449 lines
35 KiB
XML
449 lines
35 KiB
XML
<document id="0EF241B7DB28C7D4043E297869D0B1DE" ID-DOI="10.3897/phytokeys.250.136139" ID-publisher-id="136139" URI-arpha="163F7496-5C12-5455-AB48-30AD78073DF0" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1734707767354" checkinUser="pensoft" docAuthor="Sukhorukov, Alexander P., Kushunina, Maria, Nilova, Maya V., Baider, Cláudia & Sennikov, Alexander N." docDate="2024" docId="2ADB2E2400FF5EDDB9503A40C7694E01" docLanguage="en" docName="PhytoKeys 250: 21-94" docOrigin="PhytoKeys 250" docSource="https://phytokeys.pensoft.net/article/136139/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Achyranthes abyssinica Nees" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="94" masterDocId="163F74965C125455AB4830AD78073DF0" masterDocTitle="Africa and Arabia encompass a much greater species diversity in the Achyranthes aspera aggregate (Amaranthaceae, achyranthoid clade): Evidence from morphological and chorological data" masterLastPageNumber="94" masterPageNumber="21" pageNumber="21" updateTime="1734739432256" updateUser="admin">
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<mods:titleInfo id="DA9BD29396853A6BBE3850E63FB90C0E">
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<mods:title id="DEBE0713A11BA627E55A3E8F70DB895C">Africa and Arabia encompass a much greater species diversity in the Achyranthes aspera aggregate (Amaranthaceae, achyranthoid clade): Evidence from morphological and chorological data</mods:title>
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<mods:name id="5708E99A6B9140B25961AA65BA4CDE44" type="personal">
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<mods:namePart id="60E6FA30223F295A5447E74C86B38A78">Sukhorukov, Alexander P.</mods:namePart>
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<mods:nameIdentifier id="0A10DBB16EFE524AE8F5A26AD31997A6" type="ORCID">0000-0003-2220-826X</mods:nameIdentifier>
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<mods:affiliation id="D5D4D9E69F8A9849ED9272B6939AE559">Department of Higher Plants, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia & Laboratory Herbarium (TK), Tomsk State University, Lenin Ave. 36, 634050, Tomsk, Russia</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="C5CDE440DC5A8D5D422729417DE8105D">Kushunina, Maria</mods:namePart>
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<mods:nameIdentifier id="C308D6133B56331B993C218490065BC5" type="ORCID">0000-0003-2541-6494</mods:nameIdentifier>
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<mods:affiliation id="6CEEBE9A967A0A3B72DC38D1EF8E6E6D">Department of Higher Plants, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="5B255DC77E5ECB5D34B7E0A6BBBEAFF0" type="personal">
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<mods:roleTerm id="741167F4FEAB0903710DB6A725441C1C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D7CD09543BE8B33C089D68CA7048D49A">Nilova, Maya V.</mods:namePart>
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<mods:nameIdentifier id="A56B57E9A0BB0E0F641E6E7FA203AA46" type="ORCID">0000-0001-5340-5915</mods:nameIdentifier>
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<mods:affiliation id="72896A5CE13741146F7EE7AF517B4790">Department of Higher Plants, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="65F17D3EB3A37A5B54338B8A2D872EAF" type="personal">
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<mods:roleTerm id="83A52180867EAD9DE6AF2587B7117CB2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5A1F0E6E73DB62AD1D2E13783586AB3E">Baider, Cláudia</mods:namePart>
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<mods:nameIdentifier id="2771950D5EECBDAE8E75030CBE90C12C" type="ORCID">0000-0002-2203-2076</mods:nameIdentifier>
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<mods:affiliation id="863354A46B596819DCB00DF1101059C1">Department of Plant Physiology, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="4D51702661C4DE32B322777062008F82" type="personal">
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<mods:roleTerm id="DA017D393EE1368E49FFC924BC276E32">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="750414FA8A3BD2643E760817208BD08D">Sennikov, Alexander N.</mods:namePart>
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<mods:nameIdentifier id="B72E0156B1166330D9C0257FB5BFEFFA" type="ORCID">0000-0001-6664-7657</mods:nameIdentifier>
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<mods:affiliation id="1A65BB53CA4957955E1A55F56A8CFCCC">The Mauritius Herbarium, RE Vaughan Building, Agricultural Services, Ministry of Agro-Industry and Food Security, Réduit, 80835, Mauritius</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="9F2D57E223481F3121FB670AA9FAA694">text</mods:typeOfResource>
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<mods:titleInfo id="813116770BAAD346DF75332D7F757C08">
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<mods:title id="44352BA30C033661DDC2FACE45F458B1">PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="A70D2DF9CE1B0708661E346416833F8B">
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<mods:date id="39B6B06362CB116002F9F8922FAC7CEF">2024</mods:date>
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<mods:detail id="0572BDE536AAF18D09E375D62DE672FF" type="pubDate">
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<mods:number id="6B2845BEAFCE6A7200410119846B174D">2024-12-20</mods:number>
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</mods:detail>
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<mods:detail id="5FB78ECADD42A6598F3A256D1EE6A5F5" type="volume">
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<mods:number id="AAAE22BD1F03EDF2404C62187CDA0668">250</mods:number>
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</mods:detail>
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<mods:extent id="282168FE03719BA01C74415E11D1EB56" unit="page">
