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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.690.13751" ID-GBIF-Dataset="356a4c29-470b-4ca5-8503-76191b4fef6f" ID-PMC="PMC5672587" ID-Pensoft-Pub="1313-2970-690-1" ID-PubMed="29118595" ID-ZBK="C1B8D7C059E54C3A944F69F4FDE96B20" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-690-1" ModsDocOrigin="ZooKeys 690" ModsDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" checkinTime="1502920743207" checkinUser="pensoft" docAuthor="Shpeley, Danny, Hunting, Wesley &amp; Ball, George E." docDate="2017" docId="2AC8839F28679C80910F1B3E1909A023" docLanguage="en" docName="ZooKeys 690: 1-195" docOrigin="ZooKeys 690" docSource="http://dx.doi.org/10.3897/zookeys.690.13751" docTitle="Selenophorus discopunctatus Dejean" docType="treatment" docVersion="5" lastPageNumber="28" masterDocId="FE026465FF86843AE44A7B6EFF8DC422" masterDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" masterLastPageNumber="195" masterPageNumber="1" pageNumber="27" updateTime="1668164690231" updateUser="ExternalLinkService">
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<mods:title>A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography</mods:title>
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<mods:namePart>Shpeley, Danny</mods:namePart>
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<mods:namePart>Hunting, Wesley</mods:namePart>
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<mods:namePart>Ball, George E.</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2017</mods:date>
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<mods:number>690</mods:number>
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<treatment ID-GBIF-Taxon="132785205" LSID="urn:lsid:plazi:treatment:2AC8839F28679C80910F1B3E1909A023" httpUri="http://treatment.plazi.org/id/2AC8839F28679C80910F1B3E1909A023" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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<paragraph pageId="26" pageNumber="27">
<taxonomicName authority="Dejean" class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus discopunctatus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="discopunctatus">Selenophorus discopunctatus Dejean</taxonomicName>
Figs 14A, 15
<normalizedToken originalValue="AC">A-C</normalizedToken>
, 17
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="reference_group">
<paragraph pageId="26" pageNumber="27">
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus discopunctatus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="discopunctatus">Selenophorus discopunctatus</taxonomicName>
Dejean, 1829: 92. 39 specimens in
<normalizedToken originalValue="Chaudoir-Oberthür">Chaudoir-Oberthuer</normalizedToken>
Collection (MNHP) in front of following box label:
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori discopunctatus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="discopunctatus">discopunctatus</taxonomicName>
/
<normalizedToken originalValue="Forsström">Forsstroem</normalizedToken>
/ Antilles/ C. Dejean; LECTOTYPE (here selected), male, labelled
<normalizedToken originalValue="Schönherr">Schoenherr</normalizedToken>
//
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori discopunctatus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="discopunctatus">discopunctatus</taxonomicName>
Sturm Forst/ palliatus Sch mihi/ in ins. St Barthelemy // [both labels hand printed on green paper]; //LECTO// //TYPE// Ball det. '72.-
<bibRefCitation author="Gemminger, M" journalOrPublisher="Gummi, Monachii" pageId="104" pageNumber="105" title="Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae- Carabidae. E. H." year="1868">Gemminger and Harold 1868</bibRefCitation>
: 266.-
<bibRefCitation author="Putzeys, JADH" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="106" pageNumber="107" pagination="3 - 73" title="Description des Selenophorus de l'Amerique." volume="39" year="1878 a">Putzeys 1878a</bibRefCitation>
: 25.-
<bibRefCitation author="Gundlach, DJ" journalOrPublisher="Anales de la Sociedad Espanola de Historia Natural" pageId="104" pageNumber="105" pagination="259 - 344" title="Apuntes para la fauna Puerto-Riquena. Octava Parte. Neuropteros, Hemipteros, Coleopteros." volume="22" year="1894">Gundlach 1894</bibRefCitation>
: 293.-
<bibRefCitation pageId="26" pageNumber="27">Csiki 1932</bibRefCitation>
: 1197.-
