307 lines
39 KiB
XML
307 lines
39 KiB
XML
<document id="8BEB9727F647A83BFF79C058DA114942" ID-DOI="10.5281/zenodo.5174192" ID-ISSN="1942-1354" ID-Zenodo-Dep="5174192" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628335183268" checkinUser="felipe" docAuthor="Mawdsley, Jonathan R." docDate="2012" docId="2B55EA35FFE4FFCFFF31BD7FFC93FEFB" docLanguage="en" docName="InsectaMundi.0228.1-11.pdf" docOrigin="Insecta Mundi 2012 (228)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Graphipterus wahlbergi Boheman 2012, new status" docType="treatment" docVersion="4" lastPageNumber="9" masterDocId="D76C924DFFEDFFC5FFF1BC3EFFE9FFC2" masterDocTitle="A taxonomic review of the ancora species group of Graphipterus Latreille (Coleoptera: Carabidae)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="8" updateTime="1699045096987" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="66B55543E1A6EEAE652CCED47E7A4357">A taxonomic review of the ancora species group of Graphipterus Latreille (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="411D307C44568B47BF382DE1270CE330">Mawdsley, Jonathan R.</mods:namePart>
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<mods:title id="97EB1CD7DB6DF2AE17E21F17521DDDF2">Insecta Mundi</mods:title>
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<mods:part id="1918C36E1A1BC70AB29EBD5BA75FE64C">
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<mods:date id="D6580621B403FA55DDACCD2DE993CA9E">2012</mods:date>
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<mods:number id="E3413BB16912ADA56876422A555F345B">2012-04-06</mods:number>
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<mods:number id="CF42EC15295644AB9697596478A8DDDC">2012</mods:number>
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<mods:identifier id="AD4828927962518673B1387ACFA862E1" type="DOI">10.5281/zenodo.5174192</mods:identifier>
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<mods:identifier id="5052A3E9D0747B3526579D6CA4848297" type="ISSN">1942-1354</mods:identifier>
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<treatment id="2B55EA35FFE4FFCFFF31BD7FFC93FEFB" ID-DOI="http://doi.org/10.5281/zenodo.5187090" ID-Zenodo-Dep="5187090" LSID="urn:lsid:plazi:treatment:2B55EA35FFE4FFCFFF31BD7FFC93FEFB" httpUri="http://treatment.plazi.org/id/2B55EA35FFE4FFCFFF31BD7FFC93FEFB" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
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<subSubSection id="EBE608A8FFE4FFCCFF31BD7FFCD5FE98" box="[192,828,321,346]" pageId="9" pageNumber="8" type="nomenclature">
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<paragraph id="A3435B23FFE4FFCCFF31BD7FFCD5FE98" blockId="9.[192,828,321,378]" box="[192,828,321,346]" pageId="9" pageNumber="8">
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<emphasis id="91888731FFE4FFCCFF31BD7FFCD5FE98" bold="true" box="[192,828,321,346]" pageId="9" pageNumber="8">
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<taxonomicName id="64FC20A0FFE4FFCCFF31BD7FFD70FE98" authority="Boheman" authorityName="Boheman" authorityYear="2012" box="[192,665,321,346]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="wahlbergi" status="stat. nov.">
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<emphasis id="91888731FFE4FFCCFF31BD7FFDE4FE9B" bold="true" box="[192,525,321,345]" italics="true" pageId="9" pageNumber="8">Graphipterus wahlbergi</emphasis>
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Boheman
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</taxonomicName>
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,
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<taxonomicNameLabel id="8ABB3A4AFFE4FFCCFD56BD7CFCD5FE98" box="[679,828,322,346]" pageId="9" pageNumber="8">new status</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="EBE608A8FFE4FFCCFF31BD5CFE99FEBB" box="[192,368,353,378]" pageId="9" pageNumber="8" type="description">
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<paragraph id="A3435B23FFE4FFCCFF31BD5CFE99FEBB" blockId="9.[192,828,321,378]" box="[192,368,353,378]" pageId="9" pageNumber="8">
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<figureCitation id="3BC747A6FFE4FFCCFF31BD5CFE99FEBB" box="[192,368,353,378]" captionStart="Figures 10–18" captionStartId="7.[193,288,1050,1072]" captionTargetBox="[202,1410,197,1029]" captionTargetId="figure-191@7.[192,1440,197,1029]" captionTargetPageId="7" captionText="Figures 10–18. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 10) G. cordiger [G. cordiger cordiger Dejean], Gideon’s Kraal Farm, 30.38 S, 25.34 E, Northern Cape Province, RSA. 11) G. cordiger [G. cordiger hamatus Boheman], nr. Hartbeespoort Dam, North West Province, RSA. 12) G. cordiger [G. cordiger subcordiger Basilewsky, paratype], Swakopmund, Namibia. 13) G. cordiger [G. cordiger subhamatus Basilewsky, paratype], Thornybush, Limpopo Province, RSA. 14) G. cordiger [G. cordiger zambezianus Péringuey], Salisbury, Mashonaland (modern-day Harare, Zimbabwe). 15) G. wahlbergi [G. cordiger betshuana Burgeon], Grootdraai, Mpumalanga Province, RSA. 16) G. wahlbergi [G. cordiger natalicus Basilewsky, paratype], “Natal,” KwaZulu/Natal Province, RSA. 17) G. wahlbergi [G. cordiger transfugus Péringuey], Nylsvley, Limpopo Province, RSA. 18) G. wahlbergi [G. cordiger wahlbergi Boheman], Ohrigstad, Mpumalanga Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174196" httpUri="https://zenodo.org/record/5174196/files/figure.png" pageId="9" pageNumber="8">Figures 15–18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="EBE608A8FFE4FFCCFF31BD9CFE77FD58" pageId="9" pageNumber="8" type="reference_group">