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<mods:start id="B27F98297589321EEFEB95AD37DCAC88">21</mods:start>
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<mods:end id="45B1BC84BD5B86DD64D4F6218BD8368D">94</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="20D750F1C28AE1B45F6706741D803180">journal article</mods:classification>
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<mods:identifier id="28764F2776CFCCBA066BB1E27757AD5A" type="DOI">10.3897/phytokeys.250.136139</mods:identifier>
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</mods:mods>
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<treatment id="2ADB2E2400FF5EDDB9503A40C7694E01" ID-arpha="2ADB2E24-00FF-5EDD-B950-3A40C7694E01" LSID="urn:lsid:plazi:treatment:2ADB2E2400FF5EDDB9503A40C7694E01" httpUri="http://treatment.plazi.org/id/2ADB2E2400FF5EDDB9503A40C7694E01" scope_family="Amaranthaceae">
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||
<subSubSection id="D169A893FEC97237E4240C53ADE962A8" type="nomenclature">
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||
<paragraph id="6677E250FCC3363BC60300F13B30C41A">
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||
<taxonomicName id="936478B025D39F2DE99E2C7E1F53D9A7" authority="Nees" authorityName="Nees" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="abyssinica">
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<emphasis id="67BD8F06C85E29B3AA709C886437612B" italics="true">Achyranthes abyssinica</emphasis>
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||
Nees
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</taxonomicName>
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||
, Del. Sem. Hort. Vratisl. 1850: 3 (1851).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F1BFFE2B176DE04F96BC4B9D3DF7AD6B" type="reference_group">
|
||
<paragraph id="C71FBDE72DE943D5DF34CD9DC9559E2F">
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||
<treatmentCitationGroup id="5F6147073A23D6AF1001EBD1C17C4C5F">
|
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=
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<treatmentCitation id="46E745F5686B6CDD22C3F91FE74CC2D2" author="Suess." journal="Bull. Jard. Bot. Etat Bruxelles" page="56" score="291.0324320782438" volume="15" year="1938">
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<taxonomicName id="A04883119C2B8059993DADE01BB3FEC4" ID-CoL="9ZZHY" authority="Suess., Bull. Jard. Bot. Etat Bruxelles" class="Magnoliopsida" family="Amaranthaceae" form="rubella" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="form" species="aspera">
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<emphasis id="775A8B0061BA51BDDF96963773963C1C" italics="true">Achyranthes aspera f. rubella</emphasis>
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Suess.
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</taxonomicName>
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||
, Bull. Jard. Bot. État Bruxelles 15: 56 (1938)
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||
</treatmentCitation>
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.
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<typeStatus id="95781B1DADFB5A996F2171793927B966" type="lectotype">Lectotype</typeStatus>
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||
(designated here):
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||
<collectingCountry id="9A86B72DDCF2B30F43F11571059F1A0D" name="Democratic Republic of the Congo">DR Congo</collectingCountry>
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, [South Kivu Prov.,]
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<location id="527FE0834308BFE7502590D8A0549E38" LSID="urn:lsid:plazi:treatment:2ADB2E2400FF5EDDB9503A40C7694E01:527FE0834308BFE7502590D8A0549E38" country="Democratic Republic of the Congo" municipality="Lejeune" name="Kizozi">Kizozi</location>
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, 1935,
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<emphasis id="9B7FECDAAFB3035A1C2E23D35B85D74D" italics="true">
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<collectingMunicipality id="C48CAE80E2D5E5F24D2149356212FF22">Lejeune</collectingMunicipality>
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190
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||
</emphasis>
|
||
(
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||
<specimenCode id="E0B72B06183A291A9B1A463408B83179" collectionCode="BR" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR 0000008819512</specimenCode>
|
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!,
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<typeStatus id="FADEA9EB9F5DB5032A5CABFD616CF501" type="isolectotype">isolectotype</typeStatus>
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||
<specimenCode id="6C241A59C979EA7ED51356FE45C4397B" collectionCode="BR" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR 0000008819192</specimenCode>
|
||
!).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7B72934C3F525229B3753064DAC30C86">
|
||
<treatmentCitationGroup id="1EF689F9D1D7301E966B7CD5FDB42A58">
|
||
=
|
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<taxonomicName id="4B300EC6FFF0E98806A629407E6327AE" authority="Suess." authorityName="Suess." class="Magnoliopsida" family="Amaranthaceae" form="latifolia" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="form" species="aspera" variety="sicula">
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<emphasis id="59928AAF60A33907F0E2E47835C784C8" italics="true">Achyranthes aspera</emphasis>
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[var. sicula]
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||
<treatmentCitation id="F0C2183AC51FFA33A0323F91498BE9FA" author="Suess." journal="Mitt. Bot. Staatssamml. Munchen" page="70" score="385.49535180271243" volume="1" year="1951">f. latifolia Suess., Mitt. Bot. Staatssamml. München 1: 70 (1951)</treatmentCitation>
|
||
</taxonomicName>
|
||
.
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||
<typeStatus id="226B920C22305A4B6BC3ED025BBAEB64" type="holotype">Holotype</typeStatus>
|
||
:
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||
<collectingCountry id="A3C37BD60A6F337AF471D86D927644D3" name="Kenya">Kenya</collectingCountry>
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||
, pr. Forest Station in silva montana,
|
||
<quantity id="4FEF74E59FA488921973D6E83AB12A65" metricMagnitude="3" metricUnit="m" metricValue="2.35" unit="m" value="2350.0">2350 m</quantity>
|
||
,
|
||
<date id="2DE62CD4759BEDE9A8FE9085C1F6B001" value="1922-01-05">5 January 1922</date>
|
||
,
|
||
<emphasis id="EA03F2EA2861B1F6AD395A3AEC7B2B85" italics="true">R. E. Fries & Th. C. E. Fries 781</emphasis>
|
||
(
|
||
<collectionCode id="E107AE3C78154E559D8769C5C3742690" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
|
||
000243724!,
|
||
<typeStatus id="C0A533B1923F3F80CA053291D5A3BD48" type="isotype">isotypes</typeStatus>
|
||
<collectionCode id="6251C68BE435E79FC4E5C53B98C20920" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
0000008819922!, S 07-12310 – image!).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="1E2B514E6A09D4228092CB55896AF076">
|
||
<treatmentCitationGroup id="E35D0930491F5835C286F470D7B8B62E">
|
||
=
|
||
<treatmentCitation id="35D450F2E993615CAD34E5711E455A50" author="Suess." journal="Mitt. Bot. Staatssamml. Munchen" page="70" score="385.49535180271243" volume="2" year="1955">
|
||
<taxonomicName id="679A805AA3B4B8CC1969B1E5E0D4F100" ID-CoL="B298Z" authority="Suess." authorityName="Suess." class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="argentea" variety="albissima">
|
||
<emphasis id="24F3D09243EA241B0F00C87C65CC7670" italics="true">Achyranthes argentea var. albissima</emphasis>
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||
Suess.
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||
</taxonomicName>
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||
, Mitt. Bot. Staatssamml. München 2: 70 (1955)
|
||
</treatmentCitation>
|
||
.
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||
<typeStatus id="C4FDB8205C6A1EA9AC6B36378B09893E" type="lectotype">Lectotype</typeStatus>
|
||
(designated here):
|
||
<collectingCountry id="840DDBE2886562062AA84AF2E0680582" name="Kenya">Kenya</collectingCountry>
|
||
, Northern Prov., Furroli [Mount], Mt. top: granite with
|
||
<taxonomicName id="D2D5ABE76ACB9310D2FFD116AAA8B3BB" ID-CoL="8VZV4" class="Magnoliopsida" family="Oleaceae" genus="Olea" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="BF57438505B182B4404B0F548ECCFA95" italics="true">Olea</emphasis>
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||
</taxonomicName>
|
||
&
|
||
<taxonomicName id="60ABDF73161CAB5B60B94E04846B0C58" ID-CoL="56YL" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="7A95E759673590C17E98269D7FCE27AE" italics="true">Juniperus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="C3312BCD22A50361D8B367B12E568E82" metricMagnitude="3" metricUnit="m" metricValue="2.0116799999999997" unit="ft" value="6600.0">6600 ft</quantity>
|
||
, occasional,
|
||
<date id="C2ECC7749B79E333630ECD0FA2FF8848" value="1952-09-20">20 September 1952</date>
|
||
,
|
||
<emphasis id="3BC28F22A434882E1FC467E2B89DD429" italics="true">J. B. Gillett 13957</emphasis>
|
||
(
|
||
<collectionCode id="A49147334747C4079A55EAB7ACDAAAF7" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
|
||
000243722!,
|
||
<typeStatus id="E4CFE42600C9D18241486DF8F51BAB62" type="isolectotype">isolectotype</typeStatus>
|
||
<collectionCode id="782648E85EF63CCA8A0835004AAB0B1D" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
|
||
000243723!).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="1E1F9835CA7236EB0933BD8946E5C442">
|
||
<treatmentCitationGroup id="E1BCE7F78F5F38E7A36984466689C9DE">
|
||
–
|
||
<treatmentCitation id="74847191FF1B9ADA128E1E02D51FD6FD">
|
||
<taxonomicName id="BD35CF5D0707F55D4509B375AA243D15" ID-CoL="99CH" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="sicula">
|
||
<emphasis id="FD48DED843D8566F7037BAE82C1672DC" italics="true">Achyranthes sicula</emphasis>
|
||
</taxonomicName>
|
||
auct. div.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E1IBI" type="Neotype (designated here)">
|
||
<paragraph id="483553BDF2AC8BD42E7E7DEB1B24934E">
|
||
<heading id="4B797800C88A052D8305556788E8EBCB" reason="title">
|
||
<typeStatus id="CC25F7C7C1D5E55B6CF781846F64D615" type="neotype">Neotype</typeStatus>
|
||
(designated here).
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="55BCF869F617970D165D8468AE396D57">
|
||
<materialsCitation id="943757B19D25B5425B684139C551BC54" collectingDate="1838-11-12" collectionCode="BM, FI, LE, WU" collectorName="Schimper" country="Ethiopia" location="Soloda Mt. near Adwa Town" municipality="Scholoda" specimenCode="G 00688998, BR 00000083356949, K 005771057, M 0241524" specimenCount="1" stateProvince="Tigray Region" typeStatus="neotype">
|
||
<collectingCountry id="010320FE019F7F7B630CA079DA7D3C3F" name="Ethiopia">Ethiopia</collectingCountry>
|
||
, [
|
||
<collectingRegion id="8A2F91B8EFB93F27DAC40C527E1D3ADE" country="Ethiopia" name="Tigray">Tigray Region</collectingRegion>
|
||
,]
|
||
<collectingMunicipality id="9C02AFC2173BE97B803DE888834ED661">Scholoda</collectingMunicipality>
|
||
[
|
||
<location id="B6A1819F6E19DE85AE603B4D32CA4B68" LSID="urn:lsid:plazi:treatment:2ADB2E2400FF5EDDB9503A40C7694E01:B6A1819F6E19DE85AE603B4D32CA4B68" country="Ethiopia" municipality="Scholoda" name="Soloda Mt. near Adwa Town" stateProvince="Tigray Region">Soloda Mt. near Adwa Town</location>
|
||
],
|
||
<date id="E837A2F77EDE4A95787F6D1AF7FAC2C5" value="1838-11-12">
|
||
<collectingDate id="6D75255AACDA9FBC03000BDE64074708" value="1838-11-12">12 November 1838</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="4A5C63E95879C25AE558BBF4B57CC340" italics="true">
|
||
<collectorName id="9129F64DDDD8BC5AB52D51AAB2B1AB1D">Schimper</collectorName>
|
||
1144
|
||
</emphasis>
|
||
[Schimper, Iter Abyssinicum, ser. 2 (1842)] (
|
||
<collectionCode id="F57A7D80F3311D2AB5C34029CF780032" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
|
||
!,
|
||
<typeStatus id="04B77D89ED378DD73D91D489F2F233C7" type="isoneotype">isoneotypes</typeStatus>
|
||
<specimenCode id="000FFE2BB7EB7D925DDF94525975D793" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G 00688998</specimenCode>
|
||
,
|
||
<specimenCode id="2C08AE9BC2456AEE86A9500345CD2C59" collectionCode="BR" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR 00000083356949</specimenCode>
|
||
!,
|
||
<collectionCode id="F2C8F81F49BC059452BCAC924C3BCAF8" country="Italy" name="Natural History Museum" type="Herbarium">FI</collectionCode>
|
||
!,
|
||
<specimenCode id="6569C07B955ABD4AE3BAE41F4AA72AA2" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K 005771057</specimenCode>
|
||
! – central twig,
|
||
<collectionCode id="C0C6294F642BD55C083316FB5B4ABB3C" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
|
||
!,
|
||
<specimenCode id="D7BCE684380ED5A955CE54A1CE9FDEAE" collectionCode="M" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M 0241524</specimenCode>
|
||
!,
|
||
<collectionCode id="BEC74768689920959270E04C1F1CB6B8" country="USA" name="Wayland University">WU</collectionCode>
|
||
!).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ECJBI" type="description">
|
||
<paragraph id="A319F5B86A92D9C3B91D675068668944">
|
||
<heading id="A839B709455DAEC3587C6DE4FAAF639B" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="E3171FCB2988996DD2C59C908E4E7514">
|
||
(Fig.
|
||
<figureCitation id="512B97E46AAF59B0B04450B8BECB96A2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Achyranthes abyssinica A an indoor cultivated plant in vegetative stage (grown from seeds from Arusha Region, Tanzania) B flowering twig in bushy vegetation in Meru area, Arusha Region, Tanzania. Photographer: A. Sukhorukov." figureDoi="10.3897/phytokeys.250.136139.figure3" httpUri="https://binary.pensoft.net/fig/1203847">3</figureCitation>
|
||
). Annual, stout or rarely delicate herb up to 1.0 (1.5–2.0) m, not rooting at nodes, very rarely creeping at lower nodes; stem four-angled, sparsely pubescent, sometimes scrambling; lateral branches long and ± horizontally spreading; leaf petioles
|
||
<quantity id="E543B0ECAB5F323ED9417ACC1A195C4B" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" unit="mm" value="17.5" valueMax="25.0" valueMin="10.0">10–25 mm</quantity>
|
||
long, blades 20–100 (120) × 10–40 (50) mm, broadly cuneate or truncate, ovate, entire, tip acuminate, bicolored: ± green, slightly to moderately pubescent above, whitish or grey below (sparsely pubescent and green when cultivated indoor); inflorescence
|
||
<quantity id="A32CFFC0F30170AB78578463679BAE6F" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" unit="mm" value="150.0" valueMax="250.0" valueMin="50.0">50–250 mm</quantity>
|
||
long, sometimes pendulous; bract ~ 2.0 mm long; bracteoles 2.5–3.0 mm long, reflexed; perianth (3.5) 4.0–4.5 (5.0) mm long, green or purplish inside; pseudostaminodes 0.6–1.0 mm long, short, white or pink, fimbriate or not; stamens 5, with pink filaments, anthers
|
||
<quantity id="C434C707D0804021699EAFEE9CC180DA" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.4–0.7 mm</quantity>
|
||
long, pink or magenta; style (1.3) 1.8–2.5 (3.0) mm long, pink; fruit (without style) (1.3) 1.8–2.5 (3.0) mm long.
|
||
</paragraph>
|
||
<caption id="BC53A04358053B97DDD6D32026BCADA6" ID-DOI="10.3897/phytokeys.250.136139.figure3" ID-arpha="4A52B1C8-1D22-5CDB-BC85-3DADA01AA9D5" httpUri="https://binary.pensoft.net/fig/1203847" startId="F3">
|
||
<paragraph id="5A9A5BB399ED313476CF50BC9262C220">
|
||
<label id="995CDA97FE2AA0466AEF6B6D0BE56429">Figure 3.</label>
|
||
</paragraph>
|
||
<paragraph id="676C716F30ED96A4326C0A4805A8E97B">
|
||
<taxonomicName id="BB7DA22F231BF7F658ECB9AAA9D8563A" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="abyssinica">
|
||
<emphasis id="5A38E790E279E734E907138C5853E1A4" italics="true">Achyranthes abyssinica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="13AF7BA683EE997A7CA59CB8339534A9" bold="true">A</emphasis>
|
||
an indoor cultivated plant in vegetative stage (grown from seeds from Arusha Region, Tanzania)
|
||
<emphasis id="FA4B51C2E21720AA14D9ECF0670BE8F7" bold="true">B</emphasis>
|
||
flowering twig in bushy vegetation in Meru area, Arusha Region, Tanzania. Photographer: A. Sukhorukov.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EOKBI" type="Taxonomic notes">
|
||
<paragraph id="221A4A71541B43CEFE820D6E622C40AE">
|
||
<heading id="E9D9C7200B12FB2AC37A14B876ED0D5B" reason="title">Taxonomic notes.</heading>
|
||
</paragraph>
|
||
<paragraph id="7753EAAD44E2577DA59703544B227806">
|
||
Morphologically,
|
||
<taxonomicName id="3846E84C37A918B345A85043AB432F60" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="abyssinica">
|
||
<emphasis id="28A822544CDCD336C5976E280AFE47DB" italics="true">A. abyssinica</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="71534DC04831C4CCCE1ACE4A85792DE3" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="sicula">
|
||
<emphasis id="E2958BDB2E672DFE6FF9C978CF0BBD1C" italics="true">A. sicula</emphasis>
|
||
</taxonomicName>
|
||
but differs from the latter by a slightly longer perianth and style, although a few specimens from tropical Africa have the same style length as in
|
||
<taxonomicName id="5BA6324613FCF72EF577A431D6BD769D" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="sicula">
|
||
<emphasis id="8F4C88A1C03289894A2CA358CC0A146C" italics="true">A. sicula</emphasis>
|
||
</taxonomicName>
|
||
. Although these two species are very similar in morphology, as also shown in our identification key, they take different positions in the molecular phylogeny (unpublished). Both species have clearly different and non-overlapping distributions, with
|
||
<taxonomicName id="BABCC2E8CAAC4D98F6AEA1673CC871E6" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="abyssinica">
|
||
<emphasis id="0620E71EDB734C8521384242435A6F18" italics="true">A. abyssinica</emphasis>