<bibRefCitation author="Darlington, PJ Jr" journalOrPublisher="Psyche" pageId="103" pageNumber="104" pagination="66 - 131" title="New West Indian Carabidae with a list of the Cuban species." url="https://doi.org/10.1155/1934/10161" volume="41" year="1934">Darlington 1934</bibRefCitation>
: 105.-
<bibRefCitation author="Darlington, PJ Jr" journalOrPublisher="Psyche" pageId="103" pageNumber="104" url="https://doi.org/10.1155/1935/84185" year="1935 a">Darlington 1935a</bibRefCitation>
: 161.-
<bibRefCitation pageId="26" pageNumber="27">Blackwelder 1944</bibRefCitation>
: 49.-
<bibRefCitation author="Erwin, TL" journalOrPublisher="Quaestiones Entomologicae" pageId="104" pageNumber="105" pagination="350 - 466" title="Carabid beetles of the West Indies (Insects: Coleoptera): a synopsis of the genera and checklists of tribes of Caraboidea, and of the West Indian species." volume="20" year="1984">Erwin and Sims 1984</bibRefCitation>
: 440.-
<bibRefCitation pageId="26" pageNumber="27">Ball 1992</bibRefCitation>
: 85.-
<bibRefCitation author="Lorenz, W" journalOrPublisher="Lorenz, Tutzing" pageId="105" pageNumber="106" title="Systematic list of ground beetles of the World (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). W." year="1998">Lorenz 1998</bibRefCitation>
: 356.-
<bibRefCitation author="Peck, SB" journalOrPublisher="Florida Department of Agriculture and Consumer Services, Gainesville" pageId="106" pageNumber="107" title="A distributional checklist of the Beetles (Coleoptera) of Florida. Arthropods of Florida and neighboring land areas, volume 16." year="1998">Peck and Thomas 1998</bibRefCitation>
: 22.-
<bibRefCitation pageId="26" pageNumber="27">Lorenz 2005</bibRefCitation>
: 378.-
<bibRefCitation author="Peck, SB" journalOrPublisher="Florida Department of Agriculture and Consumer Services, Gainesville" pageId="106" pageNumber="107" title="A checklist of the beetles of Cuba, with data on distribution and bionomics (Insecta: Coleoptera). Arthropods of Florida and neighboring land areas, Volume 18." year="2005">Peck 2005</bibRefCitation>
: 32.-
<bibRefCitation author="Peck, SB" journalOrPublisher="Insecta Mundi" pageId="106" pageNumber="107" pagination="165 - 209" title="The beetle fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): diversity and distribution." volume="20" year="2006">Peck 2006</bibRefCitation>
: 176.-
<bibRefCitation pageId="26" pageNumber="27">Ivie et al. 2008</bibRefCitation>
: 238.-
<bibRefCitation author="Perez-Gelabert, DE" journalOrPublisher="Zootaxa" pageId="106" pageNumber="107" pagination="1 - 530" title="Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography." volume="1831" year="2008">Perez-Gelabert 2008</bibRefCitation>
: 79.-
<bibRefCitation author="Turnbow, RH" journalOrPublisher="Insecta Mundi" pageId="106" pageNumber="107" pagination="1 - 66" title="An annotated checklist of the Coleoptera (Insecta) of the Bahamas." volume="0034" year="2008">Turnbow and Thomas 2008</bibRefCitation>
: 14.-
<bibRefCitation author="Peck, SB" journalOrPublisher="Insecta Mundi" pageId="106" pageNumber="107" pagination="1 - 54" title="The diversity and distribution of the beetles (Insecta: Coleoptera) of the northern Leeward Islands, Lesser Antilles (Anguilla, Antigua, Barbuda, Nevis, Saba, St. Barthelemy, St. Eustatius, St. Kitts, and St. Martin-St. Maarten)." volume="0159" year="2011">Peck 2011</bibRefCitation>
: 13.-
<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="102" pageNumber="103" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico." url="https://doi.org/10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet 2012</bibRefCitation>
: 1137.
</paragraph>
<paragraph pageId="26" pageNumber="27">
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus cuprinus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="cuprinus">Selenophorus cuprinus</taxonomicName>
Dejean, 1829: 96. TYPE MATERIAL: not located in
<normalizedToken originalValue="Chaudoir-Oberthür">Chaudoir-Oberthuer</normalizedToken>
Collection (MNHP).-
<bibRefCitation author="Gemminger, M" journalOrPublisher="Gummi, Monachii" pageId="104" pageNumber="105" title="Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae- Carabidae. E. H." year="1868">Gemminger and Harold 1868</bibRefCitation>
: 266.-
<bibRefCitation author="Putzeys, JADH" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="106" pageNumber="107" pagination="3 - 73" title="Description des Selenophorus de l'Amerique." volume="39" year="1878 a">Putzeys 1878a</bibRefCitation>
: 25 (established the synonymy).
</paragraph>
<paragraph pageId="26" pageNumber="27">
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus aeratus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="aeratus">Selenophorus aeratus</taxonomicName>
Reiche, 1843: 142. LECTOTYPE: male, in
<normalizedToken originalValue="Chaudoir-Oberthür">Chaudoir-Oberthuer</normalizedToken>
Collection (MNHP), labelled:
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori aeratus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="aeratus">aeratus</taxonomicName>
Reiche/ Venezuela// LECTO// TYPE// [type labels hand printed, on red paper].-
<bibRefCitation author="Gemminger, M" journalOrPublisher="Gummi, Monachii" pageId="104" pageNumber="105" title="Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae- Carabidae. E. H." year="1868">Gemminger and Harold 1868</bibRefCitation>
: 265.-
<bibRefCitation author="Putzeys, JADH" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="106" pageNumber="107" pagination="3 - 73" title="Description des Selenophorus de l'Amerique." volume="39" year="1878 a">Putzeys 1878a</bibRefCitation>
: 25 (established the synonymy).
</paragraph>
<paragraph pageId="26" pageNumber="27">
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus harpaloides" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="harpaloides">Selenophorus harpaloides</taxonomicName>
Reiche, 1843: 142. LECTOTYPE: female, in
<normalizedToken originalValue="Chaudoir-Oberthür">Chaudoir-Oberthuer</normalizedToken>
Collection (MNHP), labelled
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori harpaloides" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="harpaloides">harpaloides</taxonomicName>
/ Reiche Rev./Cuv. 1843/ Caracas// LECTO// TYPE// [type labels hand printed, on red paper].-
<bibRefCitation author="Gemminger, M" journalOrPublisher="Gummi, Monachii" pageId="104" pageNumber="105" title="Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae- Carabidae. E. H." year="1868">Gemminger and Harold 1868</bibRefCitation>
: 266.-
<bibRefCitation author="Putzeys, JADH" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="106" pageNumber="107" pagination="3 - 73" title="Description des Selenophorus de l'Amerique." volume="39" year="1878 a">Putzeys 1878a</bibRefCitation>
: 25 (established the synonymy).
</paragraph>
<paragraph pageId="26" pageNumber="27">
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus subpunctatus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="subpunctatus">Selenophorus subpunctatus</taxonomicName>
Reiche, 1843: 143. LECTOTYPE: female, in
<normalizedToken originalValue="Chaudoir-Oberthür">Chaudoir-Oberthuer</normalizedToken>
Collection (MNHP), labelled:
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori subpunctatus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="subpunctatus">subpunctatus</taxonomicName>
/ Reiche Rev/ Cuv.[...illegible]//.-
<bibRefCitation author="Gemminger, M" journalOrPublisher="Gummi, Monachii" pageId="104" pageNumber="105" title="Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae- Carabidae. E. H." year="1868">Gemminger and Harold 1868</bibRefCitation>
: 267. According to the original description, the provenance of this specimen is Venezuela, near Caracas (
<bibRefCitation author="Putzeys, JADH" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="106" pageNumber="107" pagination="3 - 73" title="Description des Selenophorus de l'Amerique." volume="39" year="1878 a">Putzeys 1878a</bibRefCitation>
: 72 [entry in index]). This specimen was found among the members of
<taxonomicName family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="S. discopunctatus" order="Coleoptera" pageId="26" pageNumber="27" rank="species" species="discopunctatus">S. discopunctatus</taxonomicName>
, as recorded above, suggesting that it was regarded as conspecific with that species. However, Putzeys did not record the name in the synonymy of
<taxonomicName family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="S. discopunctatus" order="Coleoptera" pageId="26" pageNumber="27" rank="species" species="discopunctatus">S. discopunctatus</taxonomicName>
, nor did he include the name in the text of his treatment of
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Selenophorus</taxonomicName>
. We treat it here as the name of a species incertae sedis.
</paragraph>
<paragraph pageId="26" pageNumber="27">
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus chokoloskei" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="chokoloskei">Selenophorus chokoloskei</taxonomicName>
Leng, 1915: 596. Synonymy established by
<bibRefCitation author="Darlington, PJ Jr" journalOrPublisher="Psyche" pageId="103" pageNumber="104" url="https://doi.org/10.1155/1935/84185" year="1935 a">Darlington 1935a</bibRefCitation>
: 161. According to
<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="102" pageNumber="103" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico." url="https://doi.org/10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet (2012</bibRefCitation>
: 1138) location of the syntypes is unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="type area">
<paragraph pageId="26" pageNumber="27">Type area.</paragraph>
<paragraph pageId="26" pageNumber="27">
Saint
<normalizedToken originalValue="Barthélemy">Barthelemy</normalizedToken>
, Leeward Islands, Lesser Antilles.
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="diagnosis">
<paragraph pageId="26" pageNumber="27">Diagnosis.</paragraph>
<paragraph pageId="26" pageNumber="27">This species is readily separated from the other member of the discopunctatus species group by the posteriolateral impressions of pronotum, which are moderately to densely punctate, but smooth, not rugose. Additionally, apical portion of male genitalia with a prominent dorsal hook (Fig. 15A, C; cf. Fig. 15D, F).</paragraph>
</subSubSection>
<subSubSection lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="descriptive notes">
<paragraph pageId="26" pageNumber="27">Descriptive notes.</paragraph>
<paragraph lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Data for SBL in Table 1. Habitus as in Fig. 14A. Labrum with anterior margin shallowly concave; clypeus with anterior margin moderately concave. Antennae and legs testaceous to slightly darker; palpi infuscated, tip testaceous, base slightly to much darker, maxillary palpomere 3 same color as base of maxillary palpomere 4. Dorsal and ventral surfaces brunneous to dark brunneous, not quite piceous; elytral epipleuron paler than disc. Frons and disc of pronotum shiny, with isodiametric microsculpture visible at 100
<normalizedToken originalValue="×">x</normalizedToken>
, microlines more impressed in females; posteriolateral
<pageBreakToken pageId="27" pageNumber="28" start="start">impressions</pageBreakToken>
of pronotum with mesh pattern isodiametric; elytra granular, with mesh pattern isodiametric. Elytral striae impunctate, except the standard setigerous punctures in striae 2, 5 and 7. Punctures of striae 2, 5 and 7 foveate. Males with two terminal setae and females with four terminal setae near the posterior margin on sternum VII.
</paragraph>
<paragraph pageId="27" pageNumber="28">
Male genitalia. Fig. 15
<normalizedToken originalValue="AC">A-C</normalizedToken>
. Apical portion of phallic median lobe with long taper, symmetrically rounded in dorsal/ventral aspect, with prominent dorsal hook; endophallus with four long spines, approximately medial in position; lamina present, more or less banana shaped, pointed at apex.
</paragraph>
<paragraph pageId="27" pageNumber="28">
Ovipositor and female reproductive tract. Very similar to that of
<taxonomicName lsidName="S. yucatanus" pageId="27" pageNumber="28" rank="species" species="yucatanus">S. yucatanus</taxonomicName>
, Fig. 16. For details, see this topic for
<taxonomicName lsidName="S. yucatanus" pageId="27" pageNumber="28" rank="species" species="yucatanus">S. yucatanus</taxonomicName>
, below.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="distribution">
<paragraph pageId="27" pageNumber="28">Geographical distribution.</paragraph>
<paragraph pageId="27" pageNumber="28">Fig. 17. This wide-ranging species is found on most of the island groups in the West Indies, with the exception of the Greater Antillean Caymans.</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="chorological affinities and relationships">
<paragraph pageId="27" pageNumber="28">Chorological affinities and relationships.</paragraph>
<paragraph pageId="27" pageNumber="28">
The West Indian range of this widely distributed species overlaps the range of
<taxonomicName lsidName="S. yucatanus" pageId="27" pageNumber="28" rank="species" species="yucatanus">S. yucatanus</taxonomicName>
in the Lesser Antillean Grenadines. Its relationships are not postulated beyond species group membership.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="materials_examined">
<paragraph pageId="27" pageNumber="28">Material examined.</paragraph>
<paragraph pageId="27" pageNumber="28">In addition to type material, we have seen a total of 1,435 specimens (714 males, 720 females, 1 unknown). See Appendix for details.</paragraph>
<caption pageId="27" pageNumber="28">
<paragraph pageId="27" pageNumber="28">
Figure 14. Habitus digital images of
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus discopunctatus" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="discopunctatus">Selenophorus discopunctatus</taxonomicName>
species group, dorsal aspect. A
<taxonomicName lsidName="S. discopunctatus" pageId="27" pageNumber="28" rank="species" species="discopunctatus">S. discopunctatus</taxonomicName>
Dejean B
<taxonomicName lsidName="S. yucatanus" pageId="27" pageNumber="28" rank="species" species="yucatanus">S. yucatanus</taxonomicName>
Putzeys. Scale bars 5 mm.
</paragraph>
</caption>
<caption pageId="27" pageNumber="28">
<paragraph pageId="27" pageNumber="28">
Figure 15. Digital images of male genitalia of
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus discopunctatus" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="discopunctatus">Selenophorus discopunctatus</taxonomicName>
species group. A, D right lateral aspect B, E dorsal aspect C, F left lateral aspect.
<normalizedToken originalValue="AC">A-C</normalizedToken>
<taxonomicName lsidName="S. discopunctatus" pageId="27" pageNumber="28" rank="species" species="discopunctatus">S. discopunctatus</taxonomicName>
Dejean
<normalizedToken originalValue="DF">D-F</normalizedToken>
<taxonomicName lsidName="S. yucatanus" pageId="27" pageNumber="28" rank="species" species="yucatanus">S. yucatanus</taxonomicName>
Putzeys. Scale bars 1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>