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<mods:title id="5AB6A0E28FD1D819D11F2A8C30C9EEF1">A revision of the Phelisterhaemorrhous species group (Coleoptera, Histeridae, Exosternini)</mods:title>
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<mods:namePart id="1C5505672BE00F0AC3970B70065556C7">Caterino, Michael S.</mods:namePart>
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<treatment id="2C04FCB06DB3C89341686AF5C730F7EA" ID-GBIF-Taxon="157501006" LSID="urn:lsid:plazi:treatment:2C04FCB06DB3C89341686AF5C730F7EA" httpUri="http://treatment.plazi.org/id/2C04FCB06DB3C89341686AF5C730F7EA" lastPageId="11" lastPageNumber="52" pageId="9" pageNumber="50">
<subSubSection id="FA4D92AE8FBEBDB746339083D2BEE42B" pageId="9" pageNumber="50" type="nomenclature">
<paragraph id="589E76BABD630AF08C32CEE77D0143EB" pageId="9" pageNumber="50">
<taxonomicName id="EA1915C64579D6374B3B3021F6C33AAE" ID-CoL="4FZ2H" authority="JE LeConte, 1859" authorityName="JE LeConte" authorityYear="1859" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister affinis" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="affinis">
<pageBreakToken id="5B7DC17AC781256ECFBA818250EBAA73" pageId="9" pageNumber="50" start="start">Phelister</pageBreakToken>
affinis JE LeConte, 1859
</taxonomicName>
Figs 1, 3; Map 1
</paragraph>
</subSubSection>
<subSubSection id="A55CFBAA1DBC9770D1DB09B484240669" pageId="10" pageNumber="51" type="reference_group">
<paragraph id="856C0134619C6D626F7569FD09579CD1" pageId="10" pageNumber="51">
<taxonomicName id="184E42AEA6549D4D3E7790A17C555DA2" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister affinis" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="affinis">
<pageBreakToken id="5B1A04283F53492D7287F3219F1198F4" pageId="10" pageNumber="51" start="start">Phelister</pageBreakToken>
affinis
</taxonomicName>
JE LeConte, 1859: 311.
</paragraph>
<paragraph id="F12FC95AC7EA884EE18926FCDCDF841A" pageId="10" pageNumber="51">
<taxonomicName id="209F1D7739CB03A56E6540B454AEB4AE" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister simplex" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="simplex">Phelister simplex</taxonomicName>
Casey, 1916: 230;
<bibRefCitation id="62BD97C148A4F4A8B083EFCB9F548DBA" pageId="10" pageNumber="51">Mazur 1997</bibRefCitation>
.
</paragraph>
<paragraph id="1275728854CEF2F40FDC2890E222F087" pageId="10" pageNumber="51">
<taxonomicName id="78AC5187EF53B3521228E13923768CE9" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister solator" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="solator">Phelister solator</taxonomicName>
Marseul, 1861: 164;
<bibRefCitation id="4A428524ECD2A7D3835CFA571F5EC0E6" author="Marseul, SA" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="42" pageNumber="83" pagination="55 - 158" title="Descriptions d'especes nouvelles d'Histerides." volume="13" year="1870">Marseul 1870</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="893F71CB00B1E3977EA02385D262134D" pageId="10" pageNumber="51" type="type material">
<paragraph id="D65A21A201961A199AF1410338981185" pageId="10" pageNumber="51">Type material.</paragraph>
<paragraph id="5B6FE0DC522004BF02CA8D2A459F1750" pageId="10" pageNumber="51">
Neotype male, hereby designated: &quot;Tejeria, Veracruz, MEX, VII:4:41&quot; / &quot;Col. by H. Dybas&quot; / &quot;Collection R. L. Wenzel&quot; / &quot;Phelister #68 det R.Wenzel&quot; / &quot;Compared with type Phelister affinis LeC.
<normalizedToken id="4FB86621D9D788D31A12D505E5FBA948" originalValue="RLW">RLW'</normalizedToken>
51; see type notes under
<taxonomicName id="85622864EE3A0FCD7E4EC0FC604FE240" lsidName="solator" pageId="10" pageNumber="51" rank="species" species="solator">solator</taxonomicName>
&quot;; dissected by Rupert Wenzel (FMNH). The type(s) of this species
<normalizedToken id="ADB93A1D26769915FBEB1F412B0203CC" originalValue="arent">aren't</normalizedToken>
known, despite searches in the most likely repository (MCZC) and others (FMNH, CMNH, USNM), and despite the apparent fact that Wenzel studied a supposed type in 1951 (labels on specimen). Due to the extreme similarity among members of
<taxonomicName id="1E1B2D5D2A266D23C7C2C7A0A2FF534C" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="genus">Phelister</taxonomicName>
, we feel that a Neotype designation is necessary to anchor a specific concept for this species.
</paragraph>
<paragraph id="87AFC7A98D8C04FA361F4882CB0AEADF" pageId="10" pageNumber="51">
Types of synonyms. Lectotype of
<taxonomicName id="DCC09F051E45AABE0A0EC6A3C2402491" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister solator" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="solator">Phelister solator</taxonomicName>
Marseul, of undetermined sex, hereby designated: &quot;Phelister solator, Mexic.
<normalizedToken id="6CD8E7C7F1480983A3364773DD5AFBAC" originalValue="Sallé">Salle</normalizedToken>
20&quot; / &quot;Coll. Desbordes&quot; /
<normalizedToken id="F28B264C4CCA5B95D7A50168BF4CA1E5" originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
, MNHN. Lectotype of
<taxonomicName id="63C088DED3B0543E7083483D2C11D920" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister simplex" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="simplex">Phelister simplex</taxonomicName>
Casey, of undetermined sex, hereby designated: &quot;Lee Co Tex&quot; / &quot;Casey bequest 1925&quot; / &quot;TYPE USNM 38453&quot; / &quot;simplex Csy&quot;, USNM.
</paragraph>
</subSubSection>
<subSubSection id="053957C6478599DB268606647A391DED" lastPageId="11" lastPageNumber="52" pageId="10" pageNumber="51" type="description">
<paragraph id="032BAEB56A2F0387D3B5CD100A8EE1B5" pageId="10" pageNumber="51">Diagnostic description.</paragraph>
<paragraph id="4B6DD8422595B6CB753EF9479266DEBB" lastPageId="11" lastPageNumber="52" pageId="10" pageNumber="51">
Length: 1.73-2.01 mm (avg. 1.94 mm); width: 1.50-1.73 mm (avg. 1.65 mm). Body elongate-oval, widest behind humeri, humeri slightly wider than base of pronotum; body more or less uniformly piceous; entire dorsum finely punctulate, the pronotum more densely so than the elytra; frons finely punctulate, impressed along midline, supraorbital stria complete, frontal stria interrupted at sides and at middle, slightly sinuate laterally; labrum wide, distinctly emarginate apically; both mandibles with strong tooth on inner edges; pronotum usually with distinct fragments of submarginal stria in anterior corners; pronotal disk with larger punctures interspersed with finer punctures along lateral thirds; elytron with single, complete epipleural stria, outer subhumeral stria present in apical third, inner subhumeral stria absent, dorsal striae 1-4 complete, stria 5 present in apical half-two-thirds, very rarely complete, but nearly always with a basal puncture, and sutural stria obsolete in basal third, diverging from the suture anteriad; propygidium with distinct secondary punctures separated by slightly greater than their widths; pygidium more finely punctate; prosternal keel with two complete striae, finely united by anterior arch, free, diverging posteriorly, finely punctulate between in both sexes; mesoventral marginal stria complete, smooth, continued at sides by postmesocoxal stria which runs posteriad two-thirds of the distance to metepipleuron; mesometaventral stria complete, weakly crenulate to smooth, angulate mediad mesocoxa, extending posteriad to near inner corner of metacoxa; first abdominal ventrite with single, complete lateral stria; protibia with apex obliquely truncate, outer margin weakly rounded, bearing ca. six evenly spaced marginal spines; meso- and metatibiae weakly expanded to apex, mesotibia
<pageBreakToken id="B26288B294AD1B5DB3A1CD65C90C1164" pageId="11" pageNumber="52" start="start">with</pageBreakToken>
ca. five marginal spines, more prominent toward apex, metatibia with distinct spines confined to apical fourth. Aedeagus with basal piece a little over one-fourth total length; tegmen widest just beyond middle, abruptly narrowed to thin, divided apices; median lobe more than half tegmen length, proximal apodemes thin near base, thickened toward gonopore.
</paragraph>
<caption id="DC18985FBB7880E99AD441AE06D2ACC0" pageId="11" pageNumber="52">
<paragraph id="EEC43AE9A777D81376B4E3B0FC967DE5" pageId="11" pageNumber="52">
Figure 3.
<normalizedToken id="E4C071C15A2CEAD346FC7E035CFD4047" originalValue="AC">A-C</normalizedToken>
<taxonomicName id="256B8D16CE7BEA7BF01AA0F492282AD7" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister parallelisternus" order="Coleoptera" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="parallelisternus">Phelister parallelisternus</taxonomicName>
Schmidt: A Dorsal habitus B Ventral view showing prosternal and meso- and metaventral striae
<normalizedToken id="6E43635936876A7B426660CAC8999ED9" originalValue="CE">C-E</normalizedToken>
<taxonomicName id="56DED07BDB151CAAB18982E13B69B9CA" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister mobilensis" order="Coleoptera" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="mobilensis">Phelister mobilensis</taxonomicName>
Casey: C Frontal view showing complete epistomal stria D Dorsal habitus E Ventral view (SEM) showing prosternal and meso- and metaventral striae.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B988522C789E8F4B0A75AE70E530002B" pageId="11" pageNumber="52" type="remarks">
<paragraph id="5B8BD5F8D9FBA7E02D7FB5B777B0C292" pageId="11" pageNumber="52">Remarks.</paragraph>
<paragraph id="29E57050C94720A6D63962311CCA5E0D" pageId="11" pageNumber="52">
This species was previously synonymized with
<taxonomicName id="F7E2860ED821C13937F85B5A42E62A67" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister contractus" order="Coleoptera" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="contractus">Phelister contractus</taxonomicName>
Casey by
<bibRefCitation id="2E046F9D7947E4C198635B6688C1A867" pageId="11" pageNumber="52">Mazur (1997)</bibRefCitation>
, in error. Having studied its type, we instead synonymize
<taxonomicName id="5D06F0F33B62DB25E6191617036776EA" lsidName="P. contractus" pageId="11" pageNumber="52" rank="species" species="contractus">P. contractus</taxonomicName>
with
<taxonomicName id="337CA37679263CE458C8CEE330022927" lsidName="P. subrotundus" pageId="11" pageNumber="52" rank="species" species="subrotundus">P. subrotundus</taxonomicName>
Say (below).
</paragraph>
</subSubSection>
<subSubSection id="4F07CB953DC30745DD8EDAD96C560982" pageId="11" pageNumber="52" type="biology_ecology">
<paragraph id="12052FEC896F9EA60A94C30DEAA9BCE7" pageId="11" pageNumber="52">Biology.</paragraph>
<paragraph id="4B30F954E55D43B3D0634B6A0250FDDB" pageId="11" pageNumber="52">
Label data indicate rather generalist habitat preferences, having been collected in cow, horse, and gopher tortoise dung, under decayed leaves, in rotting breadfruit, in fire-scorched
<taxonomicName id="11FEA6B767A2B2AFEF04591C846AC54B" class="Liliopsida" family="Asparagaceae" genus="Yucca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Yucca" order="Asparagales" pageId="11" pageNumber="52" phylum="Tracheophyta" rank="genus">Yucca</taxonomicName>
L., and in rotten
<taxonomicName id="DE203826FE1F6FFAD1497025C84EC7E6" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Opuntia" order="Caryophyllales" pageId="11" pageNumber="52" phylum="Tracheophyta" rank="genus">Opuntia</taxonomicName>
Miller, and the species even exhibits some facultative myrmecophily, with records from nests of both
<taxonomicName id="3E76B338379150F8262B7309CBE624EA" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex" order="Hymenoptera" pageId="11" pageNumber="52" phylum="Arthropoda" rank="genus">Acromyrmex</taxonomicName>
Mayr and
<taxonomicName id="016050A51AE35D6DE0FEBA91B4421448" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Azteca" order="Hymenoptera" pageId="11" pageNumber="52" phylum="Arthropoda" rank="genus">Azteca</taxonomicName>
Forel ants.
</paragraph>
</subSubSection>
<subSubSection id="DBD541F22DA358985A292CBC4294A827" pageId="11" pageNumber="52" type="distribution">
<paragraph id="F1D607914A411C43F57B864686417719" pageId="11" pageNumber="52">Distribution.</paragraph>
<paragraph id="92A53181AF5EF06C9EA2ED1459D0F054" pageId="11" pageNumber="52">
This species occurs from Central America through Mexico, just into the south-central United States. Records: COSTA RICA: Guanacaste, Puntarenas, San
<normalizedToken id="E697F9A86200C3A5620CB0BAB9598D12" originalValue="José">Jose</normalizedToken>
; EL SALVADOR: La Libertad, San Salvador; GUATEMALA: Baja Verapaz, Escuintla, Santa Rosa; HONDURAS: Choluteca, Francisco
<normalizedToken id="67A16B6E2E35DFF207B86F214DF73B17" originalValue="Morazán">Morazan</normalizedToken>
; MEXICO: Chiapas, Colima, Hidalgo,
<normalizedToken id="7DE15AD724101CE13A01B7F76F7BD6B3" originalValue="Michoacán">Michoacan</normalizedToken>
, Morelos, Nayarit, Oaxaca, Sinaloa, Sonora, Tamaulipas, Veracruz; NICARAGUA: Granada,
<normalizedToken id="BB04BCCF718D0439CF56A402AC97E8AB" originalValue="León">Leon</normalizedToken>
, Madriz, Managua, Zelaya; USA: Oklahoma: Latimer; Texas: Cameron, Hidalgo.
</paragraph>
</subSubSection>
</treatment>
</document>