|
||
</taxonomicName>
|
||
growing in mountainous areas of East and Central Africa and
|
||
<collectingCountry id="EDDDF01E51BBECEB7E77B02EF17D218F" name="Yemen">South Arabia</collectingCountry>
|
||
, whereas
|
||
<taxonomicName id="3E53260B65505A5B838EFED8D443754D" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="sicula">
|
||
<emphasis id="A86742630B0C56335D82C92DFA78CB75" italics="true">A. sicula</emphasis>
|
||
</taxonomicName>
|
||
is a subtropical species found in areas with a Mediterranean climate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ELMBI" type="Nomenclatural notes">
|
||
<paragraph id="236E7640E6FA8D4923716B362A57339C">
|
||
<heading id="82ED38FA9C9E5399916026794042F3E6" reason="title">Nomenclatural notes.</heading>
|
||
</paragraph>
|
||
<paragraph id="45E130A7D0B3D7761725F1F87AA99369">
|
||
Unfortunately, the whereabouts of the type specimens of
|
||
<taxonomicName id="2CEE852C05639BC1B8DE0FCA6915AB27" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="abyssinica">
|
||
<emphasis id="13DA79EA139A4CE63C2D164342E4971A" italics="true">Achyranthes abyssinica</emphasis>
|
||
</taxonomicName>
|
||
in the Nees herbarium are mostly unknown because this collection had been split into numerous parts, which were sold to institutions or private individuals (
|
||
<bibRefCitation id="0F7383AF465DB6273BBD737CC4D966D7" author="Stafleu and Cowan" firstAuthor="Stafleu" refId="B106" refString="Stafleu FA, Cowan RS (1981) Taxonomic literature. A selective guide to botanical publications and collections with dates, commentaries and types, vol. 3. Bohn, Scheltema & Holkema, Utrecht; dr. W. Junk b. v., Publishers, The Hague, [xii +] 980 pp. [Regnum vegetabile 105]" year="1981">Stafleu and Cowan 1981</bibRefCitation>
|
||
). The contents of these parts are not always certain; besides, the collection was heavily affected by extensive losses in German herbaria during the Second World War. Our request to Bonn, where these type specimens were possibly kept, was unsuccessful because that Herbarium was completely destroyed during the war (Thomas Joßberger, pers. comm.). There are also no authentic specimens in
|
||
<collectionCode id="92835096EF344D6C7599922C435E945E" country="France" httpUri="http://grbio.org/cool/jm1p-44gp" name="Institut de Botanique" type="Herbarium">STR</collectionCode>
|
||
(Gisele Haan-Archipoff, pers. comm.) and
|
||
<collectionCode id="268F5C88086D3BF27995B3E604AFC17F" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
|
||
(Sukhorukov, pers. obs.), where large parts of Nees collections are housed.
|
||
</paragraph>
|
||
<paragraph id="51EE05F7FA2036C17B33D10B1864ACE2">
|
||
In the absence of any original material, we designate Schimper’s collection from
|
||
<collectingCountry id="5CDD46B99E9728CD21DBC0D713CFA59B" name="Ethiopia">Ethiopia</collectingCountry>
|
||
, which was prepared in numerous duplicates and commercially distributed to many Herbaria under the name
|
||
<taxonomicName id="0C391B2784DB32FBB21E899A8DAA0DB1" authorityName="Lam. (Schweinfurth" authorityYear="1867" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="argentea">
|
||
<emphasis id="CA385B7A76C2E5B59378F0D56D943C5E" italics="true">A. argentea</emphasis>
|
||
</taxonomicName>
|
||
, as the
|
||
<typeStatus id="965D677E80B738C62EB380D22BD9F453" type="neotype">neotype</typeStatus>
|
||
of
|
||
<taxonomicName id="2CFD19EE6529812BC37B7AC358DAD0B4" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="abyssinica">
|
||
<emphasis id="2CD36C196D3BAF4C6D7EC5EFAB55BA4C" italics="true">A. abyssinica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="683454B27FF4CA701D25EF17CF5E26F4">
|
||
The label of the
|
||
<typeStatus id="253DA1FD8BF90439F1BED680D1F7FB3B" type="isolectotype">isolectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="BE46224BD34E6E97064ABD58B0CAF020" authority="Suess." authorityName="Suess." class="Magnoliopsida" family="Amaranthaceae" form="rubella" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="form" species="aspera">
|
||
<emphasis id="34A63AEAD71E46C2FA7707FB85621953" italics="true">A. aspera f. rubella</emphasis>
|
||
Suess.
|
||
</taxonomicName>
|
||
gives no exact location information, but both specimens belong to a single gathering (
|
||
<emphasis id="7FD336D9657341F1A6D4C19545C5E056" italics="true">Lejeune 190</emphasis>
|
||
). No other specimens of the original material are known, and the type choice is therefore rather formal.
|
||
</paragraph>
|
||
<paragraph id="F0077E6698674BFFE4D8B4F5C03994F0">
|
||
Suessenguth in
|
||
<bibRefCitation id="021F79634FD2BE64C1BD95D3465E71FA" author="Suessenguth K & Merxmüller H" firstAuthor="Suessenguth" journalOrPublisher="Mitteilungen der Botanischen Staatssammlung München" pagination="69-94" refId="B111" refString="Suessenguth K, Merxmüller H (1951) Species et varietates novae vel rarae in Africa australi et centrali lectae. Mitteilungen der Botanischen Staatssammlung München 1: 69–94." title="Species et varietates novae vel rarae in Africa australi et centrali lectae." volume="1" year="1951">Suessenguth and Merxmüller (1951)</bibRefCitation>
|
||
cited the type of
|
||
<taxonomicName id="711D2AFEC4B1624223A99519E84BB994" authority="Suess." authorityName="Suess." class="Magnoliopsida" family="Amaranthaceae" form="latifolia" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="form" species="aspera">
|
||
<emphasis id="23A21EFFE77D29E46C6500158B04C648" italics="true">A. aspera f. latifolia</emphasis>
|
||
Suess.
|
||
</taxonomicName>
|
||
as kept at Kew. The duplicates in
|
||
<collectingRegion id="624D3BD0ADD9074C108CB5E7B0304CD9" country="Serbia" name="Bor">BR</collectingRegion>
|
||
and S are, therefore,
|
||
<typeStatus id="8A83205AC001276239AB04A82BF6475D" type="isotype">isotypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="19BC964A7065DE568BBC4808AD466CF5">
|
||
<taxonomicName id="80E91465911A83B0602551B37DFE9D58" authority="Suess." authorityName="Suess." class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="argentea" variety="albissima">
|
||
<emphasis id="377037A50353BC36854CFB3698F457D9" italics="true">Achyranthes argentea var. albissima</emphasis>
|
||
Suess.
|
||
</taxonomicName>
|
||
comprises mountainous plants with small white leaves. Suessenguth in
|
||
<bibRefCitation id="FFCFE48E73864BBA68A9D260081B2CC5" author="Suessenguth K & Merxmüller H" firstAuthor="Suessenguth" journalOrPublisher="Mitteilungen der Botanischen Staatssammlung München" pagination="69-94" refId="B111" refString="Suessenguth K, Merxmüller H (1951) Species et varietates novae vel rarae in Africa australi et centrali lectae. Mitteilungen der Botanischen Staatssammlung München 1: 69–94." title="Species et varietates novae vel rarae in Africa australi et centrali lectae." volume="1" year="1951">Suessenguth and Merxmüller (1951)</bibRefCitation>
|
||
designated
|
||
<specimenCount id="DD958C060C11FEB8650DCC8CFD437F19" count="2" type="generic">two specimens</specimenCount>
|
||
of a single gathering at Kew as “ typus ”. These specimens are
|
||
<typeStatus id="F7719DA9399D7A93932345DAE16733DE" type="syntype">syntypes</typeStatus>
|
||
; a
|
||
<typeStatus id="D0980B4391F938053EAD57EA0C12F6DD" type="lectotype">lectotype</typeStatus>
|
||
is designated here to restrict the typification to a single specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E2PBI" type="habitat">
|
||
<paragraph id="FBE08AD44A03D30E05E74C812F29DA5C">
|
||
<heading id="A9168FE062C68CDDFF65751168646E09" reason="title">Habitat.</heading>
|
||
</paragraph>
|
||
<paragraph id="88EFBBE972629E03A6372C2D131ED7D4">
|
||
Open or slightly shady grassy slopes, bushy vegetation, calcareous outcrops, sometimes as a weed, at elevations of
|
||
<quantity id="00157663F9B6A129DF0EE407815BE052" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="3.5" metricValueMin="1.0" unit="m" value="2250.0" valueMax="3500.0" valueMin="1000.0">1000–3500 m</quantity>
|
||
a. s. l. in arid or seasonally moist areas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EAQBI" type="distribution">
|
||
<paragraph id="1992E0DD921ADEC1B9D58D300268B1A9">
|
||
<heading id="EDFC17FF643E899000D346E1E9129D8E" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="A4180C86A85A4BB4AEDCF3E622A0B99E">
|
||
(Fig.
|
||
<figureCitation id="F19E4471DE44975CEC2E05D1FF7BC621" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution map of Achyranthes abyssinica." figureDoi="10.3897/phytokeys.250.136139.figure4" httpUri="https://binary.pensoft.net/fig/1203848">4</figureCitation>
|
||
; see also Appendix
|
||
<xref id="CE420746C2FBF5CBD6CA85F5453DDFD3" ref-type="app" rid="app1">1</xref>
|
||
).
|
||
<emphasis id="1C1BF9351E20D7BEB73F2585A26BDF89" bold="true">Africa</emphasis>
|
||
:
|
||
<collectingCountry id="708EFC49C2A3592B6C08FE1ACCE9E9BB" name="Angola">Angola</collectingCountry>
|
||
,
|
||
<collectingCountry id="33423B6809C10963F69347D6DDC98F0A" name="Burundi">Burundi</collectingCountry>
|
||
,
|
||
<collectingCountry id="7756106D97E3DEDE6FA4BA1F563FF820" name="Cameroon">Cameroon</collectingCountry>
|
||
,
|
||
<collectingCountry id="835A8FE4856ED45AD5EA7E5BE9E39D1D" name="Democratic Republic of the Congo">DR Congo</collectingCountry>
|
||
,
|
||
<collectingCountry id="342A3A2D0DF8E05A3B078DC7C7FD5C65" name="Eritrea">Eritrea</collectingCountry>
|
||
,
|
||
<collectingCountry id="8FFCDA2409B1948044CDEBC7D147898E" name="Ethiopia">Ethiopia</collectingCountry>
|
||
,
|
||
<collectingCountry id="7273A261DB7559AD574262CA645B9359" name="Guinea-Bissau">Guinea</collectingCountry>
|
||
,
|
||
<collectingCountry id="FC688E75D4910CAB101F93C5D8735ED7" name="Kenya">Kenya</collectingCountry>
|
||
,
|
||
<collectingCountry id="956AAA59AD24C5C1CF9804B4FE7E790A" name="Madagascar">Madagascar</collectingCountry>
|
||
,
|
||
<collectingCountry id="E9FAFD937B31E0A27157400BD5160ABF" name="Malawi">Malawi</collectingCountry>
|
||
,
|
||
<collectingCountry id="46B8FF22483AAB64089CF94E10F538A8" name="Nigeria">Nigeria</collectingCountry>
|
||
,
|
||
<collectingCountry id="0A6F9EC814E10629F79FAE98EA74B546" name="Rwanda">Rwanda</collectingCountry>
|
||
,
|
||
<collectingCountry id="D5F7341F4E410DF293045403987C6849" name="Somalia">Somalia</collectingCountry>
|
||
,
|
||
<collectingCountry id="846E4B24ACE5900DA6C14CD4FD099891" name="South Sudan">South Sudan</collectingCountry>
|
||
,
|
||
<collectingCountry id="FD4B60BC222FC03185315BD3615F7023" name="Sudan">Sudan</collectingCountry>
|
||
,
|
||
<collectingCountry id="9CC85AB9DD2EDFBEE9BD89B097925B61" name="Tanzania">Tanzania</collectingCountry>
|
||
,
|
||
<collectingCountry id="7F0EFA41FBF7D92A262D3DC5D79A05A4" name="Uganda">Uganda</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<caption id="6EE939CF5D39B630B3602820C027ED26" ID-DOI="10.3897/phytokeys.250.136139.figure4" ID-arpha="64FCB577-F14A-590B-A456-07EFEB291ED3" httpUri="https://binary.pensoft.net/fig/1203848" startId="F4">
|
||
<paragraph id="2048347D471397F6A566717B3D660B6B">
|
||
<label id="50A31768D2ED831790C85F070E580DCE">Figure 4.</label>
|
||
</paragraph>
|
||
<paragraph id="6292B17A1EEC93B4D885D8EB9AFEE2BB">
|
||
Distribution map of
|
||
<taxonomicName id="B8118AB12FAFF498933A523EE0F5A9A6" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="abyssinica">
|
||
<emphasis id="70EEEBD5187821D97F9590CA5A31BF93" italics="true">Achyranthes abyssinica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="97F7BE225D86C759CA7D337324D726BE">
|
||
<emphasis id="0236285607D71E1AF919FAD20B15AE76" bold="true">Arabian Peninsula</emphasis>
|
||
:
|
||
<collectingCountry id="CAFDA68833718B777A6B26B574D3875A" name="Oman">Oman</collectingCountry>
|
||
,
|
||
<collectingCountry id="282727D17191CBA7C255BB64E0670933" name="Saudi Arabia">Saudi Arabia</collectingCountry>
|
||
,
|
||
<collectingCountry id="97D359AAB5EC699D445CB1FDDA3BA5B0" name="Yemen">Yemen</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EIRBI" type="distribution">
|
||
<paragraph id="A1E2BD9504CDDBDDBD00A60A46FE6EB9">
|
||
<heading id="AA77064BEEC9421AA5A59E15720E1F93" reason="title">General distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="B9FF0900A264C4D9B3CCC6F36ACD3764">The species is only known from tropical Africa and Arabia.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |