276 lines
36 KiB
XML
276 lines
36 KiB
XML
<document id="9E0F84764293A17FDDD290DCDA6ED58E" ID-DOI="10.5281/zenodo.5174192" ID-ISSN="1942-1354" ID-Zenodo-Dep="5174192" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628335183268" checkinUser="felipe" docAuthor="Mawdsley, Jonathan R." docDate="2012" docId="2B55EA35FFE9FFC0FF31B996FA49F8D6" docLanguage="en" docName="InsectaMundi.0228.1-11.pdf" docOrigin="Insecta Mundi 2012 (228)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Graphipterus ancora Dejean 1831" docType="treatment" docVersion="3" lastPageNumber="4" masterDocId="D76C924DFFEDFFC5FFF1BC3EFFE9FFC2" masterDocTitle="A taxonomic review of the ancora species group of Graphipterus Latreille (Coleoptera: Carabidae)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1699045096987" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="F8A221DA0DDBBC2366ED740966F68253">A taxonomic review of the ancora species group of Graphipterus Latreille (Coleoptera: Carabidae)</mods:title>
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<mods:name id="90CF92F010598A9E4A2340498327FC8F" type="personal">
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<mods:namePart id="3B079E8F40C8CBE87BE3B018E5D892D9">Mawdsley, Jonathan R.</mods:namePart>
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<mods:title id="62DFAA3B59B2AD3AE3C3CFE7497BDB26">Insecta Mundi</mods:title>
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<mods:part id="6ECF07C60B55ED433E0586BFC5775428">
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<mods:date id="B99B554EF015D449272C6EAC4AB55038">2012</mods:date>
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<mods:number id="4AC4CDFA0483B563AF8C63C85BCAE406">2012-04-06</mods:number>
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<mods:number id="524AD074BEB1D3C6FCE49A7DCE8E25AE">2012</mods:number>
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<mods:number id="A8611102B75C6597AC0E98C88180493A">228</mods:number>
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<mods:identifier id="CD14C20BC1393FEF1555D43CF60661F6" type="DOI">10.5281/zenodo.5174192</mods:identifier>
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<mods:identifier id="7FD27BAB5D359A84E5635065AE798788" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="44B0DC266A6CA1FE21F147389635AA54" type="Zenodo-Dep">5174192</mods:identifier>
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<treatment id="2B55EA35FFE9FFC0FF31B996FA49F8D6" LSID="urn:lsid:plazi:treatment:2B55EA35FFE9FFC0FF31B996FA49F8D6" httpUri="http://treatment.plazi.org/id/2B55EA35FFE9FFC0FF31B996FA49F8D6" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
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<subSubSection id="EBE608A8FFE9FFC1FF31B996FDA5FA02" box="[192,588,1448,1472]" pageId="4" pageNumber="3" type="nomenclature">
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<paragraph id="A3435B23FFE9FFC1FF31B996FDA5FA02" blockId="4.[192,588,1448,1504]" box="[192,588,1448,1472]" pageId="4" pageNumber="3">
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<heading id="F80BEC4FFFE9FFC1FF31B996FDA5FA02" bold="true" box="[192,588,1448,1472]" fontSize="10" level="2" pageId="4" pageNumber="3" reason="3">
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<taxonomicName id="64FC20A0FFE9FFC1FF31B996FDA5FA02" ID-CoL="3H95J" authority="Dejean" authorityName="Dejean" authorityYear="1831" box="[192,588,1448,1472]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="ancora">
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<emphasis id="91888731FFE9FFC1FF31B996FDA5FA02" bold="true" box="[192,588,1448,1472]" pageId="4" pageNumber="3">
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<emphasis id="91888731FFE9FFC1FF31B996FE0AFA7D" bold="true" box="[192,483,1448,1472]" italics="true" pageId="4" pageNumber="3">Graphipterus ancora</emphasis>
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Dejean
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EBE608A8FFE9FFC1FF31B9F6FEC1FA22" box="[192,296,1480,1504]" pageId="4" pageNumber="3" type="description">
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<paragraph id="A3435B23FFE9FFC1FF31B9F6FEC1FA22" blockId="4.[192,588,1448,1504]" box="[192,296,1480,1504]" pageId="4" pageNumber="3">
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<figureCitation id="3BC747A6FFE9FFC1FF31B9F6FEC1FA22" box="[192,296,1480,1504]" captionStart="Figures 1–9" captionStartId="5.[192,288,1044,1066]" captionTargetBox="[192,1440,192,1024]" captionTargetId="figure-218@5.[192,1440,192,1024]" captionTargetPageId="5" captionText="Figures 1–9. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 1) G. ancora Dejean [G. ancora ancora Dejean], Hanover, Northern Cape Province, RSA. 2) G. distinctus Péringuey [G. fasciatus distinctus Péringuey], Shilouvane, Limpopo Province, RSA. 3) G. fasciatus Chaudoir [G. fasciatus fasciatus Chaudoir], Umfolozi, KwaZulu/Natal Province, RSA. 4) G. fritschi Chaudoir, Pietersburg, Limpopo Province, RSA. 5) G. westwoodi Brême [G. ancora aurantiacus Basilewsky], Bulawayo, Zimbabwe. 6) G. westwoodi [G. ancora egressus Péringuey], Ohrigstad, Mpumalanga Province, RSA. 7) G. westwoodi [G. ancora subegregius Basilewsky, paratype], Tuli District, Botswana. 8) G. westwoodi [G. ancora transitans Péringuey], Metsimaklaba, Botswana. 9) G. westwoodi [G. ancora westwoodi Brême], Swartruggens Marico, North West Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174194" httpUri="https://zenodo.org/record/5174194/files/figure.png" pageId="4" pageNumber="3">Figure 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="EBE608A8FFE9FFC1FF31BA36FDBCF902" pageId="4" pageNumber="3" type="reference_group">
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<paragraph id="A3435B23FFE9FFC1FF31BA36FDAEF982" blockId="4.[192,1440,1544,1728]" pageId="4" pageNumber="3">
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<treatmentCitationGroup id="83EC7C0DFFE9FFC1FF31BA36FDAEF982" pageId="4" pageNumber="3">
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<treatmentCitation id="225D7D32FFE9FFC1FF31BA36FD20F9E2" author="Dejean, P. F. M. A." box="[192,713,1544,1568]" page=")" pageId="4" pageNumber="3" year="1831">
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<taxonomicName id="64FC20A0FFE9FFC1FF31BA36FD20F9E2" ID-CoL="3H95J" authority="Dejean (1831: 460)" authorityName="Dejean" authorityPageNumber="460" authorityYear="1831" box="[192,713,1544,1568]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="ancora">
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<emphasis id="91888731FFE9FFC1FF31BA36FE23F9E2" box="[192,458,1544,1568]" italics="true" pageId="4" pageNumber="3">Graphipterus ancora</emphasis>
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<bibRefCitation id="C76D26D2FFE9FFC1FE27BA36FD20F9E2" author="Dejean, P. F. M. A." box="[470,713,1544,1568]" pageId="4" pageNumber="3" refId="ref7604" refString="Dejean, P. F. M. A. 1831. Species general des Coleopteres de la collection de M. le Baron Dejean, Tome V. Mequignon-Marvis Pere et Fils; Paris. viii + 883 p." type="book" year="1831">Dejean (1831: 460)</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(
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<typeStatus id="7C47E581FFE9FFC1FD2CBA36FCBBF9E2" box="[733,850,1544,1568]" pageId="4" pageNumber="3" type="holotype">Holotype</typeStatus>
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, Cap de Bonne-Espérance, Muséum National d’Histoire Naturelle, Paris).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="A3435B23FFE9FFC1FF31BA76FDAEF942" blockId="4.[192,1440,1544,1728]" pageId="4" pageNumber="3">
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<treatmentCitationGroup id="83EC7C0DFFE9FFC1FF31BA76FDAEF942" pageId="4" pageNumber="3">
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<treatmentCitation id="225D7D32FFE9FFC1FF31BA76FCEAF9A2" author="Chaudoir, M." box="[192,771,1608,1632]" page=")" pageId="4" pageNumber="3" year="1870">
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<taxonomicName id="64FC20A0FFE9FFC1FF31BA76FCEAF9A2" authority="Chaudoir (1870: 323)" authorityName="Chaudoir" authorityPageNumber="323" authorityYear="1870" box="[192,771,1608,1632]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="giganteus">
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<emphasis id="91888731FFE9FFC1FF31BA76FE03F9A2" box="[192,490,1608,1632]" italics="true" pageId="4" pageNumber="3">Graphipterus giganteus</emphasis>
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<bibRefCitation id="C76D26D2FFE9FFC1FE04BA76FCEAF9A2" author="Chaudoir, M." box="[501,771,1608,1632]" pageId="4" pageNumber="3" pagination="284 - 340" refId="ref7517" refString="Chaudoir, M. Baron de. 1870. Monographie des Graphiterides. Bulletin de la Societe Imperiale des Naturalistes de Moscou 43 (2): 284 - 340." type="journal article" year="1870">Chaudoir (1870: 323)</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(
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<typeStatus id="7C47E581FFE9FFC1FCE6BA76FC62F9A2" box="[791,907,1608,1632]" pageId="4" pageNumber="3" type="holotype">Holotype</typeStatus>
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, Bords du lac N’Gami, Muséum National d’Histoire Naturelle, Paris).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="A3435B23FFE9FFC1FF31BAB6FDBCF902" blockId="4.[192,1440,1544,1728]" pageId="4" pageNumber="3">
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<treatmentCitationGroup id="83EC7C0DFFE9FFC1FF31BAB6FDBCF902" pageId="4" pageNumber="3">
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<treatmentCitation id="225D7D32FFE9FFC1FF31BAB6FC80F962" author="Basilewsky, P." box="[192,873,1672,1696]" page="67" pageId="4" pageNumber="3" year="1977">
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<taxonomicName id="64FC20A0FFE9FFC1FF31BAB6FC80F962" ID-CoL="7H7LR" authority="Basilewsky (1977: 67" authorityName="Basilewsky" authorityYear="1977" box="[192,873,1672,1696]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="ancora" subSpecies="subgiganteus">
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<emphasis id="91888731FFE9FFC1FF31BAB6FD8BF962" box="[192,610,1672,1696]" italics="true" pageId="4" pageNumber="3">Graphipterus ancora subgiganteus</emphasis>
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<bibRefCitation id="C76D26D2FFE9FFC1FD99BAB6FC80F962" author="Basilewsky, P." box="[616,873,1672,1696]" pageId="4" pageNumber="3" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977: 67</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, 71) (
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<typeStatus id="7C47E581FFE9FFC1FC56BAB6FBF0F962" box="[935,1049,1672,1696]" pageId="4" pageNumber="3" type="holotype">Holotype</typeStatus>
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,
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<collectingRegion id="613895C1FFE9FFC1FBD3BAB6FB3AF962" box="[1058,1235,1672,1696]" country="South Africa" name="Western Cape" pageId="4" pageNumber="3">Cape Province</collectingRegion>
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, Vryburg, Musee Royale de l’Afrique Centrale)
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="EBE608A8FFE9FFC0FF31BAD6FDADFA76" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3" type="diagnosis">
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<paragraph id="A3435B23FFE9FFC0FF31BAD6FDADFA76" blockId="4.[192,1440,1768,1920]" lastBlockId="5.[192,1440,1340,1460]" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
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<emphasis id="91888731FFE9FFC1FF31BAD6FEA1F8C2" bold="true" box="[192,328,1768,1792]" pageId="4" pageNumber="3">Diagnosis</emphasis>
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. Large adults, length
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<quantity id="6404F6C6FFE9FFC1FDA0BAD6FD3DF93D" box="[593,724,1768,1792]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="4" pageNumber="3" unit="mm" value="15.5" valueMax="18.0" valueMin="13.0">13–18 mm</quantity>
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. Pattern of elytral vestiture distinctive, with a narrow dark band of pubescence along the suture and extensive areas of yellowish-grey to orange pubescence which surround an oval patch of dark pubescence on the basal half of the elytra (
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<figureCitation id="3BC747A6FFE9FFC1FB94BB16FB27F882" box="[1125,1230,1832,1856]" captionStart="Figures 1–9" captionStartId="5.[192,288,1044,1066]" captionTargetBox="[192,1440,192,1024]" captionTargetId="figure-218@5.[192,1440,192,1024]" captionTargetPageId="5" captionText="Figures 1–9. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 1) G. ancora Dejean [G. ancora ancora Dejean], Hanover, Northern Cape Province, RSA. 2) G. distinctus Péringuey [G. fasciatus distinctus Péringuey], Shilouvane, Limpopo Province, RSA. 3) G. fasciatus Chaudoir [G. fasciatus fasciatus Chaudoir], Umfolozi, KwaZulu/Natal Province, RSA. 4) G. fritschi Chaudoir, Pietersburg, Limpopo Province, RSA. 5) G. westwoodi Brême [G. ancora aurantiacus Basilewsky], Bulawayo, Zimbabwe. 6) G. westwoodi [G. ancora egressus Péringuey], Ohrigstad, Mpumalanga Province, RSA. 7) G. westwoodi [G. ancora subegregius Basilewsky, paratype], Tuli District, Botswana. 8) G. westwoodi [G. ancora transitans Péringuey], Metsimaklaba, Botswana. 9) G. westwoodi [G. ancora westwoodi Brême], Swartruggens Marico, North West Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174194" httpUri="https://zenodo.org/record/5174194/files/figure.png" pageId="4" pageNumber="3">Figure 1</figureCitation>
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). The triangular patch of lighter pubescence along the suture at the very apex of the elytra is also diagnostic of many specimens from
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<collectingCountry id="DBEB1BB3FFE9FFC1FE7ABB56FE2BF842" box="[395,450,1896,1920]" name="South Africa" pageId="4" pageNumber="3">RSA</collectingCountry>
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(specimens from
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<collectingCountry id="DBEB1BB3FFE9FFC1FD51BB56FCF3F8BD" box="[672,794,1896,1919]" name="Botswana" pageId="4" pageNumber="3">Botswana</collectingCountry>
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and
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<collectingCountry id="DBEB1BB3FFE9FFC1FCAABB56FC2FF842" box="[859,966,1896,1920]" name="Namibia" pageId="4" pageNumber="3">Namibia</collectingCountry>
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often lack the patches of dark pubescence on the apical half of the elytra). Adults can be separated from
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<taxonomicName id="64FC20A0FFE8FFC0FBF1B903FB48FA96" box="[1024,1185,1340,1364]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="westwoodi">
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<emphasis id="91888731FFE8FFC0FBF1B903FB48FA96" box="[1024,1185,1340,1364]" italics="true" pageId="5" pageNumber="4">G. westwoodi</emphasis>
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</taxonomicName>
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by the differences in elytral pubescence patterns (compare
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<figureCitation id="3BC747A6FFE8FFC0FD63B962FD13FAB6" box="[658,762,1372,1396]" captionStart="Figures 1–9" captionStartId="5.[192,288,1044,1066]" captionTargetBox="[192,1440,192,1024]" captionTargetId="figure-218@5.[192,1440,192,1024]" captionTargetPageId="5" captionText="Figures 1–9. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 1) G. ancora Dejean [G. ancora ancora Dejean], Hanover, Northern Cape Province, RSA. 2) G. distinctus Péringuey [G. fasciatus distinctus Péringuey], Shilouvane, Limpopo Province, RSA. 3) G. fasciatus Chaudoir [G. fasciatus fasciatus Chaudoir], Umfolozi, KwaZulu/Natal Province, RSA. 4) G. fritschi Chaudoir, Pietersburg, Limpopo Province, RSA. 5) G. westwoodi Brême [G. ancora aurantiacus Basilewsky], Bulawayo, Zimbabwe. 6) G. westwoodi [G. ancora egressus Péringuey], Ohrigstad, Mpumalanga Province, RSA. 7) G. westwoodi [G. ancora subegregius Basilewsky, paratype], Tuli District, Botswana. 8) G. westwoodi [G. ancora transitans Péringuey], Metsimaklaba, Botswana. 9) G. westwoodi [G. ancora westwoodi Brême], Swartruggens Marico, North West Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174194" httpUri="https://zenodo.org/record/5174194/files/figure.png" pageId="5" pageNumber="4">Figure 1</figureCitation>
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with
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<figureCitation id="3BC747A6FFE8FFC0FCB1B962FC38FAB6" box="[832,977,1372,1396]" captionStart="Figures 1–9" captionStartId="5.[192,288,1044,1066]" captionTargetBox="[192,1440,192,1024]" captionTargetId="figure-218@5.[192,1440,192,1024]" captionTargetPageId="5" captionText="Figures 1–9. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 1) G. ancora Dejean [G. ancora ancora Dejean], Hanover, Northern Cape Province, RSA. 2) G. distinctus Péringuey [G. fasciatus distinctus Péringuey], Shilouvane, Limpopo Province, RSA. 3) G. fasciatus Chaudoir [G. fasciatus fasciatus Chaudoir], Umfolozi, KwaZulu/Natal Province, RSA. 4) G. fritschi Chaudoir, Pietersburg, Limpopo Province, RSA. 5) G. westwoodi Brême [G. ancora aurantiacus Basilewsky], Bulawayo, Zimbabwe. 6) G. westwoodi [G. ancora egressus Péringuey], Ohrigstad, Mpumalanga Province, RSA. 7) G. westwoodi [G. ancora subegregius Basilewsky, paratype], Tuli District, Botswana. 8) G. westwoodi [G. ancora transitans Péringuey], Metsimaklaba, Botswana. 9) G. westwoodi [G. ancora westwoodi Brême], Swartruggens Marico, North West Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174194" httpUri="https://zenodo.org/record/5174194/files/figure.png" pageId="5" pageNumber="4">Figures 5–9</figureCitation>
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) and by the somewhat coarser elytral vestiture. Adults can also be separated from
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<taxonomicName id="64FC20A0FFE8FFC0FD17B943FC6BFA56" box="[742,898,1404,1428]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="distinctus">
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<emphasis id="91888731FFE8FFC0FD17B943FC6BFA56" box="[742,898,1404,1428]" italics="true" pageId="5" pageNumber="4">G. distinctus</emphasis>
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</taxonomicName>
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(
|
||
<figureCitation id="3BC747A6FFE8FFC0FC60B942FC13FA56" box="[913,1018,1404,1428]" captionStart="Figures 1–9" captionStartId="5.[192,288,1044,1066]" captionTargetBox="[192,1440,192,1024]" captionTargetId="figure-218@5.[192,1440,192,1024]" captionTargetPageId="5" captionText="Figures 1–9. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 1) G. ancora Dejean [G. ancora ancora Dejean], Hanover, Northern Cape Province, RSA. 2) G. distinctus Péringuey [G. fasciatus distinctus Péringuey], Shilouvane, Limpopo Province, RSA. 3) G. fasciatus Chaudoir [G. fasciatus fasciatus Chaudoir], Umfolozi, KwaZulu/Natal Province, RSA. 4) G. fritschi Chaudoir, Pietersburg, Limpopo Province, RSA. 5) G. westwoodi Brême [G. ancora aurantiacus Basilewsky], Bulawayo, Zimbabwe. 6) G. westwoodi [G. ancora egressus Péringuey], Ohrigstad, Mpumalanga Province, RSA. 7) G. westwoodi [G. ancora subegregius Basilewsky, paratype], Tuli District, Botswana. 8) G. westwoodi [G. ancora transitans Péringuey], Metsimaklaba, Botswana. 9) G. westwoodi [G. ancora westwoodi Brême], Swartruggens Marico, North West Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174194" httpUri="https://zenodo.org/record/5174194/files/figure.png" pageId="5" pageNumber="4">Figure 2</figureCitation>
|
||
) by the much larger body size and by the coarser elytral vestiture.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F7830BABFFE8FFC0FF31B82AFA87FAD2" ID-DOI="http://doi.org/10.5281/zenodo.5174194" ID-Zenodo-Dep="5174194" httpUri="https://zenodo.org/record/5174194/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,288,1044,1066]" targetBox="[192,1440,192,1024]" targetPageId="5">
|
||
<paragraph id="A3435B23FFE8FFC0FF31B82AFA87FAD2" blockId="5.[192,1441,1044,1296]" pageId="5" pageNumber="4">
|
||
<emphasis id="91888731FFE8FFC0FF31B82AFEBAFBEB" bold="true" box="[192,339,1044,1066]" pageId="5" pageNumber="4">Figures 1–9.</emphasis>
|
||
Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured.
|
||
<emphasis id="91888731FFE8FFC0FEEDB873FEDAFBA1" bold="true" box="[284,307,1101,1123]" pageId="5" pageNumber="4">1)</emphasis>
|
||
<taxonomicName id="64FC20A0FFE8FFC0FECBB870FE13FBA6" authority="Dejean" authorityName="Dejean" authorityYear="1831" box="[314,506,1102,1124]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="ancora">
|
||
<emphasis id="91888731FFE8FFC0FECBB870FE4CFBA1" box="[314,421,1102,1123]" italics="true" pageId="5" pageNumber="4">G. ancora</emphasis>
|
||
Dejean
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="64FC20A0FFE8FFC0FDF9B870FCF7FBA1" authority="ancora Dejean" authorityName="Dejean" box="[520,798,1101,1123]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="ancora" subSpecies="ancora">
|
||
<emphasis id="91888731FFE8FFC0FDF9B870FD2FFBA1" box="[520,710,1102,1123]" italics="true" pageId="5" pageNumber="4">G. ancora ancora</emphasis>
|
||
Dejean
|
||
</taxonomicName>
|
||
], Hanover, Northern Cape Province, RSA.
|
||
<emphasis id="91888731FFE8FFC0FB07B873FAE5FBA1" bold="true" box="[1270,1292,1101,1123]" pageId="5" pageNumber="4">2)</emphasis>
|
||
<taxonomicName id="64FC20A0FFE8FFC0FAE2B870FED8FB42" authority="Peringuey" authorityName="Peringuey" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="distinctus">
|
||
<emphasis id="91888731FFE8FFC0FAE2B870FA49FBA1" box="[1299,1440,1102,1123]" italics="true" pageId="5" pageNumber="4">G. distinctus</emphasis>
|
||
Péringuey
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="64FC20A0FFE8FFC0FEB0B855FD5CFB42" authority="Peringuey" authorityName="Peringuey" authorityYear="1899" box="[321,693,1130,1152]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="fasciatus" subSpecies="distinctus">
|
||
<emphasis id="91888731FFE8FFC0FEB0B855FDD1FBBD" box="[321,568,1130,1152]" italics="true" pageId="5" pageNumber="4">G. fasciatus distinctus</emphasis>
|
||
Péringuey
|
||
</taxonomicName>
|
||
], Shilouvane, Limpopo Province, RSA.
|
||
<emphasis id="91888731FFE8FFC0FB96B854FB94FB42" bold="true" box="[1127,1149,1130,1152]" pageId="5" pageNumber="4">3)</emphasis>
|
||
<taxonomicName id="64FC20A0FFE8FFC0FB74B855FA9EFB42" authority="Chaudoir" authorityName="Chaudoir" authorityYear="1870" box="[1157,1399,1130,1152]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="fasciatus">
|
||
<emphasis id="91888731FFE8FFC0FB74B855FAE0FBBD" box="[1157,1289,1130,1152]" italics="true" pageId="5" pageNumber="4">G. fasciatus</emphasis>
|
||
Chaudoir
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="64FC20A0FFE8FFC0FA76B855FE1AFB5E" authority="fasciatus Chaudoir" authorityName="Chaudoir" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="fasciatus" subSpecies="fasciatus">
|
||
<emphasis id="91888731FFE8FFC0FA76B855FE6DFB5E" italics="true" pageId="5" pageNumber="4">G. fasciatus fasciatus</emphasis>
|
||
Chaudoir
|
||
</taxonomicName>
|
||
], Umfolozi, KwaZulu/Natal Province, RSA.
|
||
<emphasis id="91888731FFE8FFC0FC4FB8B9FC3DFB5F" bold="true" box="[958,980,1159,1181]" pageId="5" pageNumber="4">4)</emphasis>
|
||
<taxonomicName id="64FC20A0FFE8FFC0FC28B8B6FAD0FB5F" authority="Chaudoir, Pietersburg" authorityName="Chaudoir, Pietersburg" box="[985,1337,1159,1181]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="fritschi">
|
||
<emphasis id="91888731FFE8FFC0FC28B8B6FBAAFB5E" box="[985,1091,1159,1181]" italics="true" pageId="5" pageNumber="4">G. fritschi</emphasis>
|
||
Chaudoir, Pietersburg
|
||
</taxonomicName>
|
||
, Limpopo Province, RSA.
|
||
<emphasis id="91888731FFE8FFC0FE9BB89AFE69FB78" bold="true" box="[362,384,1188,1210]" pageId="5" pageNumber="4">5)</emphasis>
|
||
<taxonomicName id="64FC20A0FFE8FFC0FE77B89AFD8DFB7B" authority="Breme" authorityName="Breme" box="[390,612,1188,1209]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="91888731FFE8FFC0FE77B89AFDFFFB7B" box="[390,534,1188,1209]" italics="true" pageId="5" pageNumber="4">G. westwoodi</emphasis>
|
||
Brême
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="64FC20A0FFE8FFC0FD80B89AFC04FB7B" authority="Basilewsky" authorityName="Basilewsky" authorityYear="1948" box="[625,1005,1187,1210]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="ancora" subSpecies="aurantiacus">
|
||
<emphasis id="91888731FFE8FFC0FD80B89AFC8FFB78" box="[625,870,1188,1210]" italics="true" pageId="5" pageNumber="4">G. ancora aurantiacus</emphasis>
|
||
Basilewsky
|
||
</taxonomicName>
|
||
], Bulawayo, Zimbabwe.
|
||
<emphasis id="91888731FFE8FFC0FB05B89AFAE3FB78" bold="true" box="[1268,1290,1188,1210]" pageId="5" pageNumber="4">6)</emphasis>
|
||
<taxonomicName id="64FC20A0FFE8FFC0FAE1B89AFA49FB7B" box="[1296,1440,1188,1209]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="91888731FFE8FFC0FAE1B89AFA49FB7B" box="[1296,1440,1188,1209]" italics="true" pageId="5" pageNumber="4">G. westwoodi</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="64FC20A0FFE8FFC0FF39B8FFFDFDFB14" authority="Peringuey" authorityName="Peringuey" authorityYear="1896" box="[200,532,1216,1238]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="ancora" subSpecies="egressus">
|
||
<emphasis id="91888731FFE8FFC0FF39B8FFFE7FFB14" box="[200,406,1217,1238]" italics="true" pageId="5" pageNumber="4">G. ancora egressus</emphasis>
|
||
Péringuey
|
||
</taxonomicName>
|
||
], Ohrigstad, Mpumalanga Province, RSA.
|
||
<emphasis id="91888731FFE8FFC0FC1AB8FEFBEBFB14" bold="true" box="[1003,1026,1216,1238]" pageId="5" pageNumber="4">7)</emphasis>
|
||
<taxonomicName id="64FC20A0FFE8FFC0FBFBB8FFFB75FB14" box="[1034,1180,1217,1238]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="91888731FFE8FFC0FBFBB8FFFB75FB14" box="[1034,1180,1217,1238]" italics="true" pageId="5" pageNumber="4">G. westwoodi</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="64FC20A0FFE8FFC0FB5AB8FFFEA9FB36" authority="Basilewsky" authorityName="Basilewsky" authorityYear="1977" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="ancora" subSpecies="subegregius">
|
||
<emphasis id="91888731FFE8FFC0FB5AB8FFFA49FB14" box="[1195,1440,1217,1238]" italics="true" pageId="5" pageNumber="4">G. ancora subegregius</emphasis>
|
||
Basilewsky
|
||
</taxonomicName>
|
||
, paratype], Tuli District, Botswana.
|
||
<emphasis id="91888731FFE8FFC0FD20B8E3FD00FB31" bold="true" box="[721,745,1245,1267]" pageId="5" pageNumber="4">8)</emphasis>
|
||
<taxonomicName id="64FC20A0FFE8FFC0FD1FB8E0FC96FB31" box="[750,895,1246,1267]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="91888731FFE8FFC0FD1FB8E0FC96FB31" box="[750,895,1246,1267]" italics="true" pageId="5" pageNumber="4">G. westwoodi</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="64FC20A0FFE8FFC0FC7DB8E0FB03FB31" authority="Peringuey" authorityName="Peringuey" authorityYear="1896" box="[908,1258,1245,1267]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="ancora" subSpecies="transitans">
|
||
<emphasis id="91888731FFE8FFC0FC7DB8E0FB86FB31" box="[908,1135,1246,1267]" italics="true" pageId="5" pageNumber="4">G. ancora transitans</emphasis>
|
||
Péringuey
|
||
</taxonomicName>
|
||
], Metsimaklaba, Botswana.
|
||
<emphasis id="91888731FFE8FFC0FECAB8C4FEBBFAD2" bold="true" box="[315,338,1274,1296]" pageId="5" pageNumber="4">9)</emphasis>
|
||
<taxonomicName id="64FC20A0FFE8FFC0FEA8B8C5FE02FACD" box="[345,491,1274,1296]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="91888731FFE8FFC0FEA8B8C5FE02FACD" box="[345,491,1274,1296]" italics="true" pageId="5" pageNumber="4">G. westwoodi</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="64FC20A0FFE8FFC0FE08B8C5FCDBFAD2" authority="Breme" authorityName="Breme" authorityYear="1844" box="[505,818,1274,1296]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="ancora" subSpecies="westwoodi">
|
||
<emphasis id="91888731FFE8FFC0FE08B8C5FD37FACD" box="[505,734,1274,1296]" italics="true" pageId="5" pageNumber="4">G. ancora westwoodi</emphasis>
|
||
Brême
|
||
</taxonomicName>
|
||
], Swartruggens Marico, North West Province, RSA.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="EBE608A8FFE8FFC0FF31B9E2FA49F8D6" pageId="5" pageNumber="4" type="distribution">
|
||
<paragraph id="A3435B23FFE8FFC0FF31B9E2FA49F9D6" blockId="5.[192,1440,1500,1556]" pageId="5" pageNumber="4">
|
||
<emphasis id="91888731FFE8FFC0FF31B9E2FE9FFA36" bold="true" box="[192,374,1500,1524]" pageId="5" pageNumber="4">Distribution.</emphasis>
|
||
<collectingCountry id="DBEB1BB3FFE8FFC0FE70B9E3FE16FA36" box="[385,511,1501,1524]" name="Botswana" pageId="5" pageNumber="4">Botswana</collectingCountry>
|
||
,
|
||
<collectingCountry id="DBEB1BB3FFE8FFC0FDFEB9E2FD9DFA36" box="[527,628,1500,1524]" name="Lesotho" pageId="5" pageNumber="4">Lesotho</collectingCountry>
|
||
,
|
||
<collectingCountry id="DBEB1BB3FFE8FFC0FD72B9E2FD1BFA36" box="[643,754,1500,1524]" name="Namibia" pageId="5" pageNumber="4">Namibia</collectingCountry>
|
||
,
|
||
<collectingCountry id="DBEB1BB3FFE8FFC0FCF3B9E2FBD4FA36" box="[770,1085,1500,1524]" name="South Africa" pageId="5" pageNumber="4">Republic of South Africa</collectingCountry>
|
||
(
|
||
<collectingRegion id="613895C1FFE8FFC0FBA0B9E2FAEBFA36" box="[1105,1282,1500,1524]" country="South Africa" name="Eastern Cape" pageId="5" pageNumber="4">Eastern Cape</collectingRegion>
|
||
,
|
||
<collectingRegion id="613895C1FFE8FFC0FAE3B9E3FA72FA36" box="[1298,1435,1500,1524]" country="South Africa" name="Free State" pageId="5" pageNumber="4">Free State</collectingRegion>
|
||
,
|
||
<collectingRegion id="613895C1FFE8FFC0FF31B9C2FE95F9D6" box="[192,380,1532,1556]" country="South Africa" name="Northern Cape" pageId="5" pageNumber="4">Northern Cape</collectingRegion>
|
||
,
|
||
<collectingRegion id="613895C1FFE8FFC0FE79B9C2FDFFF9D6" box="[392,534,1532,1556]" country="South Africa" name="North West" pageId="5" pageNumber="4">North West</collectingRegion>
|
||
Provinces). For a list of collecting localities see
|
||
<bibRefCitation id="C76D26D2FFE8FFC0FB91B9C2FA74F9D6" author="Basilewsky, P." box="[1120,1437,1532,1556]" pageId="5" pageNumber="4" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977: 71–72)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A3435B23FFE8FFC0FF31BA02FA49F8D6" blockId="5.[192,1440,1596,1812]" pageId="5" pageNumber="4">
|
||
<emphasis id="91888731FFE8FFC0FF31BA02FE51F996" bold="true" box="[192,440,1596,1620]" pageId="5" pageNumber="4">Taxonomic Notes.</emphasis>
|
||
I tentatively include
|
||
<taxonomicName id="64FC20A0FFE8FFC0FD35BA03FCB6F996" authorityName="Chaudoir" authorityYear="1870" box="[708,863,1597,1620]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="91888731FFE8FFC0FD35BA03FCB6F996" box="[708,863,1597,1620]" italics="true" pageId="5" pageNumber="4">G. giganteus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="64FC20A0FFE8FFC0FC6EBA03FB28F996" authorityName="Basilewsky" authorityYear="1977" box="[927,1217,1596,1620]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="ancora" subSpecies="subgiganteus">
|
||
<emphasis id="91888731FFE8FFC0FC6EBA03FB28F996" box="[927,1217,1596,1620]" italics="true" pageId="5" pageNumber="4">G. ancora subgiganteus</emphasis>
|
||
</taxonomicName>
|
||
with this species, based on the overall similarity in elytral vestiture among the three forms, especially the similar large oval patch of dark pubescence on the base of the elytra (see
|
||
<bibRefCitation id="C76D26D2FFE8FFC0FC55BA42FB4AF956" author="Basilewsky, P." box="[932,1187,1660,1684]" pageId="5" pageNumber="4" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky 1977: 70</bibRefCitation>
|
||
). The few specimens known of
|
||
<taxonomicName id="64FC20A0FFE8FFC0FEC4BAA3FE24F976" authorityName="Chaudoir" authorityYear="1870" box="[309,461,1693,1716]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="91888731FFE8FFC0FEC4BAA3FE24F976" box="[309,461,1693,1716]" italics="true" pageId="5" pageNumber="4">G. giganteus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="64FC20A0FFE8FFC0FDF9BAA3FCCFF976" authorityName="Basilewsky" authorityYear="1977" box="[520,806,1692,1716]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="ancora" subSpecies="subgiganteus">
|
||
<emphasis id="91888731FFE8FFC0FDF9BAA3FCCFF976" box="[520,806,1692,1716]" italics="true" pageId="5" pageNumber="4">G. ancora subgiganteus</emphasis>
|
||
</taxonomicName>
|
||
have much more extensive areas of pale pubescence on the apical half of the elytra than the nominate form. It is possible that these forms will ultimately be found to represent one or more separate species. Further material is needed to resolve the status of these two forms, which are known collectively from a total of
|
||
<specimenCount id="B5FA90AAFFE8FFC0FC3BBAC2FB6FF8D6" box="[970,1158,1788,1812]" pageId="5" pageNumber="4" type="generic">nine specimens</specimenCount>
|
||
(
|
||
<bibRefCitation id="C76D26D2FFE8FFC0FB64BAC2FA7BF8D6" author="Basilewsky, P." box="[1173,1426,1788,1812]" pageId="5" pageNumber="4" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky 1977: 71</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |