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<mods:title id="5960FDFF764F1887CC010CAA75B8E59A">Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense- 19. Odontocheila microptera nom. nov., a new replacement name for O. euryoides W. Horn, 1922, and lectotype designation of O. nitidicollis (Dejean, 1825) (Coleoptera: Cicindelidae)</mods:title>
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<mods:namePart id="9CA98CD6BD02AC754C709EA004F06214">Moravec, Jiří</mods:namePart>
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<mods:namePart id="1B7548E56AEF50DABA5E9D40C786C0FA">Huber, Ronald</mods:namePart>
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<paragraph id="A475EC57FFD5C81D279EFF61FDFF1891" blockId="10.[151,581,151,211]" box="[151,581,151,178]" pageId="10" pageNumber="119">
<heading id="FF3D5B3BFFD5C81D279EFF61FDFF1891" bold="true" box="[151,581,151,178]" fontSize="11" level="1" pageId="10" pageNumber="119" reason="1">
<emphasis id="96BE3045FFD5C81D279EFF61FDFF1891" bold="true" box="[151,581,151,178]" pageId="10" pageNumber="119">
<taxonomicName id="63CA97D4FFD5C81D279EFF61FE7F1892" ID-CoL="8MD7N" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[151,453,151,177]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD5C81D279EFF61FE7F1892" bold="true" box="[151,453,151,177]" italics="true" pageId="10" pageNumber="119">ODONTOcheila micrOPTera</emphasis>
</taxonomicName>
nom. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="A475EC57FFD5C81D279EFF4CFE9D18F0" blockId="10.[151,581,151,211]" box="[151,295,186,211]" pageId="10" pageNumber="119">
(
<figureCitation id="3CF1F0D2FFD5C81D2796FF4CFEA518F0" box="[159,287,186,211]" captionStart-0="FIGURES 30 – 36" captionStart-1="FIGURES 37 – 48" captionStart-2="FIGURES 49 – 57" captionStart-3="FIGURE 58" captionStartId-0="12.[151,264,1877,1899]" captionStartId-1="13.[151,264,1842,1864]" captionStartId-2="14.[151,264,1877,1899]" captionStartId-3="15.[151,250,1172,1194]" captionTargetBox-0="[161,1426,205,1840]" captionTargetBox-1="[161,1417,205,1803]" captionTargetBox-2="[158,1416,204,1812]" captionTargetBox-3="[155,1432,193,1151]" captionTargetId-0="figure@12.[151,1436,193,1856]" captionTargetId-1="figure@13.[151,1436,193,1821]" captionTargetId-2="figure@14.[151,1436,193,1856]" captionTargetId-3="figure@15.[155,1432,193,1151]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="FIGURES 30 – 36. Odontocheila microptera nom. nov. 30 – 32 — habitus: 30 — ♂, 9.5 mm, “ Brasilia ”, HT (SDEI); 31 — ♂, 11 mm, Uruguay, Tacuarembo (CCJM ex DBCN); 32 — ♀, 10.2 mm, Argentina, Estancia San Noria (BMNH); 33 – 36 — elytron: 33 — ♂, HT (SDEI); 34 — ♂, Uruguay, Tacuarembo (CCJM); 35 — ♀, ibid., (CCJM); 36 — ♀, Bahia (MFNB). Bar = 1 mm." captionText-1="FIGURES 37 – 48. Odontocheila microptera nom. nov. 37 – 40 — head: 37 — ♂, HT (SDEI); 38 — ♂, Uruguay, Tacuarembo (CCJM); 39 — ♂, ibid.; 40 — ♂, ditto, ventral view; 41 — right mandible, ♀, ibid.; 42 — maxillary palpus, ♀, ibid. (CCJM); 43 — head part, ♀, Bahia (MFNB); 44 – 48 — labrum: 44 — ♀, Bahia (MFNB); 45 — ♀, Uruguay, Tacuarembo (CCJM); 46 — ♀, Argentina, Estancia San Noria (BMNH); 47 — ♂, HT (SDEI); 48 — ♂, Uruguay, Tacuarembo (CCJM). Bars = 1 mm." captionText-2="FIGURES 49 – 57. Odontocheila microptera nom. nov. 49 – 51 — atrophic thoracic wings: 49 – 50 — ♀, Uruguay, Tacuarembo (CCJM); 51 — ♂, HT (SDEI); 52 – 54 — pronotum: 52 — ♂, HT (SDEI); 53 — ♂, Uruguay, Tacuarembo (CCJM); 54 — ♀, Argentina, Estancia San Noria (BMNH); 55 – 57 — male aedeagi: 55 — HT (SDEI); 56 – 57 — Uruguay, Tacuarembo (CCJM). Bars = 1 mm." captionText-3="FIGURE 58. One of the habitats of O. microptera nom. nov. near Tacuarembo, Uruguay." httpUri-0="https://zenodo.org/record/1044920/files/figure.png" httpUri-1="https://zenodo.org/record/1044922/files/figure.png" httpUri-2="https://zenodo.org/record/1044924/files/figure.png" httpUri-3="https://zenodo.org/record/1044926/files/figure.png" pageId="10" pageNumber="119">Figs 30–58</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="A475EC57FFD5C81D279EFEF4FAB31914" blockId="10.[151,1437,258,403]" pageId="10" pageNumber="119">
<treatmentCitationGroup id="84DACB79FFD5C81D279EFEF4FAB31914" infered="true" pageId="10" pageNumber="119">
<taxonomicName id="63CA97D4FFD5C81D279EFEF4FDE0193B" authority="W. Horn, 1922: 100" authorityName="W. Horn" authorityPageNumber="100" authorityYear="1922" box="[151,602,258,280]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="euryoides">
<emphasis id="96BE3045FFD5C81D279EFEF4FE3B193B" box="[151,385,258,280]" italics="true" pageId="10" pageNumber="119">Odontochila euryoides</emphasis>
<treatmentCitation id="256BCA46FFD5C81D2680FEF5FDE0193B" author="Horn" box="[393,602,258,280]" page="100" pageId="10" pageNumber="119" year="1922">
<bibRefCitation id="C05B91A6FFD5C81D2680FEF5FD9C193B" author="Horn" box="[393,550,258,280]" pageId="10" pageNumber="119" refString="Horn, W. (1922) Studien uber neue und alte Cicindeliden (Col.), (Neubeschreibungen, Synonymie, Faunistik). Zoologische Mededeelingen, 7, 1 - 112." type="journal article" year="1922">W. Horn, 1922</bibRefCitation>
: 100
</treatmentCitation>
</taxonomicName>
, 101, fig. 9, primary junior homonym of 
<taxonomicName id="63CA97D4FFD5C81D2305FEF4FEDA1914" ID-CoL="8LKV8" authority="W. Horn, 1906: 85" authorityName="W. Horn" authorityPageNumber="85" authorityYear="1906" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="subSpecies" species="bennigseni" subSpecies="euryoides">
<emphasis id="96BE3045FFD5C81D2305FEF4FAC2193B" box="[1036,1400,258,280]" italics="true" pageId="10" pageNumber="119">Odontocheila bennigseni euryoides</emphasis>
<treatmentCitation id="256BCA46FFD5C81D2289FEF5FEDA1914" author="Horn" page="85" pageId="10" pageNumber="119" year="1906">
<bibRefCitation id="C05B91A6FFD5C81D2289FEF5FE871914" author="Horn" pageId="10" pageNumber="119" refString="Horn, W. (1906) Odontochila bennigseni euryoides (nov. subsp.) und das System der Cicindelini. Deutsche Entomologische Zeitschrift, 1, 85 - 86." type="journal article" year="1906">W. Horn, 1906</bibRefCitation>
: 85
</treatmentCitation>
</taxonomicName>
(currently a subspecies of 
<taxonomicName id="63CA97D4FFD5C81D2578FED7FBB51914" ID-CoL="3D334" authority="(W. Horn, 1897)" authorityName="W. Horn" authorityYear="1897" baseAuthorityName="W. Horn" baseAuthorityYear="1897" box="[625,1039,289,311]" class="Insecta" family="Carabidae" genus="Euryarthron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="bennigseni">
<emphasis id="96BE3045FFD5C81D2578FED7FCD81914" box="[625,866,289,311]" italics="true" pageId="10" pageNumber="119">Euryarthron bennigseni</emphasis>
(W. Horn, 1897)
</taxonomicName>
– see “Remarks” below.
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<emphasis id="96BE3045FFD5C81D279EFEB6FEA51975" bold="true" box="[151,287,320,342]" pageId="10" pageNumber="119">Type locality</emphasis>
. “Brasilia”.
</paragraph>
</subSubSection>
<subSubSection id="ECD0BFDCFFD5C81D279EFEA8FDEA19B0" pageId="10" pageNumber="119" type="reference_group">
<paragraph id="A475EC57FFD5C81D279EFEA8FDE61957" blockId="10.[151,1437,258,403]" box="[151,604,350,372]" pageId="10" pageNumber="119">
<treatmentCitationGroup id="84DACB79FFD5C81D279EFEA8FDE61957" box="[151,604,350,372]" pageId="10" pageNumber="119">
<taxonomicName id="63CA97D4FFD5C81D279EFEA8FE9B1957" ID-CoL="8KY2T" authorityName="Laporte de Castelnau" authorityYear="1834" box="[151,289,350,372]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="96BE3045FFD5C81D279EFEA8FE9B1957" box="[151,289,350,372]" italics="true" pageId="10" pageNumber="119">Odontocheila</emphasis>
</taxonomicName>
? 
<taxonomicName id="63CA97D4FFD5C81D263BFEA8FDE21957" authority=": Wiesner 1992: 80" authorityName="Wiesner" authorityPageNumber="80" authorityYear="1992" box="[306,600,350,372]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="euryoides">
<emphasis id="96BE3045FFD5C81D263BFEA8FE2E1957" box="[306,404,350,372]" italics="true" pageId="10" pageNumber="119">euryoides</emphasis>
: 
<treatmentCitation id="256BCA46FFD5C81D26ABFEA8FDE21957" author="Wiesner" box="[418,600,350,372]" page="80" pageId="10" pageNumber="119" year="1992">
<bibRefCitation id="C05B91A6FFD5C81D26ABFEA8FD881957" author="Wiesner" box="[418,562,350,372]" pageId="10" pageNumber="119" refString="Wiesner, J. (1992) Verzeichnis der Sandlaufkafer der Welt. Checklist of the tiger beetles of the world (Coleoptera, Cicindelidae). Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">Wiesner 1992</bibRefCitation>
: 80
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A475EC57FFD5C81D279EFE8BFDEA19B0" blockId="10.[151,1437,258,403]" box="[151,592,381,403]" pageId="10" pageNumber="119">
<treatmentCitationGroup id="84DACB79FFD5C81D279EFE8BFDEA19B0" box="[151,592,381,403]" pageId="10" pageNumber="119">
<taxonomicName id="63CA97D4FFD5C81D279EFE8BFDF619B0" ID-CoL="8LKV8" authority=": Lorenz 1998 a: 35" authorityName=": Lorenz" authorityPageNumber="35" authorityYear="1998" box="[151,588,381,403]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="euryoides">
<emphasis id="96BE3045FFD5C81D279EFE8BFE3319B0" box="[151,393,381,403]" italics="true" pageId="10" pageNumber="119">Odontocheila euryoides</emphasis>
: 
<treatmentCitation id="256BCA46FFD5C81D269EFE88FDF619B0" author="Lorenz" box="[407,588,381,403]" page="35" pageId="10" pageNumber="119" year="1998">
<bibRefCitation id="C05B91A6FFD5C81D269EFE88FD9C19B0" author="Lorenz" box="[407,550,381,403]" pageId="10" pageNumber="119" refString="Lorenz, W. (1998 a) Systematic list of extant ground beetles of the world (Insecta, Coleoptera, &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Privately published, Tutzing, 502 pp." type="book" year="1998" yearSuffix="a">Lorenz 1998a</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="ECD0BFDCFFD5C81D279EFE49FD1F1C43" pageId="10" pageNumber="119" type="materials_examined">
<paragraph id="A475EC57FFD5C81D279EFE49FD471AF6" blockId="10.[151,1437,447,2021]" pageId="10" pageNumber="119">
<emphasis id="96BE3045FFD5C81D279EFE49FEF319FB" bold="true" box="[151,329,447,472]" pageId="10" pageNumber="119">
<typeStatus id="7B7152F5FFD5C81D279EFE49FF6819FB" box="[151,210,447,472]" pageId="10" pageNumber="119">Type</typeStatus>
material.
</emphasis>
<typeStatus id="7B7152F5FFD5C81D265DFE49FE0619FB" box="[340,444,447,472]" pageId="10" pageNumber="119" type="holotype">Holotype</typeStatus>
[originally of 
<taxonomicName id="63CA97D4FFD5C81D2562FE36FD4519FB" authorityName=": Lorenz" authorityYear="1998" box="[619,767,448,472]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="euryoides">
<emphasis id="96BE3045FFD5C81D2562FE36FD4519FB" box="[619,767,448,472]" italics="true" pageId="10" pageNumber="119">O. euryoides</emphasis>
</taxonomicName>
] (by monotypy), 
<specimenCount id="B2CC27DEFFD5C81D24D8FE36FC5C19F4" box="[977,998,448,471]" pageId="10" pageNumber="119" type="male">♂</specimenCount>
in 
<collectionCode id="C2DB7492FFD5C81D231BFE36FBE919FB" box="[1042,1107,448,472]" pageId="10" pageNumber="119">SDEI</collectionCode>
, labelled: “Brasilia / ex cab Dieck.” [handwritten] // “Coll. Ehlers / V. de Poll” [printed]; “ 
<typeStatus id="7B7152F5FFD5C81D246BFE12FC1B19DF" box="[866,929,484,508]" pageId="10" pageNumber="119">Type</typeStatus>
! / Dr. W. Horn” [printed] // “coll. W. Horn / 
<collectionCode id="C2DB7492FFD5C81D279EFDFEFF7C1A3C" box="[151,198,520,543]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="10" pageNumber="119">DEI</collectionCode>
Eberswalde” [printed] // “ 
<typeStatus id="7B7152F5FFD5C81D26E2FDF1FDDE1A03" box="[491,612,519,544]" pageId="10" pageNumber="119" type="holotype">Holotypus</typeStatus>
” [red, printed] // “ 
<taxonomicName id="63CA97D4FFD5C81D2426FDF1FC1B1A03" authorityName=": Lorenz" authorityYear="1998" box="[815,929,519,544]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="euryoides">euryoides</taxonomicName>
, mihi” [large green-blue collection label with black frame, handwritten] // “ 
<taxonomicName id="63CA97D4FFD5C81D26FFFDDAFCC01A67" authority="W. Horn" authorityName="W. Horn" box="[502,890,556,581]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="euryoides">Odontocheila euryoides W. Horn</taxonomicName>
<typeStatus id="7B7152F5FFD5C81D248FFDDBFC041A66" box="[902,958,557,581]" pageId="10" pageNumber="119">Type</typeStatus>
(
<collectionCode id="C2DB7492FFD5C81D24D8FDDAFBBA1A66" box="[977,1024,556,581]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="10" pageNumber="119">DEI</collectionCode>
–Eberswalde) / borrowed by D. L. Pearson / 
<date id="D074CA97FFD5C81D2618FDB9FE211A4B" box="[273,411,591,616]" pageId="10" pageNumber="119" value="1996-10-23">23.Oct.1996</date>
(drawer # 60)”[printed] // “Revision Jiří Moravec 2012: / 
<typeStatus id="7B7152F5FFD5C81D236CFDB9FB771A4B" box="[1125,1229,591,616]" pageId="10" pageNumber="119" type="holotype">Holotype</typeStatus>
(by monotypy) / 
<taxonomicName id="63CA97D4FFD5C81D279EFD82FE981AAE" authorityName="Agassiz" authorityYear="1846" box="[151,290,628,653]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="genus">Odontochila</taxonomicName>
/ 
<taxonomicName id="63CA97D4FFD5C81D2648FD82FDD01AAF" authority="W. Horn, 1922" authorityName="W. Horn" authorityYear="1922" box="[321,618,628,653]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="euryoides">
euryoides 
<bibRefCitation id="C05B91A6FFD5C81D26B0FD83FDD01AAF" author="Horn" box="[441,618,628,652]" pageId="10" pageNumber="119" refString="Horn, W. (1922) Studien uber neue und alte Cicindeliden (Col.), (Neubeschreibungen, Synonymie, Faunistik). Zoologische Mededeelingen, 7, 1 - 112." type="journal article" year="1922">W. Horn, 1922</bibRefCitation>
</taxonomicName>
” [red, printed] // “ 
<taxonomicName id="63CA97D4FFD5C81D2443FD82FC591AAE" authorityName="Laporte de Castelnau" authorityYear="1834" box="[842,995,628,653]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="genus">Odontocheila</taxonomicName>
/ 
<taxonomicName id="63CA97D4FFD5C81D230AFD82FA911AAF" authority="W. Horn, 1922" authorityName="W. Horn" authorityYear="1922" box="[1027,1323,628,653]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="euryoides">
euryoides 
<bibRefCitation id="C05B91A6FFD5C81D2375FD83FA911AAF" author="Horn" box="[1148,1323,628,652]" pageId="10" pageNumber="119" refString="Horn, W. (1922) Studien uber neue und alte Cicindeliden (Col.), (Neubeschreibungen, Synonymie, Faunistik). Zoologische Mededeelingen, 7, 1 - 112." type="journal article" year="1922">W. Horn, 1922</bibRefCitation>
</taxonomicName>
/ det Jiří Moravec 2012” [printed]. The 
<typeStatus id="7B7152F5FFD5C81D26FBFD61FDEE1A93" box="[498,596,663,688]" pageId="10" pageNumber="119" type="holotype">holotype</typeStatus>
is accompanied with separately mounted thoracic wing, labelled by Horn: “Unterflügel / von / 
<taxonomicName id="63CA97D4FFD5C81D2673FD4AFDAD1AF6" box="[378,535,700,725]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="euryoides">Od. euryoides</taxonomicName>
/ m.” [handwritten].
</paragraph>
<paragraph id="A475EC57FFD5C81D27CEFD29FD1F1C43" blockId="10.[151,1437,447,2021]" pageId="10" pageNumber="119">
<emphasis id="96BE3045FFD5C81D27CEFD29FE4D1ADB" bold="true" box="[199,503,735,760]" pageId="10" pageNumber="119">Other material examined</emphasis>
. Historical data. 1 ♀ in MFNB: “Hist. Coll. (
<taxonomicName id="63CA97D4FFD5C81D24F7FD29FBC51ADB" authorityName="" authorityYear="1758" box="[1022,1151,735,760]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
), Nr. 3642, 
<taxonomicName id="63CA97D4FFD5C81D220CFD29FEB51B3E" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nitidicollis">Odontocheila nitidicollis</taxonomicName>
/ Dej. / Bahia, Freireiss / Zool. Mus. Berlin”. 1 ♀ in BMNH: “Estancia San Noria / Rio San Javier, / Santa Fe, Argentine / G. E. Bryant / 
<date id="D074CA97FFD5C81D253AFCDEFD181B63" box="[563,674,808,832]" pageId="10" pageNumber="119" value="1912-01-12">12.I.1912</date>
”; “ 
<taxonomicName id="63CA97D4FFD5C81D25CDFCD1FCEB1B63" authorityName="Agassiz" authorityYear="1846" box="[708,849,807,832]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="genus">Odontochila</taxonomicName>
/ 
<taxonomicName id="63CA97D4FFD5C81D2460FCD1FC5B1B63" box="[873,993,807,832]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nitidicollis">nitidicollis</taxonomicName>
/ Dej. / Dr. W. Horn det, 1925”; “Pres. by / Imp. Bur. Ent. / Brit. Mus. / 1925–292”. All specimens labelled: “ 
<taxonomicName id="63CA97D4FFD5C81D24B5FCBAFBEC1B46" authorityName="Laporte de Castelnau" authorityYear="1834" box="[956,1110,844,869]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="genus">Odontocheila</taxonomicName>
/ 
<taxonomicName id="63CA97D4FFD5C81D2367FCBAFA2A1B47" authority="W. Horn, 1922" authorityName="W. Horn" authorityYear="1922" box="[1134,1424,844,869]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="euryoides">
euryoides 
<bibRefCitation id="C05B91A6FFD5C81D23EDFCBBFA2A1B47" author="Horn" box="[1252,1424,844,868]" pageId="10" pageNumber="119" refString="Horn, W. (1922) Studien uber neue und alte Cicindeliden (Col.), (Neubeschreibungen, Synonymie, Faunistik). Zoologische Mededeelingen, 7, 1 - 112." type="journal article" year="1922">W. Horn, 1922</bibRefCitation>
</taxonomicName>
), det Jiří Moravec 2012 (2014 respectively)” Recent data. 2 ♂♂, 10 ♀♀ in DBCN, 3 ♂♂, 1 ♀ in CCJM: “Uruguay / Hyw. 5, KM. 395 (N.- Tacuarembo) 
<quantity id="633241B2FFD5C81D2536FC62FD311B8F" box="[575,651,916,940]" metricMagnitude="2" metricUnit="m" metricValue="1.7899999999999998" pageId="10" pageNumber="119" unit="m" value="179.0">179 m</quantity>
, -31.684, -55.926° / D. Brzoska 
<date id="D074CA97FFD5C81D230FFC62FBC11B8E" box="[1030,1147,916,941]" pageId="10" pageNumber="119" value="2017-01-15">15-I-2017</date>
”. 2 ♂♂, 4 ♀♀ in JCDG: ibid., except for “leg. Johnnie Chong”. 4 ♂♂, 2 ♀♀ in DBCN: ibid. except for: “D. Brzoska 
<date id="D074CA97FFD5C81D23BDFC41FA931BF3" box="[1204,1321,951,976]" pageId="10" pageNumber="119" value="2017-01-16">16-I-2017</date>
”. 8 ♂♂, 5 ♀♀ in JCDG: ibid., except for: “leg. Johnnie Chong”. 3 ♂♂ in DBCN: ibid., except for: “Hyw. 26, KM. 220 (W.- Tacuarembo) 
<quantity id="633241B2FFD5C81D263DFC09FE391C34" box="[308,387,1023,1048]" metricMagnitude="2" metricUnit="m" metricValue="1.64" pageId="10" pageNumber="119" unit="m" value="164.0">164 m</quantity>
., -31.763, -56.077°, D. Brzoska 
<date id="D074CA97FFD5C81D25F3FC09FC891C34" box="[762,819,1023,1048]" bridgedPair="-" pageId="10" pageNumber="119" value="2016-01">16-I-</date>
017°”. 5 ♂♂, 2 ♀♀ in JCDG: ibid., except for “leg. Johnnie Chong”. 1 ♂, 1♀ in DBCN ibid., except for: “246 (W.- Tacuarembo) 
<quantity id="633241B2FFD5C81D2319FBD2FBE31C1F" box="[1040,1113,1060,1084]" metricMagnitude="2" metricUnit="m" metricValue="1.71" pageId="10" pageNumber="119" unit="m" value="171.0">171 m</quantity>
, 31.711, -55.866”. 2 ♂♂ in JCDG: ibid., except for: “leg. Johnnie Chong”.
</paragraph>
</subSubSection>
<subSubSection id="ECD0BFDCFFD5C81D27CEFB9AFD6B1FE3" pageId="10" pageNumber="119" type="diagnosis">
<paragraph id="A475EC57FFD5C81D27CEFB9AFBDE1E16" blockId="10.[151,1437,447,2021]" pageId="10" pageNumber="119">
<emphasis id="96BE3045FFD5C81D27CEFB9AFE7F1CA6" bold="true" box="[199,453,1132,1157]" pageId="10" pageNumber="119">Differential diagnosis</emphasis>
. 
<taxonomicName id="63CA97D4FFD5C81D26DAFB9AFD561CA7" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[467,748,1132,1156]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD5C81D26DAFB9AFD561CA7" box="[467,748,1132,1156]" italics="true" pageId="10" pageNumber="119">Odontocheila microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD5C81D25FAFB9BFCD81CA7" box="[755,866,1133,1156]" pageId="10" pageNumber="119">nom. nov.</taxonomicNameLabel>
shares the notably nitid, reddish-cupreous pronotal surface, immaculate elytra and coloration of other body portions with 
<taxonomicName id="63CA97D4FFD5C81D24D0FB66FB4F1C8B" box="[985,1269,1168,1192]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD5C81D24D0FB66FB4F1C8B" box="[985,1269,1168,1192]" italics="true" pageId="10" pageNumber="119">Odontocheila nitidicollis</emphasis>
</taxonomicName>
. However, 
<taxonomicName id="63CA97D4FFD5C81D2289FB66FEA91CEF" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD5C81D2289FB66FEA91CEF" italics="true" pageId="10" pageNumber="119">O. microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD5C81D2612FB43FE361CEF" box="[283,396,1205,1228]" pageId="10" pageNumber="119">nom. nov.</taxonomicNameLabel>
can be immediately distinguished by its uniquely mutually ovoid elytra in both sexes (
<figureCitation id="3CF1F0D2FFD5C81D2263FB42FF5B1CD3" captionStart="FIGURES 30 – 36" captionStartId="12.[151,264,1877,1899]" captionTargetBox="[161,1426,205,1840]" captionTargetId="figure@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 30 – 36. Odontocheila microptera nom. nov. 30 – 32 — habitus: 30 — ♂, 9.5 mm, “ Brasilia ”, HT (SDEI); 31 — ♂, 11 mm, Uruguay, Tacuarembo (CCJM ex DBCN); 32 — ♀, 10.2 mm, Argentina, Estancia San Noria (BMNH); 33 – 36 — elytron: 33 — ♂, HT (SDEI); 34 — ♂, Uruguay, Tacuarembo (CCJM); 35 — ♀, ibid., (CCJM); 36 — ♀, Bahia (MFNB). Bar = 1 mm." httpUri="https://zenodo.org/record/1044920/files/figure.png" pageId="10" pageNumber="119">Figs 30–36</figureCitation>
). Its male labrum is distinctly bicolored with large ivory-yellow anterior area (
<figureCitation id="3CF1F0D2FFD5C81D2342FB21FB701CD3" box="[1099,1226,1239,1264]" captionStart="FIGURES 37 – 48" captionStartId="13.[151,264,1842,1864]" captionTargetBox="[161,1417,205,1803]" captionTargetId="figure@13.[151,1436,193,1821]" captionTargetPageId="13" captionText="FIGURES 37 – 48. Odontocheila microptera nom. nov. 37 – 40 — head: 37 — ♂, HT (SDEI); 38 — ♂, Uruguay, Tacuarembo (CCJM); 39 — ♂, ibid.; 40 — ♂, ditto, ventral view; 41 — right mandible, ♀, ibid.; 42 — maxillary palpus, ♀, ibid. (CCJM); 43 — head part, ♀, Bahia (MFNB); 44 – 48 — labrum: 44 — ♀, Bahia (MFNB); 45 — ♀, Uruguay, Tacuarembo (CCJM); 46 — ♀, Argentina, Estancia San Noria (BMNH); 47 — ♂, HT (SDEI); 48 — ♂, Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044922/files/figure.png" pageId="10" pageNumber="119">Figs 47–48</figureCitation>
) and has 4-dentate anterior margin which is truncate or shallowly emarginated in middle (lacking median tooth) with the anterolateral and anterior teeth in the same level, giving the male labrum almost transverse shape, in contrast to the predominantly black male labrum with 5-dentate anterior margin in 
<taxonomicName id="63CA97D4FFD5C81D249DFAB2FB951D7F" box="[916,1071,1348,1372]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD5C81D249DFAB2FB951D7F" box="[916,1071,1348,1372]" italics="true" pageId="10" pageNumber="119">O. nitidicollis</emphasis>
</taxonomicName>
. In contrast to the entirely black female labrum in 
<taxonomicName id="63CA97D4FFD5C81D2655FA9EFE4F1DA3" box="[348,501,1384,1408]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD5C81D2655FA9EFE4F1DA3" box="[348,501,1384,1408]" italics="true" pageId="10" pageNumber="119">O. nitidicollis</emphasis>
</taxonomicName>
, the female labrum in 
<taxonomicName id="63CA97D4FFD5C81D25F9FA9EFC371D5C" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[752,909,1384,1407]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD5C81D25F9FA9EFC371D5C" box="[752,909,1384,1407]" italics="true" pageId="10" pageNumber="119">O. microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD5C81D249AFA9EFBB81D5C" box="[915,1026,1384,1407]" pageId="10" pageNumber="119">nom. nov.</taxonomicNameLabel>
is either entirely black or bicoloured and may resemble the bicoloured labrum in females of the species-group related to 
<taxonomicName id="63CA97D4FFD5C81D233DFA7AFB161D87" authorityName="Klug" authorityYear="1834" box="[1076,1196,1420,1444]" class="Insecta" family="Carabidae" genus="Cicindela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="rutilans">
<emphasis id="96BE3045FFD5C81D233DFA7AFB161D87" box="[1076,1196,1420,1444]" italics="true" pageId="10" pageNumber="119">O. rutilans</emphasis>
</taxonomicName>
, including 
<taxonomicName id="63CA97D4FFD5C81D2220FA7AFA261D87" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[1321,1436,1420,1444]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="fulgens">
<emphasis id="96BE3045FFD5C81D2220FA7AFA261D87" box="[1321,1436,1420,1444]" italics="true" pageId="10" pageNumber="119">O. fulgens</emphasis>
</taxonomicName>
(but see annotation in the “Variability” below). Moreover, the terminal palpomeres of maxillary palpi are in 
<taxonomicName id="63CA97D4FFD5C81D2288FA46FEA91DCF" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD5C81D2288FA46FEA91DCF" italics="true" pageId="10" pageNumber="119">O. microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD5C81D2615FA23FE2A1DCF" box="[284,400,1493,1516]" pageId="10" pageNumber="119">nom. nov.</taxonomicNameLabel>
slightly but notably narrower (
<figureCitation id="3CF1F0D2FFD5C81D25F5FA22FC171DCF" box="[764,941,1492,1517]" captionStart="FIGURES 37 – 48" captionStartId="13.[151,264,1842,1864]" captionTargetBox="[161,1417,205,1803]" captionTargetId="figure@13.[151,1436,193,1821]" captionTargetPageId="13" captionText="FIGURES 37 – 48. Odontocheila microptera nom. nov. 37 – 40 — head: 37 — ♂, HT (SDEI); 38 — ♂, Uruguay, Tacuarembo (CCJM); 39 — ♂, ibid.; 40 — ♂, ditto, ventral view; 41 — right mandible, ♀, ibid.; 42 — maxillary palpus, ♀, ibid. (CCJM); 43 — head part, ♀, Bahia (MFNB); 44 – 48 — labrum: 44 — ♀, Bahia (MFNB); 45 — ♀, Uruguay, Tacuarembo (CCJM); 46 — ♀, Argentina, Estancia San Noria (BMNH); 47 — ♂, HT (SDEI); 48 — ♂, Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044922/files/figure.png" pageId="10" pageNumber="119">Figs 38–40, 42</figureCitation>
) than the almost spatulate-dilated terminal palpomeres in males of 
<taxonomicName id="63CA97D4FFD5C81D26B2FA0EFDE61E33" box="[443,604,1528,1552]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD5C81D26B2FA0EFDE61E33" box="[443,604,1528,1552]" italics="true" pageId="10" pageNumber="119">O. nitidicollis</emphasis>
</taxonomicName>
(
<figureCitation id="3CF1F0D2FFD5C81D2579FA01FC921E33" box="[624,808,1527,1552]" captionStart="FIGURES 10 – 22" captionStartId="8.[151,264,1814,1836]" captionTargetBox="[154,1424,204,1775]" captionTargetId="figure@8.[151,1436,193,1792]" captionTargetPageId="8" captionText="FIGURES 10 – 22. Odontocheila nitidicollis Dejean. 10 – 12 — head: 10 — ♂, LT (MNHN); 11 — ♂, Paraguay, Estancia Terrado (CCJM); 12 — ♀, Venezuela, Canaima NP (CPVP); 13 – 18 — labrum: 13 —— ♂, LT (MNHN); 14 — ♂, Paraguay, Estancia Terrado (CCJM); 15 — ♂, Bolivia, Concepcion (CCJM); 16 — ♂, ex Dejean-Chaudoir, PLT (MNHN); 17 — ♀, Brazil, Barra do Carças (CMHP); 18 — ♀, Venezuela, Canaima NP (CPVP); 19 — maxillary palpus, Paraguay, Estancia Terrado (CCJM); 20 – 21 — antennal scape: 20 — ♂, Paraguay, Estancia Terrado (CCJM); 21 — ♂, Bolivia, Concepcion (CCJM); 22 — thoracic wing, ♀, ibid., CCJM. Bars = 1 mm." httpUri="https://zenodo.org/record/1044916/files/figure.png" pageId="10" pageNumber="119">Figs 10–12, 19</figureCitation>
), and also the antennal scape (
<figureCitation id="3CF1F0D2FFD5C81D23ABFA01FA931E33" box="[1186,1321,1527,1552]" captionStart="FIGURES 10 – 22" captionStartId="8.[151,264,1814,1836]" captionTargetBox="[154,1424,204,1775]" captionTargetId="figure@8.[151,1436,193,1792]" captionTargetPageId="8" captionText="FIGURES 10 – 22. Odontocheila nitidicollis Dejean. 10 – 12 — head: 10 — ♂, LT (MNHN); 11 — ♂, Paraguay, Estancia Terrado (CCJM); 12 — ♀, Venezuela, Canaima NP (CPVP); 13 – 18 — labrum: 13 —— ♂, LT (MNHN); 14 — ♂, Paraguay, Estancia Terrado (CCJM); 15 — ♂, Bolivia, Concepcion (CCJM); 16 — ♂, ex Dejean-Chaudoir, PLT (MNHN); 17 — ♀, Brazil, Barra do Carças (CMHP); 18 — ♀, Venezuela, Canaima NP (CPVP); 19 — maxillary palpus, Paraguay, Estancia Terrado (CCJM); 20 – 21 — antennal scape: 20 — ♂, Paraguay, Estancia Terrado (CCJM); 21 — ♂, Bolivia, Concepcion (CCJM); 22 — thoracic wing, ♀, ibid., CCJM. Bars = 1 mm." httpUri="https://zenodo.org/record/1044916/files/figure.png" pageId="10" pageNumber="119">Figs 20–21</figureCitation>
) is in 
<taxonomicName id="63CA97D4FFD5C81D2288FA0EFEB51E17" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD5C81D2288FA0EFEB51E17" italics="true" pageId="10" pageNumber="119">O. nitidicollis</emphasis>
</taxonomicName>
slightly more dilated (but in both species never distinctly spatulate-dilated).
</paragraph>
<paragraph id="A475EC57FFD5C81D27CEF9B6FF791EE7" blockId="10.[151,1437,447,2021]" pageId="10" pageNumber="119">
Because of its uniquely atrophic thoracic wings which are firmly connate with the soft thoracic tissue (
<figureCitation id="3CF1F0D2FFD5C81D2235F9C9FA351E7B" box="[1340,1423,1599,1624]" captionStart="FIGURES 49 – 57" captionStartId="14.[151,264,1877,1899]" captionTargetBox="[158,1416,204,1812]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURES 49 – 57. Odontocheila microptera nom. nov. 49 – 51 — atrophic thoracic wings: 49 – 50 — ♀, Uruguay, Tacuarembo (CCJM); 51 — ♂, HT (SDEI); 52 – 54 — pronotum: 52 — ♂, HT (SDEI); 53 — ♂, Uruguay, Tacuarembo (CCJM); 54 — ♀, Argentina, Estancia San Noria (BMNH); 55 – 57 — male aedeagi: 55 — HT (SDEI); 56 – 57 — Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044924/files/figure.png" pageId="10" pageNumber="119">Fig. 49</figureCitation>
), 
<taxonomicName id="63CA97D4FFD5C81D279EF992FE8F1E5F" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[151,309,1636,1660]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD5C81D279EF992FE8F1E5F" box="[151,309,1636,1660]" italics="true" pageId="10" pageNumber="119">O. microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD5C81D2635F993FE171E5F" box="[316,429,1637,1660]" pageId="10" pageNumber="119">nom. nov.</taxonomicNameLabel>
, is the only flightless species of 
<taxonomicName id="63CA97D4FFD5C81D2411F992FC0A1E5F" authorityName="Laporte de Castelnau" authorityYear="1834" box="[792,944,1636,1660]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="96BE3045FFD5C81D2411F992FC0A1E5F" box="[792,944,1636,1660]" italics="true" pageId="10" pageNumber="119">Odontocheila</emphasis>
</taxonomicName>
; the total length of the atrophic wings (
<figureCitation id="3CF1F0D2FFD5C81D2263F992FF641E83" captionStart="FIGURES 49 – 57" captionStartId="14.[151,264,1877,1899]" captionTargetBox="[158,1416,204,1812]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURES 49 – 57. Odontocheila microptera nom. nov. 49 – 51 — atrophic thoracic wings: 49 – 50 — ♀, Uruguay, Tacuarembo (CCJM); 51 — ♂, HT (SDEI); 52 – 54 — pronotum: 52 — ♂, HT (SDEI); 53 — ♂, Uruguay, Tacuarembo (CCJM); 54 — ♀, Argentina, Estancia San Noria (BMNH); 55 – 57 — male aedeagi: 55 — HT (SDEI); 56 – 57 — Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044924/files/figure.png" pageId="10" pageNumber="119">Figs 50–51</figureCitation>
) does not surpass the elytral third, in contrast to the normally developed, long wings in 
<taxonomicName id="63CA97D4FFD5C81D23CDF97EFAE41E83" box="[1220,1374,1672,1696]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD5C81D23CDF97EFAE41E83" box="[1220,1374,1672,1696]" italics="true" pageId="10" pageNumber="119">O. nitidicollis</emphasis>
</taxonomicName>
(
<figureCitation id="3CF1F0D2FFD5C81D2267F971FF0D1EE7" captionStart="FIGURES 10 – 22" captionStartId="8.[151,264,1814,1836]" captionTargetBox="[154,1424,204,1775]" captionTargetId="figure@8.[151,1436,193,1792]" captionTargetPageId="8" captionText="FIGURES 10 – 22. Odontocheila nitidicollis Dejean. 10 – 12 — head: 10 — ♂, LT (MNHN); 11 — ♂, Paraguay, Estancia Terrado (CCJM); 12 — ♀, Venezuela, Canaima NP (CPVP); 13 – 18 — labrum: 13 —— ♂, LT (MNHN); 14 — ♂, Paraguay, Estancia Terrado (CCJM); 15 — ♂, Bolivia, Concepcion (CCJM); 16 — ♂, ex Dejean-Chaudoir, PLT (MNHN); 17 — ♀, Brazil, Barra do Carças (CMHP); 18 — ♀, Venezuela, Canaima NP (CPVP); 19 — maxillary palpus, Paraguay, Estancia Terrado (CCJM); 20 – 21 — antennal scape: 20 — ♂, Paraguay, Estancia Terrado (CCJM); 21 — ♂, Bolivia, Concepcion (CCJM); 22 — thoracic wing, ♀, ibid., CCJM. Bars = 1 mm." httpUri="https://zenodo.org/record/1044916/files/figure.png" pageId="10" pageNumber="119">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="A475EC57FFD5C81D27CEF939FD6B1FE3" blockId="10.[151,1437,447,2021]" pageId="10" pageNumber="119">
The aedeagus apex (
<figureCitation id="3CF1F0D2FFD5C81D26A6F939FD8A1ECB" box="[431,560,1743,1768]" captionStart="FIGURES 49 – 57" captionStartId="14.[151,264,1877,1899]" captionTargetBox="[158,1416,204,1812]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURES 49 – 57. Odontocheila microptera nom. nov. 49 – 51 — atrophic thoracic wings: 49 – 50 — ♀, Uruguay, Tacuarembo (CCJM); 51 — ♂, HT (SDEI); 52 – 54 — pronotum: 52 — ♂, HT (SDEI); 53 — ♂, Uruguay, Tacuarembo (CCJM); 54 — ♀, Argentina, Estancia San Noria (BMNH); 55 – 57 — male aedeagi: 55 — HT (SDEI); 56 – 57 — Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044924/files/figure.png" pageId="10" pageNumber="119">Figs 55–57</figureCitation>
) in 
<taxonomicName id="63CA97D4FFD5C81D2556F926FD451EC4" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[607,767,1744,1767]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD5C81D2556F926FD451EC4" box="[607,767,1744,1767]" italics="true" pageId="10" pageNumber="119">O. microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD5C81D240EF927FCC21ECB" box="[775,888,1745,1768]" pageId="10" pageNumber="119">nom. nov.</taxonomicNameLabel>
is a rather thick, dorsally directed hook, but the hook is on its ventral side rounded, thus more similar to the aedeagus apex of 
<taxonomicName id="63CA97D4FFD5C81D24F0F902FBCB1F2F" authorityName="Klug" authorityYear="1834" box="[1017,1137,1780,1804]" class="Insecta" family="Carabidae" genus="Cicindela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="rutilans">
<emphasis id="96BE3045FFD5C81D24F0F902FBCB1F2F" box="[1017,1137,1780,1804]" italics="true" pageId="10" pageNumber="119">O. rutilans</emphasis>
</taxonomicName>
. Nevertheless, as in others of the species-group related to 
<taxonomicName id="63CA97D4FFD5C81D26FAF8EEFDD71F13" authorityName="Klug" authorityYear="1834" box="[499,621,1816,1840]" class="Insecta" family="Carabidae" genus="Cicindela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="rutilans">
<emphasis id="96BE3045FFD5C81D26FAF8EEFDD71F13" box="[499,621,1816,1840]" italics="true" pageId="10" pageNumber="119">O. rutilans</emphasis>
</taxonomicName>
and 
<taxonomicName id="63CA97D4FFD5C81D25ADF8EEFC091F13" authority="(Klug, 1834)" authorityName="Klug" authorityYear="1834" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[676,947,1815,1840]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="fulgens">
<emphasis id="96BE3045FFD5C81D25ADF8EEFCA21F13" box="[676,792,1816,1840]" italics="true" pageId="10" pageNumber="119">O. fulgens</emphasis>
(Klug, 1834)
</taxonomicName>
, the shape in syntopic adults may vary, and in one male of 
<taxonomicName id="63CA97D4FFD5C81D2644F8CAFE551F77" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[333,495,1852,1876]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD5C81D2644F8CAFE551F77" box="[333,495,1852,1876]" italics="true" pageId="10" pageNumber="119">O. microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD5C81D26F0F8CBFDD61F77" box="[505,620,1853,1876]" pageId="10" pageNumber="119">nom. nov.</taxonomicNameLabel>
from 
<collectingRegion id="660E22B5FFD5C81D25BEF8CAFCFE1F76" box="[695,836,1852,1877]" country="Uruguay" name="Tacuarembo" pageId="10" pageNumber="119">Tacuarembo</collectingRegion>
, 
<collectingCountry id="DCDDACC7FFD5C81D245AF8CBFC0F1F76" box="[851,949,1853,1877]" name="Uruguay" pageId="10" pageNumber="119">Uruguay</collectingCountry>
the aedeagus apex (
<figureCitation id="3CF1F0D2FFD5C81D23ACF8CAFB401F76" box="[1189,1274,1852,1877]" captionStart="FIGURES 49 – 57" captionStartId="14.[151,264,1877,1899]" captionTargetBox="[158,1416,204,1812]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURES 49 – 57. Odontocheila microptera nom. nov. 49 – 51 — atrophic thoracic wings: 49 – 50 — ♀, Uruguay, Tacuarembo (CCJM); 51 — ♂, HT (SDEI); 52 – 54 — pronotum: 52 — ♂, HT (SDEI); 53 — ♂, Uruguay, Tacuarembo (CCJM); 54 — ♀, Argentina, Estancia San Noria (BMNH); 55 – 57 — male aedeagi: 55 — HT (SDEI); 56 – 57 — Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044924/files/figure.png" pageId="10" pageNumber="119">Fig. 56</figureCitation>
) is somewhat similar to that in some of the aedeagi in 
<emphasis id="96BE3045FFD5C81D255DF896FD4F1F5B" box="[596,757,1888,1912]" italics="true" pageId="10" pageNumber="119">
<taxonomicName id="63CA97D4FFD5C81D255DF896FD4B1F5B" box="[596,753,1888,1912]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nitidicollis">O. nitidicollis</taxonomicName>
.
</emphasis>
In turn, the aedeagus apex of the male of 
<taxonomicName id="63CA97D4FFD5C81D23CCF896FAE51F5B" box="[1221,1375,1888,1912]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD5C81D23CCF896FAE51F5B" box="[1221,1375,1888,1912]" italics="true" pageId="10" pageNumber="119">O. nitidicollis</emphasis>
</taxonomicName>
from Paraguayan Estancia Terrado (
<figureCitation id="3CF1F0D2FFD5C81D26FAF872FDF31FBF" box="[499,585,1924,1949]" captionStart="FIGURES 23 – 29" captionStartId="9.[151,264,1770,1792]" captionTargetBox="[166,1419,206,1695]" captionTargetId="figure@9.[151,1436,193,1748]" captionTargetPageId="9" captionText="FIGURES 23 – 29. Odontocheila nitidicollis Dejean. 23 – 24 — pronotum: 23 — ♂, LT (MNHN); 24 — ♂, ex Dejean-Chaudoir, PLT (MNHN); 25 – 29 — male aedeagi: 25 — LT (MNHN); 26 a—ex Dejean-Chaudoir, PLT (MNHN); 26 b—ditto, ventral view; 27 — Brazil, Barra do Carças (CMHP); 28 — Bolivia, Santa Cruz, Alta Vista (COSJ); 29 — showing internal sac, Bolivia, Reyes (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044918/files/figure.png" pageId="10" pageNumber="119">Fig. 28</figureCitation>
), as well as that from Bolivian Alta Vista (
<figureCitation id="3CF1F0D2FFD5C81D2332F872FB2A1FBF" box="[1083,1168,1924,1949]" captionStart="FIGURES 23 – 29" captionStartId="9.[151,264,1770,1792]" captionTargetBox="[166,1419,206,1695]" captionTargetId="figure@9.[151,1436,193,1748]" captionTargetPageId="9" captionText="FIGURES 23 – 29. Odontocheila nitidicollis Dejean. 23 – 24 — pronotum: 23 — ♂, LT (MNHN); 24 — ♂, ex Dejean-Chaudoir, PLT (MNHN); 25 – 29 — male aedeagi: 25 — LT (MNHN); 26 a—ex Dejean-Chaudoir, PLT (MNHN); 26 b—ditto, ventral view; 27 — Brazil, Barra do Carças (CMHP); 28 — Bolivia, Santa Cruz, Alta Vista (COSJ); 29 — showing internal sac, Bolivia, Reyes (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044918/files/figure.png" pageId="10" pageNumber="119">Fig. 29</figureCitation>
), is almost of the same shape as in most males of 
<taxonomicName id="63CA97D4FFD5C81D26B4F85FFDE01FE3" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[445,602,1961,1984]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD5C81D26B4F85FFDE01FE3" box="[445,602,1961,1984]" italics="true" pageId="10" pageNumber="119">O. microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD5C81D2568F85FFD6B1FE3" box="[609,721,1961,1984]" pageId="10" pageNumber="119">nom. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="ECD0BFDCFFD5C81C27CEF83AFC501E96" lastPageId="11" lastPageNumber="120" pageId="10" pageNumber="119" type="description">
<paragraph id="A475EC57FFD5C81C27CEF83AFC6B18F6" blockId="10.[151,1437,447,2021]" lastBlockId="11.[151,1437,151,2038]" lastPageId="11" lastPageNumber="120" pageId="10" pageNumber="119">
<emphasis id="96BE3045FFD5C81D27CEF83AFED71FC6" bold="true" box="[199,365,1996,2021]" pageId="10" pageNumber="119">Redescription</emphasis>
. Body (
<figureCitation id="3CF1F0D2FFD5C81D26C1F83AFDF31FC7" box="[456,585,1996,2021]" captionStart="FIGURES 30 – 36" captionStartId="12.[151,264,1877,1899]" captionTargetBox="[161,1426,205,1840]" captionTargetId="figure@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 30 – 36. Odontocheila microptera nom. nov. 30 – 32 — habitus: 30 — ♂, 9.5 mm, “ Brasilia ”, HT (SDEI); 31 — ♂, 11 mm, Uruguay, Tacuarembo (CCJM ex DBCN); 32 — ♀, 10.2 mm, Argentina, Estancia San Noria (BMNH); 33 – 36 — elytron: 33 — ♂, HT (SDEI); 34 — ♂, Uruguay, Tacuarembo (CCJM); 35 — ♀, ibid., (CCJM); 36 — ♀, Bahia (MFNB). Bar = 1 mm." httpUri="https://zenodo.org/record/1044920/files/figure.png" pageId="10" pageNumber="119">Figs 30–32</figureCitation>
) medium-sized (of a variable size independent of sex) 9.40–11.4 (HT 9.50) mm long, 2.90–3.80 (HT 3.20) mm wide, dark coppery or more vividly reddish-cupreous except for shiny reddishcupreous pronotum or also head, with faint green-blue lateral iridescence.
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFF29FBDB18D4" blockId="11.[151,1437,151,2038]" box="[199,1121,223,248]" pageId="11" pageNumber="120">
Head (
<figureCitation id="3CF1F0D2FFD4C81C2618FF29FE2B18DB" box="[273,401,223,248]" captionStart="FIGURES 10 – 22" captionStartId="8.[151,264,1814,1836]" captionTargetBox="[154,1424,204,1775]" captionTargetId="figure@8.[151,1436,193,1792]" captionTargetPageId="8" captionText="FIGURES 10 – 22. Odontocheila nitidicollis Dejean. 10 – 12 — head: 10 — ♂, LT (MNHN); 11 — ♂, Paraguay, Estancia Terrado (CCJM); 12 — ♀, Venezuela, Canaima NP (CPVP); 13 – 18 — labrum: 13 —— ♂, LT (MNHN); 14 — ♂, Paraguay, Estancia Terrado (CCJM); 15 — ♂, Bolivia, Concepcion (CCJM); 16 — ♂, ex Dejean-Chaudoir, PLT (MNHN); 17 — ♀, Brazil, Barra do Carças (CMHP); 18 — ♀, Venezuela, Canaima NP (CPVP); 19 — maxillary palpus, Paraguay, Estancia Terrado (CCJM); 20 – 21 — antennal scape: 20 — ♂, Paraguay, Estancia Terrado (CCJM); 21 — ♂, Bolivia, Concepcion (CCJM); 22 — thoracic wing, ♀, ibid., CCJM. Bars = 1 mm." httpUri="https://zenodo.org/record/1044916/files/figure.png" pageId="11" pageNumber="120">Figs 10–12</figureCitation>
) shaped and coloured as in 
<taxonomicName id="63CA97D4FFD4C81C25C1FF16FCD818DB" box="[712,866,224,248]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD4C81C25C1FF16FCD818DB" box="[712,866,224,248]" italics="true" pageId="11" pageNumber="120">O. nitidicollis</emphasis>
</taxonomicName>
, width 
<quantity id="633241B2FFD4C81C24BEFF29FBE718D4" box="[951,1117,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="3.075" metricValueMax="3.35" metricValueMin="2.8" pageId="11" pageNumber="120" unit="mm" value="3.075" valueMax="3.35" valueMin="2.8">2.80–3.35 mm</quantity>
.
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFEF3FBD71963" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
Frons, vertex genae and clypeus coloured and with surface sculptures as in 
<taxonomicName id="63CA97D4FFD4C81C232CFEF2FB7B193F" box="[1061,1217,260,284]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD4C81C232CFEF2FB7B193F" box="[1061,1217,260,284]" italics="true" pageId="11" pageNumber="120">O. nitidicollis</emphasis>
</taxonomicName>
, but vertex usually less convex in middle, sometimes almost flat or with shallow anteromedian impression.
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFEBAFE161A43" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
Labrum 4-setose, often strongly sexually dimorphic in coloration; male labrum (
<figureCitation id="3CF1F0D2FFD4C81C236CFEBAFB5D1947" box="[1125,1255,332,357]" captionStart="FIGURES 37 – 48" captionStartId="13.[151,264,1842,1864]" captionTargetBox="[161,1417,205,1803]" captionTargetId="figure@13.[151,1436,193,1821]" captionTargetPageId="13" captionText="FIGURES 37 – 48. Odontocheila microptera nom. nov. 37 – 40 — head: 37 — ♂, HT (SDEI); 38 — ♂, Uruguay, Tacuarembo (CCJM); 39 — ♂, ibid.; 40 — ♂, ditto, ventral view; 41 — right mandible, ♀, ibid.; 42 — maxillary palpus, ♀, ibid. (CCJM); 43 — head part, ♀, Bahia (MFNB); 44 – 48 — labrum: 44 — ♀, Bahia (MFNB); 45 — ♀, Uruguay, Tacuarembo (CCJM); 46 — ♀, Argentina, Estancia San Noria (BMNH); 47 — ♂, HT (SDEI); 48 — ♂, Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044922/files/figure.png" pageId="11" pageNumber="120">Figs 47–48</figureCitation>
) predominantly ivory-white (ochre-yellow in old specimens) except for narrow, black basolateral area, rather short, length 
<quantity id="633241B2FFD4C81C2254FE99FF46198F" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="11" pageNumber="120" unit="mm" value="0.675" valueMax="0.75" valueMin="0.6">0.60– 0.75 mm</quantity>
, width 
<quantity id="633241B2FFD4C81C2647FE62FE48198F" box="[334,498,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="11" pageNumber="120" unit="mm" value="1.25" valueMax="1.4" valueMin="1.1">1.10–1.40 mm</quantity>
, basolateral and anterolateral teeth subacute or acute, anterior margin 4-dentate with anterolateral and anterior teeth in same level giving the labrum nearly transverse shape, truncate or shallowly emarginate in middle, lacking median tooth; female labrum (
<figureCitation id="3CF1F0D2FFD4C81C244CFE2AFC7C19D6" box="[837,966,476,501]" captionStart="FIGURES 37 – 48" captionStartId="13.[151,264,1842,1864]" captionTargetBox="[161,1417,205,1803]" captionTargetId="figure@13.[151,1436,193,1821]" captionTargetPageId="13" captionText="FIGURES 37 – 48. Odontocheila microptera nom. nov. 37 – 40 — head: 37 — ♂, HT (SDEI); 38 — ♂, Uruguay, Tacuarembo (CCJM); 39 — ♂, ibid.; 40 — ♂, ditto, ventral view; 41 — right mandible, ♀, ibid.; 42 — maxillary palpus, ♀, ibid. (CCJM); 43 — head part, ♀, Bahia (MFNB); 44 – 48 — labrum: 44 — ♀, Bahia (MFNB); 45 — ♀, Uruguay, Tacuarembo (CCJM); 46 — ♀, Argentina, Estancia San Noria (BMNH); 47 — ♂, HT (SDEI); 48 — ♂, Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044922/files/figure.png" pageId="11" pageNumber="120">Figs 44–46</figureCitation>
) variably either entirely metallic black or with testaceous anteromedian or whole anterior area, but always with black basal area expanded to the middle, much longer, length 
<quantity id="633241B2FFD4C81C2689FDD2FD9F1A1F" box="[384,549,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="11" pageNumber="120" unit="mm" value="1.2" valueMax="1.25" valueMin="1.15">1.15–1.25 mm</quantity>
, width 
<quantity id="633241B2FFD4C81C2570FDD2FCA51A1F" box="[633,799,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="11" pageNumber="120" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.40–1.50 mm</quantity>
, its tridentate median lobe with protruding median tooth of rather variable length.
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFD9AFBFB1AEF" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
Mandibles (
<figureCitation id="3CF1F0D2FFD4C81C2642FD9AFE711AA7" box="[331,459,620,645]" captionStart="FIGURES 37 – 48" captionStartId="13.[151,264,1842,1864]" captionTargetBox="[161,1417,205,1803]" captionTargetId="figure@13.[151,1436,193,1821]" captionTargetPageId="13" captionText="FIGURES 37 – 48. Odontocheila microptera nom. nov. 37 – 40 — head: 37 — ♂, HT (SDEI); 38 — ♂, Uruguay, Tacuarembo (CCJM); 39 — ♂, ibid.; 40 — ♂, ditto, ventral view; 41 — right mandible, ♀, ibid.; 42 — maxillary palpus, ♀, ibid. (CCJM); 43 — head part, ♀, Bahia (MFNB); 44 – 48 — labrum: 44 — ♀, Bahia (MFNB); 45 — ♀, Uruguay, Tacuarembo (CCJM); 46 — ♀, Argentina, Estancia San Noria (BMNH); 47 — ♂, HT (SDEI); 48 — ♂, Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044922/files/figure.png" pageId="11" pageNumber="120">Figs 37–41</figureCitation>
) shaped and coloured as in 
<taxonomicName id="63CA97D4FFD4C81C25F7FD9AFC221AA7" box="[766,920,620,644]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD4C81C25F7FD9AFC221AA7" box="[766,920,620,644]" italics="true" pageId="11" pageNumber="120">O. nitidicollis</emphasis>
</taxonomicName>
, but often paler (faded to brownish-testaceous in old specimens, such as in HT), and with similar variability in shape of inner teeth, but the third tooth is more often and also in left mandible notably smaller than the second and fourth (
<figureCitation id="3CF1F0D2FFD4C81C24E9FD42FB891AEF" box="[992,1075,692,717]" captionStart="FIGURES 37 – 48" captionStartId="13.[151,264,1842,1864]" captionTargetBox="[161,1417,205,1803]" captionTargetId="figure@13.[151,1436,193,1821]" captionTargetPageId="13" captionText="FIGURES 37 – 48. Odontocheila microptera nom. nov. 37 – 40 — head: 37 — ♂, HT (SDEI); 38 — ♂, Uruguay, Tacuarembo (CCJM); 39 — ♂, ibid.; 40 — ♂, ditto, ventral view; 41 — right mandible, ♀, ibid.; 42 — maxillary palpus, ♀, ibid. (CCJM); 43 — head part, ♀, Bahia (MFNB); 44 – 48 — labrum: 44 — ♀, Bahia (MFNB); 45 — ♀, Uruguay, Tacuarembo (CCJM); 46 — ♀, Argentina, Estancia San Noria (BMNH); 47 — ♂, HT (SDEI); 48 — ♂, Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044922/files/figure.png" pageId="11" pageNumber="120">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFD21FDEB1B37" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
Palpi (
<figureCitation id="3CF1F0D2FFD4C81C261BFD21FE7B1AD3" box="[274,449,727,752]" captionStart="FIGURES 37 – 48" captionStartId="13.[151,264,1842,1864]" captionTargetBox="[161,1417,205,1803]" captionTargetId="figure@13.[151,1436,193,1821]" captionTargetPageId="13" captionText="FIGURES 37 – 48. Odontocheila microptera nom. nov. 37 – 40 — head: 37 — ♂, HT (SDEI); 38 — ♂, Uruguay, Tacuarembo (CCJM); 39 — ♂, ibid.; 40 — ♂, ditto, ventral view; 41 — right mandible, ♀, ibid.; 42 — maxillary palpus, ♀, ibid. (CCJM); 43 — head part, ♀, Bahia (MFNB); 44 – 48 — labrum: 44 — ♀, Bahia (MFNB); 45 — ♀, Uruguay, Tacuarembo (CCJM); 46 — ♀, Argentina, Estancia San Noria (BMNH); 47 — ♂, HT (SDEI); 48 — ♂, Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044922/files/figure.png" pageId="11" pageNumber="120">Figs 38–40, 42</figureCitation>
) as in 
<emphasis id="96BE3045FFD4C81C251BFD2EFD0F1AD3" box="[530,693,728,752]" italics="true" pageId="11" pageNumber="120">
<taxonomicName id="63CA97D4FFD4C81C251BFD2EFD0B1AD3" box="[530,689,728,752]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidicollis">O. nitidicollis</taxonomicName>
,
</emphasis>
but terminal palpomeres of maxillary in both sexes notably less distinctly dilated, width 
<quantity id="633241B2FFD4C81C26AEFD0AFDF71B37" box="[423,589,764,789]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="11" pageNumber="120" unit="mm" value="0.16" valueMax="0.17" valueMin="0.15">0.15–0.17 mm</quantity>
.
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFCD6FAD11B5C" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
Antennae (
<figureCitation id="3CF1F0D2FFD4C81C264BFCE9FE7E1B1B" box="[322,452,799,824]" captionStart="FIGURES 30 – 36" captionStartId="12.[151,264,1877,1899]" captionTargetBox="[161,1426,205,1840]" captionTargetId="figure@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 30 – 36. Odontocheila microptera nom. nov. 30 – 32 — habitus: 30 — ♂, 9.5 mm, “ Brasilia ”, HT (SDEI); 31 — ♂, 11 mm, Uruguay, Tacuarembo (CCJM ex DBCN); 32 — ♀, 10.2 mm, Argentina, Estancia San Noria (BMNH); 33 – 36 — elytron: 33 — ♂, HT (SDEI); 34 — ♂, Uruguay, Tacuarembo (CCJM); 35 — ♀, ibid., (CCJM); 36 — ♀, Bahia (MFNB). Bar = 1 mm." httpUri="https://zenodo.org/record/1044920/files/figure.png" pageId="11" pageNumber="120">Figs 30–32</figureCitation>
, 
<figureCitation id="3CF1F0D2FFD4C81C26D9FCD6FDAD1B1B" box="[464,535,800,824]" captionStart="FIGURES 37 – 48" captionStartId="13.[151,264,1842,1864]" captionTargetBox="[161,1417,205,1803]" captionTargetId="figure@13.[151,1436,193,1821]" captionTargetPageId="13" captionText="FIGURES 37 – 48. Odontocheila microptera nom. nov. 37 – 40 — head: 37 — ♂, HT (SDEI); 38 — ♂, Uruguay, Tacuarembo (CCJM); 39 — ♂, ibid.; 40 — ♂, ditto, ventral view; 41 — right mandible, ♀, ibid.; 42 — maxillary palpus, ♀, ibid. (CCJM); 43 — head part, ♀, Bahia (MFNB); 44 – 48 — labrum: 44 — ♀, Bahia (MFNB); 45 — ♀, Uruguay, Tacuarembo (CCJM); 46 — ♀, Argentina, Estancia San Noria (BMNH); 47 — ♂, HT (SDEI); 48 — ♂, Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044922/files/figure.png" pageId="11" pageNumber="120">37–39</figureCitation>
) shaped and coloured as in 
<emphasis id="96BE3045FFD4C81C245AFCD6FC4F1B1B" box="[851,1013,800,824]" italics="true" pageId="11" pageNumber="120">
<taxonomicName id="63CA97D4FFD4C81C245AFCD6FC4A1B1B" box="[851,1008,800,824]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidicollis">O. nitidicollis</taxonomicName>
,
</emphasis>
but notably shorter, in male reaching only elytral third, in female only elytral shoulders, coloration in old specimens usually faded (antennomeres 
<quantity id="633241B2FFD4C81C225CFCB2FA261B7E" box="[1365,1436,836,861]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="11" pageNumber="120" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
HT faded to pale mahogany with only apices metallic-black); scape moderately dilated, width up to 
<quantity id="633241B2FFD4C81C23F1FC9EFADC1B5C" box="[1272,1382,872,896]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="120" unit="mm" value="0.28">0. 28 mm</quantity>
.
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFC7AFD931C33" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
Thorax. Pronotum (
<figureCitation id="3CF1F0D2FFD4C81C26A1FC7AFD961B86" box="[424,556,908,933]" captionStart="FIGURES 49 – 57" captionStartId="14.[151,264,1877,1899]" captionTargetBox="[158,1416,204,1812]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURES 49 – 57. Odontocheila microptera nom. nov. 49 – 51 — atrophic thoracic wings: 49 – 50 — ♀, Uruguay, Tacuarembo (CCJM); 51 — ♂, HT (SDEI); 52 – 54 — pronotum: 52 — ♂, HT (SDEI); 53 — ♂, Uruguay, Tacuarembo (CCJM); 54 — ♀, Argentina, Estancia San Noria (BMNH); 55 – 57 — male aedeagi: 55 — HT (SDEI); 56 – 57 — Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044924/files/figure.png" pageId="11" pageNumber="120">Figs 52–54</figureCitation>
) coloured and with surface sculpture as in 
<taxonomicName id="63CA97D4FFD4C81C232AFC7AFB7A1B87" box="[1059,1216,908,932]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD4C81C232AFC7AFB7A1B87" box="[1059,1216,908,932]" italics="true" pageId="11" pageNumber="120">O. nitidicollis</emphasis>
</taxonomicName>
, but the transverse rugae on pronotal disc are sometimes, particularly in male (also in HT) denser and more wavy; in male slightly longer than wide, length 2.00– 
<quantity id="633241B2FFD4C81C26FCFC22FDE71BCF" box="[501,605,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="11" pageNumber="120" unit="mm" value="2.3">2.30 mm</quantity>
, width 
<quantity id="633241B2FFD4C81C25BAFC22FCEF1BCF" box="[691,853,980,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.025" metricValueMax="2.15" metricValueMin="1.9" pageId="11" pageNumber="120" unit="mm" value="2.025" valueMax="2.15" valueMin="1.9">1.90-2.15 mm</quantity>
, in female as long as wide or slightly wider, 
<quantity id="633241B2FFD4C81C2254FC22FF411C2C" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="11" pageNumber="120" unit="mm" value="2.25" valueMax="2.3" valueMin="2.2">2.20– 2.30 mm</quantity>
long, 
<quantity id="633241B2FFD4C81C2648FC0EFE5E1C2C" box="[321,484,1016,1040]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="11" pageNumber="120" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.20–2.40 mm</quantity>
wide.
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFBEAFC7E1C83" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
Elytra (
<figureCitation id="3CF1F0D2FFD4C81C2615FBEAFE271C16" box="[284,413,1052,1077]" captionStart="FIGURES 30 – 36" captionStartId="12.[151,264,1877,1899]" captionTargetBox="[161,1426,205,1840]" captionTargetId="figure@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 30 – 36. Odontocheila microptera nom. nov. 30 – 32 — habitus: 30 — ♂, 9.5 mm, “ Brasilia ”, HT (SDEI); 31 — ♂, 11 mm, Uruguay, Tacuarembo (CCJM ex DBCN); 32 — ♀, 10.2 mm, Argentina, Estancia San Noria (BMNH); 33 – 36 — elytron: 33 — ♂, HT (SDEI); 34 — ♂, Uruguay, Tacuarembo (CCJM); 35 — ♀, ibid., (CCJM); 36 — ♀, Bahia (MFNB). Bar = 1 mm." httpUri="https://zenodo.org/record/1044920/files/figure.png" pageId="11" pageNumber="120">Figs 33–36</figureCitation>
) mutually notably ovoid, their outer margin markedly arched, length 5.70–7.00 mm, with widely arcuate humeri; anteapical angles widely arched towards apices which are in both sexes mostly almost acute with only small emargination towards distinct sutural spine; elytral surface punctate, but coloration slightly less variable and generally darker; whitish elytral maculation entirely absent.
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFB5AFDA21CE6" blockId="11.[151,1437,151,2038]" box="[199,536,1196,1221]" pageId="11" pageNumber="120">
Abdomen as in 
<taxonomicName id="63CA97D4FFD4C81C267EFB5AFDA81CE7" box="[375,530,1196,1220]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD4C81C267EFB5AFDA81CE7" box="[375,530,1196,1220]" italics="true" pageId="11" pageNumber="120">O. nitidicollis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFB27FBD71CCB" blockId="11.[151,1437,151,2038]" box="[199,1133,1231,1257]" pageId="11" pageNumber="120">
Legs as in 
<taxonomicName id="63CA97D4FFD4C81C2636FB26FE601CCB" box="[319,474,1232,1256]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD4C81C2636FB26FE601CCB" box="[319,474,1232,1256]" italics="true" pageId="11" pageNumber="120">O. nitidicollis</emphasis>
</taxonomicName>
, the black coloration in old specimens (HT) usually faded.
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFB02FDED1D54" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
Aedeagus (
<figureCitation id="3CF1F0D2FFD4C81C2641FB02FE761D2E" box="[328,460,1268,1293]" captionStart="FIGURES 49 – 57" captionStartId="14.[151,264,1877,1899]" captionTargetBox="[158,1416,204,1812]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURES 49 – 57. Odontocheila microptera nom. nov. 49 – 51 — atrophic thoracic wings: 49 – 50 — ♀, Uruguay, Tacuarembo (CCJM); 51 — ♂, HT (SDEI); 52 – 54 — pronotum: 52 — ♂, HT (SDEI); 53 — ♂, Uruguay, Tacuarembo (CCJM); 54 — ♀, Argentina, Estancia San Noria (BMNH); 55 – 57 — male aedeagi: 55 — HT (SDEI); 56 – 57 — Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044924/files/figure.png" pageId="11" pageNumber="120">Figs 55–57</figureCitation>
) moderately voluminous in middle, 
<quantity id="633241B2FFD4C81C2473FB02FB9B1D2F" box="[890,1057,1268,1293]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="11" pageNumber="120" unit="mm" value="3.3" valueMax="3.5" valueMin="3.1">3.10–3.50 mm</quantity>
long, 
<quantity id="633241B2FFD4C81C2379FB02FAAD1D2F" box="[1136,1303,1268,1293]" metricMagnitude="-4" metricUnit="m" metricValue="7.749999999999999" metricValueMax="8.5" metricValueMin="7.0" pageId="11" pageNumber="120" unit="mm" value="0.7749999999999999" valueMax="0.85" valueMin="0.7">0.70–0.85 mm</quantity>
wide, with simply hooked apex which is ventrally rounded, rarely subquadrate (
<figureCitation id="3CF1F0D2FFD4C81C24A9FAE1FC4C1D13" box="[928,1014,1303,1328]" captionStart="FIGURES 49 – 57" captionStartId="14.[151,264,1877,1899]" captionTargetBox="[158,1416,204,1812]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURES 49 – 57. Odontocheila microptera nom. nov. 49 – 51 — atrophic thoracic wings: 49 – 50 — ♀, Uruguay, Tacuarembo (CCJM); 51 — ♂, HT (SDEI); 52 – 54 — pronotum: 52 — ♂, HT (SDEI); 53 — ♂, Uruguay, Tacuarembo (CCJM); 54 — ♀, Argentina, Estancia San Noria (BMNH); 55 – 57 — male aedeagi: 55 — HT (SDEI); 56 – 57 — Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044924/files/figure.png" pageId="11" pageNumber="120">Fig. 56</figureCitation>
); internal sac as in 
<taxonomicName id="63CA97D4FFD4C81C23D8FAEEFAD61D0C" box="[1233,1388,1303,1327]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD4C81C23D8FAEEFAD61D0C" box="[1233,1388,1303,1327]" italics="true" pageId="11" pageNumber="120">O. nitidicollis</emphasis>
</taxonomicName>
and most other species of the species-group related to 
<taxonomicName id="63CA97D4FFD4C81C25C6FACDFCF31D70" authorityName="Klug" authorityYear="1834" box="[719,841,1339,1363]" class="Insecta" family="Carabidae" genus="Cicindela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="rutilans">
<emphasis id="96BE3045FFD4C81C25C6FACDFCF31D70" box="[719,841,1339,1363]" italics="true" pageId="11" pageNumber="120">O. rutilans</emphasis>
</taxonomicName>
with large, normally shaped voluminous base of the flagellum (partly visible in 
<figureCitation id="3CF1F0D2FFD4C81C26C0FAA8FDF11D54" box="[457,587,1374,1399]" captionStart="FIGURES 49 – 57" captionStartId="14.[151,264,1877,1899]" captionTargetBox="[158,1416,204,1812]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURES 49 – 57. Odontocheila microptera nom. nov. 49 – 51 — atrophic thoracic wings: 49 – 50 — ♀, Uruguay, Tacuarembo (CCJM); 51 — ♂, HT (SDEI); 52 – 54 — pronotum: 52 — ♂, HT (SDEI); 53 — ♂, Uruguay, Tacuarembo (CCJM); 54 — ♀, Argentina, Estancia San Noria (BMNH); 55 – 57 — male aedeagi: 55 — HT (SDEI); 56 – 57 — Uruguay, Tacuarembo (CCJM). Bars = 1 mm." httpUri="https://zenodo.org/record/1044924/files/figure.png" pageId="11" pageNumber="120">Figs 55, 56</figureCitation>
).
</paragraph>
<paragraph id="A475EC57FFD4C81C27CEFA77FC501E96" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
<emphasis id="96BE3045FFD4C81C27CEFA77FEFF1DB9" bold="true" box="[199,325,1409,1434]" pageId="11" pageNumber="120">Variability</emphasis>
. Apart from that emphasized in the “Differential diagnosis” and the “Redescription” above, the most important is the variable coloration of the female labrum which is either entirely black, or distinctly bicoloured, thus similar to the bicoloured labrum in most females of 
<taxonomicName id="63CA97D4FFD4C81C24A9FA3FFBAE1DC2" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[928,1044,1481,1505]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="fulgens">
<emphasis id="96BE3045FFD4C81C24A9FA3FFBAE1DC2" box="[928,1044,1481,1505]" italics="true" pageId="11" pageNumber="120">O. fulgens</emphasis>
</taxonomicName>
. Although 
<taxonomicName id="63CA97D4FFD4C81C239DFA3FFAB21DC2" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[1172,1288,1481,1505]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="fulgens">
<emphasis id="96BE3045FFD4C81C239DFA3FFAB21DC2" box="[1172,1288,1481,1505]" italics="true" pageId="11" pageNumber="120">O. fulgens</emphasis>
</taxonomicName>
immediately differs by its whitish elytral maculation, it is important to rectify here the error by 
<bibRefCitation id="C05B91A6FFD4C81C2335FA1AFB531E26" author="Rivalier" box="[1084,1257,1516,1541]" pageId="11" pageNumber="120" refString="Rivalier, E. (1969) Demembrement du genre Odontochila (Col. Cicindelidae) et revision des principales especes. Annales de la Societe entomologique de France, Nouvelle Serie, 5 (1), 195 - 237." type="journal article" year="1969">Rivalier (1969)</bibRefCitation>
who stated that one of the distinguishing characters of 
<taxonomicName id="63CA97D4FFD4C81C2558F9E6FD7C1E0B" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[593,710,1552,1576]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="fulgens">
<emphasis id="96BE3045FFD4C81C2558F9E6FD7C1E0B" box="[593,710,1552,1576]" italics="true" pageId="11" pageNumber="120">O. fulgens</emphasis>
</taxonomicName>
is metallic black female labrum, and such misleading error was adopted by 
<bibRefCitation id="C05B91A6FFD4C81C2617F9C2FDAB1E68" box="[286,529,1587,1612]" pageId="11" pageNumber="120" refString="Pearson, D. L., Guerra, J. F. &amp; Brzoska, D. W. (1999) The Tiger beetles of Bolivia: their Identification, Distribution and Natural History (Coleoptera: Cicindelidae). Contributions on Entomology, International, 3 (4), 379 - 524." type="journal article">Pearson et al. (1999)</bibRefCitation>
. In fact, the labrum of the female 
<typeStatus id="7B7152F5FFD4C81C24A9F9C5FBB81E6F" box="[928,1026,1587,1612]" pageId="11" pageNumber="120" type="holotype">holotype</typeStatus>
(MFNB) of 
<taxonomicName id="63CA97D4FFD4C81C2391F9C5FAB41E68" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[1176,1294,1587,1611]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="fulgens">
<emphasis id="96BE3045FFD4C81C2391F9C5FAB41E68" box="[1176,1294,1587,1611]" italics="true" pageId="11" pageNumber="120">O. fulgens</emphasis>
</taxonomicName>
is distinctly bicolored, as also are the labra of majority of the other examined females of 
<taxonomicName id="63CA97D4FFD4C81C233BF9A1FB101E4D" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[1074,1194,1622,1646]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="fulgens">
<emphasis id="96BE3045FFD4C81C233BF9A1FB101E4D" box="[1074,1194,1622,1646]" italics="true" pageId="11" pageNumber="120">O. fulgens</emphasis>
</taxonomicName>
from 
<collectingCountry id="DCDDACC7FFD4C81C23F1F9A0FADD1E4C" box="[1272,1383,1622,1647]" name="Argentina" pageId="11" pageNumber="120">Argentina</collectingCountry>
and 
<collectingRegion id="660E22B5FFD4C81C279EF98DFEBA1EB0" box="[151,256,1659,1683]" country="Paraguay" name="Asuncion" pageId="11" pageNumber="120">
<collectingCountry id="DCDDACC7FFD4C81C279EF98DFEBA1EB0" box="[151,256,1659,1683]" name="Paraguay" pageId="11" pageNumber="120">Paraguay</collectingCountry>
</collectingRegion>
, except for females from Rio Salado, 
<collectingCountry id="DCDDACC7FFD4C81C25C7F98CFC871EB0" box="[718,829,1658,1683]" name="Argentina" pageId="11" pageNumber="120">Argentina</collectingCountry>
(IRSNB), where the female labrum is prevailingly metallic black with only testaceous median area which is often very narrow.
</paragraph>
</subSubSection>
<subSubSection id="ECD0BFDCFFD4C81C27CEF937FF591EDE" pageId="11" pageNumber="120" type="etymology">
<paragraph id="A475EC57FFD4C81C27CEF937FF591EDE" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
<emphasis id="96BE3045FFD4C81C27CEF937FEFC1EF9" bold="true" box="[199,326,1729,1754]" pageId="11" pageNumber="120">Etymology</emphasis>
. The species name 
<taxonomicName id="63CA97D4FFD4C81C253FF934FD0B1EFA" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[566,689,1730,1753]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="microptera">
<emphasis id="96BE3045FFD4C81C253FF934FD0B1EFA" box="[566,689,1730,1753]" italics="true" pageId="11" pageNumber="120">microptera</emphasis>
</taxonomicName>
= having small or rudimentary wings, is derived from ancient Greek.
</paragraph>
</subSubSection>
<subSubSection id="ECD0BFDCFFD4C81827CEF8F1FB6C1EC4" lastPageId="15" lastPageNumber="124" pageId="11" pageNumber="120" type="biology_ecology">
<paragraph id="A475EC57FFD4C81C27CEF8F1FEB41FD5" blockId="11.[151,1437,151,2038]" pageId="11" pageNumber="120">
<emphasis id="96BE3045FFD4C81C27CEF8F1FE4B1F03" bold="true" box="[199,497,1799,1824]" pageId="11" pageNumber="120">Biology and distribution</emphasis>
. This species was hitherto known (under the name 
<taxonomicName id="63CA97D4FFD4C81C235DF8FEFB5D1F03" box="[1108,1255,1800,1824]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="euryoides">
<emphasis id="96BE3045FFD4C81C235DF8FEFB5D1F03" box="[1108,1255,1800,1824]" italics="true" pageId="11" pageNumber="120">O. euryoides</emphasis>
</taxonomicName>
) only from the holotype labelled “Brasilia”. 
<bibRefCitation id="C05B91A6FFD4C81C26EEF8DAFDD41F67" author="Horn" box="[487,622,1836,1860]" pageId="11" pageNumber="120" refString="Horn, W. (1922) Studien uber neue und alte Cicindeliden (Col.), (Neubeschreibungen, Synonymie, Faunistik). Zoologische Mededeelingen, 7, 1 - 112." type="journal article" year="1922">Horn (1922</bibRefCitation>
, 
<bibRefCitation id="C05B91A6FFD4C81C2575F8DAFD0D1F67" author="Horn" box="[636,695,1836,1860]" pageId="11" pageNumber="120" refString="Horn, W. (1910) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P., Genera Insectorum, Fascicule 82 B, pp. 105 - 208, pls. 6 - 15." type="journal article" year="1910">1910</bibRefCitation>
, 
<bibRefCitation id="C05B91A6FFD4C81C25CCF8DDFD441F67" author="Horn" box="[709,766,1835,1860]" pageId="11" pageNumber="120" refString="Horn, W. (1926) Carabidae, Cicindelinae (Pars 86). In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Junk, Berlin, 345 pp." type="book" year="1926">1926</bibRefCitation>
) mentioned only Brazil as its occurrence. Only two other historical specimens were identified in collections within this revision of the genus by the first author: one of them the female (MFNB) from Bahia, Brazil, the other the female (BMNH) from the Estancia San Noria, Santa Fe, Argentina, both cited above. Although the female (BMNH) from Argentina was previously labelled by Horn as 
<taxonomicName id="63CA97D4FFD4C81C2288F860FEB51FF2" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD4C81C2288F860FEB51FF2" italics="true" pageId="11" pageNumber="120">O. nitidicollis</emphasis>
</taxonomicName>
, he later (
<bibRefCitation id="C05B91A6FFD4C81C2683F84CFDB31FF1" author="Horn" box="[394,521,1978,2002]" pageId="11" pageNumber="120" refString="Horn, W. (1928) Sur quelques especes interessantes du genre Odontochila appartenantes a la faune de l'Argentine. Revista de la Sociedad Entomologica Argentina, 7, 75 - 76." type="journal article" year="1928">Horn 1928</bibRefCitation>
) mentioned a possible occurrence of this species (under 
<taxonomicName id="63CA97D4FFD4C81C23AEF84CFA801FF2" box="[1191,1338,1977,2001]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="euryoides">
<emphasis id="96BE3045FFD4C81C23AEF84CFA801FF2" box="[1191,1338,1977,2001]" italics="true" pageId="11" pageNumber="120">O. euryoides</emphasis>
</taxonomicName>
) also in Argentina.
</paragraph>
<caption id="F0B5BCDFFFD3C81B279EF8A3FADD1F8B" httpUri="https://zenodo.org/record/1044920/files/figure.png" pageId="12" pageNumber="121" targetBox="[161,1426,205,1840]" targetPageId="12">
<paragraph id="A475EC57FFD3C81B279EF8A3FADD1F8B" blockId="12.[151,1436,1877,1960]" pageId="12" pageNumber="121">
<emphasis id="96BE3045FFD3C81B279EF8A3FEF51F48" bold="true" box="[151,335,1877,1899]" pageId="12" pageNumber="121">FIGURES 30–36</emphasis>
. 
<taxonomicName id="63CA97D4FFD3C81B2654F8A3FDDA1F48" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[349,608,1877,1899]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD3C81B2654F8A3FDDA1F48" box="[349,608,1877,1899]" italics="true" pageId="12" pageNumber="121">Odontocheila microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD3C81B256EF8A0FD751F48" box="[615,719,1878,1899]" pageId="12" pageNumber="121">nom. nov.</taxonomicNameLabel>
30–32—habitus: 30—♂, 9.5 mm, “Brasilia”, HT (SDEI); 31—♂, 11 mm, Uruguay, Tacuarembo (CCJM ex DBCN); 32—♀, 10.2 mm, Argentina, Estancia San Noria (BMNH); 33–36—elytron: 33—♂, HT (SDEI); 34—♂, Uruguay, Tacuarembo (CCJM); 35—♀, ibid., (CCJM); 36—♀, Bahia (MFNB). Bar = 1 mm.
</paragraph>
</caption>
<caption id="F0B5BCDFFFD2C81A279EF8C4FA951F87" httpUri="https://zenodo.org/record/1044922/files/figure.png" pageId="13" pageNumber="122" targetBox="[161,1417,205,1803]" targetPageId="13">
<paragraph id="A475EC57FFD2C81A279EF8C4FA951F87" blockId="13.[151,1436,1842,1956]" pageId="13" pageNumber="122">
<emphasis id="96BE3045FFD2C81A279EF8C4FEE81F6B" bold="true" box="[151,338,1842,1864]" pageId="13" pageNumber="122">FIGURES 37–48</emphasis>
. 
<taxonomicName id="63CA97D4FFD2C81A266AF8C4FDD21F6B" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[355,616,1842,1864]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="122" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD2C81A266AF8C4FDD21F6B" box="[355,616,1842,1864]" italics="true" pageId="13" pageNumber="122">Odontocheila microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD2C81A257AF8C5FD661F6B" box="[627,732,1843,1864]" pageId="13" pageNumber="122">nom. nov.</taxonomicNameLabel>
37–40—head: 37—♂, HT (SDEI); 38—♂, Uruguay, Tacuarembo (CCJM); 39—♂, ibid.; 40—♂, ditto, ventral view; 41—right mandible, ♀, ibid.; 42—maxillary palpus, ♀, ibid. (CCJM); 43— head part, ♀, Bahia (MFNB); 44–48—labrum: 44—♀, Bahia (MFNB); 45—♀, Uruguay, Tacuarembo (CCJM); 46—♀, Argentina, Estancia San Noria (BMNH); 47—♂, HT (SDEI); 48—♂, Uruguay, Tacuarembo (CCJM). Bars = 1 mm.
</paragraph>
</caption>
<caption id="F0B5BCDFFFD1C819279EF8A3FE981FE4" httpUri="https://zenodo.org/record/1044924/files/figure.png" pageId="14" pageNumber="123" targetBox="[158,1416,204,1812]" targetPageId="14">
<paragraph id="A475EC57FFD1C819279EF8A3FE981FE4" blockId="14.[151,1436,1877,1991]" pageId="14" pageNumber="123">
<emphasis id="96BE3045FFD1C819279EF8A3FEEA1F48" bold="true" box="[151,336,1877,1899]" pageId="14" pageNumber="123">FIGURES 49–57</emphasis>
. 
<taxonomicName id="63CA97D4FFD1C8192669F8A3FDDE1F48" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[352,612,1877,1899]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="123" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD1C8192669F8A3FDDE1F48" box="[352,612,1877,1899]" italics="true" pageId="14" pageNumber="123">Odontocheila microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD1C8192564F8A0FD6C1F48" box="[621,726,1878,1899]" pageId="14" pageNumber="123">nom. nov.</taxonomicNameLabel>
49–51—atrophic thoracic wings: 49–50—♀, Uruguay, Tacuarembo (CCJM); 51—♂, HT (SDEI); 52–54—pronotum: 52—♂, HT (SDEI); 53—♂, Uruguay, Tacuarembo (CCJM); 54—♀, Argentina, Estancia San Noria (BMNH); 55–57—male aedeagi: 55—HT (SDEI); 56–57—Uruguay, Tacuarembo (CCJM). Bars = 1 mm.
</paragraph>
</caption>
<caption id="F0B5BCDFFFD0C818279EFB62FB9A1C89" box="[151,1056,1172,1195]" httpUri="https://zenodo.org/record/1044926/files/figure.png" pageId="15" pageNumber="124" targetBox="[155,1432,193,1151]" targetPageId="15">
<paragraph id="A475EC57FFD0C818279EFB62FB9A1C89" blockId="15.[151,1056,1172,1195]" box="[151,1056,1172,1195]" pageId="15" pageNumber="124">
<emphasis id="96BE3045FFD0C818279EFB62FEA51C89" bold="true" box="[151,287,1172,1194]" pageId="15" pageNumber="124">FIGURE 58.</emphasis>
One of the habitats of 
<taxonomicName id="63CA97D4FFD0C818250EFB63FD2C1C88" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[519,662,1173,1195]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD0C818250EFB63FD2C1C88" box="[519,662,1173,1195]" italics="true" pageId="15" pageNumber="124">O. microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD0C8182595FB63FCBB1C89" box="[668,769,1173,1194]" pageId="15" pageNumber="124">nom. nov.</taxonomicNameLabel>
near Tacuarembo, Uruguay.
</paragraph>
</caption>
<paragraph id="A475EC57FFD0C81827CEFB21FD311DA3" blockId="15.[151,1437,1239,2020]" pageId="15" pageNumber="124">
Therefore, the most surprising is that this flightless species was recently discovered by the third author and his collecting companion Johnnie Chong, in north-central 
<collectingCountry id="DCDDACC7FFD0C818243BFB0AFC2F1D37" box="[818,917,1276,1300]" name="Uruguay" pageId="15" pageNumber="124">Uruguay</collectingCountry>
, in the three adjoining localities north of 
<collectingRegion id="660E22B5FFD0C818279EFAE9FE9B1D1B" box="[151,289,1311,1336]" country="Uruguay" name="Tacuarembo" pageId="15" pageNumber="124">Tacuarembo</collectingRegion>
(cited above), in a good number of 
<specimenCount id="B2CC27DEFFD0C81825CFFAE9FC971D1B" box="[710,813,1311,1336]" pageId="15" pageNumber="124" type="adult">55 adults</specimenCount>
of both sexes. Nevertheless, it remains a great enigma how this flightless species could penetrate from the Atlantic Rainforest of north- and southeastern 
<collectingCountry id="DCDDACC7FFD0C8182205FAB5FAEA1D7F" box="[1292,1360,1347,1372]" name="Brazil" pageId="15" pageNumber="124">Brazil</collectingCountry>
to the very distant country, as well as to 
<collectingCountry id="DCDDACC7FFD0C818251DFA91FD3D1DA3" box="[532,647,1383,1408]" name="Argentina" pageId="15" pageNumber="124">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="A475EC57FFD0C81827CEFA7DFB6C1EC4" blockId="15.[151,1437,1239,2020]" pageId="15" pageNumber="124">
The flightless adults run and hide in the vegetation and due to their atrophic thoracic wings, this species was obviously forced out of the vegetative cover. It is one of the few 
<emphasis id="96BE3045FFD0C8182476FA59FBB01DE4" box="[895,1034,1455,1479]" italics="true" pageId="15" pageNumber="124">Odotocheila</emphasis>
that can survive in 
<collectingCountry id="DCDDACC7FFD0C81823E4FA46FAF51DEB" box="[1261,1359,1456,1480]" name="Uruguay" pageId="15" pageNumber="124">Uruguay</collectingCountry>
where probably most of the native forest has been removed and replaced with pine and 
<taxonomicName id="63CA97D4FFD0C8182321FA25FB1E1DC8" box="[1064,1188,1491,1515]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="124" phylum="Tracheophyta" rank="genus">
<emphasis id="96BE3045FFD0C8182321FA25FB1E1DC8" box="[1064,1188,1491,1515]" italics="true" pageId="15" pageNumber="124">Eucalyptus</emphasis>
</taxonomicName>
, as also in the area of the locality of 
<taxonomicName id="63CA97D4FFD0C8182630FA0EFE6D1E2C" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[313,471,1528,1551]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="microptera">
<emphasis id="96BE3045FFD0C8182630FA0EFE6D1E2C" box="[313,471,1528,1551]" italics="true" pageId="15" pageNumber="124">O. microptera</emphasis>
</taxonomicName>
. The biotope is partly similar to that of 
<taxonomicName id="63CA97D4FFD0C818249FFA0EFB8A1E2C" box="[918,1072,1527,1551]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="96BE3045FFD0C818249FFA0EFB8A1E2C" box="[918,1072,1527,1551]" italics="true" pageId="15" pageNumber="124">O. nitidicollis</emphasis>
</taxonomicName>
, which is, however, a very good flyer, and therefore may occupy somewhat more various biotopes. Unlike most species of 
<taxonomicName id="63CA97D4FFD0C81823BCF9EDFAF71E10" authorityName="Laporte de Castelnau" authorityYear="1834" box="[1205,1357,1563,1587]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="96BE3045FFD0C81823BCF9EDFAF71E10" box="[1205,1357,1563,1587]" italics="true" pageId="15" pageNumber="124">Odontocheila</emphasis>
</taxonomicName>
which usually occur along forest trails and more shaded areas, these two species inhabit more open and grassy areas, but 
<taxonomicName id="63CA97D4FFD0C818279EF992FE8F1E58" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[151,309,1636,1659]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD0C818279EF992FE8F1E58" box="[151,309,1636,1659]" italics="true" pageId="15" pageNumber="124">O. microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD0C8182635F992FE161E58" box="[316,428,1636,1659]" pageId="15" pageNumber="124">nom. nov.</taxonomicNameLabel>
was mostly found in heavier grassy places. At one of the three Uruguayan localities with pine trees (
<figureCitation id="3CF1F0D2FFD0C818261DF971FED21E83" box="[276,360,1671,1696]" captionStart="FIGURE 58" captionStartId="15.[151,250,1172,1194]" captionTargetBox="[155,1432,193,1151]" captionTargetId="figure@15.[155,1432,193,1151]" captionTargetPageId="15" captionText="FIGURE 58. One of the habitats of O. microptera nom. nov. near Tacuarembo, Uruguay." httpUri="https://zenodo.org/record/1044926/files/figure.png" pageId="15" pageNumber="124">Fig. 58</figureCitation>
), the adults were found among grass, but not under the trees; in the other two places they were found in heavy grassy areas along the shoulders of highways where no trees were present. It also may be significant that no other tiger beetle species was found in the habitats together with 
<taxonomicName id="63CA97D4FFD0C81824CBF926FBDA1EC4" authorityName="Moravec &amp; Huber &amp; Brzoska" authorityYear="2017" box="[962,1120,1744,1767]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="microptera" status="nom. nov.">
<emphasis id="96BE3045FFD0C81824CBF926FBDA1EC4" box="[962,1120,1744,1767]" italics="true" pageId="15" pageNumber="124">O. microptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8D8D8D3EFFD0C818236FF926FB6C1EC4" box="[1126,1238,1744,1767]" pageId="15" pageNumber="124">nom. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="ECD0BFDCFFD0C80727CEF905FF62198E" lastPageId="16" lastPageNumber="125" pageId="15" pageNumber="124" type="discussion">
<paragraph id="A475EC57FFD0C81827CEF905FD671FC7" blockId="15.[151,1437,1239,2020]" pageId="15" pageNumber="124">
<emphasis id="96BE3045FFD0C81827CEF905FE8E1F2F" bold="true" box="[199,308,1779,1804]" pageId="15" pageNumber="124">Remarks</emphasis>
. 
<bibRefCitation id="C05B91A6FFD0C818264DF902FE681F2F" author="Horn" box="[324,466,1780,1804]" pageId="15" pageNumber="124" refString="Horn, W. (1922) Studien uber neue und alte Cicindeliden (Col.), (Neubeschreibungen, Synonymie, Faunistik). Zoologische Mededeelingen, 7, 1 - 112." type="journal article" year="1922">Horn (1922)</bibRefCitation>
described 
<taxonomicName id="63CA97D4FFD0C8182558F905FCDA1F28" box="[593,864,1779,1803]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="euryoides">
<emphasis id="96BE3045FFD0C8182558F905FCDA1F28" box="[593,864,1779,1803]" italics="true" pageId="15" pageNumber="124">Odontocheila euryoides</emphasis>
</taxonomicName>
despite the existence of 
<taxonomicName id="63CA97D4FFD0C8182388F905FEBE1F0C" authorityName="W. Horn" authorityYear="1906" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="subSpecies" species="bennigseni" subSpecies="euryoides">
<emphasis id="96BE3045FFD0C8182388F905FEBE1F0C" italics="true" pageId="15" pageNumber="124">Odontocheila bennigseni euryoides</emphasis>
</taxonomicName>
described previously by himself (
<bibRefCitation id="C05B91A6FFD0C8182589F8EEFD411F13" author="Horn" box="[640,763,1815,1840]" pageId="15" pageNumber="124" refString="Horn, W. (1906) Odontochila bennigseni euryoides (nov. subsp.) und das System der Cicindelini. Deutsche Entomologische Zeitschrift, 1, 85 - 86." type="journal article" year="1906">Horn 1906</bibRefCitation>
) from 
<collectingCountry id="DCDDACC7FFD0C8182441F8E1FC0A1F13" box="[840,944,1815,1840]" name="Tanzania" pageId="15" pageNumber="124">Tanzania</collectingCountry>
, and he kept this African taxon in the genus 
<taxonomicName id="63CA97D4FFD0C818279EF8CDFE951F70" authorityName="Laporte de Castelnau" authorityYear="1834" box="[151,303,1851,1875]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="96BE3045FFD0C818279EF8CDFE951F70" box="[151,303,1851,1875]" italics="true" pageId="15" pageNumber="124">Odontocheila</emphasis>
</taxonomicName>
also later (
<bibRefCitation id="C05B91A6FFD0C81826CFF8CAFDF71F77" author="Horn" box="[454,589,1852,1876]" pageId="15" pageNumber="124" refString="Horn, W. (1910) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P., Genera Insectorum, Fascicule 82 B, pp. 105 - 208, pls. 6 - 15." type="journal article" year="1910">Horn 1910</bibRefCitation>
, 
<bibRefCitation id="C05B91A6FFD0C8182568F8CDFD201F77" author="Horn" box="[609,666,1851,1876]" pageId="15" pageNumber="124" refString="Horn, W. (1926) Carabidae, Cicindelinae (Pars 86). In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Junk, Berlin, 345 pp." type="book" year="1926">1926</bibRefCitation>
) in his wide concept both of the subtribe Odontocheilina (as Odontochilina) and of the genus 
<taxonomicName id="63CA97D4FFD0C8182507F8A9FD1C1F54" authorityName="Laporte de Castelnau" authorityYear="1834" box="[526,678,1887,1911]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="96BE3045FFD0C8182507F8A9FD1C1F54" box="[526,678,1887,1911]" italics="true" pageId="15" pageNumber="124">Odontocheila</emphasis>
</taxonomicName>
(as 
<taxonomicName id="63CA97D4FFD0C81825D1F8A9FCDE1F54" authorityName="Agassiz" authorityYear="1846" box="[728,868,1887,1911]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="96BE3045FFD0C81825D1F8A9FCDE1F54" box="[728,868,1887,1911]" italics="true" pageId="15" pageNumber="124">Odontochila</emphasis>
</taxonomicName>
) which in his sense also comprised some African and Asian taxa. 
<taxonomicName id="63CA97D4FFD0C818265AF875FC991FBF" authority="W. Horn, 1897" authorityName="W. Horn" authorityYear="1897" box="[339,803,1923,1948]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="bennigseni">
<emphasis id="96BE3045FFD0C818265AF875FDD41FB8" box="[339,622,1923,1947]" italics="true" pageId="15" pageNumber="124">Odontocheila bennigseni</emphasis>
W. Horn, 1897
</taxonomicName>
was first transferred to the African genus 
<taxonomicName id="63CA97D4FFD0C8182218F875FE901FE3" authority="Guerin, 1849" authorityName="Guerin" authorityYear="1849" class="Insecta" family="Carabidae" genus="Euryarthron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="96BE3045FFD0C8182218F875FA261FB8" box="[1297,1436,1923,1947]" italics="true" pageId="15" pageNumber="124">Euryarthron</emphasis>
Guerin, 1849
</taxonomicName>
by 
<bibRefCitation id="C05B91A6FFD0C818265DF851FDB81FE3" author="Rivalier" box="[340,514,1959,1984]" pageId="15" pageNumber="124" refString="Rivalier, E. (1957) Demembrement du genre Cicindela Linne, III. Faune Africano-Malgache. Revue Francaise d'Entomologie, 24, 312 - 342." type="journal article" year="1957">Rivalier (1957)</bibRefCitation>
, but without mentioning its subspecies 
<taxonomicName id="63CA97D4FFD0C81824B2F851FB921F9C" box="[955,1064,1959,1983]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="euryoides">
<emphasis id="96BE3045FFD0C81824B2F851FB921F9C" box="[955,1064,1959,1983]" italics="true" pageId="15" pageNumber="124">euryoides</emphasis>
</taxonomicName>
that was mentioned as a member of 
<taxonomicName id="63CA97D4FFD0C81827BCF83DFEFA1FC0" authorityName="Guerin" authorityYear="1849" box="[181,320,1995,2019]" class="Insecta" family="Carabidae" genus="Euryarthron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="96BE3045FFD0C81827BCF83DFEFA1FC0" box="[181,320,1995,2019]" italics="true" pageId="15" pageNumber="124">Euryarthron</emphasis>
</taxonomicName>
for the first time by 
<bibRefCitation id="C05B91A6FFD0C8182521F83DFD621FC7" author="Wiesner" box="[552,728,1995,2020]" pageId="15" pageNumber="124" refString="Wiesner, J. (1992) Verzeichnis der Sandlaufkafer der Welt. Checklist of the tiger beetles of the world (Coleoptera, Cicindelidae). Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">Wiesner (1992)</bibRefCitation>
.
</paragraph>
<paragraph id="A475EC57FFCFC80727CEFF61FAE218F6" blockId="16.[151,1436,151,429]" pageId="16" pageNumber="125">
Thus both 
<taxonomicName id="63CA97D4FFCFC807264DFF6EFCB71893" authority="W. Horn, 1906" authorityName="W. Horn" authorityYear="1906" box="[324,781,151,176]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="125" phylum="Arthropoda" rank="subSpecies" species="bennigseni" subSpecies="euryoides">
<emphasis id="96BE3045FFCFC807264DFF6EFDE31893" box="[324,601,152,176]" italics="true" pageId="16" pageNumber="125">O. bennigseni euryoides</emphasis>
<bibRefCitation id="C05B91A6FFCFC807256AFF6EFCB71893" author="Horn" box="[611,781,151,176]" pageId="16" pageNumber="125" refString="Horn, W. (1906) Odontochila bennigseni euryoides (nov. subsp.) und das System der Cicindelini. Deutsche Entomologische Zeitschrift, 1, 85 - 86." type="journal article" year="1906">W. Horn, 1906</bibRefCitation>
</taxonomicName>
and its junior primary homonym 
<taxonomicName id="63CA97D4FFCFC807239EFF6EFF7518F7" authority="W. Horn, 1922" authorityName="W. Horn" authorityYear="1922" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="125" phylum="Arthropoda" rank="species" species="euryoides">
<emphasis id="96BE3045FFCFC807239EFF6EFA931893" box="[1175,1321,152,176]" italics="true" pageId="16" pageNumber="125">O. euryoides</emphasis>
<bibRefCitation id="C05B91A6FFCFC807223AFF6EFF7518F7" author="Horn" pageId="16" pageNumber="125" refString="Horn, W. (1922) Studien uber neue und alte Cicindeliden (Col.), (Neubeschreibungen, Synonymie, Faunistik). Zoologische Mededeelingen, 7, 1 - 112." type="journal article" year="1922">W. Horn, 1922</bibRefCitation>
</taxonomicName>
were in use congeneric until 1992. The homonymy was overlooked also by 
<bibRefCitation id="C05B91A6FFCFC807232FFF4BFB7418F6" author="Lorenz" box="[1062,1230,188,213]" pageId="16" pageNumber="125" refString="Lorenz, W. (1998 a) Systematic list of extant ground beetles of the world (Insecta, Coleoptera, &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Privately published, Tutzing, 502 pp." type="book" year="1998" yearSuffix="a">Lorenz (1998a</bibRefCitation>
, 
<bibRefCitation id="C05B91A6FFCFC80723DAFF4AFB5818F6" author="Lorenz" box="[1235,1250,188,213]" pageId="16" pageNumber="125" refString="Lorenz, W. (1998 b) Nomina Carabidarum: a directory of the scientific names of ground beetles (Insecta, Coleoptera, &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Privately published, Tutzing, 937 pp." type="book" year="1998" yearSuffix="b">b</bibRefCitation>
, 
<bibRefCitation id="C05B91A6FFCFC80723E4FF4AFA8318F6" author="Lorenz" box="[1261,1337,188,213]" pageId="16" pageNumber="125" refString="Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta, Coleoptera, &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Wolfgang Lorenz, Tutzing, iv + 530 pp." type="book" year="2005" yearSuffix="a">2005a</bibRefCitation>
. 
<bibRefCitation id="C05B91A6FFCFC8072235FF4AFAF118F6" author="Lorenz" box="[1340,1355,188,213]" pageId="16" pageNumber="125" refString="Lorenz, W. (2005 b) Nomina Carabidarum: a directory of the scientific names of ground beetles (Insecta, Coleoptera, &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Wolfgang Lorenz, Tutzing, Germany, 937 pp." type="book" year="2005" yearSuffix="b">b</bibRefCitation>
).
</paragraph>
<paragraph id="A475EC57FFCFC80727CEFF29FF62198E" blockId="16.[151,1436,151,429]" pageId="16" pageNumber="125">
According to Article 23.9.5 (
<bibRefCitation id="C05B91A6FFCFC8072504FF16FD2E18DB" author="ICZN" box="[525,660,224,248]" pageId="16" pageNumber="125" refString="ICZN (1999) International Code of Zoological Nomenclature, fourth edition, adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
), when a species-group name in use is a junior primary homonym of another species-group name also in use, but the names apply to taxa not considered congeneric after 1899, the author must not automatically replace the junior homonym. However, in this case both homonyms were considered congeneric after 1899, so the junior name is permanently invalid (Article 57.2). Since there is no available synonym for 
<taxonomicName id="63CA97D4FFCFC8072625FE86FD5D19AB" authority="W. Horn. 1922" authorityName="W. Horn." authorityYear="1922" box="[300,743,368,392]" class="Insecta" family="Carabidae" genus="Odontocheila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="125" phylum="Arthropoda" rank="species" species="euryoides">
<emphasis id="96BE3045FFCFC8072625FE86FD8219AB" box="[300,568,368,392]" italics="true" pageId="16" pageNumber="125">Odontocheila euryoides</emphasis>
<bibRefCitation id="C05B91A6FFCFC8072536FE86FD5D19AB" author="Horn" box="[575,743,368,392]" pageId="16" pageNumber="125" refString="Horn, W. (1922) Studien uber neue und alte Cicindeliden (Col.), (Neubeschreibungen, Synonymie, Faunistik). Zoologische Mededeelingen, 7, 1 - 112." type="journal article" year="1922">W. Horn. 1922</bibRefCitation>
</taxonomicName>
, the name must be replaced by a new substitute name (Article 60.3).
</paragraph>
</subSubSection>
</treatment>
</document>