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<paragraph id="A3435B23FFE4FFCCFF31BD9CFE02FE18" blockId="9.[192,1440,418,666]" pageId="9" pageNumber="8">
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<treatmentCitationGroup id="83EC7C0DFFE4FFCCFF31BD9CFE02FE18" pageId="9" pageNumber="8">
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<treatmentCitation id="225D7D32FFE4FFCCFF31BD9CFD00FE78" author="Boheman, C." box="[192,745,418,442]" page=")" pageId="9" pageNumber="8" year="1848">
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<taxonomicName id="64FC20A0FFE4FFCCFF31BD9CFD00FE78" authority="Boheman (1848: 60)" authorityName="Boheman" authorityPageNumber="60" authorityYear="1848" box="[192,745,418,442]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="wahlbergi">
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<emphasis id="91888731FFE4FFCCFF31BD9CFE01FE78" box="[192,488,418,442]" italics="true" pageId="9" pageNumber="8">Graphipterus wahlbergi</emphasis>
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<bibRefCitation id="C76D26D2FFE4FFCCFE00BD9CFD00FE78" author="Boheman, C." box="[497,745,418,442]" pageId="9" pageNumber="8" refId="ref7411" refString="Boheman, C. 1848. Insecta Caffraria, annis 1838 - 1845 a J. A. Wahlberg collecta, pars 1 fasc. 1. Officina Norstedtiana; Stockholm. 626 p." type="book" year="1848">Boheman (1848: 60)</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(
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<typeStatus id="7C47E581FFE4FFCCFD0BBD9CFC85FE78" box="[762,876,418,442]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
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, in Caffraria interiore, Naturhistoriska Riksmuseet, Stockholm).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="A3435B23FFE4FFCCFF31BDDCFE85FDD8" blockId="9.[192,1440,418,666]" pageId="9" pageNumber="8">
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<treatmentCitation id="225D7D32FFE4FFCCFF31BDDCFCE1FE38" author="Peringuey, L." box="[192,776,482,506]" page="297" pageId="9" pageNumber="8" year="1896">
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<taxonomicName id="64FC20A0FFE4FFCCFF31BDDCFCE1FE38" authority="Peringuey (1896: 297" authorityName="Peringuey" authorityYear="1896" box="[192,776,482,506]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="transfugus">
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<emphasis id="91888731FFE4FFCCFF31BDDCFE1BFE38" box="[192,498,482,506]" italics="true" pageId="9" pageNumber="8">Graphipterus transfugus</emphasis>
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<bibRefCitation id="C76D26D2FFE4FFCCFE0DBDDCFCE1FE38" author="Peringuey, L." box="[508,776,482,506]" pageId="9" pageNumber="8" pagination="99 - 623" refId="ref8046" refString="Peringuey, L. 1896. Descriptive catalogue of the Coleoptera of South Africa, part II. Transactions of the South African Philosophical Society 7: 99 - 623." type="journal article" year="1896">Péringuey (1896: 297</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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, 324) (
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<typeStatus id="7C47E581FFE4FFCCFCACBDDCFC26FE38" box="[861,975,482,506]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
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, Transvaal, Makapan, South African Museum).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="A3435B23FFE4FFCCFF31BE1CFD99FD98" blockId="9.[192,1440,418,666]" pageId="9" pageNumber="8">
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<treatmentCitationGroup id="83EC7C0DFFE4FFCCFF31BE1CFD99FD98" pageId="9" pageNumber="8">
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<treatmentCitation id="225D7D32FFE4FFCCFF31BE1CFC82FDF8" author="Burgeon, L." box="[192,875,546,570]" page=")" pageId="9" pageNumber="8" year="1929">
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<taxonomicName id="64FC20A0FFE4FFCCFF31BE1CFC82FDF8" authority="Burgeon (1929: 294)" authorityName="Burgeon" authorityPageNumber="294" authorityYear="1929" box="[192,875,546,570]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="cordiger" subSpecies="betshuana">
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<emphasis id="91888731FFE4FFCCFF31BE1CFD89FDF8" box="[192,608,546,570]" italics="true" pageId="9" pageNumber="8">Graphipterus cordiger betshuana</emphasis>
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<bibRefCitation id="C76D26D2FFE4FFCCFD9ABE1CFC82FDF8" author="Burgeon, L." box="[619,875,546,570]" pageId="9" pageNumber="8" pagination="273 - 351" refId="ref7492" refString="Burgeon, L. 1929. Monographie du genre Graphipterus Latr. Annales de la Societe Entomologique du Belgique 69: 273 - 351." type="journal article" year="1929">Burgeon (1929: 294)</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
(
|
||
<typeStatus id="7C47E581FFE4FFCCFC8EBE1CFC1BFDF8" box="[895,1010,546,570]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
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||
, Serue, Zoologisches Museum der Humboldt-Universität, Berlin).
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</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="A3435B23FFE4FFCCFF31BE5CFE77FD58" blockId="9.[192,1440,418,666]" pageId="9" pageNumber="8">
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<treatmentCitationGroup id="83EC7C0DFFE4FFCCFF31BE5CFE77FD58" pageId="9" pageNumber="8">
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<treatmentCitation id="225D7D32FFE4FFCCFF31BE5CFCBFFDBB" author="Basilewsky, P." box="[192,854,610,634]" page="77" pageId="9" pageNumber="8" year="1977">
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||
<taxonomicName id="64FC20A0FFE4FFCCFF31BE5CFCBFFDBB" ID-CoL="7H7MC" authority="Basilewsky (1977: 77" authorityName="Basilewsky" authorityYear="1977" box="[192,854,610,634]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="cordiger" subSpecies="natalicus">
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<emphasis id="91888731FFE4FFCCFF31BE5CFDA0FDB8" box="[192,585,610,634]" italics="true" pageId="9" pageNumber="8">Graphipterus cordiger natalicus</emphasis>
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||
<bibRefCitation id="C76D26D2FFE4FFCCFDA0BE5CFCBFFDBB" author="Basilewsky, P." box="[593,854,610,634]" pageId="9" pageNumber="8" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977: 77</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, 87–88) (
|
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<typeStatus id="7C47E581FFE4FFCCFC34BE5CFBDEFDB8" box="[965,1079,610,634]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
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||
, Natal, Mfongosi, South African Museum).
|
||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="EBE608A8FFE4FFCCFF31BEFFFE51FBFB" pageId="9" pageNumber="8" type="diagnosis">
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<paragraph id="A3435B23FFE4FFCCFF31BEFFFE51FBFB" blockId="9.[192,1440,705,1082]" pageId="9" pageNumber="8">
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||
<emphasis id="91888731FFE4FFCCFF31BEFFFEB9FD1B" bold="true" box="[192,336,705,729]" pageId="9" pageNumber="8">Diagnosis.</emphasis>
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Smaller adults, length
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<quantity id="6404F6C6FFE4FFCCFD81BEFFFD1DFD1B" box="[624,756,705,729]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="9" pageNumber="8" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">11–15 mm</quantity>
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. Pattern of elytral vestiture distinctive (
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<figureCitation id="3BC747A6FFE4FFCCFB11BEFCFA7BFD1B" box="[1248,1426,705,730]" captionStart="Figures 10–18" captionStartId="7.[193,288,1050,1072]" captionTargetBox="[202,1410,197,1029]" captionTargetId="figure-191@7.[192,1440,197,1029]" captionTargetPageId="7" captionText="Figures 10–18. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 10) G. cordiger [G. cordiger cordiger Dejean], Gideon’s Kraal Farm, 30.38 S, 25.34 E, Northern Cape Province, RSA. 11) G. cordiger [G. cordiger hamatus Boheman], nr. Hartbeespoort Dam, North West Province, RSA. 12) G. cordiger [G. cordiger subcordiger Basilewsky, paratype], Swakopmund, Namibia. 13) G. cordiger [G. cordiger subhamatus Basilewsky, paratype], Thornybush, Limpopo Province, RSA. 14) G. cordiger [G. cordiger zambezianus Péringuey], Salisbury, Mashonaland (modern-day Harare, Zimbabwe). 15) G. wahlbergi [G. cordiger betshuana Burgeon], Grootdraai, Mpumalanga Province, RSA. 16) G. wahlbergi [G. cordiger natalicus Basilewsky, paratype], “Natal,” KwaZulu/Natal Province, RSA. 17) G. wahlbergi [G. cordiger transfugus Péringuey], Nylsvley, Limpopo Province, RSA. 18) G. wahlbergi [G. cordiger wahlbergi Boheman], Ohrigstad, Mpumalanga Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174196" httpUri="https://zenodo.org/record/5174196/files/figure.png" pageId="9" pageNumber="8">Figures 15–18</figureCitation>
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), with contrasting patches of yellowish-grey and black pubescence, the latter forming a narrow band along suture and a single broad transverse, nearly rectangular patch extending onto the disc but not attaining the lateral margin. The color of the lighter pubescence ranges from yellowish-grey to orange. This species has long been confused with
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<taxonomicName id="64FC20A0FFE4FFCCFD3DBF7CFCBFFC98" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[716,854,834,858]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cordiger">
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<emphasis id="91888731FFE4FFCCFD3DBF7CFCBFFC98" box="[716,854,834,858]" italics="true" pageId="9" pageNumber="8">G. cordiger</emphasis>
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||
</taxonomicName>
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||
and in fact was placed in synonymy with that species by
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||
<bibRefCitation id="C76D26D2FFE4FFCCFEB7BF5CFDEDFCB8" author="Burgeon, L." box="[326,516,866,890]" pageId="9" pageNumber="8" pagination="273 - 351" refId="ref7492" refString="Burgeon, L. 1929. Monographie du genre Graphipterus Latr. Annales de la Societe Entomologique du Belgique 69: 273 - 351." type="journal article" year="1929">Burgeon (1929)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFDB4BF5CFCC3FCB8" author="Basilewsky, P." box="[581,810,866,890]" pageId="9" pageNumber="8" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977)</bibRefCitation>
|
||
. In
|
||
<taxonomicName id="64FC20A0FFE4FFCCFCAABF5CFC0DFCB8" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[859,996,866,890]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cordiger">
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||
<emphasis id="91888731FFE4FFCCFCAABF5CFC0DFCB8" box="[859,996,866,890]" italics="true" pageId="9" pageNumber="8">G. cordiger</emphasis>
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||
</taxonomicName>
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||
, however, the median elytral patch of dark pubescence is broadly rounded or lobed on the disc, and the anterior band of pale pubescence has a distinct triangular projection onto the disc (
|
||
<figureCitation id="3BC747A6FFE4FFCCFCDABF9CFC08FC78" box="[811,993,930,954]" captionStart="Figures 10–18" captionStartId="7.[193,288,1050,1072]" captionTargetBox="[202,1410,197,1029]" captionTargetId="figure-191@7.[192,1440,197,1029]" captionTargetPageId="7" captionText="Figures 10–18. Adult habitus images of specimens of species in the ancora species group, from the TMSA collection. Specimen identifications by P. Basilewsky are given in brackets after the current valid name for the specimen figured. 10) G. cordiger [G. cordiger cordiger Dejean], Gideon’s Kraal Farm, 30.38 S, 25.34 E, Northern Cape Province, RSA. 11) G. cordiger [G. cordiger hamatus Boheman], nr. Hartbeespoort Dam, North West Province, RSA. 12) G. cordiger [G. cordiger subcordiger Basilewsky, paratype], Swakopmund, Namibia. 13) G. cordiger [G. cordiger subhamatus Basilewsky, paratype], Thornybush, Limpopo Province, RSA. 14) G. cordiger [G. cordiger zambezianus Péringuey], Salisbury, Mashonaland (modern-day Harare, Zimbabwe). 15) G. wahlbergi [G. cordiger betshuana Burgeon], Grootdraai, Mpumalanga Province, RSA. 16) G. wahlbergi [G. cordiger natalicus Basilewsky, paratype], “Natal,” KwaZulu/Natal Province, RSA. 17) G. wahlbergi [G. cordiger transfugus Péringuey], Nylsvley, Limpopo Province, RSA. 18) G. wahlbergi [G. cordiger wahlbergi Boheman], Ohrigstad, Mpumalanga Province, RSA." figureDoi="http://doi.org/10.5281/zenodo.5174196" httpUri="https://zenodo.org/record/5174196/files/figure.png" pageId="9" pageNumber="8">Figures 10–14</figureCitation>
|
||
). Along with
|
||
<taxonomicName id="64FC20A0FFE4FFCCFB76BF9CFAE6FC78" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[1159,1295,930,954]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cordiger">
|
||
<emphasis id="91888731FFE4FFCCFB76BF9CFAE6FC78" box="[1159,1295,930,954]" italics="true" pageId="9" pageNumber="8">G. cordiger</emphasis>
|
||
</taxonomicName>
|
||
, this is one of the few species of
|
||
<taxonomicName id="64FC20A0FFE4FFCCFE34BFFCFD83FC18" authorityName="Latreille" authorityYear="1802" box="[453,618,962,986]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="91888731FFE4FFCCFE34BFFCFD83FC18" box="[453,618,962,986]" italics="true" pageId="9" pageNumber="8">Graphipterus</emphasis>
|
||
</taxonomicName>
|
||
in which the second metatibial spur is arcuate and not spatulate (
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFF39BFDCFE73FC38" author="Basilewsky, P." box="[200,410,994,1018]" pageId="9" pageNumber="8" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky 1977</bibRefCitation>
|
||
). The shape of the patches of dark setae and the coloration of the lighter setae varies between populations and was used by
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFD65B83CFCB9FBD8" author="Burgeon, L." box="[660,848,1026,1050]" pageId="9" pageNumber="8" pagination="273 - 351" refId="ref7492" refString="Burgeon, L. 1929. Monographie du genre Graphipterus Latr. Annales de la Societe Entomologique du Belgique 69: 273 - 351." type="journal article" year="1929">Burgeon (1929)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFC7DB83CFB82FBD8" author="Basilewsky, P." box="[908,1131,1026,1050]" pageId="9" pageNumber="8" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977)</bibRefCitation>
|
||
as the basis for recognizing subspecific taxa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EBE608A8FFE4FFCFFF31B85FFC93FEFB" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="distribution">
|
||
<paragraph id="A3435B23FFE4FFCCFF31B85FFDE8FB78" blockId="9.[192,1440,1121,1210]" pageId="9" pageNumber="8">
|
||
<emphasis id="91888731FFE4FFCCFF31B85FFE9DFBBB" bold="true" box="[192,372,1121,1145]" pageId="9" pageNumber="8">Distribution.</emphasis>
|
||
<collectingCountry id="DBEB1BB3FFE4FFCCFE88B85CFE1FFBBB" box="[377,502,1122,1145]" name="Botswana" pageId="9" pageNumber="8">Botswana</collectingCountry>
|
||
,
|
||
<collectingCountry id="DBEB1BB3FFE4FFCCFDF0B85CFCC0FBB8" box="[513,809,1122,1146]" name="South Africa" pageId="9" pageNumber="8">Republic of South Africa</collectingCountry>
|
||
(
|
||
<collectingRegion id="613895C1FFE4FFCCFCC6B85CFC4DFBB8" box="[823,932,1122,1146]" country="South Africa" name="Gauteng" pageId="9" pageNumber="8">Gauteng</collectingRegion>
|
||
, KwaZulu/Natal,
|
||
<collectingRegion id="613895C1FFE4FFCCFB8DB85CFB03FBB8" box="[1148,1258,1122,1146]" country="South Africa" name="Limpopo" pageId="9" pageNumber="8">Limpopo</collectingRegion>
|
||
,
|
||
<collectingRegion id="613895C1FFE4FFCCFB04B85CFA75FBB8" box="[1269,1436,1122,1146]" country="South Africa" name="Mpumalanga" pageId="9" pageNumber="8">Mpumalanga</collectingRegion>
|
||
,
|
||
<collectingRegion id="613895C1FFE4FFCCFF31B8BCFE92FB58" box="[192,379,1154,1178]" country="South Africa" name="Northern Cape" pageId="9" pageNumber="8">Northern Cape</collectingRegion>
|
||
, and
|
||
<collectingRegion id="613895C1FFE4FFCCFE4AB8BCFDA0FB5B" box="[443,585,1154,1178]" country="South Africa" name="North West" pageId="9" pageNumber="8">North West</collectingRegion>
|
||
Provinces),
|
||
<collectingCountry id="DBEB1BB3FFE4FFCCFD2FB8BCFC88FB58" box="[734,865,1154,1178]" name="Swaziland" pageId="9" pageNumber="8">Swaziland</collectingCountry>
|
||
,
|
||
<collectingCountry id="DBEB1BB3FFE4FFCCFC9DB8BCFC06FB58" box="[876,1007,1154,1178]" name="Zimbabwe" pageId="9" pageNumber="8">Zimbabwe</collectingCountry>
|
||
. For a list of collecting localities see
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFF31B89CFE14FB78" author="Basilewsky, P." box="[192,509,1186,1210]" pageId="9" pageNumber="8" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977: 82–88)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A3435B23FFE4FFCCFF31B8DFFAB2F898" blockId="9.[192,1440,1249,1882]" pageId="9" pageNumber="8">
|
||
<emphasis id="91888731FFE4FFCCFF31B8DFFE50FB38" bold="true" box="[192,441,1249,1274]" pageId="9" pageNumber="8">Taxonomic Notes.</emphasis>
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFE30B8DCFD69FB38" author="Burgeon, L." box="[449,640,1250,1274]" pageId="9" pageNumber="8" pagination="273 - 351" refId="ref7492" refString="Burgeon, L. 1929. Monographie du genre Graphipterus Latr. Annales de la Societe Entomologique du Belgique 69: 273 - 351." type="journal article" year="1929">Burgeon (1929)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFD31B8DCFC4BFB38" author="Basilewsky, P." box="[704,930,1250,1274]" pageId="9" pageNumber="8" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977)</bibRefCitation>
|
||
placed this species in synonymy with
|
||
<taxonomicName id="64FC20A0FFE4FFCCFA75B8DCFECDFAD8" baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cordiger">
|
||
<emphasis id="91888731FFE4FFCCFA75B8DCFECDFAD8" italics="true" pageId="9" pageNumber="8">G. cordiger</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFE93B93CFDABFAD8" author="Basilewsky, P." box="[354,578,1282,1306]" pageId="9" pageNumber="8" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977)</bibRefCitation>
|
||
recognized a total of nine subspecific taxa under that single polytypic species. Application of the polytypic species concept to this group of taxa is not straightforward, as shown by the fact that Basilewsky could only offer a partial key to subspecies (
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFB82B97CFA87FA98" author="Basilewsky, P." box="[1139,1390,1346,1370]" pageId="9" pageNumber="8" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky 1977: 78</bibRefCitation>
|
||
, 80) and a map (
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFEBEB95CFDC8FAB8" author="Basilewsky, P." box="[335,545,1378,1402]" pageId="9" pageNumber="8" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky 1977</bibRefCitation>
|
||
: Carte 9) which shows clear areas of overlap between many of the alleged subspecific forms (e.g. between
|
||
<emphasis id="91888731FFE4FFCCFDB0B9BCFD14FA58" box="[577,765,1410,1434]" italics="true" pageId="9" pageNumber="8">G. c. betshuana</emphasis>
|
||
and
|
||
<emphasis id="91888731FFE4FFCCFCCBB9BCFBE6FA58" box="[826,1039,1410,1434]" italics="true" pageId="9" pageNumber="8">G. c. subhamatus</emphasis>
|
||
, between
|
||
<emphasis id="91888731FFE4FFCCFB7BB9BCFAAEFA58" box="[1162,1351,1410,1434]" italics="true" pageId="9" pageNumber="8">G. c. betshuana</emphasis>
|
||
and
|
||
<emphasis id="91888731FFE4FFCCFA75B9BCFE93FA78" italics="true" pageId="9" pageNumber="8">
|
||
G. c.
|
||
<taxonomicName id="64FC20A0FFE4FFCCFF2AB99CFE93FA78" authorityName="Peringuey" authorityYear="1892" box="[219,378,1442,1466]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="zambezianus">zambezianus</taxonomicName>
|
||
</emphasis>
|
||
, and between
|
||
<emphasis id="91888731FFE4FFCCFDDAB99CFD3CFA78" box="[555,725,1442,1466]" italics="true" pageId="9" pageNumber="8">
|
||
G. c.
|
||
<taxonomicName id="64FC20A0FFE4FFCCFD98B99CFD3CFA78" authorityName="Boheman" authorityYear="1848" box="[617,725,1442,1466]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="hamatus">hamatus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="91888731FFE4FFCCFCE2B99CFC3CFA78" box="[787,981,1442,1466]" italics="true" pageId="9" pageNumber="8">
|
||
G. c.
|
||
<taxonomicName id="64FC20A0FFE4FFCCFCA0B99CFC3CFA78" authorityName="Peringuey" authorityYear="1896" box="[849,981,1442,1466]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="transfugus">transfugus</taxonomicName>
|
||
</emphasis>
|
||
). Furthermore, several pairs of putative subspecies have identical patterns of elytral pubescence and can be separated only by collecting locality (e.g.
|
||
<emphasis id="91888731FFE4FFCCFEADB9DCFE17FA38" box="[348,510,1506,1530]" italics="true" pageId="9" pageNumber="8">
|
||
G. c.
|
||
<taxonomicName id="64FC20A0FFE4FFCCFE6BB9DCFE17FA38" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[410,510,1506,1530]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cordiger">cordiger</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="91888731FFE4FFCCFDCAB9DCFCE1FA38" box="[571,776,1506,1530]" italics="true" pageId="9" pageNumber="8">G. c. subcordiger</emphasis>
|
||
,
|
||
<emphasis id="91888731FFE4FFCCFCE6B9DCFC28FA38" box="[791,961,1506,1530]" italics="true" pageId="9" pageNumber="8">
|
||
G. c.
|
||
<taxonomicName id="64FC20A0FFE4FFCCFCA4B9DCFC28FA38" authorityName="Boheman" authorityYear="1848" box="[853,961,1506,1530]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="hamatus">hamatus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="91888731FFE4FFCCFC0EB9DCFB3DFA38" box="[1023,1236,1506,1530]" italics="true" pageId="9" pageNumber="8">G. c. subhamatus</emphasis>
|
||
,
|
||
<emphasis id="91888731FFE4FFCCFB12B9DCFA49FA38" box="[1251,1440,1506,1530]" italics="true" pageId="9" pageNumber="8">G. c. betshuana</emphasis>
|
||
and
|
||
<emphasis id="91888731FFE4FFCCFF04BA3CFE5AF9D8" box="[245,435,1538,1562]" italics="true" pageId="9" pageNumber="8">
|
||
<taxonomicName id="64FC20A0FFE4FFCCFF04BA3CFE42F9D8" box="[245,427,1538,1562]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="cordiger" subSpecies="wahlbergi">G. c. wahlbergi</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
<bibRefCitation id="C76D26D2FFE4FFCCFE48BA3CFD63F9D8" author="Basilewsky, P." box="[441,650,1538,1562]" pageId="9" pageNumber="8" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky 1977</bibRefCitation>
|
||
). As discussed above under
|
||
<taxonomicName id="64FC20A0FFE4FFCCFC2BBA3CFB88F9D8" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[986,1121,1538,1562]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cordiger">
|
||
<emphasis id="91888731FFE4FFCCFC2BBA3CFB88F9D8" box="[986,1121,1538,1562]" italics="true" pageId="9" pageNumber="8">G. cordiger</emphasis>
|
||
</taxonomicName>
|
||
, many of these difficulties can be resolved by dividing this single polytypic taxon into two distinct species, one with a rounded, lobe-shaped median patch of pubescence on the elytra, the other with a transverse, rectangular median patch of pubescence on the elytra. The observed areas of overlap between the subspecies always involve a form with a lobe-shaped pubescence patch on the elytra (
|
||
<emphasis id="91888731FFE4FFCCFC67BABCFEC5F978" italics="true" pageId="9" pageNumber="8">
|
||
G. c. subhamatus, G. c.
|
||
<taxonomicName id="64FC20A0FFE4FFCCFB4ABABCFAB4F958" authorityName="Peringuey" authorityYear="1892" box="[1211,1373,1666,1690]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="zambezianus">zambezianus</taxonomicName>
|
||
, G. c.
|
||
<taxonomicName id="64FC20A0FFE4FFCCFF31BA9CFEC5F978" authorityName="Boheman" authorityYear="1848" box="[192,300,1698,1722]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="hamatus">hamatus</taxonomicName>
|
||
</emphasis>
|
||
) and a form with a transverse pubescence patch on the elytra (
|
||
<emphasis id="91888731FFE4FFCCFBB4BA9CFEEDF918" italics="true" pageId="9" pageNumber="8">
|
||
G. c. betshuana, G. c.
|
||
<taxonomicName id="64FC20A0FFE4FFCCFAA7BA9CFEEDF918" authorityName="Peringuey" authorityYear="1896" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="transfugus">transfugus</taxonomicName>
|
||
</emphasis>
|
||
). The forms with identical patterns of pubescence can be interpreted as disjunct forms of either
|
||
<taxonomicName id="64FC20A0FFE4FFCCFF31BADCFEA1F938" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[192,328,1762,1786]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cordiger">
|
||
<emphasis id="91888731FFE4FFCCFF31BADCFEA1F938" box="[192,328,1762,1786]" italics="true" pageId="9" pageNumber="8">G. cordiger</emphasis>
|
||
</taxonomicName>
|
||
(in the case of
|
||
<emphasis id="91888731FFE4FFCCFDF2BADCFD4FF938" box="[515,678,1762,1786]" italics="true" pageId="9" pageNumber="8">
|
||
G. c.
|
||
<taxonomicName id="64FC20A0FFE4FFCCFDB3BADCFD4FF938" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[578,678,1762,1786]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cordiger">cordiger</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="91888731FFE4FFCCFD15BADCFC5BF938" box="[740,946,1762,1786]" italics="true" pageId="9" pageNumber="8">G. c. subcordiger</emphasis>
|
||
and in the case of
|
||
<emphasis id="91888731FFE4FFCCFB6ABADCFAAFF938" box="[1179,1350,1762,1786]" italics="true" pageId="9" pageNumber="8">
|
||
G. c.
|
||
<taxonomicName id="64FC20A0FFE4FFCCFB2BBADCFAAFF938" authorityName="Boheman" authorityYear="1848" box="[1242,1350,1762,1786]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="hamatus">hamatus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="91888731FFE4FFCCFA75BADCFE9BF8D8" italics="true" pageId="9" pageNumber="8">G. c. subhamatus</emphasis>
|
||
) or
|
||
<taxonomicName id="64FC20A0FFE4FFCCFE52BB3CFDABF8D8" box="[419,578,1794,1818]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="wahlbergi">
|
||
<emphasis id="91888731FFE4FFCCFE52BB3CFDABF8D8" box="[419,578,1794,1818]" italics="true" pageId="9" pageNumber="8">G. wahlbergi</emphasis>
|
||
</taxonomicName>
|
||
(in the case of
|
||
<emphasis id="91888731FFE4FFCCFD0DBB3CFC53F8D8" box="[764,954,1794,1818]" italics="true" pageId="9" pageNumber="8">G. c. betshuana</emphasis>
|
||
and
|
||
<taxonomicName id="64FC20A0FFE4FFCCFC09BB3CFB58F8D8" box="[1016,1201,1794,1818]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="cordiger" subSpecies="wahlbergi">
|
||
<emphasis id="91888731FFE4FFCCFC09BB3CFB58F8D8" box="[1016,1201,1794,1818]" italics="true" pageId="9" pageNumber="8">G. c. wahlbergi</emphasis>
|
||
</taxonomicName>
|
||
). It is possible that even more taxa will be split from
|
||
<taxonomicName id="64FC20A0FFE4FFCCFD93BB1CFD03F8F8" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[610,746,1826,1850]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cordiger">
|
||
<emphasis id="91888731FFE4FFCCFD93BB1CFD03F8F8" box="[610,746,1826,1850]" italics="true" pageId="9" pageNumber="8">G. cordiger</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="64FC20A0FFE4FFCCFCD8BB1CFC21F8F8" box="[809,968,1826,1850]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="wahlbergi">
|
||
<emphasis id="91888731FFE4FFCCFCD8BB1CFC21F8F8" box="[809,968,1826,1850]" italics="true" pageId="9" pageNumber="8">G. wahlbergi</emphasis>
|
||
</taxonomicName>
|
||
when additional material is available for study and the boundaries between the different geographic forms are studied in more detail.
|
||
</paragraph>
|
||
<paragraph id="A3435B23FFE7FFCFFF31BCFCFC93FEFB" blockId="10.[192,1440,194,314]" pageId="10" pageNumber="9">
|
||
<taxonomicName id="64FC20A0FFE7FFCFFF31BCFCFE01FF18" box="[192,488,194,218]" class="Insecta" family="Carabidae" genus="Graphipterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="wahlbergi">
|
||
<emphasis id="91888731FFE7FFCFFF31BCFCFE01FF18" box="[192,488,194,218]" italics="true" pageId="10" pageNumber="9">Graphipterus wahlbergi</emphasis>
|
||
</taxonomicName>
|
||
is generally distributed throughout the central and northern portions of the
|
||
<collectingCountry id="DBEB1BB3FFE7FFCFFF31BCDCFE18FF38" box="[192,497,226,250]" name="South Africa" pageId="10" pageNumber="9">Republic of South Africa</collectingCountry>
|
||
, where it can be locally abundant (
|
||
<bibRefCitation id="C76D26D2FFE7FFCFFC53BCDCFB4BFF38" author="Basilewsky, P." box="[930,1186,226,250]" pageId="10" pageNumber="9" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky 1977: 82</bibRefCitation>
|
||
). Populations of this species extend into adjacent portions of
|
||
<collectingCountry id="DBEB1BB3FFE7FFCFFD52BD3CFCF4FEDB" box="[675,797,258,281]" name="Botswana" pageId="10" pageNumber="9">Botswana</collectingCountry>
|
||
and
|
||
<collectingCountry id="DBEB1BB3FFE7FFCFFCA6BD3CFC3FFED8" box="[855,982,258,282]" name="Zimbabwe" pageId="10" pageNumber="9">Zimbabwe</collectingCountry>
|
||
(
|
||
<bibRefCitation id="C76D26D2FFE7FFCFFC15BD3CFB5FFED8" author="Basilewsky, P." box="[996,1206,258,282]" pageId="10" pageNumber="9" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky 1977</bibRefCitation>
|
||
: Carte 9). For a full synonymy for this species, see
|
||
<bibRefCitation id="C76D26D2FFE7FFCFFDC8BD1CFC9CFEFB" author="Basilewsky, P." box="[569,885,289,314]" pageId="10" pageNumber="9" pagination="1 - 472" refId="ref7374" refString="Basilewsky, P. 1977. Revision du genre Graphipterus Latreille (Coleoptera Carabidae). Musee Royal de l'Afrique Centrale, Annales, Serie in- 8 o, Sciences Zoologiques 221: 1 - 472." type="journal article" year="1977">Basilewsky (1977: 76–77)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |