417 lines
52 KiB
XML
417 lines
52 KiB
XML
<document id="310482386DAAEAD0EBBAD426F784AE59" ID-CLB-Dataset="22904" ID-DOI="10.5281/zenodo.3760976" ID-GBIF-Dataset="aaaa853e-f89e-4090-81d4-a4406eebfc53" ID-ISBN="978-2-85653-822-7" ID-ISSN="1243-4442" ID-Zenodo-Dep="3760976" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1587481772657" checkinUser="felipe" docAuthor="Baba, Keiji" docDate="2018" docId="2A1C87B5FDAF4F56FF1ADC4AFA897ACD" docLanguage="en" docName="MemoiresMuseumNationalHistoireNaturelle.30.212.1-612.pdf.imd" docOrigin="Tropical Deep-Sea Benthos (Mémoires du Muséum national d'Histoire naturelle 212)" docStyle="DocumentStyle:18F5342D935943D6DB33581BB9A2E502.4:MemMusNatHistNatu.2010-.journal_article" docStyleId="18F5342D935943D6DB33581BB9A2E502" docStyleName="MemMusNatHistNatu.2010-.journal_article" docStyleVersion="4" docTitle="Heteroptychus Baba 2018, n. gen." docType="treatment" docVersion="10" lastPageNumber="572" masterDocId="D625FFCDFF964D6DFFF1DF73FFCF7B06" masterDocTitle="Chirostylidae of the Western and Central Pacific: Uroptychus and a new genus (Crustacea: Decapoda: Anomura)" masterLastPageNumber="612" masterPageNumber="1" pageNumber="570" updateTime="1698834427920" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="E787A6A4FF96D566906E9272E40CBB61">
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<mods:title id="226D42AF7D74F9EF7E0EF921352F9220">Chirostylidae of the Western and Central Pacific: Uroptychus and a new genus (Crustacea: Decapoda: Anomura)</mods:title>
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<mods:name id="73D9D6F1FE3B021DEF33D55D6A8DCFB0" type="personal">
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<mods:namePart id="D86E7195DE85F49DEBB804FFDFE4A5BA">Baba, Keiji</mods:namePart>
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<mods:affiliation id="1DB578BF4F46426FAABD1270D8F4B2B2">Kumamoto University Faculty of Education 2 - 40 - 1 Kurokami Kumamoto 860 - 8555 Japan kbaba. kumamoto @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="6F542ECD31BF7FBCD060AA786A69B228" type="email">kbaba.kumamoto@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="3CB85C3D84986BB3A27D18453E9CE73F">text</mods:typeOfResource>
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<mods:title id="7063E7E937E92114293297A1BB01C83C">Mémoires du Muséum national d'Histoire naturelle</mods:title>
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<mods:part id="D03C4EAAD5354C3BFD1A567A9F160547">
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<mods:date id="7F7B326D3F43174A6144F5BA629F74C0">2018</mods:date>
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<mods:number id="710E5531EEC300F80BFC9093E14DB8F8">2018-09-30</mods:number>
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<mods:title id="676DBF45697281A885320BFB00E00B93">Tropical Deep-Sea Benthos</mods:title>
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<mods:detail id="80A3623719B9C50C7B1EB441BB759B4F" type="volume">
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<mods:number id="9D2E558ECF36DDF885CBE99F30B41C6F">212</mods:number>
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<mods:identifier id="C284CD09334C844FCAE0D257B1520643" type="DOI">10.5281/zenodo.3760976</mods:identifier>
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<treatment id="2A1C87B5FDAF4F56FF1ADC4AFA897ACD" ID-GBIF-Taxon="163726716" LSID="urn:lsid:plazi:treatment:2A1C87B5FDAF4F56FF1ADC4AFA897ACD" httpUri="http://treatment.plazi.org/id/2A1C87B5FDAF4F56FF1ADC4AFA897ACD" lastPageId="571" lastPageNumber="572" pageId="569" pageNumber="570">
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<subSubSection id="EAAF6528FDAF4F54FF1ADC4AFE3A7852" box="[235,501,825,852]" pageId="569" pageNumber="570" type="nomenclature">
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<paragraph id="A20A36A3FDAF4F54FF1ADC4AFE3A7852" blockId="569.[235,501,825,852]" box="[235,501,825,852]" pageId="569" pageNumber="570">
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<heading id="F94281CFFDAF4F54FF1ADC4AFE3A7852" box="[235,501,825,852]" fontSize="10" level="2" pageId="569" pageNumber="570" reason="2">
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<taxonomicName id="65B54D20FDAF4F54FF1ADC4AFE657854" ID-CoL="acdc69d8-e033-42f5-a851-d66576bb5acb" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[235,426,825,850]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="90C1EAB1FDAF4F54FF1ADC4AFE657854" bold="true" box="[235,426,825,850]" italics="true" pageId="569" pageNumber="570">HETEROPTYCHUS</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8BF257CAFDAF4F54FE41DC4FFE3A7852" box="[432,501,828,852]" pageId="569" pageNumber="570" rank="genus">n. gen.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EAAF6528FDAF4F54FF3DDC0CFD177F07" pageId="569" pageNumber="570" type="etymology">
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<paragraph id="A20A36A3FDAF4F54FF3DDC0CFEF378BD" blockId="569.[204,1436,895,955]" pageId="569" pageNumber="570">
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<emphasis id="90C1EAB1FDAF4F54FF3DDC0CFE997890" bold="true" box="[204,342,895,918]" pageId="569" pageNumber="570">ETYMOLOGY„</emphasis>
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From the Greek
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<emphasis id="90C1EAB1FDAF4F54FDEFDCF2FDA8789F" box="[542,615,897,921]" italics="true" pageId="569" pageNumber="570">heteros</emphasis>
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(different), plus the last syllables of
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<taxonomicName id="65B54D20FDAF4F54FC2DDCF2FB9D789F" box="[988,1106,897,921]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="genus">
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<emphasis id="90C1EAB1FDAF4F54FC2DDCF2FB9D789F" box="[988,1106,897,921]" italics="true" pageId="569" pageNumber="570">Uroptychus</emphasis>
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</taxonomicName>
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(
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<emphasis id="90C1EAB1FDAF4F54FB90DCF2FB61789F" box="[1121,1198,897,921]" italics="true" pageId="569" pageNumber="570">ptychos</emphasis>
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= plate). The gender is masculine.
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</paragraph>
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<paragraph id="A20A36A3FDAF4F54FF3DDC9AFD177F07" blockId="569.[204,728,1001,1025]" box="[204,728,1001,1025]" pageId="569" pageNumber="570">
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<typeStatus id="7D0E8801FDAF4F54FF3DDC9AFECF7F07" box="[204,256,1001,1025]" pageId="569" pageNumber="570">Type</typeStatus>
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species.
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<taxonomicName id="65B54D20FDAF4F54FEADDC9AFD1C7F07" authority="Benedict, 1902" authorityName="Benedict" authorityYear="1902" baseAuthorityName="Benedict" baseAuthorityYear="1902" box="[348,723,1001,1025]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="species" species="scambus">
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<emphasis id="90C1EAB1FDAF4F54FEADDC9AFDFD7F07" box="[348,562,1001,1025]" italics="true" pageId="569" pageNumber="570">Uroptychus scambus</emphasis>
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<bibRefCitation id="C6244B52FDAF4F54FDC6DC9AFD1C7F07" author="Benedict" box="[567,723,1001,1025]" firstAuthor="Benedict" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="569" pageNumber="570" pagination="243 - 334" part="26" refId="ref324766" refString="BENEDICT J. E. 1902 - Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species. Proceedings of the Biological Society of Washington 26: 243 - 334." title="Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species" type="journal article" year="1902">Benedict, 1902</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="EAAF6528FDAF4F54FF3DDB5EFDE17E5D" pageId="569" pageNumber="570" type="diagnosis">
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<paragraph id="A20A36A3FDAF4F54FF3DDB5EFDE17E5D" blockId="569.[204,1436,1069,1372]" pageId="569" pageNumber="570">
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<emphasis id="90C1EAB1FDAF4F54FF3DDB5EFE877F42" bold="true" box="[204,328,1069,1092]" pageId="569" pageNumber="570">DIAGNOSIS„</emphasis>
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Carapace dorsal surface smooth and glabrous, lateral margin with anterolateral spine only, rarely with 1 or 2 processes along branchial region. Rostrum narrowly or broadly triangular. Pterygostomian flap very low on posterior half, height of posterior half 0.1-0.3 × that of anterior half. Sternal plastron different in sexes, posterior margin in females strongly excavated, with median parts of sternites 5-7 absorbed into sternite 4 (left and right parts of sternites 5-7 discontinuous, interrupted by loss of median parts). Antennal scale articulated or fused with article 2, not reaching distal end of article 4, articles 4 and 5 unarmed. P1 ischium with anterior dorsal process lobe-like or spiniform, posterior process usually lobe-like (rarely obsolescent). P4 very short, especially carpus 0.3-0.5 × length of P3 carpus. Distal two articles of P2-4 with long prehensile margins thickly fringed with setae, dactyli with slender spines perpendicular to flexor margin. G1 and G2 present.
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</paragraph>
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</subSubSection>
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<subSubSection id="EAAF6528FDAF4F57FF3DDAF4FAB17F40" lastPageId="570" lastPageNumber="571" pageId="569" pageNumber="570" type="discussion">
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<paragraph id="A20A36A3FDAF4F54FF3DDAF4FDA17D48" blockId="569.[204,1437,1415,1788]" pageId="569" pageNumber="570">
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<emphasis id="90C1EAB1FDAF4F54FF3DDAF4FEF97E98" bold="true" box="[204,310,1415,1438]" pageId="569" pageNumber="570">REMARKS„</emphasis>
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The new genus is established on the basis of the following characters: the female sternal plastron is strongly excavated on the posterior margin, with sternites 5-7 medially discontinuous; the pterygostomian flap is very low in the posterior half (the posterior height at most 0.3 times the anterior height, but usually slightly lower in the majority of
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<taxonomicName id="65B54D20FDAF4F54FF16DA82FE927D0F" box="[231,349,1521,1545]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="genus">
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<emphasis id="90C1EAB1FDAF4F54FF16DA82FE927D0F" box="[231,349,1521,1545]" italics="true" pageId="569" pageNumber="570">Uroptychus</emphasis>
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</taxonomicName>
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species); and the P4 carpus is very short, at most half as long as the P3 carpus, whereas it is subequal to, slightly shorter or rarely longer in
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<taxonomicName id="65B54D20FDAF4F54FDDFD960FD6B7D2D" box="[558,676,1555,1579]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="genus">
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<emphasis id="90C1EAB1FDAF4F54FDDFD960FD6B7D2D" box="[558,676,1555,1579]" italics="true" pageId="569" pageNumber="570">Uroptychus</emphasis>
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</taxonomicName>
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. In addition, it is noteworthy that the antenna is inclined laterally or posteriorly in most of preserved specimens.
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</paragraph>
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<paragraph id="A20A36A3FDAF4F54FF1BD92AFA417DB0" blockId="569.[204,1437,1415,1788]" pageId="569" pageNumber="570">
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The present proposal of the new genus is supported by molecular data that showed that
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<taxonomicName id="65B54D20FDAF4F54FBA8D92AFB017D77" box="[1113,1230,1625,1649]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="genus">
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<emphasis id="90C1EAB1FDAF4F54FBA8D92AFB017D77" box="[1113,1230,1625,1649]" italics="true" pageId="569" pageNumber="570">Uroptychus</emphasis>
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</taxonomicName>
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is not monophyletic (
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<bibRefCitation id="C6244B52FDAF4F54FF25D90FFE667D92" author="Roterman" box="[212,425,1659,1684]" etAl="et al." firstAuthor="Roterman" journalOrPublisher="Proceedings of the Royal Society B" pageId="569" pageNumber="570" pagination="20130718" part="280" refId="ref326231" refString="ROTERMAN C. N., COPLEY J. T., LINSE K. T., TYLER P. A. & ROGERS A. D. 2013 - The biogeography of the yeti crabs (Kiwaidae) with notes on the phylogeny of the Chirostyloidea (Decapoda: Anomura). Proceedings of the Royal Society B 280 (1764): 20130718." title="The biogeography of the yeti crabs (Kiwaidae) with notes on the phylogeny of the Chirostyloidea (Decapoda: Anomura)" type="journal article" year="2013">
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Roterman
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<emphasis id="90C1EAB1FDAF4F54FEB1D90FFEBD7D95" box="[320,370,1659,1683]" italics="true" pageId="569" pageNumber="570">et al.</emphasis>
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2013
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</bibRefCitation>
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;
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<bibRefCitation id="C6244B52FDAF4F54FE40D908FD197D92" author="Bracken-Grissom" box="[433,726,1659,1684]" etAl="et al." firstAuthor="Bracken-Grissom" journalOrPublisher="BMC Evolutionary Biology" pageId="569" pageNumber="570" pagination="128" part="13" refId="ref324846" refString="BRACKEN-GRISSOM H. D., CANNON M. E., CABEZAS P., FELDMANN R. M., SCHWEITZER C. E., AHYONG S. T., FELDER D. L., LEMAITRE R. & CRANDALL K. A. 2013 - A comprehensive and integrative reconstruction of evolutionary history for Anomura (Crustacea: Decapoda). BMC Evolutionary Biology 13: 128." title="A comprehensive and integrative reconstruction of evolutionary history for Anomura (Crustacea: Decapoda)" type="journal article" year="2013">
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Bracken-Grissom
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<emphasis id="90C1EAB1FDAF4F54FD9DD90FFD527D95" box="[620,669,1659,1683]" italics="true" pageId="569" pageNumber="570">et al.</emphasis>
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2013
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</bibRefCitation>
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). Apparently
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<taxonomicName id="65B54D20FDAF4F54FCADD90FFC1D7D95" authorityName="Benedict" authorityYear="1902" baseAuthorityName="Benedict" baseAuthorityYear="1902" box="[860,978,1659,1683]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="species" species="scambus">
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<emphasis id="90C1EAB1FDAF4F54FCADD90FFC1D7D95" box="[860,978,1659,1683]" italics="true" pageId="569" pageNumber="570">U. scambus</emphasis>
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</taxonomicName>
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resides in a clade separate from other species of
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<taxonomicName id="65B54D20FDAF4F54FF16D9EDFE927DB0" box="[231,349,1694,1718]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="genus">
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<emphasis id="90C1EAB1FDAF4F54FF16D9EDFE927DB0" box="[231,349,1694,1718]" italics="true" pageId="569" pageNumber="570">Uroptychus</emphasis>
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</taxonomicName>
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species, although the specific identity of
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<taxonomicName id="65B54D20FDAF4F54FCF7D9ECFCB27DB0" authorityName="Benedict" authorityYear="1902" baseAuthorityName="Benedict" baseAuthorityYear="1902" box="[774,893,1694,1718]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="species" species="scambus">
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<emphasis id="90C1EAB1FDAF4F54FCF7D9ECFCB27DB0" box="[774,893,1694,1718]" italics="true" pageId="569" pageNumber="570">U. scambus</emphasis>
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</taxonomicName>
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reported by these papers remains to be confirmed.
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</paragraph>
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<paragraph id="A20A36A3FDAF4F57FF1BD9B2FC387967" blockId="569.[204,1437,1415,1788]" lastBlockId="570.[174,1407,238,1094]" lastPageId="570" lastPageNumber="571" pageId="569" pageNumber="570">
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The unique shape of the sternal plastron, different in sexes in
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<taxonomicName id="65B54D20FDAF4F54FC90D9B2FBF87DDF" authorityName="Benedict" authorityYear="1902" baseAuthorityName="Benedict" baseAuthorityYear="1902" box="[865,1079,1729,1753]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="species" species="scambus">
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<emphasis id="90C1EAB1FDAF4F54FC90D9B2FBF87DDF" box="[865,1079,1729,1753]" italics="true" pageId="569" pageNumber="570">Uroptychus scambus</emphasis>
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</taxonomicName>
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, was first mentioned by
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<bibRefCitation id="C6244B52FDAF4F54FAC3D9B2FEDE7DFA" author="Baba" bookContentInfo="311 pp." etAl="et al." firstAuthor="Baba" journalOrPublisher="National Taiwan Ocean University Keelung" pageId="569" pageNumber="570" refId="ref324467" refString="BABA K., MACPHERSON E., LIN C. - W. & CHAN T. - Y. 2009 - Crustacean Fauna of Taiwan: Squat Lobsters (Chirostylidae and Galatheidae). National Taiwan Ocean University Keelung, ix + 311 pp." title="Crustacean Fauna of Taiwan: Squat Lobsters (Chirostylidae and Galatheidae)" type="book" year="2009">
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Baba
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<emphasis id="90C1EAB1FDAF4F54FA9AD9B1FA587DDF" box="[1387,1431,1729,1753]" italics="true" pageId="569" pageNumber="570">et al</emphasis>
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. (2009)
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</bibRefCitation>
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. This is also applicable to
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<taxonomicName id="65B54D20FDAF4F54FDD3D997FCEE7DFA" authority="Benedict, 1902" authorityName="Benedict" authorityYear="1902" box="[546,801,1763,1788]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="569" pageNumber="570" phylum="Arthropoda" rank="species" species="brevis">
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<emphasis id="90C1EAB1FDAF4F54FDD3D997FDB07DFD" box="[546,639,1763,1787]" italics="true" pageId="569" pageNumber="570">U. brevis</emphasis>
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||
<bibRefCitation id="C6244B52FDAF4F54FD77D990FCEE7DFA" author="Benedict" box="[646,801,1763,1788]" firstAuthor="Benedict" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="569" pageNumber="570" pagination="243 - 334" part="26" refId="ref324766" refString="BENEDICT J. E. 1902 - Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species. Proceedings of the Biological Society of Washington 26: 243 - 334." title="Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species" type="journal article" year="1902">Benedict, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
from the western Atlantic Ocean, the features confirmed by examination of the type material (
|
||
<typeStatus id="7D0E8801FDAC4F57FDFBDF9DFDA77A00" box="[522,616,238,262]" pageId="570" pageNumber="571" type="holotype">holotype</typeStatus>
|
||
, ovigerous female, poc
|
||
<quantity id="654D9B46FDAC4F57FCA6DF9DFC687A00" box="[855,935,238,262]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="570" pageNumber="571" unit="mm" value="5.2">5.2 mm</quantity>
|
||
, USNM 20566) of that species.
|
||
<taxonomicName id="65B54D20FDAC4F57FB1ADF9DFEC87A2F" baseAuthorityName="Benedict" baseAuthorityYear="1902" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="scambus">
|
||
<emphasis id="90C1EAB1FDAC4F57FB1ADF9DFEC87A2F" italics="true" pageId="570" pageNumber="571">Heteroptychus scambus</emphasis>
|
||
</taxonomicName>
|
||
is definitely a deep-sea species and has been believed to show a wide distribution from the western Indian Ocean to the western Pacific. However, molecular analyses (L. Corbari, personal comm.) suggest that there are some cryptic species, so that an attempt was made to detect morphological differences. Accordingly, slight differences that had been regarded as individual variations were validated as discriminating characters: the presence or absence of articulation between the antennal scale and article 2 and in the carapace shape, especially the rostrum and anterolateral spine. These characters delineate nine species including six new species, but no clear differences are found on other characters in all the material examined. However, there exist unstable characters in
|
||
<taxonomicName id="65B54D20FDAC4F57FC93DE91FC137AFF" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[866,988,481,505]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="claudeae" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAC4F57FC93DE91FC137AFF" box="[866,988,481,505]" italics="true" pageId="570" pageNumber="571">H. claudeae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAC4F57FC13DE92FBD67AFE" box="[994,1049,481,504]" pageId="570" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="90C1EAB1FDAC4F57FBD2DE91FB8E7AFF" box="[1059,1089,482,505]" italics="true" pageId="570" pageNumber="571">i.e.</emphasis>
|
||
the presence or absence of the lateral orbital spine, and the presence or absence of median processes of the anterior margin of sternite 3. In addition, color differences are found in morphologically identical specimens from
|
||
<collectingCountry id="DAA27633FDAC4F57FC6ADD55FC287938" box="[923,999,550,574]" name="Taiwan" pageId="570" pageNumber="571">Taiwan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="C6244B52FDAC4F57FC02DD55FB567938" author="Baba" bookContentInfo="311 pp." box="[1011,1177,550,574]" etAl="et al." firstAuthor="Baba" journalOrPublisher="National Taiwan Ocean University Keelung" pageId="570" pageNumber="571" refId="ref324467" refString="BABA K., MACPHERSON E., LIN C. - W. & CHAN T. - Y. 2009 - Crustacean Fauna of Taiwan: Squat Lobsters (Chirostylidae and Galatheidae). National Taiwan Ocean University Keelung, ix + 311 pp." title="Crustacean Fauna of Taiwan: Squat Lobsters (Chirostylidae and Galatheidae)" type="book" year="2009">
|
||
Baba
|
||
<emphasis id="90C1EAB1FDAC4F57FBDDDD54FB907938" box="[1068,1119,550,574]" italics="true" pageId="570" pageNumber="571">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
). It is not unlikely that this species may contain cryptic species, for which extensive studies are required.
|
||
</paragraph>
|
||
<paragraph id="A20A36A3FDAC4F57FF3DDD1FFBFA7F07" blockId="570.[174,1407,238,1094]" pageId="570" pageNumber="571">
|
||
Two classical names pose a problem for establishing this new genus because of the brief species descriptions:
|
||
<taxonomicName id="65B54D20FDAC4F57FAFADD1FFE8B79A0" authority="Benedict, 1902" authorityName="Benedict" authorityYear="1902" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="scambus">
|
||
<emphasis id="90C1EAB1FDAC4F57FAFADD1FFAB27985" box="[1291,1405,619,643]" italics="true" pageId="570" pageNumber="571">U. scambus</emphasis>
|
||
<bibRefCitation id="C6244B52FDAC4F57FF5FDDFDFE8B79A0" author="Benedict" box="[174,324,654,678]" firstAuthor="Benedict" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="570" pageNumber="571" pagination="243 - 334" part="26" refId="ref324766" refString="BENEDICT J. E. 1902 - Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species. Proceedings of the Biological Society of Washington 26: 243 - 334." title="Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species" type="journal article" year="1902">Benedict, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="65B54D20FDAC4F57FE85DDFCFD3C79A0" authority="MacGilchrist, 1905" authorityName="MacGilchrist" authorityYear="1905" baseAuthorityName="MacGilchrist" baseAuthorityYear="1905" box="[372,755,654,678]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="glyphodactylus">
|
||
<emphasis id="90C1EAB1FDAC4F57FE85DDFCFDE779A0" box="[372,552,654,678]" italics="true" pageId="570" pageNumber="571">U. glyphodactylus</emphasis>
|
||
MacGilchrist, 1905
|
||
</taxonomicName>
|
||
. Examination of the
|
||
<typeStatus id="7D0E8801FDAC4F57FC31DDFDFBD479A0" box="[960,1051,654,678]" pageId="570" pageNumber="571" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="65B54D20FDAC4F57FBC9DDFDFAC579A0" authorityName="Benedict" authorityYear="1902" baseAuthorityName="Benedict" baseAuthorityYear="1902" box="[1080,1290,654,678]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="scambus">
|
||
<emphasis id="90C1EAB1FDAC4F57FBC9DDFDFAC579A0" box="[1080,1290,654,678]" italics="true" pageId="570" pageNumber="571">Uroptychus scambus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="DAA27633FDAC4F57FAB4DDFCFAB179A0" box="[1349,1406,655,678]" name="Japan" pageId="570" pageNumber="571">Japan</collectingCountry>
|
||
now in the collection of the Smithsonian Institution has elaborated on its systematic status. However, the type material of
|
||
<taxonomicName id="65B54D20FDAC4F57FF5FDDA7FEAA79ED" authorityName="MacGilchrist" authorityYear="1905" baseAuthorityName="MacGilchrist" baseAuthorityYear="1905" box="[174,357,723,747]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="glyphodactylus">
|
||
<emphasis id="90C1EAB1FDAC4F57FF5FDDA7FEAA79ED" box="[174,357,723,747]" italics="true" pageId="570" pageNumber="571">U. glyphodactylus</emphasis>
|
||
</taxonomicName>
|
||
from the Andaman Sea now housed in the collection of the Zoological Survey of
|
||
<collectingCountry id="DAA27633FDAC4F57FB5ADDA0FB2F79ED" box="[1195,1248,723,747]" name="India" pageId="570" pageNumber="571">India</collectingCountry>
|
||
could not have been examined, despite repeated queries to the director of the institution. Some of the morphological features of this species that were not described in MacGilchrist (1905) are interpreted from the illustrations given by Alcock & MacGilchrist (1905: pl. 70, fig. 4, pl. 71, figs 1, 1a, 1b, 1c, 1d) albeit not in sufficient detail: the antennal scale in the figure appears to be articulated with article 2. As shown in the key to species provided below, these two species key out in the same couplet. Given that these species are known from disjunct localities, it is not unlikely that they are different. However, they are treated as synonymous for the time being.
|
||
<taxonomicName id="65B54D20FDAC4F57FD92DCD0FC0A78BA" authority="Kensley, 1981" authorityName="Kensley" authorityYear="1981" box="[611,965,931,956]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="edwardi">
|
||
<emphasis id="90C1EAB1FDAC4F57FD92DCD0FCFC78BD" box="[611,819,931,955]" italics="true" pageId="570" pageNumber="571">Uroptychus edwardi</emphasis>
|
||
<bibRefCitation id="C6244B52FDAC4F57FCC9DCD0FC0A78BA" author="Kensley" box="[824,965,931,956]" firstAuthor="Kensley" journalOrPublisher="Annals of the South African Museum" pageId="570" pageNumber="571" pagination="49 - 78" part="83" refId="ref324876" refString="KENSLEY B. 1981 - The South African Museum's Meiring Naude cruises. Part 12. Crustacea Decapoda of the 1977 1978 1979 cruises. Annals of the South African Museum 83: 49 - 78." title="The South African Museum's Meiring Naude cruises. Part 12. Crustacea Decapoda of the 1977 1978 1979 cruises." type="book chapter" year="1981">Kensley, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
from off the east coast of
|
||
<collectingCountry id="DAA27633FDAC4F57FB3FDCD0FA9D78BD" box="[1230,1362,931,955]" name="South Africa" pageId="570" pageNumber="571">South Africa</collectingCountry>
|
||
was synonymized with
|
||
<taxonomicName id="65B54D20FDAC4F57FE80DCB4FE2878D8" authorityName="Benedict" authorityYear="1902" baseAuthorityName="Benedict" baseAuthorityYear="1902" box="[369,487,966,990]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="scambus">
|
||
<emphasis id="90C1EAB1FDAC4F57FE80DCB4FE2878D8" box="[369,487,966,990]" italics="true" pageId="570" pageNumber="571">U. scambus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="C6244B52FDAC4F57FDEBDCB5FD4278D8" author="Baba" bookContentInfo="203 pp." box="[538,653,966,990]" firstAuthor="Baba" journalOrPublisher="Researches on Crustacea Special Number 2" pageId="570" pageNumber="571" refId="ref323864" refString="BABA K. 1988 - Chirostylid and Galatheid crustaceans (Decapoda: Anomura) of the " Albatross " Philippine Expedition 1907 - 1910. Researches on Crustacea Special Number 2: 203 pp." title="Chirostylid and Galatheid crustaceans (Decapoda: Anomura) of the " Albatross " Philippine Expedition 1907 - 1910" type="book" year="1988">Baba, 1988</bibRefCitation>
|
||
), but it is validated in this paper, on the basis of differences in the rostrum and anterolateral spine of the carapace (see below under the remarks of
|
||
<taxonomicName id="65B54D20FDAC4F57FC64DC99FC247F07" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[917,1003,1001,1025]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="colini" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAC4F57FC64DC99FC247F07" box="[917,1003,1001,1025]" italics="true" pageId="570" pageNumber="571">H. colini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAC4F57FC00DC9AFBE67F07" box="[1009,1065,1001,1025]" pageId="570" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A20A36A3FDAC4F57FF3DDB78FAB17F40" blockId="570.[174,1407,238,1094]" pageId="570" pageNumber="571">
|
||
The genus now contains nine species, including six new species: eight from the Indo-West Pacific and one from the western Atlantic. Females of three species have not been collected (
|
||
<taxonomicName id="65B54D20FDAC4F57FCB8DB5CFC077F40" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[841,968,1070,1094]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="apophysis" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAC4F57FCB8DB5CFC077F40" box="[841,968,1070,1094]" italics="true" pageId="570" pageNumber="571">H. apophysis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAC4F57FC3DDB5CFBCF7F40" box="[972,1024,1071,1094]" pageId="570" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="65B54D20FDAC4F57FBF8DB5CFB937F40" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[1033,1116,1070,1094]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="colini" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAC4F57FBF8DB5CFB937F40" box="[1033,1116,1070,1094]" italics="true" pageId="570" pageNumber="571">H. colini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAC4F57FB91DB5CFB5B7F40" box="[1120,1172,1071,1094]" pageId="570" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="65B54D20FDAC4F57FB33DB5CFAF67F40" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[1218,1337,1070,1094]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="lemaitrei" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAC4F57FB33DB5CFAF67F40" box="[1218,1337,1070,1094]" italics="true" pageId="570" pageNumber="571">H. lemaitrei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAC4F57FACCDB5CFABD7F40" box="[1341,1394,1070,1094]" pageId="570" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSection id="CB3A2F48FDAC4F56FD27DBE6FA897ACD" lastPageId="571" lastPageNumber="572" pageId="570" pageNumber="571" type="key">
|
||
<subSubSection id="EAAF6528FDAC4F57FD27DBE6FC9A7FAD" box="[726,853,1173,1196]" pageId="570" pageNumber="571" type="discussion">
|
||
<paragraph id="A20A36A3FDAC4F57FD27DBE6FC9A7FAD" blockId="570.[726,853,1173,1196]" box="[726,853,1173,1196]" pageId="570" pageNumber="571">
|
||
<heading id="F94281CFFDAC4F57FD27DBE6FC9A7FAD" allCaps="true" bold="true" box="[726,853,1173,1196]" fontSize="9" level="3" pageId="570" pageNumber="571" reason="3">
|
||
<emphasis id="90C1EAB1FDAC4F57FD27DBE6FC9A7FAD" bold="true" box="[726,853,1173,1196]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FD27DBE6FD387FAD" bold="true" box="[726,759,1173,1196]" pageId="570" pageNumber="571">KEY</emphasis>
|
||
TO SPECIES
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EAAF6528FDAC4F56FEF4DBABFA897ACD" lastPageId="571" lastPageNumber="572" pageId="570" pageNumber="571" type="key">
|
||
<key id="D6246192FDAC4F56FEF4DBABFA897ACD" lastPageId="571" lastPageNumber="572" pageId="570" pageNumber="571">
|
||
<keyStep id="19412E06FDAC4F57FEF4DBABFAE87E08" pageId="570" pageNumber="571">
|
||
<paragraph id="A20A36A3FDAC4F57FEF4DBABFAE87FE8" blockId="570.[261,1320,1240,1724]" box="[261,1319,1240,1264]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4DBABFAE87FE8" box="[261,1319,1240,1264]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4DBABFEDB7FE8" bold="true" box="[261,276,1240,1262]" pageId="570" pageNumber="571">1.</emphasis>
|
||
Antennal scale articulated with antennal article 2 .....................................................................................................
|
||
<emphasis id="90C1EAB1FDAC4F57FAECDBABFAE87FE8" bold="true" box="[1309,1319,1240,1262]" pageId="570" pageNumber="571">2</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A20A36A3FDAC4F57FEF4DB8BFAE87E08" blockId="570.[261,1320,1240,1724]" box="[261,1319,1272,1296]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4DB8BFAE87E08" box="[261,1319,1272,1296]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4DB8BFEDE7E08" bold="true" box="[261,273,1272,1294]" pageId="570" pageNumber="571">–</emphasis>
|
||
Antennal scale fused with antennal article 2 ................................................................................................................
|
||
<emphasis id="90C1EAB1FDAC4F57FAECDB8BFAE87E08" bold="true" box="[1309,1319,1272,1294]" pageId="570" pageNumber="571">7</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="19412E06FDAC4F57FEF4DA54FAE87E5B" pageId="570" pageNumber="571">
|
||
<paragraph id="A20A36A3FDAC4F57FEF4DA54FAE77E39" blockId="570.[261,1320,1240,1724]" box="[261,1320,1319,1344]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4DA54FAE77E39" box="[261,1320,1319,1344]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4DA54FEDB7E3B" bold="true" box="[261,276,1319,1341]" pageId="570" pageNumber="571">2.</emphasis>
|
||
Branchial margin with 2 processes ...............................................................................................
|
||
<taxonomicName id="65B54D20FDAC4F57FB83DA59FB217E39" ID-CoL="935fdff5-73a9-478e-a238-1d4b46863ef6" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[1138,1262,1322,1343]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="apophysis" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAC4F57FB83DA59FB217E39" box="[1138,1262,1322,1343]" italics="true" pageId="570" pageNumber="571">H. apophysis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAC4F57FB05DA59FAE77E39" box="[1268,1320,1322,1343]" pageId="570" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A20A36A3FDAC4F57FEF4DA34FAE87E5B" blockId="570.[261,1320,1240,1724]" box="[261,1319,1351,1376]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4DA34FAE87E5B" box="[261,1319,1351,1376]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4DA34FEDE7E5B" bold="true" box="[261,273,1351,1373]" pageId="570" pageNumber="571">–</emphasis>
|
||
Branchial margin smooth ..................................................................................................................................................
|
||
<emphasis id="90C1EAB1FDAC4F57FAECDA34FAE87E5B" bold="true" box="[1309,1319,1351,1373]" pageId="570" pageNumber="571">3</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="19412E06FDAC4F57FEF4DA04FAE87EAB" pageId="570" pageNumber="571">
|
||
<paragraph id="A20A36A3FDAC4F57FEF4DA04FAE77E88" blockId="570.[261,1320,1240,1724]" box="[261,1320,1399,1423]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4DA04FAE77E88" box="[261,1320,1399,1423]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4DA04FEDB7E8B" bold="true" box="[261,276,1399,1421]" pageId="570" pageNumber="571">3.</emphasis>
|
||
Anterolateral spine of carapace slightly overreaching apex of rostrum .............................
|
||
<taxonomicName id="65B54D20FDAC4F57FB99DA0AFB217E88" ID-CoL="69a1e42b-47aa-43cc-bb67-372d50a20d5c" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[1128,1262,1401,1422]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="anouchkae" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAC4F57FB99DA0AFB217E88" box="[1128,1262,1401,1422]" italics="true" pageId="570" pageNumber="571">H. anouchkae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAC4F57FB05DA0AFAE77E88" box="[1268,1320,1401,1422]" pageId="570" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A20A36A3FDAC4F57FEF4DAE4FAE87EAB" blockId="570.[261,1320,1240,1724]" box="[261,1319,1431,1455]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4DAE4FAE87EAB" box="[261,1319,1431,1455]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4DAE4FEDE7EAB" bold="true" box="[261,273,1431,1453]" pageId="570" pageNumber="571">–</emphasis>
|
||
Anterolateral spine of carapace not overreaching apex of rostrum ........................................................................
|
||
<emphasis id="90C1EAB1FDAC4F57FAECDAE4FAE87EAB" bold="true" box="[1309,1319,1431,1453]" pageId="570" pageNumber="571">4</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="19412E06FDAC4F57FEF4DAB5FAE87D1A" pageId="570" pageNumber="571">
|
||
<paragraph id="A20A36A3FDAC4F57FEF4DAB5FAE87EF8" blockId="570.[261,1320,1240,1724]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4DAB5FAE87EF8" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4DAB5FEDB7EDA" bold="true" box="[261,276,1478,1500]" pageId="570" pageNumber="571">4.</emphasis>
|
||
Rostrum not reaching apex of eye..................................................................................................................................... ..................................................................
|
||
<taxonomicName id="65B54D20FDAC4F57FD97DA9AFCB17EF8" ID-CoL="04312531-654f-4446-a571-83e5de305a55" authority="(Benedict, 1902)" authorityName="Benedict" authorityYear="1902" baseAuthorityName="Benedict" baseAuthorityYear="1902" box="[614,894,1512,1534]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="scambus">
|
||
<emphasis id="90C1EAB1FDAC4F57FD97DA9AFD187EFB" box="[614,727,1512,1534]" italics="true" pageId="570" pageNumber="571">H. scambus</emphasis>
|
||
(
|
||
<bibRefCitation id="C6244B52FDAC4F57FD15DA9BFCB87EF8" author="Benedict" box="[740,887,1512,1534]" firstAuthor="Benedict" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="570" pageNumber="571" pagination="243 - 334" part="26" refId="ref324766" refString="BENEDICT J. E. 1902 - Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species. Proceedings of the Biological Society of Washington 26: 243 - 334." title="Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species" type="journal article" year="1902">Benedict, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="65B54D20FDAC4F57FC6EDA9AFAE87EF8" ID-CoL="4ae48b1b-6c4b-470f-9d92-d75fb7a9fd44" authority="(MacGilchrist, 1905))" authorityName="MacGilchrist" authorityYear="1905" baseAuthorityName="MacGilchrist" baseAuthorityYear="1905" box="[927,1319,1512,1534]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="glyphodactylus">
|
||
<emphasis id="90C1EAB1FDAC4F57FC6EDA9AFB817EFB" box="[927,1102,1512,1534]" italics="true" pageId="570" pageNumber="571">H. glyphodactylus</emphasis>
|
||
(MacGilchrist, 1905))
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A20A36A3FDAC4F57FEF4D975FAE87D1A" blockId="570.[261,1320,1240,1724]" box="[261,1319,1542,1566]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4D975FAE87D1A" box="[261,1319,1542,1566]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4D975FEDE7D1A" bold="true" box="[261,273,1542,1564]" pageId="570" pageNumber="571">–</emphasis>
|
||
Rostrum overreaching eye ................................................................................................................................................
|
||
<emphasis id="90C1EAB1FDAC4F57FAECD975FAE87D1A" bold="true" box="[1309,1319,1542,1564]" pageId="570" pageNumber="571">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="19412E06FDAC4F57FEF4D946FAE87D6D" pageId="570" pageNumber="571">
|
||
<paragraph id="A20A36A3FDAC4F57FEF4D946FAE77D4A" blockId="570.[261,1320,1240,1724]" box="[261,1320,1589,1613]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4D946FAE77D4A" box="[261,1320,1589,1613]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4D946FEDB7D4D" bold="true" box="[261,276,1589,1611]" pageId="570" pageNumber="571">5.</emphasis>
|
||
Rostrum broad triangular, distally blunt ...........................................................................................
|
||
<taxonomicName id="65B54D20FDAC4F57FB7FD94BFB217D4A" ID-CoL="3d5d715e-0bfe-4570-a2cc-6bf5ecbc1bff" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[1166,1262,1591,1613]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="paulae" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAC4F57FB7FD94BFB217D4A" box="[1166,1262,1591,1613]" italics="true" pageId="570" pageNumber="571">H. paulae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAC4F57FB05D944FAE77D4A" box="[1268,1320,1591,1612]" pageId="570" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A20A36A3FDAC4F57FEF4D926FAE87D6D" blockId="570.[261,1320,1240,1724]" box="[261,1319,1621,1645]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4D926FAE87D6D" box="[261,1319,1621,1645]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4D926FEDE7D6D" bold="true" box="[261,273,1621,1643]" pageId="570" pageNumber="571">–</emphasis>
|
||
Rostrum narrow triangular or distally spiciform ........................................................................................................
|
||
<emphasis id="90C1EAB1FDAC4F57FAECD926FAE87D6D" bold="true" box="[1309,1319,1621,1643]" pageId="570" pageNumber="571">6</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="A20A36A3FDAC4F57FEF4D9F7FAE77D9A" blockId="570.[261,1320,1240,1724]" box="[261,1320,1668,1692]" pageId="570" pageNumber="571">
|
||
<keyStep id="19412E06FDAC4F57FEF4D9F7FAE77D9A" box="[261,1320,1668,1692]" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4D9F7FAE77D9A" box="[261,1320,1668,1692]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4D9F7FEDB7D9C" bold="true" box="[261,276,1668,1690]" pageId="570" pageNumber="571">6.</emphasis>
|
||
Rostrum narrow triangular. Sternite 3 with pair of median processes .........................................
|
||
<taxonomicName id="65B54D20FDAC4F57FB6DD9F4FB217D9D" ID-CoL="20fcdabe-c0f7-4caa-a90b-63edb3f017ab" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[1180,1262,1670,1692]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="colini" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAC4F57FB6DD9F4FB217D9D" box="[1180,1262,1670,1692]" italics="true" pageId="570" pageNumber="571">H. colini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAC4F57FB05D9F4FAE77D9A" box="[1268,1320,1671,1692]" pageId="570" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</keyStep>
|
||
</paragraph>
|
||
<paragraph id="A20A36A3FDAC4F56FEF4D9D7FA8A7A1A" blockId="570.[261,1320,1240,1724]" lastBlockId="571.[291,1350,229,460]" lastPageId="571" lastPageNumber="572" pageId="570" pageNumber="571">
|
||
<keyLead id="19449596FDAC4F57FEF4D9D7FAE87DBA" box="[261,1319,1700,1724]" pageId="570" pageNumber="571">
|
||
<emphasis id="90C1EAB1FDAC4F57FEF4D9D7FEDE7DBC" bold="true" box="[261,273,1700,1722]" pageId="570" pageNumber="571">–</emphasis>
|
||
Rostrum spiciform. Sternite 3 with median notch, with no median processes ....
|
||
<taxonomicName id="65B54D20FDAC4F57FBD1D9D4FAE87DBA" ID-CoL="d2bbc6dd-d7ca-4675-a0a0-89d54d079451" authority="(Kensley, 1981)" baseAuthorityName="Kensley" baseAuthorityYear="1981" box="[1056,1319,1702,1724]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="570" pageNumber="571" phylum="Arthropoda" rank="species" species="edwardi">
|
||
<emphasis id="90C1EAB1FDAC4F57FBD1D9D4FB427DBD" box="[1056,1165,1702,1724]" italics="true" pageId="570" pageNumber="571">H. edwardi</emphasis>
|
||
(
|
||
<bibRefCitation id="C6244B52FDAC4F57FB69D9D5FAEF7DBA" author="Kensley" box="[1176,1312,1702,1724]" firstAuthor="Kensley" journalOrPublisher="Annals of the South African Museum" pageId="570" pageNumber="571" pagination="49 - 78" part="83" refId="ref324876" refString="KENSLEY B. 1981 - The South African Museum's Meiring Naude cruises. Part 12. Crustacea Decapoda of the 1977 1978 1979 cruises. Annals of the South African Museum 83: 49 - 78." title="The South African Museum's Meiring Naude cruises. Part 12. Crustacea Decapoda of the 1977 1978 1979 cruises." type="book chapter" year="1981">Kensley, 1981</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
<keyStep id="19412E06FDAD4F56FED2DF96FA8A7A1A" pageId="571" pageNumber="572">
|
||
<keyLead id="19449596FDAD4F56FED2DF96FA8A7A1A" pageId="571" pageNumber="572">
|
||
<emphasis id="90C1EAB1FDAD4F56FED2DF96FEFC7BFD" bold="true" box="[291,307,229,251]" pageId="571" pageNumber="572">7.</emphasis>
|
||
Rostrum and anterolateral spines distally blunt. P1 ischium with posterior dorsal lobe obsolescent ............. .............................................................................
|
||
<taxonomicName id="65B54D20FDAD4F56FD35DE74FC0E7A1B" ID-CoL="86664ab0-7fb5-46f4-9379-d1b20f3e9f06" authority="(Benedict, 1902)" baseAuthorityName="Benedict" baseAuthorityYear="1902" box="[708,961,263,285]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" kingdom="Animalia" order="Decapoda" pageId="571" pageNumber="572" phylum="Arthropoda" rank="species" species="brevis">
|
||
<emphasis id="90C1EAB1FDAD4F56FD35DE74FCD57A1A" box="[708,794,263,284]" italics="true" pageId="571" pageNumber="572">H. brevis</emphasis>
|
||
(
|
||
<bibRefCitation id="C6244B52FDAD4F56FCD7DE74FC757A1B" author="Benedict" box="[806,954,263,285]" firstAuthor="Benedict" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="571" pageNumber="572" pagination="243 - 334" part="26" refId="ref324766" refString="BENEDICT J. E. 1902 - Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species. Proceedings of the Biological Society of Washington 26: 243 - 334." title="Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species" type="journal article" year="1902">Benedict, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
[Western Atlantic;
|
||
<typeStatus id="7D0E8801FDAD4F56FB70DE74FB177A1B" box="[1153,1240,263,285]" pageId="571" pageNumber="572" type="holotype">holotype</typeStatus>
|
||
examined]
|
||
</keyLead>
|
||
</keyStep>
|
||
</paragraph>
|
||
<paragraph id="A20A36A3FDAD4F56FED2DE56FA897A5D" blockId="571.[291,1350,229,460]" pageId="571" pageNumber="572">
|
||
<keyLead id="19449596FDAD4F56FED2DE56FA897A5D" pageId="571" pageNumber="572">
|
||
<emphasis id="90C1EAB1FDAD4F56FED2DE56FEE07A3D" bold="true" box="[291,303,293,315]" pageId="571" pageNumber="572">–</emphasis>
|
||
Rostrum and anterolateral spines sharply pointed. P1 ischium with posterior dorsal lobe overhanging basis ...........................................................................................................................................................................................
|
||
<emphasis id="90C1EAB1FDAD4F56FACDDE36FA897A5D" bold="true" box="[1340,1350,325,347]" pageId="571" pageNumber="572">8</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyStep id="19412E06FDAD4F56FED2DE07FA897ACD" pageId="571" pageNumber="572">
|
||
<paragraph id="A20A36A3FDAD4F56FED2DE07FA897A8D" blockId="571.[291,1350,229,460]" box="[291,1350,372,396]" pageId="571" pageNumber="572">
|
||
<keyLead id="19449596FDAD4F56FED2DE07FA897A8D" box="[291,1350,372,396]" pageId="571" pageNumber="572">
|
||
<emphasis id="90C1EAB1FDAD4F56FED2DE07FEFC7A8C" bold="true" box="[291,307,372,394]" pageId="571" pageNumber="572">8.</emphasis>
|
||
Carapace lateral margin smooth. Rostrum narrow, more or less depressed distally ...........
|
||
<taxonomicName id="65B54D20FDAD4F56FB69DE04FAC37A8D" ID-CoL="cca83fdc-0a7b-4d2e-b141-df05f0fe23f4" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[1176,1292,374,396]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="571" pageNumber="572" phylum="Arthropoda" rank="species" species="claudeae" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAD4F56FB69DE04FAC37A8D" box="[1176,1292,374,396]" italics="true" pageId="571" pageNumber="572">H. claudeae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAD4F56FAE3DE05FA897A8D" box="[1298,1350,374,395]" pageId="571" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A20A36A3FDAD4F56FED2DEE7FA897ACD" blockId="571.[291,1350,229,460]" pageId="571" pageNumber="572">
|
||
<keyLead id="19449596FDAD4F56FED2DEE7FA897ACD" pageId="571" pageNumber="572">
|
||
<emphasis id="90C1EAB1FDAD4F56FED2DEE7FEE07AAC" bold="true" box="[291,303,404,426]" pageId="571" pageNumber="572">–</emphasis>
|
||
Carapace lateral margin with small protuberance at anterior end of branchial margin. Rostrum spiniform. .....................................................................................................................................................................
|
||
<taxonomicName id="65B54D20FDAD4F56FB66DEC4FAC37ACD" ID-CoL="afe5ddff-c763-4e0d-9b1c-5ae26d8d0545" authority="Baba, 2018" authorityName="Baba" authorityYear="2018" box="[1175,1292,438,460]" class="Malacostraca" family="Chirostylidae" genus="Heteroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="571" pageNumber="572" phylum="Arthropoda" rank="species" species="lemaitrei" status="sp. nov.">
|
||
<emphasis id="90C1EAB1FDAD4F56FB66DEC4FAC37ACD" box="[1175,1292,438,460]" italics="true" pageId="571" pageNumber="572">H. lemaitrei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BF257CAFDAD4F56FAE3DEC5FA897ACD" box="[1298,1350,438,459]" pageId="571" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |