,1649]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="103" pageNumber="104" unit="mm" value="2.0">2 mm</quantity>
wide, 16 chaetigers; right parapodium of chaetiger 3, and left parapodia of chaetigers 3, 8, 9, and 14 previously removed (three ones removed by Pettibone in a small vial, others missing); a slightly lateral anteroventral dissection for observing pharynx features previously made. Body brownish, venter with two wide longitudinal brown bands (
<figureCitation id="3D8B2A77FF84734BFB8BF936F0C467D4" box="[1087,1190,1731,1758]" captionStart="FIGURE 61" captionStartId="104.[151,250,1204,1229]" captionTargetBox="[173,1416,184,1171]" captionTargetId="figure@104.[170,1418,181,1177]" captionTargetPageId="104" captionText="FIGURE 61. Paraleocrates djankarensis (Augener &amp; Pettibone in Pettibone, 1970), new combination, holotype (ZMA V. Pol. 534.3). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 3, left parapodium, anterior view, MGS (inset: lower neurochaetal blade). E. Chaetiger 8, left parapodium, anterior view, MGS (insets: lower neurochaetal blades). Scale bars: A: 1.4 mm, B: 0.6 mm, C: 0.8 mm, D: 0.2 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672674" httpUri="https://zenodo.org/record/3672674/files/figure.png" pageId="103" pageNumber="104">Fig. 61C</figureCitation>
), leaving a paler midventral region (purple in recently preserved specimens after Horst).
</paragraph>
<paragraph id="A50F36F2FF84734BFF73F8FEF6DE66BF" blockId="103.[151,1437,1442,1973]" pageId="103" pageNumber="104">
Prostomium slightly wider than long, slightly wider anteriorly (
<figureCitation id="3D8B2A77FF84734BFC21F8FEF79E662C" box="[917,1020,1803,1830]" captionStart="FIGURE 61" captionStartId="104.[151,250,1204,1229]" captionTargetBox="[173,1416,184,1171]" captionTargetId="figure@104.[170,1418,181,1177]" captionTargetPageId="104" captionText="FIGURE 61. Paraleocrates djankarensis (Augener &amp; Pettibone in Pettibone, 1970), new combination, holotype (ZMA V. Pol. 534.3). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 3, left parapodium, anterior view, MGS (inset: lower neurochaetal blade). E. Chaetiger 8, left parapodium, anterior view, MGS (insets: lower neurochaetal blades). Scale bars: A: 1.4 mm, B: 0.6 mm, C: 0.8 mm, D: 0.2 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672674" httpUri="https://zenodo.org/record/3672674/files/figure.png" pageId="103" pageNumber="104">Fig. 61B</figureCitation>
). Lateral antennae with ceratophores distinct, left one corrugated, right one missing, slightly longer than left palp (right palp distorted, directed laterally); palpophore 23 times longer than palpostyles. Median antenna missing, scar inserted centrally, between eyes. Eyes barely pigmented, round, anterior ones twice larger and slightly more distant to each other than posterior ones. Lateral peristomial tubercles round, as long as wide.
</paragraph>
<caption id="F1CF667AFF8B7344FF23FB41F07B6432" ID-DOI="http://doi.org/10.5281/zenodo.3672674" ID-Zenodo-Dep="3672674" httpUri="https://zenodo.org/record/3672674/files/figure.png" pageId="104" pageNumber="105" startId="104.[151,250,1204,1229]" targetBox="[173,1416,184,1171]" targetPageId="104">
<paragraph id="A50F36F2FF8B7344FF23FB41F07B6432" blockId="104.[151,1437,1204,1337]" pageId="104" pageNumber="105">
<emphasis id="97C4EAE0FF8B7344FF23FB41F57E65C7" bold="true" box="[151,284,1204,1229]" pageId="104" pageNumber="105">FIGURE 61</emphasis>
.
<taxonomicName id="62B04D71FF8B7344FE99FB41F06965C6" authority="(Augener &amp; Pettibone in Pettibone, 1970)" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Augener &amp; Pettibone in Pettibone" baseAuthorityYear="1970" box="[301,1035,1204,1229]" class="Polychaeta" family="Hesionidae" genus="Paraleocrates" kingdom="Animalia" order="Phyllodocida" pageId="104" pageNumber="105" phylum="Annelida" rank="species" species="djankarensis" status="comb. nov.">
<emphasis id="97C4EAE0FF8B7344FE99FB41F62565C6" box="[301,583,1204,1228]" italics="true" pageId="104" pageNumber="105">Paraleocrates djankarensis</emphasis>
(Augener &amp; Pettibone
<emphasis id="97C4EAE0FF8B7344FCF5FB40F73765C6" box="[833,853,1205,1228]" italics="true" pageId="104" pageNumber="105">in</emphasis>
<bibRefCitation id="C1214B03FF8B7344FCEAFB41F06665C6" author="Pettibone" box="[862,1028,1204,1229]" firstAuthor="Pettibone" pageId="104" pageNumber="105" pagination="199 - 270" refId="ref88038" refString="Pettibone, M. H. (1970) Polychaeta Errantia of the Siboga Expedition, 4. Some additional polychaetes of the Polynoidae, Hesionidae, Nereidae, Goniadidae, Eunicidae, and Onuphidae, selected as new species by the late Dr. Hermann Augener with remarks on other related species. Siboga Expeditie Monographie, 24 (1 d), 199 - 270." type="journal article" year="1970">Pettibone, 1970</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="8CF7579BFF8B7344FBADFB40F0AC65C7" box="[1049,1230,1204,1229]" pageId="104" pageNumber="105" rank="species">new combination</taxonomicNameLabel>
, holotype (ZMA V. Pol. 534.3). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 3, left parapodium, anterior view, MGS (inset: lower neurochaetal blade). E. Chaetiger 8, left parapodium, anterior view, MGS (insets: lower neurochaetal blades). Scale bars: A: 1.4 mm, B: 0.6 mm, C: 0.8 mm, D: 0.2 mm, E: 0.4 mm.
</paragraph>
</caption>
<paragraph id="A50F36F2FF8B7344FF73FA7EF03F64E7" blockId="104.[151,1437,1419,2021]" pageId="104" pageNumber="105">Nuchal organs lobes horizontal C-shaped, completely concealed by anterior margin of tentacular belt; lateral ciliated bands visible dorsally. Tentacular cirri almost completely missing, one ventral cirrus without tip reaching chaetiger 3. Lateral cushions low, projected, most tripartite; longitudinal striae visible.</paragraph>
<paragraph id="A50F36F2FF8B7344FF73FA02F59D6753" blockId="104.[151,1437,1419,2021]" pageId="104" pageNumber="105">Pharynx not exposed; jaws observed through previous dissection. Lateral vesicles not seen. Anterior margin with about 20 irregular constrictions. Dorsal and ventral jaws single, fang-shaped brownish, exposed, tapered, ventral jaw smaller than dorsal one.</paragraph>
<paragraph id="A50F36F2FF8B7344FF73F996F0FE6777" blockId="104.[151,1437,1419,2021]" box="[199,1180,1635,1661]" pageId="104" pageNumber="105">Peristomial dorsolateral tubercles as long as wide, slightly projected anteriorly, smooth.</paragraph>
<paragraph id="A50F36F2FF8B7344FF73F972F565665C" blockId="104.[151,1437,1419,2021]" pageId="104" pageNumber="105">
Dorsal cirri broken, some remaining shorter than body width. Chaetigers 13 without notochaetae (
<figureCitation id="3D8B2A77FF8B7344FA92F972F1ED67AB" box="[1318,1423,1670,1697]" captionStart="FIGURE 61" captionStartId="104.[151,250,1204,1229]" captionTargetBox="[173,1416,184,1171]" captionTargetId="figure@104.[170,1418,181,1177]" captionTargetPageId="104" captionText="FIGURE 61. Paraleocrates djankarensis (Augener &amp; Pettibone in Pettibone, 1970), new combination, holotype (ZMA V. Pol. 534.3). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 3, left parapodium, anterior view, MGS (inset: lower neurochaetal blade). E. Chaetiger 8, left parapodium, anterior view, MGS (insets: lower neurochaetal blades). Scale bars: A: 1.4 mm, B: 0.6 mm, C: 0.8 mm, D: 0.2 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672674" httpUri="https://zenodo.org/record/3672674/files/figure.png" pageId="104" pageNumber="105">Fig. 61D</figureCitation>
); notochaetae present along chaetigers 416, about 30 per bundle, delicate, arranged in bundles, notochaetal surface smooth. Notacicular lobes tapered, blunt; neuracicular lobes projected, blunt, as long as wide (
<figureCitation id="3D8B2A77FF8B7344FB7BF93AF15567E3" box="[1231,1335,1742,1769]" captionStart="FIGURE 61" captionStartId="104.[151,250,1204,1229]" captionTargetBox="[173,1416,184,1171]" captionTargetId="figure@104.[170,1418,181,1177]" captionTargetPageId="104" captionText="FIGURE 61. Paraleocrates djankarensis (Augener &amp; Pettibone in Pettibone, 1970), new combination, holotype (ZMA V. Pol. 534.3). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 3, left parapodium, anterior view, MGS (inset: lower neurochaetal blade). E. Chaetiger 8, left parapodium, anterior view, MGS (insets: lower neurochaetal blades). Scale bars: A: 1.4 mm, B: 0.6 mm, C: 0.8 mm, D: 0.2 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672674" httpUri="https://zenodo.org/record/3672674/files/figure.png" pageId="104" pageNumber="105">Fig. 61E</figureCitation>
). Ventral cirri surpassing neurochaetal lobes. Neurochaetae about 30 per bundle, decreasing in size ventrally, blades unidentate, 465 times longer than wide, guards projected beyond tooth as long aristae, tooth slightly falcate (
<figureCitation id="3D8B2A77FF8B7344FA9BF8E2F4C8665F" captionStart="FIGURE 61" captionStartId="104.[151,250,1204,1229]" captionTargetBox="[173,1416,184,1171]" captionTargetId="figure@104.[170,1418,181,1177]" captionTargetPageId="104" captionText="FIGURE 61. Paraleocrates djankarensis (Augener &amp; Pettibone in Pettibone, 1970), new combination, holotype (ZMA V. Pol. 534.3). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 3, left parapodium, anterior view, MGS (inset: lower neurochaetal blade). E. Chaetiger 8, left parapodium, anterior view, MGS (insets: lower neurochaetal blades). Scale bars: A: 1.4 mm, B: 0.6 mm, C: 0.8 mm, D: 0.2 mm, E: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.3672674" httpUri="https://zenodo.org/record/3672674/files/figure.png" pageId="104" pageNumber="105">Fig. 61D, E</figureCitation>
, insets).
</paragraph>
<paragraph id="A50F36F2FF8B7344FF73F8AAF7286697" blockId="104.[151,1437,1419,2021]" pageId="104" pageNumber="105">Posterior region tapered, almost without cirri. Prepygidial segment with dorsal cirrophores 45 thicker than ventral ones. Pygidium with anus terminal, anal cirri missing.</paragraph>
<paragraph id="A50F36F2FF8B7344FF73F852F5EC66CB" blockId="104.[151,1437,1419,2021]" box="[199,398,1959,1985]" pageId="104" pageNumber="105">Oocytes not seen.</paragraph>
<paragraph id="A50F36F2FF8B7345FF73F83FF1656014" blockId="104.[151,1437,1419,2021]" lastBlockId="105.[151,1437,152,683]" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105">
<emphasis id="97C4EAE0FF8B7344FF73F83FF55A66EF" bold="true" box="[199,312,1994,2021]" pageId="104" pageNumber="105">Variation</emphasis>
.
<typeStatus id="7A0B8850FF8B7344FEFCF83EF5C866EF" box="[328,426,1995,2021]" pageId="104" pageNumber="105" type="paratype">Paratype</typeStatus>
(ZMA V. Pol. 2104) damaged, soft, brownish. Body
<quantity id="62489B17FF8B7344FBBAF83FF03D66EF" box="[1038,1119,1994,2021]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="104" pageNumber="105" unit="mm" value="35.0">35 mm</quantity>
long,
<quantity id="62489B17FF8B7344FB1DF83EF08E66EF" box="[1193,1260,1995,2021]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="104" pageNumber="105" unit="mm" value="3.0">3 mm</quantity>
wide, 16 chae- tigers; cirri and many chaetae missing; left parapodia of chaetigers 4 and 6, and right parapodia of chaetigers 9 and 10 previously removed. An anteroventral dissection for observing pharynx features previously made. Notopodia from chaetiger 4, most notochaetae missing, a few remaining, smooth. Neuropodia with a few chaetae remaining (715), blades unidentate, most straight, a few bent dorsally, guards projected far beyond tip of tooth.
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FF8A7345FF73FEDCF14C638C" pageId="105" pageNumber="106" type="discussion">
<paragraph id="A50F36F2FF8A7345FF73FEDCF116606C" blockId="105.[151,1437,152,683]" pageId="105" pageNumber="106">
<emphasis id="97C4EAE0FF8A7345FF73FEDCF5566049" bold="true" box="[199,308,297,323]" pageId="105" pageNumber="106">Remarks</emphasis>
.
<taxonomicName id="62B04D71FF8A7345FEF5FEDDF0296049" authority="(Augener &amp; Pettibone in Pettibone, 1970)" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Augener &amp; Pettibone in Pettibone" baseAuthorityYear="1970" box="[321,1099,296,323]" class="Polychaeta" family="Hesionidae" genus="Paraleocrates" kingdom="Animalia" order="Phyllodocida" pageId="105" pageNumber="106" phylum="Annelida" rank="species" species="djankarensis">
<emphasis id="97C4EAE0FF8A7345FEF5FEDDF6166048" box="[321,628,296,322]" italics="true" pageId="105" pageNumber="106">Paraleocrates djankarensis</emphasis>
(Augener &amp; Pettibone
<emphasis id="97C4EAE0FF8A7345FCC2FEDCF7EE6048" box="[886,908,297,322]" italics="true" pageId="105" pageNumber="106">in</emphasis>
<bibRefCitation id="C1214B03FF8A7345FC26FEDDF0206049" author="Pettibone" box="[914,1090,296,323]" firstAuthor="Pettibone" pageId="105" pageNumber="106" pagination="199 - 270" refId="ref88038" refString="Pettibone, M. H. (1970) Polychaeta Errantia of the Siboga Expedition, 4. Some additional polychaetes of the Polynoidae, Hesionidae, Nereidae, Goniadidae, Eunicidae, and Onuphidae, selected as new species by the late Dr. Hermann Augener with remarks on other related species. Siboga Expeditie Monographie, 24 (1 d), 199 - 270." type="journal article" year="1970">Pettibone, 1970</bibRefCitation>
)
</taxonomicName>
is newly combined into
<taxonomicName id="62B04D71FF8A7345FAEFFEDCF49C606C" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" class="Polychaeta" family="Hesionidae" genus="Paraleocrates" kingdom="Animalia" order="Phyllodocida" pageId="105" pageNumber="104" phylum="Annelida" rank="genus" status="gen. nov.">
<emphasis id="97C4EAE0FF8A7345FAEFFEDCF49C606C" italics="true" pageId="105" pageNumber="106">Paraleocrates</emphasis>
</taxonomicName>
by having unidentate blades, as opposed to bidentate blades present in
<taxonomicName id="62B04D71FF8A7345FBA8FEB8F0E8606C" authorityName="Kinberg" authorityYear="1866" box="[1052,1162,333,358]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="105" pageNumber="106" phylum="Annelida" rank="genus">
<emphasis id="97C4EAE0FF8A7345FBA8FEB8F0E8606C" box="[1052,1162,333,358]" italics="true" pageId="105" pageNumber="106">Leocrates</emphasis>
</taxonomicName>
, as herein restricted.
</paragraph>
<paragraph id="A50F36F2FF8A7345FF73FE85F6DC6310" blockId="105.[151,1437,152,683]" pageId="105" pageNumber="106">
<taxonomicName id="62B04D71FF8A7345FF73FE85F5986080" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Augener &amp; Pettibone in Pettibone" baseAuthorityYear="1970" box="[199,506,368,394]" class="Polychaeta" family="Hesionidae" genus="Paraleocrates" kingdom="Animalia" order="Phyllodocida" pageId="105" pageNumber="106" phylum="Annelida" rank="species" species="djankarensis">
<emphasis id="97C4EAE0FF8A7345FF73FE85F5986080" box="[199,506,368,394]" italics="true" pageId="105" pageNumber="106">Paraleocrates djankarensis</emphasis>
</taxonomicName>
differs from the other species in the genus,
<taxonomicName id="62B04D71FF8A7345FC6FFE84F1E26081" authority="(Pettibone, 1970)" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Pettibone" baseAuthorityYear="1970" box="[987,1408,368,395]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="105" pageNumber="106" phylum="Annelida" rank="species" species="wesenberglundae" status="comb. nov.">
<emphasis id="97C4EAE0FF8A7345FC6FFE84F0DB6080" box="[987,1209,368,394]" italics="true" pageId="105" pageNumber="106">P. wesenberglundae</emphasis>
(
<bibRefCitation id="C1214B03FF8A7345FB72FE85F1156081" author="Pettibone" box="[1222,1399,368,395]" firstAuthor="Pettibone" pageId="105" pageNumber="106" pagination="199 - 270" refId="ref88038" refString="Pettibone, M. H. (1970) Polychaeta Errantia of the Siboga Expedition, 4. Some additional polychaetes of the Polynoidae, Hesionidae, Nereidae, Goniadidae, Eunicidae, and Onuphidae, selected as new species by the late Dr. Hermann Augener with remarks on other related species. Siboga Expeditie Monographie, 24 (1 d), 199 - 270." type="journal article" year="1970">Pettibone, 1970</bibRefCitation>
)
</taxonomicName>
<emphasis id="97C4EAE0FF8A7345FA32FE84F4BD60A5" bold="true" pageId="105" pageNumber="106">
<taxonomicNameLabel id="8CF7579BFF8A7345FA32FE84F4BD60A5" pageId="105" pageNumber="106" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
from the Gulf of
<collectingCountry id="DDA77662FF8A7345FE10FE60F58960A5" box="[420,491,405,431]" name="Oman" pageId="105" pageNumber="106">Oman</collectingCountry>
, in the size of eyes and in the
<typeStatus id="7A0B8850FF8A7345FC8CFE60F70A60A5" box="[824,872,405,431]" pageId="105" pageNumber="106">type</typeStatus>
of neurochaetal blades. In
<taxonomicName id="62B04D71FF8A7345FB21FE60F15C60A4" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Augener &amp; Pettibone in Pettibone" baseAuthorityYear="1970" box="[1173,1342,404,430]" class="Polychaeta" family="Hesionidae" genus="Paraleocrates" kingdom="Animalia" order="Phyllodocida" pageId="105" pageNumber="106" phylum="Annelida" rank="species" species="djankarensis">
<emphasis id="97C4EAE0FF8A7345FB21FE60F15C60A4" box="[1173,1342,404,430]" italics="true" pageId="105" pageNumber="106">P. djankarensis</emphasis>
</taxonomicName>
anterior eyes are twice larger than posterior ones, and neurochaetal blades are unidentate, with teeth round and have long, aristate guards, whereas in
<taxonomicName id="62B04D71FF8A7345FE7DFE28F6CA60FC" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Pettibone" baseAuthorityYear="1970" box="[457,680,476,502]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="105" pageNumber="106" phylum="Annelida" rank="species" species="wesenberglundae">
<emphasis id="97C4EAE0FF8A7345FE7DFE28F6CA60FC" box="[457,680,476,502]" italics="true" pageId="105" pageNumber="106">P. wesenberglundae</emphasis>
</taxonomicName>
eyes are of similar size, and neurochaetal blades have no guards at all, but their teeth are tapered but without aristae.
</paragraph>
<paragraph id="A50F36F2FF8A7345FF73FDD1F14C638C" blockId="105.[151,1437,152,683]" pageId="105" pageNumber="106">
The
<typeStatus id="7A0B8850FF8A7345FF4DFDD0F53B6335" box="[249,345,549,575]" pageId="105" pageNumber="106" type="paratype">paratype</typeStatus>
has neurochaetal blades slightly different because instead of having slightly falcate tooth, as in the
<typeStatus id="7A0B8850FF8A7345FF23FDBDF49E6368" box="[151,252,584,610]" pageId="105" pageNumber="106" type="holotype">holotype</typeStatus>
, it is rather straight, and even bent dorsally; because of the poor condition of the specimen, no other differences were found and consequently it is regarded as conspecific pending the finding of better specimens.
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FF8A7345FF73FD65F0DE63A0" box="[199,1212,656,683]" pageId="105" pageNumber="106" type="distribution">
<paragraph id="A50F36F2FF8A7345FF73FD65F0DE63A0" blockId="105.[151,1437,152,683]" box="[199,1212,656,683]" pageId="105" pageNumber="106">
<emphasis id="97C4EAE0FF8A7345FF73FD65F53B63A1" bold="true" box="[199,345,656,683]" pageId="105" pageNumber="106">Distribution</emphasis>
.
<collectingCountry id="DDA77662FF8A7345FED3FD65F58763A0" box="[359,485,656,682]" name="Philippines" pageId="105" pageNumber="106">Philippines</collectingCountry>
and
<collectingCountry id="DDA77662FF8A7345FDA8FD65F6EE63A0" box="[540,652,656,682]" name="Indonesia" pageId="105" pageNumber="106">Indonesia</collectingCountry>
, in mixed or muddy bottoms at
<quantity id="62489B17FF8A7345FC44FD65F00D63A1" box="[1008,1135,656,683]" metricMagnitude="2" metricUnit="m" metricValue="3.025" metricValueMax="3.3" metricValueMin="2.75" pageId="105" pageNumber="106" unit="m" value="302.5" valueMax="330.0" valueMin="275.0">275330 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>