625 lines
78 KiB
XML
625 lines
78 KiB
XML
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<mods:title id="1041A9EA63D37A80B365BB0A73A9EC43">Ɯandinus (Ɯandinus) trailini sp. n. from Ethiopia (Scorpiones: Scorpionidae), with data on localities and life strategy</mods:title>
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<mods:namePart id="AD4A0DDE84B957779CFBFB513DF1330A">František Kovařík</mods:namePart>
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<mods:title id="A22F0B2116808D1A481C4D976729B159">Euscorpius</mods:title>
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<mods:date id="2234331C037A818D5FB33BD805E73450">2013</mods:date>
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<mods:number id="5B210EB44F3D44D6665CD74BBA961CF3">163</mods:number>
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<treatment id="2A0587C5FF81A94CC0DEF9E8FEE1FA68" ID-DOI="http://doi.org/10.5281/zenodo.5744009" ID-Zenodo-Dep="5744009" LSID="urn:lsid:plazi:treatment:2A0587C5FF81A94CC0DEF9E8FEE1FA68" httpUri="http://treatment.plazi.org/id/2A0587C5FF81A94CC0DEF9E8FEE1FA68" lastPageId="11" lastPageNumber="10" pageId="2" pageNumber="1">
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<subSubSection id="EAB66558FF81A945C0DEF9E8FA99F955" box="[924,1347,1659,1685]" pageId="2" pageNumber="1" type="nomenclature">
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<paragraph id="A21336D3FF81A945C0DEF9E8FA99F955" blockId="2.[838,1432,1659,1777]" box="[924,1347,1659,1685]" pageId="2" pageNumber="1">
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<heading id="F95B81BFFF81A945C0DEF9E8FA99F955" box="[924,1347,1659,1685]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1" reason="2">
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<emphasis id="90D8EAC1FF81A945C0DEF9E8FA99F955" bold="true" box="[924,1347,1659,1685]" pageId="2" pageNumber="1">
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<taxonomicName id="65AC4D50FF81A945C0DEF9E8FB23F955" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[924,1273,1659,1685]" class="Arachnida" family="Euscorpiidae" genus="Mandinus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov." subGenus="Mandinus">
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<emphasis id="90D8EAC1FF81A945C0DEF9E8FBCAF955" bold="true" box="[924,1040,1659,1685]" italics="true" pageId="2" pageNumber="1">Ɯandinus</emphasis>
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(
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<emphasis id="90D8EAC1FF81A945C763F9E8FB4FF955" bold="true" box="[1057,1173,1659,1685]" italics="true" pageId="2" pageNumber="1">Ɯandinus</emphasis>
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)
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<emphasis id="90D8EAC1FF81A945C7E4F9E8FB23F955" bold="true" box="[1190,1273,1659,1685]" italics="true" pageId="2" pageNumber="1">trailini</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8BEB57BAFF81A945C643F9E8FA99F955" box="[1281,1347,1659,1685]" pageId="2" pageNumber="1" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EAB66558FF81A945C751F908FA8BF931" pageId="2" pageNumber="1" type="description">
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<paragraph id="A21336D3FF81A945C751F908FA8BF931" blockId="2.[838,1432,1659,1777]" pageId="2" pageNumber="1">
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(
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<figureCitation id="3A972A56FF81A945C759F908FB5BF973" box="[1051,1153,1691,1715]" captionStart="Figures 1–8" captionStartId="3.[192,278,1795,1819]" captionTargetBox="[222,1387,166,1747]" captionTargetId="figure-96@3.[212,1416,165,1765]" captionTargetPageId="3" captionText="Figures 1–8: 1–6: Pandinus trailini sp. n., ♂ holotype. 1–2. Dorsal and ventral views. 3. Pectinal area. 4. Metasoma and telson, lateral view. 5–6. Tarsomere II of 4th leg, lateral and ventral views. 7: Pandinus ugandaensis Kovařík, 2011, ♂ holotype, tarsomere II of 4th leg. 8: Pandinus smithi (Pocock, 1897), ♂ (107 mm), Ethiopia, 55 km of Degehbur, 07°49'27.2"N 43°41'56.3"E, 1053 m a.s.l., FKCP, tarsomere II of 4th leg." pageId="2" pageNumber="1">Figs. 1–6</figureCitation>
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,
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<figureCitation id="3A972A56FF81A945C7CFF908FB1EF973" box="[1165,1220,1691,1715]" captionStart-0="Figures 9–13" captionStart-1="Figures 14–20" captionStart-2="Figures 21–22" captionStart-3="Figures 23–24" captionStart-4="Figures 25–27" captionStart-5="Figures 28–29" captionStart-6="Figures 30–33" captionStart-7="Figures 34–35" captionStart-8="Figure 36" captionStartId-0="4.[192,278,1854,1878]" captionStartId-1="6.[192,278,1464,1488]" captionStartId-2="7.[192,278,1865,1889]" captionStartId-3="8.[192,278,1862,1886]" captionStartId-4="9.[192,278,1833,1857]" captionStartId-5="10.[192,278,1864,1888]" captionStartId-6="12.[192,278,1857,1881]" captionStartId-7="13.[192,278,1870,1894]" captionStartId-8="14.[192,267,1843,1867]" captionTargetBox-0="[227,1401,188,1797]" captionTargetBox-1="[250,1407,176,1425]" captionTargetBox-2="[191,1441,191,1835]" captionTargetBox-3="[191,1441,191,1830]" captionTargetBox-4="[213,1417,167,1805]" captionTargetBox-5="[211,1417,167,1833]" captionTargetBox-6="[191,1441,167,1827]" captionTargetBox-7="[191,1441,167,1838]" captionTargetBox-8="[196,1437,196,1802]" captionTargetId-0="figure-52@4.[214,1416,188,1824]" captionTargetId-1="figure-246@6.[192,1440,168,1435]" captionTargetId-2="figure-34@7.[192,1440,192,1835]" captionTargetId-3="figure-42@8.[192,1440,192,1829]" captionTargetId-4="figure-65@9.[214,1416,168,1804]" captionTargetId-5="figure-59@10.[212,1416,168,1832]" captionTargetId-6="figure-58@12.[192,1440,168,1827]" captionTargetId-7="figure-40@13.[192,1440,168,1837]" captionTargetId-8="figure-43@14.[192,1440,192,1813]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionText-0="Figures 9–13: Pandinus trailini sp. n., ♀ (116 mm) paratype. 9–10. Dorsal and ventral views. 11. Pectinal area. 12. Chela dorsoexternal view. 13. Metasoma and telson, lateral view." captionText-1="Figures 14–20: Pandinus trailini sp. n., ♂ holotype, trichobothrial pattern. 14. Chela dorsal. 15. Chela external. 16. Chela ventral. 17. Patella dorsal. 18. Patella external. 19. Patella ventral. 20. Femur dorsal." captionText-2="Figures 21–22: Pandinus trailini sp. n., ♂ holotype. 21. Shortly before adulthood (sixth) ecdysis. 22. One day after adulthood ecdysis." captionText-3="Figures 23–24: Pandinus trailini sp. n. 23. Male holotype one month after adulthood ecdysis. 24. Adult female paratype at Locality No. 13EC (Figs. 34–35)." captionText-4="Figures 25–27: Pandinus trailini sp. n., type locality, Ethiopia, Oromia State, Arsi Province, 06°56'06"N 40°41'23"E, 1693 m a.s.l. (Locality No. 13EA). 27. Red arrow points to the spot where a female paratype was found together with 10 juveniles. The overturned rock under which the female was found is to the right." captionText-5="Figures 28–29: Pandinus trailini sp. n., Female paratype at type locality. Well apparent are remains of the prey consumed by the female and juveniles. Fig. 29 shows one of the 10 juveniles found with their mother. This juvenile is after the fourth ecdysis." captionText-6="Figures 30–33: Pandinus trailini sp. n. 30–31. Locality No. 13EB, Oromia State, Arsi Province, 06°54'19"N 40°50'08"E, 1708 m a.s.l. Specimens were found under large rocks apparent in Fig. 31. 32–33. Juvenile after the fifth ecdysis and its exuvia." captionText-7="Figures 34–35: Pandinus trailini sp. n., locality No. 13EC, Oromia State, Arsi Province, Sof Omar, 06°54'19"N 40°51'04"E, 1200 m a.s.l. Most specimens were found in rock fissures." captionText-8="Figure 36: Distribution map of the subgenus Pandinus. Localities of P. imperator and P. gambiensis are after Prendini (2004: 255, fig. 13), localities of other species are after Kovařík (2011)." pageId="2" pageNumber="1">9–36</figureCitation>
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)
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<emphasis id="90D8EAC1FF81A945C004F928FA8BF931" bold="true" pageId="2" pageNumber="1">
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http://zoobank.org/
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<uri id="D63D3AD1FF81A945C760F928FA8BF931" pageId="2" pageNumber="1">
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urn:lsid:zoobank.org:act:
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<uuid id="D60A0C06FF81A945C604F928FA8BF931" pageId="2" pageNumber="1">E01BC A9A-F57C-4EC5-B4F8-7034BBF1898B</uuid>
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</paragraph>
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</subSubSection>
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<subSubSection id="EAB66558FF81A942C07DF884FCC1FC40" lastPageId="5" lastPageNumber="4" pageId="2" pageNumber="1" type="materials_examined">
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<paragraph id="A21336D3FF81A945C07DF884FA42F88D" blockId="2.[831,1441,1814,1869]" pageId="2" pageNumber="1">
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<materialsCitation id="12C43C8EFF81A945C07DF884FC68F88D" collectionCode="TYPE" country="Ethiopia" pageId="2" pageNumber="1" stateProvince="Oromia State" typeStatus="TYPE">
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<collectionCode id="C4BDAE16FF81A945C07DF884FCADF8ED" box="[831,887,1815,1838]" pageId="2" pageNumber="1">
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<typeStatus id="7D178871FF81A945C07DF884FCADF8ED" box="[831,887,1815,1838]" pageId="2" pageNumber="1">TYPE</typeStatus>
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</collectionCode>
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||
LOCALITY AND
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||
<typeStatus id="7D178871FF81A945C76FF889FBB8F8ED" box="[1069,1122,1818,1837]" pageId="2" pageNumber="1">
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<collectionCode id="C4BDAE16FF81A945C76FF889FBB8F8ED" box="[1069,1122,1818,1837]" pageId="2" pageNumber="1">TYPE</collectionCode>
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</typeStatus>
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DEPOSITORY.
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||
<collectingCountry id="DABB7643FF81A945C640F885FA85F8EE" box="[1282,1375,1814,1838]" name="Ethiopia" pageId="2" pageNumber="1" value="Ethiopia">Ethiopia</collectingCountry>
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||
,
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<collectingRegion id="6068F831FF81A945C62CF885FC68F88D" country="Ethiopia" name="Oromiya" pageId="2" pageNumber="1">Oromia State</collectingRegion>
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</materialsCitation>
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||
,
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||
<materialsCitation id="12C43C8EFF81A945C089F8A6FA4EF88D" box="[971,1428,1845,1869]" country="Ethiopia" latitude="6.935" longitude="40.689724" pageId="2" pageNumber="1" stateProvince="Arsi Province">
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<collectingRegion id="6068F831FF81A945C089F8A6FBB7F88D" box="[971,1133,1845,1869]" pageId="2" pageNumber="1">Arsi Province</collectingRegion>
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,
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||
<geoCoordinate id="C7985014FF81A945C7C7F8A6FAD9F88D" box="[1157,1283,1845,1869]" degrees="06" direction="north" minutes="56" orientation="latitude" pageId="2" pageNumber="1" precision="15" seconds="06" value="6.935">06°56'06"N</geoCoordinate>
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<geoCoordinate id="C7985014FF81A945C655F8A6FA4EF88D" box="[1303,1428,1845,1869]" degrees="40" direction="east" minutes="41" orientation="longitude" pageId="2" pageNumber="1" precision="15" seconds="23" value="40.689724">40°41'23"E</geoCoordinate>
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</materialsCitation>
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,
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||
</paragraph>
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||
<caption id="F6D3665BFF80A944C382F890FD23F8A5" pageId="3" pageNumber="2" startId="3.[192,278,1795,1819]" targetBox="[222,1387,166,1747]" targetPageId="3">
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<paragraph id="A21336D3FF80A944C382F890FD23F8A5" blockId="3.[192,1441,1795,1893]" pageId="3" pageNumber="2">
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||
<emphasis id="90D8EAC1FF80A944C382F890FE5CF8DA" bold="true" box="[192,390,1795,1819]" pageId="3" pageNumber="2">Figures 1–8: 1–6:</emphasis>
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<taxonomicName id="65AC4D50FF80A944C2D2F896FDEFF8DB" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[400,565,1797,1819]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
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||
<emphasis id="90D8EAC1FF80A944C2D2F896FDEFF8DB" box="[400,565,1797,1819]" italics="true" pageId="3" pageNumber="2">Pandinus trailini</emphasis>
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</taxonomicName>
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<emphasis id="90D8EAC1FF80A944C17DF896FDA3F8DA" bold="true" box="[575,633,1797,1818]" pageId="3" pageNumber="2">
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||
<taxonomicNameLabel id="8BEB57BAFF80A944C17DF896FDA3F8DA" box="[575,633,1797,1818]" pageId="3" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, ♂ holotype.
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<emphasis id="90D8EAC1FF80A944C045F896FCF1F8DA" bold="true" box="[775,811,1797,1818]" pageId="3" pageNumber="2">1–2</emphasis>
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. Dorsal and ventral views.
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<emphasis id="90D8EAC1FF80A944C700F896FB94F8DA" bold="true" box="[1090,1102,1797,1818]" pageId="3" pageNumber="2">3</emphasis>
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. Pectinal area.
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<emphasis id="90D8EAC1FF80A944C7A9F896FB2DF8DA" bold="true" box="[1259,1271,1797,1818]" pageId="3" pageNumber="2">4</emphasis>
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. Metasoma and telson, lateral view.
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<emphasis id="90D8EAC1FF80A944C2CBF88DFE77F8F3" bold="true" box="[393,429,1822,1843]" pageId="3" pageNumber="2">5–6</emphasis>
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. Tarsomere II of 4th leg, lateral and ventral views.
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<emphasis id="90D8EAC1FF80A944C0DCF88DFC68F8F3" bold="true" box="[926,946,1822,1843]" pageId="3" pageNumber="2">7:</emphasis>
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||
<taxonomicName id="65AC4D50FF80A944C0FAF88DFAFEF8F4" authority="Kovarik, 2011" authorityName="Kovarik" authorityYear="2011" box="[952,1316,1822,1844]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="ugandaensis">
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<emphasis id="90D8EAC1FF80A944C0FAF88DFB4BF8F4" box="[952,1169,1822,1844]" italics="true" pageId="3" pageNumber="2">Pandinus ugandaensis</emphasis>
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Kovařík, 2011
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</taxonomicName>
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, ♂ holotype, tarsomere II of 4th leg.
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<emphasis id="90D8EAC1FF80A944C286F8A4FE02F88C" bold="true" box="[452,472,1847,1868]" pageId="3" pageNumber="2">8:</emphasis>
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||
<taxonomicName id="65AC4D50FF80A944C2A7F8A5FCF5F88C" authority="(Pocock, 1897)" baseAuthorityName="Pocock" baseAuthorityYear="1897" box="[485,815,1846,1868]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="smithi">
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<emphasis id="90D8EAC1FF80A944C2A7F8A5FD52F88C" box="[485,648,1846,1868]" italics="true" pageId="3" pageNumber="2">Pandinus smithi</emphasis>
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(Pocock, 1897)
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</taxonomicName>
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, ♂ (107 mm), Ethiopia, 55 km of Degehbur, 07°49'27.2"N 43°41'56.3"E, 1053 m a.s.l., FKCP, tarsomere II of 4th leg.
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||
</paragraph>
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||
</caption>
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||
<caption id="F6D3665BFF87A943C382F8ADFD2CF8AD" pageId="4" pageNumber="3" startId="4.[192,278,1854,1878]" targetBox="[227,1401,188,1797]" targetPageId="4">
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<paragraph id="A21336D3FF87A943C382F8ADFD2CF8AD" blockId="4.[192,1440,1854,1902]" pageId="4" pageNumber="3">
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||
<emphasis id="90D8EAC1FF87A943C382F8ADFE84F896" bold="true" box="[192,350,1854,1878]" pageId="4" pageNumber="3">Figures 9–13:</emphasis>
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<taxonomicName id="65AC4D50FF87A943C225F8ACFDD7F895" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[359,525,1855,1877]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
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<emphasis id="90D8EAC1FF87A943C225F8ACFDD7F895" box="[359,525,1855,1877]" italics="true" pageId="4" pageNumber="3">Pandinus trailini</emphasis>
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||
</taxonomicName>
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||
<emphasis id="90D8EAC1FF87A943C154F8ACFD95F894" bold="true" box="[534,591,1855,1876]" pageId="4" pageNumber="3">
|
||
<taxonomicNameLabel id="8BEB57BAFF87A943C154F8ACFD95F894" box="[534,591,1855,1876]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
, ♀ (116 mm) paratype.
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||
<emphasis id="90D8EAC1FF87A943C004F8ACFCADF894" bold="true" box="[838,887,1855,1876]" pageId="4" pageNumber="3">9–10</emphasis>
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||
. Dorsal and ventral views.
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||
<emphasis id="90D8EAC1FF87A943C7CFF8ACFB7FF894" bold="true" box="[1165,1189,1855,1876]" pageId="4" pageNumber="3">11</emphasis>
|
||
. Pectinal area.
|
||
<emphasis id="90D8EAC1FF87A943C603F8ACFA83F894" bold="true" box="[1345,1369,1855,1876]" pageId="4" pageNumber="3">12</emphasis>
|
||
. Chela dorsoexternal view.
|
||
<emphasis id="90D8EAC1FF87A943C2C1F8CBFE41F8AD" bold="true" box="[387,411,1880,1901]" pageId="4" pageNumber="3">13</emphasis>
|
||
. Metasoma and telson, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A21336D3FF86A942C382FF55FEB1FF3C" blockId="5.[192,801,198,252]" pageId="5" pageNumber="4">
|
||
<materialsCitation id="12C43C8EFF86A942C382FF55FD89FF1E" box="[192,595,198,222]" collectionCode="EA" country="Ethiopia" pageId="5" pageNumber="4">
|
||
<quantity id="65549B36FF86A942C382FF55FECDFF1D" box="[192,279,198,222]" metricMagnitude="3" metricUnit="m" metricValue="1.693" pageId="5" pageNumber="4" unit="m" value="1693.0">1693 m</quantity>
|
||
a.s.l. (Locality No. 13
|
||
<collectionCode id="C4BDAE16FF86A942C160FF55FD90FF1E" box="[546,586,198,222]" pageId="5" pageNumber="4">EA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="12C43C8EFF86A942C127FF55FEBDFF3C" collectionCode="FKCP" country="Ethiopia" pageId="5" pageNumber="4">
|
||
the authoŕs collection (
|
||
<collectionCode id="C4BDAE16FF86A942C255FF77FEBAFF3C" box="[279,352,228,252]" pageId="5" pageNumber="4">FKCP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A21336D3FF86A942C382FEB1FE4EFD6A" blockId="5.[192,802,289,896]" pageId="5" pageNumber="4">
|
||
<materialsCitation id="12C43C8EFF86A942C382FEB1FDA1FEF9" box="[192,635,289,314]" collectionCode="TYPE" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Oromia State" typeStatus="TYPE">
|
||
<collectionCode id="C4BDAE16FF86A942C382FEB1FF23FEF9" box="[192,249,290,313]" pageId="5" pageNumber="4">
|
||
<typeStatus id="7D178871FF86A942C382FEB1FF23FEF9" box="[192,249,290,313]" pageId="5" pageNumber="4">TYPE</typeStatus>
|
||
</collectionCode>
|
||
MATERIAL.
|
||
<emphasis id="90D8EAC1FF86A942C238FEB1FE01FEFA" bold="true" box="[378,475,290,314]" pageId="5" pageNumber="4">
|
||
<collectingCountry id="DABB7643FF86A942C238FEB1FE01FEFA" box="[378,475,290,314]" name="Ethiopia" pageId="5" pageNumber="4" value="Ethiopia">Ethiopia</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="6068F831FF86A942C2A8FEB2FDA1FEF9" box="[490,635,289,313]" country="Ethiopia" name="Oromiya" pageId="5" pageNumber="4">Oromia State</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="12C43C8EFF86A942C1C5FEB2FD55FE98" country="Ethiopia" latitude="6.935" longitude="40.689724" pageId="5" pageNumber="4" stateProvince="Arsi Province">
|
||
<collectingRegion id="6068F831FF86A942C1C5FEB2FCC4FEF9" box="[647,798,289,313]" pageId="5" pageNumber="4">Arsi Province</collectingRegion>
|
||
,
|
||
<geoCoordinate id="C7985014FF86A942C382FED3FEE4FE98" box="[192,318,320,344]" degrees="06" direction="north" minutes="56" orientation="latitude" pageId="5" pageNumber="4" precision="15" seconds="06" value="6.935">06°56'06"N</geoCoordinate>
|
||
<geoCoordinate id="C7985014FF86A942C20FFED3FE10FE98" box="[333,458,320,344]" degrees="40" direction="east" minutes="41" orientation="longitude" pageId="5" pageNumber="4" precision="15" seconds="23" value="40.689724">40°41'23"E</geoCoordinate>
|
||
,
|
||
<quantity id="65549B36FF86A942C29CFED3FDEDFE98" box="[478,567,320,344]" metricMagnitude="3" metricUnit="m" metricValue="1.693" pageId="5" pageNumber="4" unit="m" value="1693.0">1693 m</quantity>
|
||
a.s.l. (
|
||
</materialsCitation>
|
||
<materialsCitation id="12C43C8EFF86A942C1D2FED3FED3FEB7" collectionCode="EA" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No">
|
||
<collectingRegion id="6068F831FF86A942C1D2FED3FCC4FE98" box="[656,798,320,344]" pageId="5" pageNumber="4">Locality No</collectingRegion>
|
||
. 13
|
||
<collectionCode id="C4BDAE16FF86A942C39BFECCFEDBFEB7" box="[217,257,351,375]" pageId="5" pageNumber="4">EA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="12C43C8EFF86A942C25BFECCFDEFFE74" collectingDate="2013-06-24" collectionCode="V" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No" typeStatus="holotype">
|
||
<date id="D6121013FF86A942C25BFECCFE57FEB7" box="[281,397,351,375]" pageId="5" pageNumber="4" value="2012-07-05">5.VII.2012</date>
|
||
,
|
||
<specimenCount id="B4AAFD5AFF86A942C2E2FECCFE1BFEB7" box="[416,449,351,375]" pageId="5" pageNumber="4" type="male">1♂</specimenCount>
|
||
(
|
||
<typeStatus id="7D178871FF86A942C291FECCFDE9FEB7" box="[467,563,351,375]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
) and its exuvia from
|
||
<date id="D6121013FF86A942C382FEEDFEF3FE56" box="[192,297,382,406]" pageId="5" pageNumber="4" value="2013-01-20">20.I.2013</date>
|
||
, leg.
|
||
<collectionCode id="C4BDAE16FF86A942C22BFEEDFEA5FE55" box="[361,383,382,405]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
|
||
. Trailin,
|
||
<date id="D6121013FF86A942C2A5FEEDFDBEFE56" box="[487,612,382,406]" pageId="5" pageNumber="4" value="2013-06-24">
|
||
24.
|
||
<collectionCode id="C4BDAE16FF86A942C14AFEEDFDFDFE56" box="[520,551,382,406]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="4">VI</collectionCode>
|
||
.2013
|
||
</date>
|
||
,
|
||
<specimenCount id="B4AAFD5AFF86A942C131FEEDFD57FE56" box="[627,653,382,406]" pageId="5" pageNumber="4" type="female">1♀</specimenCount>
|
||
10 juvs after 3rd and 4th ecdyses (
|
||
<typeStatus id="7D178871FF86A942C280FE0FFDF4FE74" box="[450,558,412,436]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="12C43C8EFF86A942C10BFE0FFE71FE13" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No">leg. F. Kovařík, J. Plíšková, P. Novák</materialsCitation>
|
||
;
|
||
<materialsCitation id="12C43C8EFF86A942C281FE28FDBBFE13" box="[451,609,443,467]" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Oromia State">
|
||
<collectingRegion id="6068F831FF86A942C281FE28FDBBFE13" box="[451,609,443,467]" country="Ethiopia" name="Oromiya" pageId="5" pageNumber="4">Oromia State</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="12C43C8EFF86A942C138FE28FD55FE31" country="Ethiopia" latitude="6.9052777" longitude="40.835556" pageId="5" pageNumber="4" stateProvince="Arsi Province">
|
||
<collectingRegion id="6068F831FF86A942C138FE28FCC7FE13" box="[634,797,443,467]" pageId="5" pageNumber="4">Arsi Province</collectingRegion>
|
||
,
|
||
<geoCoordinate id="C7985014FF86A942C382FE4AFEE7FE31" box="[192,317,473,497]" degrees="06" direction="north" minutes="54" orientation="latitude" pageId="5" pageNumber="4" precision="15" seconds="19" value="6.9052777">06°54'19"N</geoCoordinate>
|
||
<geoCoordinate id="C7985014FF86A942C20FFE4AFE11FE31" box="[333,459,473,497]" degrees="40" direction="east" minutes="50" orientation="longitude" pageId="5" pageNumber="4" precision="15" seconds="08" value="40.835556">40°50'08"E</geoCoordinate>
|
||
,
|
||
<quantity id="65549B36FF86A942C29CFE4AFDEDFE31" box="[478,567,473,497]" metricMagnitude="3" metricUnit="m" metricValue="1.708" pageId="5" pageNumber="4" unit="m" value="1708.0">1708 m</quantity>
|
||
a.s.l. (
|
||
</materialsCitation>
|
||
<materialsCitation id="12C43C8EFF86A942C1D2FE4AFED2FDD0" collectionCode="EB" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No">
|
||
<collectingRegion id="6068F831FF86A942C1D2FE4AFCC4FE31" box="[656,798,473,497]" pageId="5" pageNumber="4">Locality No</collectingRegion>
|
||
. 13
|
||
<collectionCode id="C4BDAE16FF86A942C398FE6BFEDAFDD0" box="[218,256,504,528]" pageId="5" pageNumber="4">EB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="12C43C8EFF86A942C25AFE6BFD9AFDD0" box="[280,576,504,528]" collectingDate="2013-06-24" collectionCode="VI" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No">
|
||
<date id="D6121013FF86A942C25AFE6BFE4EFDD0" box="[280,404,504,528]" pageId="5" pageNumber="4" value="2013-06-24">
|
||
24.
|
||
<collectionCode id="C4BDAE16FF86A942C27AFE6BFE8CFDD0" box="[312,342,504,528]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="4">VI</collectionCode>
|
||
.2013
|
||
</date>
|
||
,
|
||
<specimenCount id="B4AAFD5AFF86A942C2E6FE6BFE1BFDD0" box="[420,449,504,528]" pageId="5" pageNumber="4" type="female">4♀</specimenCount>
|
||
1im.12juvs
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="12C43C8EFF86A942C10DFE6BFD12FDD0" box="[591,712,504,528]" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No" typeStatus="paratype">
|
||
(
|
||
<typeStatus id="7D178871FF86A942C114FE6BFD18FDD0" box="[598,706,504,528]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="12C43C8EFF86A942C19BFE6BFD5AFDEF" collectionCode="V" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No">
|
||
leg. F. Kovařík, J. Plíšková, P. Novák,
|
||
<collectionCode id="C4BDAE16FF86A942C158FD8BFDEAFDEF" box="[538,560,536,559]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
|
||
. Socha
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="12C43C8EFF86A942C1CEFD84FCC7FDEF" box="[652,797,535,559]" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Oromia State">
|
||
<collectingRegion id="6068F831FF86A942C1CEFD84FCC7FDEF" box="[652,797,535,559]" country="Ethiopia" name="Oromiya" pageId="5" pageNumber="4">Oromia State</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="12C43C8EFF86A942C382FDA5FEECFDAC" country="Ethiopia" latitude="6.9052777" longitude="40.85111" pageId="5" pageNumber="4" stateProvince="Arsi Province">
|
||
<collectingRegion id="6068F831FF86A942C382FDA5FE8DFD8E" box="[192,343,566,590]" pageId="5" pageNumber="4">Arsi Province</collectingRegion>
|
||
, Sof Omar,
|
||
<geoCoordinate id="C7985014FF86A942C29EFDA5FD80FD8E" box="[476,602,566,590]" degrees="06" direction="north" minutes="54" orientation="latitude" pageId="5" pageNumber="4" precision="15" seconds="19" value="6.9052777">06°54'19"N</geoCoordinate>
|
||
<geoCoordinate id="C7985014FF86A942C120FDA5FD05FD8E" box="[610,735,566,590]" degrees="40" direction="east" minutes="51" orientation="longitude" pageId="5" pageNumber="4" precision="15" seconds="04" value="40.85111">40°51'04"E</geoCoordinate>
|
||
,
|
||
<quantity id="65549B36FF86A942C1AEFDA5FF0EFDAC" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="4" unit="m" value="1200.0">1200 m</quantity>
|
||
a.s.l. (
|
||
</materialsCitation>
|
||
<materialsCitation id="12C43C8EFF86A942C274FDC7FDF2FDAC" box="[310,552,596,620]" collectionCode="EC" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No">
|
||
<collectingRegion id="6068F831FF86A942C274FDC7FE13FDAC" box="[310,457,596,620]" pageId="5" pageNumber="4">Locality No</collectingRegion>
|
||
. 13
|
||
<collectionCode id="C4BDAE16FF86A942C2B8FDC7FDFAFDAC" box="[506,544,596,620]" pageId="5" pageNumber="4">EC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="12C43C8EFF86A942C103FDC7FEF8FD4B" collectingDate="2013-06-24" collectionCode="VI" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No">
|
||
<date id="D6121013FF86A942C103FDC7FD3DFDAC" box="[577,743,596,620]" pageId="5" pageNumber="4" value="2013-06-24" valueMax="2013-06-25" valueMin="2013-06-24">
|
||
24.-25.
|
||
<collectionCode id="C4BDAE16FF86A942C1C9FDC7FD70FDAC" box="[651,682,596,620]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="4">VI</collectionCode>
|
||
.2013
|
||
</date>
|
||
,
|
||
<specimenCount id="B4AAFD5AFF86A942C043FDC7FCFBFDAC" box="[769,801,596,620]" pageId="5" pageNumber="4" type="female">3♀</specimenCount>
|
||
1im.
|
||
<specimenCount id="B4AAFD5AFF86A942C3ADFDE0FEF8FD4B" box="[239,290,627,651]" pageId="5" pageNumber="4" type="generic">1juv</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="12C43C8EFF86A942C26CFDE0FD88FD4B" box="[302,594,627,651]" collectionCode="UV" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No" typeStatus="paratype">
|
||
(
|
||
<typeStatus id="7D178871FF86A942C277FDE0FE44FD4B" box="[309,414,627,651]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode id="C4BDAE16FF86A942C2F7FDE0FE07FD4B" box="[437,477,627,651]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="5" pageNumber="4">UV</collectionCode>
|
||
detection)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="12C43C8EFF86A942C11CFDE0FE55FD6A" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No">leg. F. Kovařík, J. Plíšková, P. Novák</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A21336D3FF86A942C3B2FD22FCC1FC40" blockId="5.[192,802,289,896]" pageId="5" pageNumber="4">
|
||
<materialsCitation id="12C43C8EFF86A942C3B2FD22FDFDFCE4" collectionCode="FKCP" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No" typeStatus="type">
|
||
Most of adult specimens are in 75% alcohol, only
|
||
<specimenCount id="B4AAFD5AFF86A942C382FD43FE98FD27" box="[192,322,719,743]" pageId="5" pageNumber="4" type="female">two females</specimenCount>
|
||
from
|
||
<collectingRegion id="6068F831FF86A942C2C1FD5CFE27FD27" box="[387,509,719,743]" pageId="5" pageNumber="4">locality No</collectingRegion>
|
||
. 13EC and all juveniles are still alive. All
|
||
<typeStatus id="7D178871FF86A942C227FD7DFE49FCC6" box="[357,403,750,774]" pageId="5" pageNumber="4">type</typeStatus>
|
||
specimens are in
|
||
<collectionCode id="C4BDAE16FF86A942C121FD7DFD7CFCC6" box="[611,678,750,774]" pageId="5" pageNumber="4">FKCP</collectionCode>
|
||
collection. Other females and juveniles from
|
||
</materialsCitation>
|
||
<materialsCitation id="12C43C8EFF86A942C16DFC9FFCCCFC40" country="Ethiopia" pageId="5" pageNumber="4" stateProvince="Locality No" typeStatus="types">
|
||
<collectingRegion id="6068F831FF86A942C16DFC9FFD70FCE4" box="[559,682,780,804]" pageId="5" pageNumber="4">locality No</collectingRegion>
|
||
. 13EC that were found associated with the
|
||
<typeStatus id="7D178871FF86A942C154FCBFFD94FC84" box="[534,590,812,836]" pageId="5" pageNumber="4">types</typeStatus>
|
||
but not included in the
|
||
<typeStatus id="7D178871FF86A942C3ACFCD9FEC6FCA2" box="[238,284,842,866]" pageId="5" pageNumber="4">type</typeStatus>
|
||
series are kept alive and cared for by Jana Plíšková, Pavel Novák, Vít Socha, and Vladimír Trailin
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EAB66558FF86A942C382FC35FF32FC3B" pageId="5" pageNumber="4" type="etymology">
|
||
<paragraph id="A21336D3FF86A942C382FC35FF32FC3B" blockId="5.[192,802,934,1019]" pageId="5" pageNumber="4">ETYMOLOGY. Named after Vladimír Trailin, a Czech herpetologist and my friend, who visited Ethiopia with me.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EAB66558FF86A942C382FBB2FD89F9A0" pageId="5" pageNumber="4" type="diagnosis">
|
||
<paragraph id="A21336D3FF86A942C382FBB2FD89F9A0" blockId="5.[192,802,1056,1632]" pageId="5" pageNumber="4">
|
||
DIAGNOSIS. Total length
|
||
<quantity id="65549B36FF86A942C29BFBB3FDBBFBF8" box="[473,609,1056,1080]" metricMagnitude="-1" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.16" metricValueMin="0.98" pageId="5" pageNumber="4" unit="mm" value="107.0" valueMax="116.0" valueMin="98.0">98–116 mm</quantity>
|
||
. Color uniformly brown to reddish black, only chela reddish brown. Chelicerae brown, reticulate, with black fingers and anterior margin. External trichobothria on patella number 17–19 (5–6
|
||
<emphasis id="90D8EAC1FF86A942C237FB08FE54FB73" box="[373,398,1179,1203]" italics="true" pageId="5" pageNumber="4">eb</emphasis>
|
||
, 3
|
||
<emphasis id="90D8EAC1FF86A942C2F4FB08FE03FB73" box="[438,473,1179,1203]" italics="true" pageId="5" pageNumber="4">esb</emphasis>
|
||
, 2
|
||
<emphasis id="90D8EAC1FF86A942C140FB0FFDFBFB73" box="[514,545,1180,1203]" italics="true" pageId="5" pageNumber="4">em</emphasis>
|
||
, 1
|
||
<emphasis id="90D8EAC1FF86A942C10BFB0FFDBCFB73" box="[585,614,1180,1203]" italics="true" pageId="5" pageNumber="4">est</emphasis>
|
||
, 6–7
|
||
<emphasis id="90D8EAC1FF86A942C1E8FB0FFD67FB73" box="[682,701,1180,1203]" italics="true" pageId="5" pageNumber="4">et</emphasis>
|
||
); ventral trichobothria on chela number 16–18. Carapace lacking carinae and with very fine sparse granules. Dorsal carinae on first throught fourth metasomal segments terminate in a larger tooth most conspicuous on fourth segment. Spination formula of tarsomere II = 6/4: 6-7/4: 7-8/4-5: 7-8/4-5. Tarsomere II with 2 spines on inclined anteroventral surface. Pectinal teeth number 16–20. Pedipalps densely hirsute, mainly chela. Granules on dorsal surface of chela conical, conspicuous but not pointed. External surface of chela with conical granules in anterior part and without carinae. Length to depth ratio of 4th metasomal segment = 1.73–2.02. Male has more pronounced tooth on movable finger of pedipalp and slightly larger telson than female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EAB66558FF86A94CC382F914FEE1FA68" lastPageId="11" lastPageNumber="10" pageId="5" pageNumber="4" type="description">
|
||
<paragraph id="A21336D3FF86A942C382F914FCFBF8B4" blockId="5.[192,802,1670,1908]" pageId="5" pageNumber="4">
|
||
DESCRIPTION. The adult male
|
||
<typeStatus id="7D178871FF86A942C14BF915FDBCF95E" box="[521,614,1670,1694]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
is
|
||
<quantity id="65549B36FF86A942C1CBF915FD39F95E" box="[649,739,1670,1694]" metricMagnitude="-1" metricUnit="m" metricValue="1.0699999999999998" pageId="5" pageNumber="4" unit="mm" value="107.0">107 mm</quantity>
|
||
long, and female
|
||
<typeStatus id="7D178871FF86A942C202F936FE7CF97D" box="[320,422,1701,1725]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
|
||
are
|
||
<quantity id="65549B36FF86A942C29AF937FD81F97C" box="[472,603,1700,1724]" metricMagnitude="-1" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.16" metricValueMin="0.98" pageId="5" pageNumber="4" unit="mm" value="107.0" valueMax="116.0" valueMin="98.0">98–116 mm</quantity>
|
||
long. The habitus is shown in
|
||
<figureCitation id="3A972A56FF86A942C21FF950FE11F91B" box="[349,459,1731,1755]" captionStart="Figures 1–8" captionStartId="3.[192,278,1795,1819]" captionTargetBox="[222,1387,166,1747]" captionTargetId="figure-96@3.[212,1416,165,1765]" captionTargetPageId="3" captionText="Figures 1–8: 1–6: Pandinus trailini sp. n., ♂ holotype. 1–2. Dorsal and ventral views. 3. Pectinal area. 4. Metasoma and telson, lateral view. 5–6. Tarsomere II of 4th leg, lateral and ventral views. 7: Pandinus ugandaensis Kovařík, 2011, ♂ holotype, tarsomere II of 4th leg. 8: Pandinus smithi (Pocock, 1897), ♂ (107 mm), Ethiopia, 55 km of Degehbur, 07°49'27.2"N 43°41'56.3"E, 1053 m a.s.l., FKCP, tarsomere II of 4th leg." pageId="5" pageNumber="4">
|
||
Figs.
|
||
<date id="D6121013FF86A942C2E1F950FE11F91B" box="[419,459,1731,1755]" pageId="5" pageNumber="4" value="2010-02-01" valueMax="2010-09-01" valueMin="2010-02-01">1–2</date>
|
||
</figureCitation>
|
||
and
|
||
<figureCitation id="3A972A56FF86A942C151F950FD91F91B" box="[531,587,1731,1755]" captionStart="Figures 9–13" captionStartId="4.[192,278,1854,1878]" captionTargetBox="[227,1401,188,1797]" captionTargetId="figure-52@4.[214,1416,188,1824]" captionTargetPageId="4" captionText="Figures 9–13: Pandinus trailini sp. n., ♀ (116 mm) paratype. 9–10. Dorsal and ventral views. 11. Pectinal area. 12. Chela dorsoexternal view. 13. Metasoma and telson, lateral view." pageId="5" pageNumber="4">9–10</figureCitation>
|
||
. For position and distribution of trichobothria of pedipalps see
|
||
<figureCitation id="3A972A56FF86A942C1F8F971FF04F8D8" captionStart="Figures 14–20" captionStartId="6.[192,278,1464,1488]" captionTargetBox="[250,1407,176,1425]" captionTargetId="figure-246@6.[192,1440,168,1435]" captionTargetPageId="6" captionText="Figures 14–20: Pandinus trailini sp. n., ♂ holotype, trichobothrial pattern. 14. Chela dorsal. 15. Chela external. 16. Chela ventral. 17. Patella dorsal. 18. Patella external. 19. Patella ventral. 20. Femur dorsal." pageId="5" pageNumber="4">Figs. 14– 20</figureCitation>
|
||
. External trichobothria on the patella number 17–19 (5–6
|
||
<emphasis id="90D8EAC1FF86A942C3BBF88CFEC8F8F7" box="[249,274,1823,1847]" italics="true" pageId="5" pageNumber="4">eb</emphasis>
|
||
, 3
|
||
<emphasis id="90D8EAC1FF86A942C274F88CFE83F8F7" box="[310,345,1823,1847]" italics="true" pageId="5" pageNumber="4">esb</emphasis>
|
||
, 2
|
||
<emphasis id="90D8EAC1FF86A942C23FF8B3FE46F8F7" box="[381,412,1824,1847]" italics="true" pageId="5" pageNumber="4">em</emphasis>
|
||
, 1
|
||
<emphasis id="90D8EAC1FF86A942C2FDF8B3FE07F8F7" box="[447,477,1824,1847]" italics="true" pageId="5" pageNumber="4">est</emphasis>
|
||
, 6–7
|
||
<emphasis id="90D8EAC1FF86A942C159F8B3FDF4F8F7" box="[539,558,1824,1847]" italics="true" pageId="5" pageNumber="4">et</emphasis>
|
||
); ventral trichobothria on the chela number 16–18. The male has a more pronounced tooth on the movable finger of pedipalp
|
||
</paragraph>
|
||
<paragraph id="A21336D3FF86A942C07DFF55FA46F8B4" blockId="5.[830,1448,198,1908]" pageId="5" pageNumber="4">
|
||
(
|
||
<figureCitation id="3A972A56FF86A942C004FF55FC4FFF1E" box="[838,917,198,222]" captionStart="Figures 14–20" captionStartId="6.[192,278,1464,1488]" captionTargetBox="[250,1407,176,1425]" captionTargetId="figure-246@6.[192,1440,168,1435]" captionTargetPageId="6" captionText="Figures 14–20: Pandinus trailini sp. n., ♂ holotype, trichobothrial pattern. 14. Chela dorsal. 15. Chela external. 16. Chela ventral. 17. Patella dorsal. 18. Patella external. 19. Patella ventral. 20. Femur dorsal." pageId="5" pageNumber="4">Fig. 14</figureCitation>
|
||
versus
|
||
<figureCitation id="3A972A56FF86A942C0ABFF55FBE2FF1E" box="[1001,1080,198,222]" captionStart="Figures 9–13" captionStartId="4.[192,278,1854,1878]" captionTargetBox="[227,1401,188,1797]" captionTargetId="figure-52@4.[214,1416,188,1824]" captionTargetPageId="4" captionText="Figures 9–13: Pandinus trailini sp. n., ♀ (116 mm) paratype. 9–10. Dorsal and ventral views. 11. Pectinal area. 12. Chela dorsoexternal view. 13. Metasoma and telson, lateral view." pageId="5" pageNumber="4">Fig. 12</figureCitation>
|
||
) and a slightly larger telson than the female (
|
||
<figureCitation id="3A972A56FF86A942C0FFFF77FC21FF3C" box="[957,1019,228,252]" captionStart="Figures 1–8" captionStartId="3.[192,278,1795,1819]" captionTargetBox="[222,1387,166,1747]" captionTargetId="figure-96@3.[212,1416,165,1765]" captionTargetPageId="3" captionText="Figures 1–8: 1–6: Pandinus trailini sp. n., ♂ holotype. 1–2. Dorsal and ventral views. 3. Pectinal area. 4. Metasoma and telson, lateral view. 5–6. Tarsomere II of 4th leg, lateral and ventral views. 7: Pandinus ugandaensis Kovařík, 2011, ♂ holotype, tarsomere II of 4th leg. 8: Pandinus smithi (Pocock, 1897), ♂ (107 mm), Ethiopia, 55 km of Degehbur, 07°49'27.2"N 43°41'56.3"E, 1053 m a.s.l., FKCP, tarsomere II of 4th leg." pageId="5" pageNumber="4">Fig. 4</figureCitation>
|
||
versus
|
||
<figureCitation id="3A972A56FF86A942C70FFF77FB46FF3C" box="[1101,1180,228,252]" captionStart="Figures 9–13" captionStartId="4.[192,278,1854,1878]" captionTargetBox="[227,1401,188,1797]" captionTargetId="figure-52@4.[214,1416,188,1824]" captionTargetPageId="4" captionText="Figures 9–13: Pandinus trailini sp. n., ♀ (116 mm) paratype. 9–10. Dorsal and ventral views. 11. Pectinal area. 12. Chela dorsoexternal view. 13. Metasoma and telson, lateral view." pageId="5" pageNumber="4">Fig. 12</figureCitation>
|
||
). COLORATION (
|
||
<figureCitation id="3A972A56FF86A942C0A7FE90FB82FEDB" box="[997,1112,259,283]" captionStart="Figures 1–8" captionStartId="3.[192,278,1795,1819]" captionTargetBox="[222,1387,166,1747]" captionTargetId="figure-96@3.[212,1416,165,1765]" captionTargetPageId="3" captionText="Figures 1–8: 1–6: Pandinus trailini sp. n., ♂ holotype. 1–2. Dorsal and ventral views. 3. Pectinal area. 4. Metasoma and telson, lateral view. 5–6. Tarsomere II of 4th leg, lateral and ventral views. 7: Pandinus ugandaensis Kovařík, 2011, ♂ holotype, tarsomere II of 4th leg. 8: Pandinus smithi (Pocock, 1897), ♂ (107 mm), Ethiopia, 55 km of Degehbur, 07°49'27.2"N 43°41'56.3"E, 1053 m a.s.l., FKCP, tarsomere II of 4th leg." pageId="5" pageNumber="4">Figs. 1–2</figureCitation>
|
||
). The carapace, mesosoma, metasoma and femur and patella of pedipalp are uniformly brown to reddish black. The chela is reddish brown and its fingers are black. The legs and telson are grayish brown to black. The chelicerae are brown, reticulate, with black fingers and anterior margins. CARAPACE. The carapace lacks carinae but has a deep sagittal furrow with a forked, Y-shaped furrow on each side in the posterior part. The surface is granulated by very fine, sparse granules, but the anterior part is smooth, without granules. The anteromedial margin is strongly concave. Present are a pair of median eyes and three lateral eyes on each side, with a furrow behind the lateral eyes. The distance ratio of the pair of median eyes from the anterior and the posterior margin of the carapace is, respectively, 0.56 and 0.44. MESOSOMA. The tergites are finely granulated, each with an incomplete sagittal carina and symmetrical shallow furrows. The sternites are smooth, lack carinae and each bears two pronounced furrows that reach neither anterior nor posterior margins. The pectinal teeth number 16–20. The pectines have three marginal and three middle lamellae, all with numerous reddish setae. The characteristic fulcra are long and bear 12 to 20 white setae on the tip. METASOMA AND TELSON (
|
||
<figureCitation id="3A972A56FF86A942C72CFB91FB6FFBDA" box="[1134,1205,1026,1050]" captionStart="Figures 1–8" captionStartId="3.[192,278,1795,1819]" captionTargetBox="[222,1387,166,1747]" captionTargetId="figure-96@3.[212,1416,165,1765]" captionTargetPageId="3" captionText="Figures 1–8: 1–6: Pandinus trailini sp. n., ♂ holotype. 1–2. Dorsal and ventral views. 3. Pectinal area. 4. Metasoma and telson, lateral view. 5–6. Tarsomere II of 4th leg, lateral and ventral views. 7: Pandinus ugandaensis Kovařík, 2011, ♂ holotype, tarsomere II of 4th leg. 8: Pandinus smithi (Pocock, 1897), ♂ (107 mm), Ethiopia, 55 km of Degehbur, 07°49'27.2"N 43°41'56.3"E, 1053 m a.s.l., FKCP, tarsomere II of 4th leg." pageId="5" pageNumber="4">Fig. 4</figureCitation>
|
||
). The first throught fourth segments bear eight carinae. The first throught third segments bear smooth ventral and lateral carinae; the fourth segment bears these carinae with several smooth teeth. The dorsal carinae on the first throught fourth segments terminate in a larger tooth most conspicuous on the fourth segment. The fifth segment bears five carinae and a row of granules on the lateral surfaces, which may form an incomplete carina. All carinae on the fifth metasomal segment bear strong granules. The surface between the carinae is granulate on the fifth segment and smooth on the other segments. The densely hirsute telson is bulbous (male), with aculeus shorter than the vesicle. The surface of the telson is unevenly granulated and bears incomplete carinae. LEGS. The legs are smooth, without carinae and granules, and unevenly hirsute. Tarsomere Iis hirsute. Tarsomere II has two spines on the inclined anteroventral surface. The spination formula of tarsomere II is 6/4: 6- 7/4: 7-8/4-5: 7-8/4-5 (
|
||
<figureCitation id="3A972A56FF86A942C76AF9DBFB56F9A0" box="[1064,1164,1608,1632]" captionStart="Figures 1–8" captionStartId="3.[192,278,1795,1819]" captionTargetBox="[222,1387,166,1747]" captionTargetId="figure-96@3.[212,1416,165,1765]" captionTargetPageId="3" captionText="Figures 1–8: 1–6: Pandinus trailini sp. n., ♂ holotype. 1–2. Dorsal and ventral views. 3. Pectinal area. 4. Metasoma and telson, lateral view. 5–6. Tarsomere II of 4th leg, lateral and ventral views. 7: Pandinus ugandaensis Kovařík, 2011, ♂ holotype, tarsomere II of 4th leg. 8: Pandinus smithi (Pocock, 1897), ♂ (107 mm), Ethiopia, 55 km of Degehbur, 07°49'27.2"N 43°41'56.3"E, 1053 m a.s.l., FKCP, tarsomere II of 4th leg." pageId="5" pageNumber="4">Figs. 5–6</figureCitation>
|
||
). PEDIPALPS. The pedipalps are densely hirsute, mainly the chela (
|
||
<figureCitation id="3A972A56FF86A942C0FDF915FB96F95E" box="[959,1100,1670,1694]" captionStart="Figures 14–20" captionStartId="6.[192,278,1464,1488]" captionTargetBox="[250,1407,176,1425]" captionTargetId="figure-246@6.[192,1440,168,1435]" captionTargetPageId="6" captionText="Figures 14–20: Pandinus trailini sp. n., ♂ holotype, trichobothrial pattern. 14. Chela dorsal. 15. Chela external. 16. Chela ventral. 17. Patella dorsal. 18. Patella external. 19. Patella ventral. 20. Femur dorsal." pageId="5" pageNumber="4">Figs. 14–16</figureCitation>
|
||
). The femur and patella are bumpy with several large granules. The femur bears four carinae composed of several large, round granules, only the externoventral carina is smooth.The patella bears four to five smooth and incomplete carinae without granules. Two large and several small granules are only on the external surface of the patella. The chela (
|
||
<figureCitation id="3A972A56FF86A942C60BF8ADFA7AF896" box="[1353,1440,1854,1878]" captionStart="Figures 9–13" captionStartId="4.[192,278,1854,1878]" captionTargetBox="[227,1401,188,1797]" captionTargetId="figure-52@4.[214,1416,188,1824]" captionTargetPageId="4" captionText="Figures 9–13: Pandinus trailini sp. n., ♀ (116 mm) paratype. 9–10. Dorsal and ventral views. 11. Pectinal area. 12. Chela dorsoexternal view. 13. Metasoma and telson, lateral view." pageId="5" pageNumber="4">Figs. 12</figureCitation>
|
||
and
|
||
<figureCitation id="3A972A56FF86A942C02EF8CFFC52F8B4" box="[876,904,1884,1908]" captionStart="Figures 14–20" captionStartId="6.[192,278,1464,1488]" captionTargetBox="[250,1407,176,1425]" captionTargetId="figure-246@6.[192,1440,168,1435]" captionTargetPageId="6" captionText="Figures 14–20: Pandinus trailini sp. n., ♂ holotype, trichobothrial pattern. 14. Chela dorsal. 15. Chela external. 16. Chela ventral. 17. Patella dorsal. 18. Patella external. 19. Patella ventral. 20. Femur dorsal." pageId="5" pageNumber="4">14</figureCitation>
|
||
) has a lobe and lacks carinae; its dorsal surface
|
||
</paragraph>
|
||
<caption id="F6D3665BFF85A941C382FA2BFC30FA28" pageId="6" pageNumber="5" startId="6.[192,278,1464,1488]" targetBox="[250,1407,176,1425]" targetPageId="6">
|
||
<paragraph id="A21336D3FF85A941C382FA2BFC30FA28" blockId="6.[192,1441,1464,1512]" pageId="6" pageNumber="5">
|
||
<emphasis id="90D8EAC1FF85A941C382FA2BFEB1FA10" bold="true" box="[192,363,1464,1488]" pageId="6" pageNumber="5">Figures 14–20:</emphasis>
|
||
<taxonomicName id="65AC4D50FF85A941C23FFA2AFDF9FA0F" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[381,547,1465,1487]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF85A941C23FFA2AFDF9FA0F" box="[381,547,1465,1487]" italics="true" pageId="6" pageNumber="5">Pandinus trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF85A941C16EFA29FDBEFA0F" bold="true" box="[556,612,1466,1487]" pageId="6" pageNumber="5">
|
||
<taxonomicNameLabel id="8BEB57BAFF85A941C16EFA29FDBEFA0F" box="[556,612,1466,1487]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♂ holotype, trichobothrial pattern.
|
||
<emphasis id="90D8EAC1FF85A941C08CFA29FC3CFA0F" bold="true" box="[974,998,1466,1487]" pageId="6" pageNumber="5">14</emphasis>
|
||
. Chela dorsal.
|
||
<emphasis id="90D8EAC1FF85A941C73CFA29FB4CFA0F" bold="true" box="[1150,1174,1466,1487]" pageId="6" pageNumber="5">15</emphasis>
|
||
. Chela external.
|
||
<emphasis id="90D8EAC1FF85A941C600FA29FA80FA0F" bold="true" box="[1346,1370,1466,1487]" pageId="6" pageNumber="5">16</emphasis>
|
||
. Chela ventral.
|
||
<emphasis id="90D8EAC1FF85A941C24DFA41FEFDFA27" bold="true" box="[271,295,1490,1511]" pageId="6" pageNumber="5">17</emphasis>
|
||
. Patella dorsal.
|
||
<emphasis id="90D8EAC1FF85A941C283FA41FE03FA27" bold="true" box="[449,473,1490,1511]" pageId="6" pageNumber="5">18</emphasis>
|
||
. Patella external.
|
||
<emphasis id="90D8EAC1FF85A941C1C7FA41FD47FA27" bold="true" box="[645,669,1490,1511]" pageId="6" pageNumber="5">19</emphasis>
|
||
. Patella ventral.
|
||
<emphasis id="90D8EAC1FF85A941C07DFA41FC8DFA27" bold="true" box="[831,855,1490,1511]" pageId="6" pageNumber="5">20</emphasis>
|
||
. Femur dorsal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A21336D3FF85A941C382F98FFD16F92C" blockId="6.[192,802,1564,1895]" pageId="6" pageNumber="5">bears conical granules that are conspicuous but not pointed; its external surface bears conical granules only in the anterior part. The dentate margins of the movable and fixed fingers are armed with two parallel rows of denticles extending the entire length of the finger, without external and internal granules but with larger granules which indicate subrows on both fingers.</paragraph>
|
||
<paragraph id="A21336D3FF85A941C382F960FC64F8CA" blockId="6.[192,802,1564,1895]" lastBlockId="6.[831,1441,1564,1895]" pageId="6" pageNumber="5">
|
||
MEASUREMENTS IN MM. Male
|
||
<typeStatus id="7D178871FF85A941C167F961FD5CF8CA" box="[549,646,1778,1802]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
. Total length 107; carapace length 13.8, width 14.3; metasoma and telson length 46.8; first metasomal segment length 6.5, width 6.3, depth 4.8; second metasomal segment length 7.0, width 5.5, depth 4.6; third metasomal segment length 7.9, width 5.2, depth 4.7; fourth metasomal segment length 8.5, width 4.7, depth 4.9; fifth metasomal segment length 11.3, width 4.4, depth 4.4; telson length 10.3; telson width 5.6; pedipalp femur length 10.5, width 4.7; pedipalp patella length 10.4, width 5.2; chela length 22.3; manus width 13.3; movable finger length 13.5.
|
||
</paragraph>
|
||
<paragraph id="A21336D3FF85A94CC07DF882FDB2FE74" blockId="6.[831,1441,1564,1895]" lastBlockId="11.[192,802,198,436]" lastPageId="11" lastPageNumber="10" pageId="6" pageNumber="5">
|
||
Female
|
||
<typeStatus id="7D178871FF85A941C0D6F881FC29F8EA" box="[916,1011,1810,1834]" pageId="6" pageNumber="5" type="paratype">paratype</typeStatus>
|
||
. Total length 116; carapace length 15.7, width 16.8; metasoma and telson length 52.6; first metasomal segment length 6.5, width 6.2, depth 4.7; second metasomal segment length 7.1, width 5.5, depth 4.6; third metasomal segment length 7.5, width 5.0, depth 4.5; fourth metasomal segment length 8.7, width 4.7, depth 4.3; fifth metasomal segment length 11.4, width 4.3, depth 4.5; telson length 11.4; telson width 4.8; pedipalp femur length 10.7, width 5.1; pedipalp patella length 11.5, width 5.7; chela length 24.2; manus width 14.5; movable finger length 15.3.
|
||
</paragraph>
|
||
<caption id="F6D3665BFF84A940C382F8DAFED4F8B9" pageId="7" pageNumber="6" startId="7.[192,278,1865,1889]" targetBox="[191,1441,191,1835]" targetPageId="7">
|
||
<paragraph id="A21336D3FF84A940C382F8DAFED4F8B9" blockId="7.[192,1440,1865,1913]" pageId="7" pageNumber="6">
|
||
<emphasis id="90D8EAC1FF84A940C382F8DAFEB3F8A1" bold="true" box="[192,361,1865,1889]" pageId="7" pageNumber="6">Figures 21–22:</emphasis>
|
||
<taxonomicName id="65AC4D50FF84A940C232F8D8FDCEF8A1" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[368,532,1867,1889]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF84A940C232F8D8FDCEF8A1" box="[368,532,1867,1889]" italics="true" pageId="7" pageNumber="6">Pandinus trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF84A940C15EF8D8FD89F8A0" bold="true" box="[540,595,1867,1888]" pageId="7" pageNumber="6">
|
||
<taxonomicNameLabel id="8BEB57BAFF84A940C15EF8D8FD89F8A0" box="[540,595,1867,1888]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♂ holotype.
|
||
<emphasis id="90D8EAC1FF84A940C19BF8D8FD2BF8A0" bold="true" box="[729,753,1867,1888]" pageId="7" pageNumber="6">21</emphasis>
|
||
. Shortly before adulthood (sixth) ecdysis.
|
||
<emphasis id="90D8EAC1FF84A940C7D2F8D8FB72F8A0" bold="true" box="[1168,1192,1867,1888]" pageId="7" pageNumber="6">22</emphasis>
|
||
. One day after adulthood ecdysis.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F6D3665BFF8BA94FC382F8D5FDDEF8B6" pageId="8" pageNumber="7" startId="8.[192,278,1862,1886]" targetBox="[191,1441,191,1830]" targetPageId="8">
|
||
<paragraph id="A21336D3FF8BA94FC382F8D5FDDEF8B6" blockId="8.[192,1440,1862,1910]" pageId="8" pageNumber="7">
|
||
<emphasis id="90D8EAC1FF8BA94FC382F8D5FEB1F89E" bold="true" box="[192,363,1862,1886]" pageId="8" pageNumber="7">Figures 23–24:</emphasis>
|
||
<taxonomicName id="65AC4D50FF8BA94FC236F8D4FDC0F89D" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[372,538,1863,1885]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF8BA94FC236F8D4FDC0F89D" box="[372,538,1863,1885]" italics="true" pageId="8" pageNumber="7">Pandinus trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF8BA94FC160F8D4FDA1F89C" bold="true" box="[546,635,1863,1884]" pageId="8" pageNumber="7">
|
||
<taxonomicNameLabel id="8BEB57BAFF8BA94FC160F8D4FD81F89C" box="[546,603,1863,1884]" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
|
||
23
|
||
</emphasis>
|
||
. Male holotype one month after adulthood ecdysis.
|
||
<emphasis id="90D8EAC1FF8BA94FC7C6F8D4FB46F89C" bold="true" box="[1156,1180,1863,1884]" pageId="8" pageNumber="7">24</emphasis>
|
||
. Adult female paratype at Locality No. 13EC (Figs. 34–35).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F6D3665BFF8AA94EC382F8BAFCECF8B2" pageId="9" pageNumber="8" startId="9.[192,278,1833,1857]" targetBox="[213,1417,167,1805]" targetPageId="9">
|
||
<paragraph id="A21336D3FF8AA94EC382F8BAFCECF8B2" blockId="9.[192,1439,1833,1906]" pageId="9" pageNumber="8">
|
||
<emphasis id="90D8EAC1FF8AA94EC382F8BAFEB0F881" bold="true" box="[192,362,1833,1857]" pageId="9" pageNumber="8">Figures 25–27:</emphasis>
|
||
<taxonomicName id="65AC4D50FF8AA94EC232F8B9FDCFF880" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[368,533,1834,1856]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF8AA94EC232F8B9FDCFF880" box="[368,533,1834,1856]" italics="true" pageId="9" pageNumber="8">Pandinus trailini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BEB57BAFF8AA94EC15EF8B8FD88F880" box="[540,594,1835,1856]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="90D8EAC1FF8AA94EC15EF8B8FD96F880" bold="true" box="[540,588,1835,1856]" pageId="9" pageNumber="8">sp. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, type locality, Ethiopia, Oromia State, Arsi Province, 06°56'06"N 40°41'23"E, 1693 m a.s.l. (Locality No. 13EA).
|
||
<emphasis id="90D8EAC1FF8AA94EC289F8D0FE39F898" bold="true" box="[459,483,1859,1880]" pageId="9" pageNumber="8">27</emphasis>
|
||
. Red arrow points to the spot where a female paratype was found together with 10 juveniles. The overturned rock under which the female was found is to the right.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F6D3665BFF89A94DC382F8DBFA4BF8B8" pageId="10" pageNumber="9" startId="10.[192,278,1864,1888]" targetBox="[211,1417,167,1833]" targetPageId="10">
|
||
<paragraph id="A21336D3FF89A94DC382F8DBFA4BF8B8" blockId="10.[192,1440,1864,1912]" pageId="10" pageNumber="9">
|
||
<emphasis id="90D8EAC1FF89A94DC382F8DBFEB0F8A0" bold="true" box="[192,362,1864,1888]" pageId="10" pageNumber="9">Figures 28–29:</emphasis>
|
||
<taxonomicName id="65AC4D50FF89A94DC233F8DAFDCCF89F" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[369,534,1865,1887]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF89A94DC233F8DAFDCCF89F" box="[369,534,1865,1887]" italics="true" pageId="10" pageNumber="9">Pandinus trailini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8BEB57BAFF89A94DC15FF8DAFD89F89F" box="[541,595,1865,1887]" pageId="10" pageNumber="9" rank="species">
|
||
<emphasis id="90D8EAC1FF89A94DC15FF8DAFD94F89E" bold="true" box="[541,590,1865,1886]" pageId="10" pageNumber="9">sp. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, Female paratype at type locality. Well apparent are remains of the prey consumed by the female and juveniles. Fig. 29 shows one of the 10 juveniles found with their mother. This juvenile is after the fourth ecdysis.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A21336D3FF88A94CC382FE4AFEE1FA68" blockId="11.[192,802,473,1448]" pageId="11" pageNumber="10">
|
||
AFFINITIES. The described features distinguish
|
||
<taxonomicName id="65AC4D50FF88A94CC1FBFE49FCFBFE31" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[697,801,473,497]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF88A94CC1FBFE49FCFBFE31" box="[697,801,473,497]" italics="true" pageId="11" pageNumber="10">P. trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF88A94CC382FE6AFF25FDD1" bold="true" box="[192,255,505,529]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="8BEB57BAFF88A94CC382FE6AFF25FDD1" box="[192,255,505,529]" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from all other species of the subgenus
|
||
<taxonomicName id="65AC4D50FF88A94CC1FFFE6BFE82FDEF" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="90D8EAC1FF88A94CC1FFFE6BFCF8FDD0" box="[701,802,504,528]" italics="true" pageId="11" pageNumber="10">Pandinus</emphasis>
|
||
<bibRefCitation id="C63D4B22FF88A94CC382FD84FE82FDEF" author="THORELL" box="[192,344,535,559]" journalOrPublisher="On the classification of scorpions. Annals and Magazine of Natural History" pageId="11" pageNumber="10" pagination="1 - 15" part="4" refId="ref5158" refString="THORELL, T. 1876. On the classification of scorpions. Annals and Magazine of Natural History, 4 (17): 1 - 15." title="T" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="65AC4D50FF88A94CC227FD8BFE15FDEF" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[357,463,535,559]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF88A94CC227FD8BFE15FDEF" box="[357,463,535,559]" italics="true" pageId="11" pageNumber="10">P. trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF88A94CC29BFD84FDCDFDEF" bold="true" box="[473,535,535,559]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="8BEB57BAFF88A94CC29BFD84FDCDFDEF" box="[473,535,535,559]" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is characterized by 6/4: 6-7/4: 7-8/4-5: 7-8/4-5 spination formula of tarsomere II and sexual dimorphism, when the male has a more pronounced tooth on the movable finger of pedipalp than the female. These two characters distinguish
|
||
<taxonomicName id="65AC4D50FF88A94CC1F4FD00FCFBFD6A" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[694,801,658,682]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF88A94CC1F4FD00FCFBFD6A" box="[694,801,658,682]" italics="true" pageId="11" pageNumber="10">P. trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF88A94CC382FD22FEDAFD09" bold="true" box="[192,256,689,713]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="8BEB57BAFF88A94CC382FD22FEDAFD09" box="[192,256,689,713]" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from
|
||
<taxonomicName id="65AC4D50FF88A94CC20BFD22FD5CFD08" authority="Pocock, 1899" authorityName="Pocock" authorityYear="1899" box="[329,646,688,712]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="gambiensis">
|
||
<emphasis id="90D8EAC1FF88A94CC20BFD22FE39FD08" box="[329,483,688,712]" italics="true" pageId="11" pageNumber="10">P. gambiensis</emphasis>
|
||
Pocock, 1899
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="65AC4D50FF88A94CC1D4FD22FE4CFD27" authority="(C. L. Koch, 1841)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1841" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="imperator">
|
||
<emphasis id="90D8EAC1FF88A94CC1D4FD22FCFBFD08" box="[662,801,689,712]" italics="true" pageId="11" pageNumber="10">P. imperator</emphasis>
|
||
(C. L. Koch, 1841)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="65AC4D50FF88A94CC297FD43FCFBFD27" authority="Kovarik, 2011" authorityName="Kovarik" authorityYear="2011" box="[469,801,719,743]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ugandaensis">
|
||
<emphasis id="90D8EAC1FF88A94CC297FD43FDA1FD27" box="[469,635,719,743]" italics="true" pageId="11" pageNumber="10">P. ugandaensis</emphasis>
|
||
Kovařík, 2011
|
||
</taxonomicName>
|
||
and indicate its close relationship with
|
||
<taxonomicName id="65AC4D50FF88A94CC1F7FD7CFEBDFCE4" authority="(Pocock, 1897)" baseAuthorityName="Pocock" baseAuthorityYear="1897" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="smithi">
|
||
<emphasis id="90D8EAC1FF88A94CC1F7FD7CFCFBFCC6" box="[693,801,750,774]" italics="true" pageId="11" pageNumber="10">P. smithi</emphasis>
|
||
(Pocock, 1897)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="65AC4D50FF88A94CC2E3FC9EFD09FCE4" authority="(Pocock, 1896)" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[417,723,780,804]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="phillipsii">
|
||
<emphasis id="90D8EAC1FF88A94CC2E3FC9EFDC5FCE4" box="[417,543,780,804]" italics="true" pageId="11" pageNumber="10">P. phillipsii</emphasis>
|
||
(Pocock, 1896)
|
||
</taxonomicName>
|
||
, which however can be easily distinguished from
|
||
<taxonomicName id="65AC4D50FF88A94CC1CDFCBFFD23FC83" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[655,761,811,835]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF88A94CC1CDFCBFFD23FC83" box="[655,761,811,835]" italics="true" pageId="11" pageNumber="10">P. trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF88A94CC040FCBFFF0FFCA2" bold="true" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="8BEB57BAFF88A94CC040FCBFFF0FFCA2" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by their yellow legs (see figs.
|
||
<quantity id="65549B36FF88A94CC112FCD9FD6FFCA2" box="[592,693,842,866]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.302" metricValueMin="2.54" pageId="11" pageNumber="10" unit="in" value="11.5" valueMax="13.0" valueMin="10.0">10–13 in</quantity>
|
||
Kovařík, 2011: 5). If this character is omitted, then of these two species
|
||
<taxonomicName id="65AC4D50FF88A94CC256FC1BFEA6FC5F" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[276,380,903,927]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF88A94CC256FC1BFEA6FC5F" box="[276,380,903,927]" italics="true" pageId="11" pageNumber="10">P. trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF88A94CC2C1FC14FE64FC5F" bold="true" box="[387,446,903,927]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="8BEB57BAFF88A94CC2C1FC14FE64FC5F" box="[387,446,903,927]" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
appears to be closer to
|
||
<taxonomicName id="65AC4D50FF88A94CC1FEFC1BFCC1FC5F" baseAuthorityName="Pocock" baseAuthorityYear="1897" box="[700,795,903,927]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="smithi">
|
||
<emphasis id="90D8EAC1FF88A94CC1FEFC1BFCC1FC5F" box="[700,795,903,927]" italics="true" pageId="11" pageNumber="10">P. smithi</emphasis>
|
||
</taxonomicName>
|
||
, which however has the tarsomere II of legs with three spines on the inclined anteroventral surface (
|
||
<figureCitation id="3A972A56FF88A94CC1ECFC57FD28FC1C" box="[686,754,964,988]" captionStart="Figures 1–8" captionStartId="3.[192,278,1795,1819]" captionTargetBox="[222,1387,166,1747]" captionTargetId="figure-96@3.[212,1416,165,1765]" captionTargetPageId="3" captionText="Figures 1–8: 1–6: Pandinus trailini sp. n., ♂ holotype. 1–2. Dorsal and ventral views. 3. Pectinal area. 4. Metasoma and telson, lateral view. 5–6. Tarsomere II of 4th leg, lateral and ventral views. 7: Pandinus ugandaensis Kovařík, 2011, ♂ holotype, tarsomere II of 4th leg. 8: Pandinus smithi (Pocock, 1897), ♂ (107 mm), Ethiopia, 55 km of Degehbur, 07°49'27.2"N 43°41'56.3"E, 1053 m a.s.l., FKCP, tarsomere II of 4th leg." pageId="11" pageNumber="10">Fig. 8</figureCitation>
|
||
).
|
||
<taxonomicName id="65AC4D50FF88A94CC049FC56FED0FC3B" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF88A94CC049FC56FED0FC3B" italics="true" pageId="11" pageNumber="10">P. trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF88A94CC251FC70FE88FC3B" bold="true" box="[275,338,995,1019]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="8BEB57BAFF88A94CC251FC70FE88FC3B" box="[275,338,995,1019]" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has only two spines, in place of the third spine there is only a spiniform seta (
|
||
<figureCitation id="3A972A56FF88A94CC125FB91FD77FBDA" box="[615,685,1026,1050]" captionStart="Figures 1–8" captionStartId="3.[192,278,1795,1819]" captionTargetBox="[222,1387,166,1747]" captionTargetId="figure-96@3.[212,1416,165,1765]" captionTargetPageId="3" captionText="Figures 1–8: 1–6: Pandinus trailini sp. n., ♂ holotype. 1–2. Dorsal and ventral views. 3. Pectinal area. 4. Metasoma and telson, lateral view. 5–6. Tarsomere II of 4th leg, lateral and ventral views. 7: Pandinus ugandaensis Kovařík, 2011, ♂ holotype, tarsomere II of 4th leg. 8: Pandinus smithi (Pocock, 1897), ♂ (107 mm), Ethiopia, 55 km of Degehbur, 07°49'27.2"N 43°41'56.3"E, 1053 m a.s.l., FKCP, tarsomere II of 4th leg." pageId="11" pageNumber="10">Fig. 5</figureCitation>
|
||
). The last species of the genus is
|
||
<taxonomicName id="65AC4D50FF88A94CC2F5FBB2FD08FBF8" authority="Kovarik, 2011" authorityName="Kovarik" authorityYear="2011" box="[439,722,1056,1080]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="mazuchi">
|
||
<emphasis id="90D8EAC1FF88A94CC2F5FBB2FDF4FBF8" box="[439,558,1056,1080]" italics="true" pageId="11" pageNumber="10">P. mazuchi</emphasis>
|
||
Kovařík, 2011
|
||
</taxonomicName>
|
||
, whose sexual dimorphism remains unknown. This species can be distinguished from
|
||
<taxonomicName id="65AC4D50FF88A94CC2F6FBCCFDC7FBB6" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" box="[436,541,1118,1142]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF88A94CC2F6FBCCFDC7FBB6" box="[436,541,1118,1142]" italics="true" pageId="11" pageNumber="10">P. trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF88A94CC167FBCDFDB8FBB6" bold="true" box="[549,610,1118,1142]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="8BEB57BAFF88A94CC167FBCDFDB8FBB6" box="[549,610,1118,1142]" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by having an additional trichobothrium
|
||
<emphasis id="90D8EAC1FF88A94CC280FBEEFE01FB54" box="[450,475,1149,1172]" italics="true" pageId="11" pageNumber="10">ea</emphasis>
|
||
(fig. 5a in Kovařík, 2011: 3), different dorsal granulation of the chela (fig. 5a in Kovařík, 2011: 3 versus
|
||
<figureCitation id="3A972A56FF88A94CC285FB29FDCFFB12" box="[455,533,1210,1234]" captionStart="Figures 9–13" captionStartId="4.[192,278,1854,1878]" captionTargetBox="[227,1401,188,1797]" captionTargetId="figure-52@4.[214,1416,188,1824]" captionTargetPageId="4" captionText="Figures 9–13: Pandinus trailini sp. n., ♀ (116 mm) paratype. 9–10. Dorsal and ventral views. 11. Pectinal area. 12. Chela dorsoexternal view. 13. Metasoma and telson, lateral view." pageId="11" pageNumber="10">Fig. 12</figureCitation>
|
||
), sparse pubescence, and having only 10 ventral trichobothria on the chela.
|
||
<taxonomicName id="65AC4D50FF88A94CC048FB4AFED0FACF" authority="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com, 2013" authorityName="Editor & Fet & University & fet & marshall.edu & Editor & Soleglad & soleglad & znet.com" authorityYear="2013" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="trailini" status="sp. nov.">
|
||
<emphasis id="90D8EAC1FF88A94CC048FB4AFED0FACF" italics="true" pageId="11" pageNumber="10">P. trailini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90D8EAC1FF88A94CC25AFB64FE80FACF" bold="true" box="[280,346,1271,1295]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="8BEB57BAFF88A94CC25AFB64FE80FACF" box="[280,346,1271,1295]" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has 16–18 ventral trichobothria on the chela (
|
||
<figureCitation id="3A972A56FF88A94CC24DFA85FEBCFAEE" box="[271,358,1302,1326]" captionStart="Figures 14–20" captionStartId="6.[192,278,1464,1488]" captionTargetBox="[250,1407,176,1425]" captionTargetId="figure-246@6.[192,1440,168,1435]" captionTargetPageId="6" captionText="Figures 14–20: Pandinus trailini sp. n., ♂ holotype, trichobothrial pattern. 14. Chela dorsal. 15. Chela external. 16. Chela ventral. 17. Patella dorsal. 18. Patella external. 19. Patella ventral. 20. Femur dorsal." pageId="11" pageNumber="10">Fig. 16</figureCitation>
|
||
), which is the highest number in the subgenus
|
||
<taxonomicName id="65AC4D50FF88A94CC272FAA7FE4FFA8C" authorityName="Thorell" authorityYear="1876" box="[304,405,1332,1356]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="90D8EAC1FF88A94CC272FAA7FE4FFA8C" box="[304,405,1332,1356]" italics="true" pageId="11" pageNumber="10">Pandinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="65AC4D50FF88A94CC2E8FAA6FD9CFA8C" authorityName="Pocock" authorityYear="1899" box="[426,582,1332,1356]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="gambiensis">
|
||
<emphasis id="90D8EAC1FF88A94CC2E8FAA6FD9CFA8C" box="[426,582,1332,1356]" italics="true" pageId="11" pageNumber="10">P. gambiensis</emphasis>
|
||
</taxonomicName>
|
||
has chela with 10 ventral trichobothria,
|
||
<taxonomicName id="65AC4D50FF88A94CC2EFFAC7FDE2FAAB" baseAuthorityName="C. L. Koch" baseAuthorityYear="1841" box="[429,568,1364,1387]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="imperator">
|
||
<emphasis id="90D8EAC1FF88A94CC2EFFAC7FDE2FAAB" box="[429,568,1364,1387]" italics="true" pageId="11" pageNumber="10">P. imperator</emphasis>
|
||
</taxonomicName>
|
||
has 9–14,
|
||
<taxonomicName id="65AC4D50FF88A94CC18DFAC7FF2FFA4A" baseAuthorityName="Pocock" baseAuthorityYear="1896" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="phillipsii">
|
||
<emphasis id="90D8EAC1FF88A94CC18DFAC7FF2FFA4A" italics="true" pageId="11" pageNumber="10">P. phillipsii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="65AC4D50FF88A94CC26FFAE0FE57FA4A" baseAuthorityName="Pocock" baseAuthorityYear="1897" box="[301,397,1394,1418]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="smithi">
|
||
<emphasis id="90D8EAC1FF88A94CC26FFAE0FE57FA4A" box="[301,397,1394,1418]" italics="true" pageId="11" pageNumber="10">P. smithi</emphasis>
|
||
</taxonomicName>
|
||
have 13–15, and
|
||
<taxonomicName id="65AC4D50FF88A94CC113FAE0FD2FFA4A" authorityName="Kovarik" authorityYear="2011" box="[593,757,1394,1418]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ugandaensis">
|
||
<emphasis id="90D8EAC1FF88A94CC113FAE0FD2FFA4A" box="[593,757,1394,1418]" italics="true" pageId="11" pageNumber="10">P. ugandaensis</emphasis>
|
||
</taxonomicName>
|
||
has only 8 or 9.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |