467 lines
70 KiB
XML
467 lines
70 KiB
XML
<document ID-CLB-Dataset="6046" ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.59674" ID-GBIF-Dataset="3054b650-b609-4439-aff3-e7b6623c6807" ID-PMC="PMC8222241" ID-Pensoft-Pub="1313-2970-1044-229" ID-Pensoft-UUID="0B5B533D7CBA50819E2AEC264FE6D977" ID-PubMed="34183879" ID-ZooBank="7EC23192F85D4D2AA31E2694A59CB014" ModsDocID="1313-2970-1044-229" checkinTime="1623875428746" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2021" docId="29E435758D6D5438AF0935206D2E7210" docLanguage="en" docName="ZooKeys 1044: 229-268" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.59674" docTitle="Paratachys terryli Liebherr 2021, sp. nov." docType="treatment" docUuid="9B7B5D1A-8243-4BBE-A95F-DE1477915432" docUuidSource="ZooBank" docVersion="6" id="0B5B533D7CBA50819E2AEC264FE6D977" lastPageNumber="229" masterDocId="0B5B533D7CBA50819E2AEC264FE6D977" masterDocTitle="Hawaiian Paratachys Casey (Coleoptera, Carabidae): small beetles of sodden summits, stony streams, and stygian voids" masterLastPageNumber="268" masterPageNumber="229" pageNumber="229" updateTime="1732754002844" updateUser="ExternalLinkService">
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<mods:titleInfo id="59DBEA54E2B8BFD4E9F51EDA334DE5CF">
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<mods:title id="DF7A296BDF43C058963BB547DFAA22CF">Hawaiian Paratachys Casey (Coleoptera, Carabidae): small beetles of sodden summits, stony streams, and stygian voids</mods:title>
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<mods:name id="EA3344E7FD545C2F717F3621ECA34DF3" type="personal">
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<mods:namePart id="8EF8EE38B33682A193901AEFFB34EC2A">Liebherr, James K.</mods:namePart>
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<mods:nameIdentifier id="B62CA29AF284CBA9CBF326B048F1ED46" type="ORCID">https://orcid.org/0000-0001-9831-884X</mods:nameIdentifier>
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<mods:affiliation id="409ECEBF370FD2C8BB8E4D94D5422654">Department of Entomology, John H. and Anna B. Comstock Hall, 129 Garden Ave., Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:nameIdentifier id="CA8B7AE269C9733636126E3AA7E774A3" type="email">jkl5@cornell.edu</mods:nameIdentifier>
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<mods:title id="4C2CCC50E3C99FB4787DEAFDC548D083">ZooKeys</mods:title>
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<mods:part id="344548AF1F44CEB1951AA5F04D7DCB73">
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<mods:date id="219E9CC374202148F20ED5C97EC0B6B9">2021</mods:date>
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<mods:number id="4D16722D7134D6EFBEB7D3F85421A0A9">2021-06-16</mods:number>
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<mods:number id="1DCC20BF90C382566B858BFA803B28B4">1044</mods:number>
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<mods:url id="1F2EFAF8AD99C6B331862C11A5D2B7CD">http://dx.doi.org/10.3897/zookeys.1044.59674</mods:url>
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<mods:classification id="DF6AEC5D41B8FFA084B2AFB10CB42779">journal article</mods:classification>
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<mods:identifier id="6CB6926E08550B88D6BDE5DA3BE7389E" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.59674</mods:identifier>
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<treatment id="29E435758D6D5438AF0935206D2E7210" ID-GBIF-Taxon="182754101" LSID="urn:lsid:zoobank.org:act:9B7B5D1A-8243-4BBE-A95F-DE1477915432" httpUri="http://treatment.plazi.org/id/29E435758D6D5438AF0935206D2E7210" lastPageNumber="229" pageId="0" pageNumber="229" scope_family="Carabidae" scope_order="Coleoptera">
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<subSubSection id="CB6485BF0B37770C62CC6989B32F1300" pageId="0" pageNumber="229" type="nomenclature">
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<paragraph id="2B4C7BEB7C79952F598278FEE65668DB" pageId="0" pageNumber="229">
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<taxonomicName id="104153D5A43F904D3AFD46F3D25A264B" ID-CoL="8MJS7" LSID="http://zoobank.org/9B7B5D1A-8243-4BBE-A95F-DE1477915432" authority="Liebherr, 2021" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys terryli" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="terryli" status="sp. nov.">Paratachys terryli</taxonomicName>
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<taxonomicNameLabel id="C3387E2045F66CBC1EED8BE0F93E8FF7" pageId="0" pageNumber="229">sp. nov.</taxonomicNameLabel>
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<figureCitation id="924C1143BE4ED8264EB3F10F0D4606E1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">Figures 1</figureCitation>
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<figureCitation id="97C5ECBBE2AEF8EAAFD7EA8D9C89FFAC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Right metathoracic flight wing, dorsal view A Paratachys terryli, macrophthalmic female; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B Tachys oahuensis Blackburn, female; Oahu: Honolulu, Public Health Dept. light trap, x- 1965, J. W. Beardsley (BPBM). Wing vein terminology follows Kukalova-Peck and Lawrence (1993) except for the oblongum cell of Forbes (1922). From front to back of wing, veins, and cells relevant to the text include the Radial Cell (RC). Radius Anterior (RA), Radius Posterior (RP), Oblongum cell (O), Media Posterior (MP), Cubitus Anterior (CuA), Anal Anterior (AA), and Anal Posterior (AP)." figureDoi="10.3897/zookeys.1044.59674.figure2" httpUri="https://binary.pensoft.net/fig/553678" pageId="0" pageNumber="229">, 2A</figureCitation>
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<figureCitation id="F5407ACDC7DA4B0FBB79281329BEF449" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paratachys spp., male aedeagus, i. e., median lobe and associated smaller right paramere and larger left paramere (right lateral view) A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. arcanicola lectotype slide; " Tachys arcanicola Blackburn Type ♂ E. C. Z. (permanent slide, Euparal?, left paramere missing; NHMUK). " C P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) D-F P. aaa; Hawai'i I D Puna, Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC) E Kau, Ocean View, Kipuka Kanohina System, 21 - xi- 2017, lot HI 00179 (CUIC) F South Hilo, Kaumana Cave, 19 - iii- 2019, lot HI 00942 (CUIC); disassociated left paramere below, ectal view." figureDoi="10.3897/zookeys.1044.59674.figure3" httpUri="https://binary.pensoft.net/fig/553679" pageId="0" pageNumber="229">, 3A</figureCitation>
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<figureCitation id="4D932F7976EBAC866432275A54A9EF4E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paratachys spp., left female gonocoxa, ventral view A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) C P. aaa; Hawai'i I.; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC). Abbreviations: afs, apical fringe seta; ans, apical nematiform setae; des, dorsal ensiform seta; gc 1, basal gonocoxite; gc 2, apical gonocoxite; les, lateral ensiform setae." figureDoi="10.3897/zookeys.1044.59674.figure4" httpUri="https://binary.pensoft.net/fig/553680" pageId="0" pageNumber="229">, 4A</figureCitation>
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<figureCitation id="2E459FE82FBDA4F07B597D4F3C4D1303" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distributional records for Paratachys terryli." figureDoi="10.3897/zookeys.1044.59674.figure5" httpUri="https://binary.pensoft.net/fig/553681" pageId="0" pageNumber="229">, 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="272758821DECBC72417697A19CEAC624" pageId="0" pageNumber="229" type="reference_group">
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<paragraph id="A2C5BA5BC5DAACB75ED2EED076739A21" pageId="0" pageNumber="229">
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<taxonomicName id="68B7E1B6CFAFB2917B3CAE542C69EDE9" authority="Liebherr, 2021" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="genus" status="sp. nov.">Paratachys</taxonomicName>
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sp. "USA: Hawaii",
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<bibRefCitation id="9675E329106B68A20C8971115C679E09" DOI="https://doi.org/10.1016/j.ympev.2018.11.006" author="Maddison, DR" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="229" pagination="151 - 176" refId="B71" refString="Maddison, DR, Kanda, K, Boyd, OF, Faille, A, Porch, N, Erwin, TL, Roig-Junent, S, 2019. Phylogeny of the beetle supertribe Trechitae (Coleoptera: Carabidae): unexpected clades, isolated lineages, and morphological convergence. Molecular Phylogenetics and Evolution 132: 151 - 176, DOI: https://doi.org/10.1016/j.ympev.2018.11.006" title="Phylogeny of the beetle supertribe Trechitae (Coleoptera: Carabidae): unexpected clades, isolated lineages, and morphological convergence." url="https://doi.org/10.1016/j.ympev.2018.11.006" volume="132" year="2019">Maddison et al. (2019</bibRefCitation>
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: 162).
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</paragraph>
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||
</subSubSection>
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<subSubSection id="A2969F3C0302C16AA503B2E0CF8934CC" pageId="0" pageNumber="229" type="type material">
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<paragraph id="270914ECAACAB08E636A99C90196379C" pageId="0" pageNumber="229">Type material.</paragraph>
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<paragraph id="52D6C95DE4FBFAC8651721EB8916BF99" pageId="0" pageNumber="229">
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<emphasis id="FB24192AADE5EF308002B1F9817BC704" bold="true" italics="true" pageId="0" pageNumber="229">Holotype</emphasis>
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male (point-mounted, CUIC): HI: NaPali-Kona / For. Res. Kawaikoi Str. / @ Alakai Swp. Tr. 16-V- / 1991 el. 1120 m under / rocks J.K. Liebherr // HOLOTYPE ♂ / Paratachys / terryli / J. K. Liebherr 2020 (black-margined red label).
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</paragraph>
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<paragraph id="2B3FCB9E28092E8627F6B47E2CBF8053" pageId="0" pageNumber="229">
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<emphasis id="9988F0C40FA3FA0AB4EF526D9B529772" bold="true" pageId="0" pageNumber="229">
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<emphasis id="FC7A9359573C6C600F3B02F927964B57" italics="true" pageId="0" pageNumber="229">Paratypes</emphasis>
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. Kauai: Halelea F. R.
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</emphasis>
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: Namolokama Mtn., Waioli Str., litter, sift, ohia/ferns,
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<geoCoordinate id="183B8BDC57FE7BB34F9E090366F46B1A" degrees="22" direction="north" minutes="08.00" orientation="latitude" precision="9" value="22.133333">22°08.00'N</geoCoordinate>
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,
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<geoCoordinate id="1579B1D35A244692B171C41D62369633" degrees="159" direction="west" minutes="29.85" orientation="longitude" precision="9" value="-159.4975">159°29.85'W</geoCoordinate>
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, 1340 m el., 21-v-2005, lot 1 Liebherr (BPBM, 2; CUIC, 3), lot 08, Liebherr (CUIC, 1),
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<geoCoordinate id="E52CEBD4DD6E1C81BD676FCFCC67EA12" degrees="22" direction="north" minutes="08.38" orientation="latitude" precision="9" value="22.139668">22°08.38'N</geoCoordinate>
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,
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<geoCoordinate id="52407BCC1D31C4E43F3F9CCA611916B7" degrees="159" direction="west" minutes="30.18" orientation="longitude" precision="9" value="-159.503">159°30.18'W</geoCoordinate>
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, 1305 m el., 23-v-2005, lot 1, Liebherr (BPBM, 2; CUIC, 3 NMNH, 2).
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<emphasis id="2461B242E70683764D790C5468805BB4" bold="true" pageId="0" pageNumber="229">NaPali-Kona F. R.</emphasis>
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: Alakai Swamp Tr. crossing Kawaikoi Str.,
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<geoCoordinate id="64AD54662D627117189F83F101551C32" degrees="22" direction="north" minutes="08.97" orientation="latitude" precision="9" value="22.1495">22°08.97'N</geoCoordinate>
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,
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<geoCoordinate id="E4B1D26034B252160040E5C10D2469EA" degrees="159" direction="west" minutes="36.95" orientation="longitude" precision="9" value="-159.61583">159°36.95'W</geoCoordinate>
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, 1130 m el., 14-v-1991, stop #91-24, Kavanaugh (CAS, 1), stop #91-26B (CAS, 1), Alakai Swamp Tr. E of Kawaikoi Str., leaf litter, sift/Berlese extraction,
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<geoCoordinate id="0426B3148FA8B16F9EB47B7920B25DCA" degrees="22" direction="north" minutes="08.85" orientation="latitude" precision="9" value="22.1475">22°08.85'N</geoCoordinate>
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,
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<geoCoordinate id="FB8394A23B0D625AEC697206E67C1CBD" degrees="159" direction="west" minutes="36.52" orientation="longitude" precision="9" value="-159.60867">159°36.52'W</geoCoordinate>
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, 1230 m el., 17-v-2005, lot 4, Liebherr (CUIC, 1), litter, sift/double boiler extraction,
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<geoCoordinate id="D638BD419C2C5331C6C5ABD9AC43A556" degrees="22" direction="north" minutes="08.85" orientation="latitude" precision="9" value="22.1475">22°08.85'N</geoCoordinate>
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,
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<geoCoordinate id="A07640A53DCF22AECC8B3437ACAF8CAB" degrees="159" direction="west" minutes="36.52" orientation="longitude" precision="9" value="-159.60867">159°36.52'W</geoCoordinate>
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, 1215 m el., 17-v-2005, lot 9, Liebherr (CUIC, 1; OSAC, 1); Mohihi Rdg. Tr., ohia forest litter, sift, moss,
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<geoCoordinate id="A6A0DB8560F6DF92B3D663B38BD30CA8" degrees="22" direction="north" minutes="06.83" orientation="latitude" precision="9" value="22.113832">22°06.83'N</geoCoordinate>
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,
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<geoCoordinate id="7CE6069E50AA84A67662B2C3590DEA65" degrees="159" direction="west" minutes="34.01" orientation="longitude" precision="9" value="-159.56683">159°34.01'W</geoCoordinate>
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, 1270 m el., 25-v-2005, lot 5, Liebherr (CUIC, 1); Pihea Trail, sifting leaf litter,
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<geoCoordinate id="13C7D3D7406F563DE2F11CC284E94250" degrees="22" direction="north" minutes="08.849" orientation="latitude" precision="1" value="22.147484">22°08.849'N</geoCoordinate>
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,
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<geoCoordinate id="348C21521D3801A05D0042BC9E01772A" degrees="159" direction="west" minutes="37.889" orientation="longitude" precision="1" value="-159.63148">159°37.889'W</geoCoordinate>
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, 1218 m el., 26-vii-2015, Toledano & Olivieri (CTVR, 1), 29-vii-2015, Toledano & Olivieri (CTVR, 2), 1123 m, 29-vii-2015, Toledano & Olivieri (NHMUK, 2; CTVR, 3; CUIC, 2).
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</paragraph>
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</subSubSection>
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<subSubSection id="B97D50A8D2F544A68157C931FF561293" pageId="0" pageNumber="229" type="diagnosis">
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<paragraph id="F852F46F1DFBEBA73BC4C92139D8C62C" pageId="0" pageNumber="229">Diagnosis.</paragraph>
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<paragraph id="25EB261DB110AFD5F17D93E75C3A86A9" pageId="0" pageNumber="229">
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This species shares elongate elytra (EL/MEW = 1.45-1.50; Fig.
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<figureCitation id="3C2C4B5BBFF6090737AE5EB6390A9BA5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1A</figureCitation>
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) with
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<taxonomicName id="8A571A796678ADE52E5B330124BA3F09" lsidName="P. perkinsi" pageId="0" pageNumber="229" rank="species" species="perkinsi">
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||
<emphasis id="86B1976A11E7BB664897F924C9782F2B" italics="true" pageId="0" pageNumber="229">P. perkinsi</emphasis>
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||
</taxonomicName>
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||
(Fig.
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<figureCitation id="C08545C4D1F3994FB95C90E875ABC00E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paratachys spp., dorsal habitus view A P. arcanicola paralectotype female; Oahu: Honolulu (Bates collection, box 389) B P. perkinsi holotype female; Moloka'i, July 1894 (R. C. L. Perkins lot 170)." figureDoi="10.3897/zookeys.1044.59674.figure6" httpUri="https://binary.pensoft.net/fig/553682" pageId="0" pageNumber="229">6B</figureCitation>
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) and
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||
<taxonomicName id="D74E152ED0A83D3D5A1A3F0BD617B0BD" lsidName="P. haleakalae" pageId="0" pageNumber="229" rank="species" species="haleakalae">
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<emphasis id="D76EB2A4487D729F248D5569EB3F34F3" italics="true" pageId="0" pageNumber="229">P. haleakalae</emphasis>
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</taxonomicName>
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(Fig.
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<figureCitation id="2D8E4C31C3E6D7DBA6B2E42C80DE696F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Paratachys haleakalae A dorsal habitus view B mentum and associated mouthparts, ventral view C larger compound eye of female, lateral view D smaller compound eye of male; lateral view E stenopterous left metathoracic flight wing of female (elytron removed), dorsal view." figureDoi="10.3897/zookeys.1044.59674.figure8" httpUri="https://binary.pensoft.net/fig/553684" pageId="0" pageNumber="229">8A</figureCitation>
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||
), but the pronotal lateral margins are less sinuate than those of
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<taxonomicName id="768551DDE80DB1E12DC88F4045753367" lsidName="P. perkinsi" pageId="0" pageNumber="229" rank="species" species="perkinsi">
|
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<emphasis id="33E793CEBF12F79BB4D2E8230C70374A" italics="true" pageId="0" pageNumber="229">P. perkinsi</emphasis>
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</taxonomicName>
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, and more sinuate than those of
|
||
<taxonomicName id="214FCF33CBCDC552FB5B4558C5C24F1F" lsidName="P. haleakalae" pageId="0" pageNumber="229" rank="species" species="haleakalae">
|
||
<emphasis id="15790FAEDFFE42BF0083B277964EBC67" italics="true" pageId="0" pageNumber="229">P. haleakalae</emphasis>
|
||
</taxonomicName>
|
||
. The apical recurrent elytral groove is distinctly and narrowly impressed, with a well-defined, medially curved anterior terminus, and the elytra are truncate apically as evidenced by the nearly straight apical groove connecting the sutural and the recurrent grooves. As with
|
||
<taxonomicName id="99147E86733F1AE9F25E951C61D02ED1" lsidName="P. haleakalae" pageId="0" pageNumber="229" rank="species" species="haleakalae">
|
||
<emphasis id="A2792E8C3439C23C044E0D1A55EF372F" italics="true" pageId="0" pageNumber="229">P. haleakalae</emphasis>
|
||
</taxonomicName>
|
||
, elytral interneurs 1-3 are clearly impressed on the disc, but interneur 4 is traceable though discontinuous in this species, opposed to obsolete as in
|
||
<taxonomicName id="048241FA1B837336A4A12A35C7CDD5B7" lsidName="P. haleakalae" pageId="0" pageNumber="229" rank="species" species="haleakalae">
|
||
<emphasis id="BAB4010EC0237E693263E06E49BCDF88" italics="true" pageId="0" pageNumber="229">P. haleakalae</emphasis>
|
||
</taxonomicName>
|
||
. Standardized body length is 2.2-2.4 mm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="33D3921C7100344DEA9A791834C352AC" pageId="0" pageNumber="229" type="description">
|
||
<paragraph id="F2AB5D113998BFFFD4313F195CCD5E8D" pageId="0" pageNumber="229">Description.</paragraph>
|
||
<paragraph id="F03A89BA7192E2FA10B761B1DDC94D7E" pageId="0" pageNumber="229">
|
||
<emphasis id="6B03D457358D12C4776471AF06487D77" bold="true" italics="true" pageId="0" pageNumber="229">Head</emphasis>
|
||
robust, frontal grooves shallow, convergent posterad clypeus, divergent to frontal lateral margin at frontoclypeal suture just anterad antennal articulation, broadly, slightly elevated laterally to position of anterior supraorbital seta; eyes variable, from large, macrophthalmic with 12 ommatidia crossed by horizontal diameter and 15 ommatidia crossed by vertical diameter (Fig.
|
||
<figureCitation id="6EC62A4C28F0A461C13ED24402791458" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1C</figureCitation>
|
||
), to very small with dimensions of four ommatidia by five ommatidia horizontally and vertically (Fig.
|
||
<figureCitation id="E1204A9DA368994083B2479D1A379BC7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1I</figureCitation>
|
||
), with continuous variation of dimensions between those extremes (Fig.
|
||
<figureCitation id="B517ADB42F7F4B58A4FCA1387FC725AF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1D-H</figureCitation>
|
||
; see Variation section below); antennae moderately elongate, antennomere 9 ovoid, length twice diameter; labral anterior margin undulated, slightly incurved each side of midline, six-setose; penultimate maxillary palpomere broadened apically, apical palpomere a narrow spindle (Fig.
|
||
<figureCitation id="36B4CDDFD9FF132EFC11474A2D77BF23" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1B, E</figureCitation>
|
||
).
|
||
<emphasis id="1CF4A5BC8B0151FD10F700769DC6B830" bold="true" italics="true" pageId="0" pageNumber="229">Prothorax</emphasis>
|
||
transverse, MPW/PL = 1.35-1.41, base moderately constricted with lateral margins sinuate anterad right hind angles, MPW/BPW = 1.21-1.34; pronotal median base depressed, longitudinally wrinkled, unmargined; basal margin beaded each side posterad laterobasal depression which is deepest just laterad broadly triangular median base; pronotal lateral margin beaded, depression mesad bead narrow but broad enough so that a row of sculpticells can be observed lining the groove; pronotal median impression finely incised, disc flat, anterior transverse impression obsolete, not evident medially or toward slightly protruded, narrowly rounded front angles.
|
||
<emphasis id="09964EF570EC0573F336934F20F9D465" bold="true" italics="true" pageId="0" pageNumber="229">Elytra</emphasis>
|
||
subquadrate, lateral margins evenly convex from humeri to apex, maximum width approximately midlength; basal groove present laterad position of fifth interneur, groove convexly joined to lateral marginal depression; lateral marginal depression reflexed, of equal breadth from seta Eo2 to subapical sinuation.
|
||
<emphasis id="DFFE4CA2D7D94323CD6F03E7DEA0740A" bold="true" italics="true" pageId="0" pageNumber="229">Pterothorax</emphasis>
|
||
elongate, mesepisternal depression smooth posterad juncture with mesosternum, depression deepest and broadest just dorsad mesocoxal cavity; metepisternum elongate, lateral length twice maximal width; metathoracic flight wings polymorphic, 18 of 19 individuals in type series with wings reduced to a vestigial stub (Fig.
|
||
<figureCitation id="36994B5BBDB2899E795DFB31E97C3817" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1J</figureCitation>
|
||
), and 1 female with fully developed flight wings (Fig.
|
||
<figureCitation id="DFD2188FC450C45774284E8C54D97087" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Right metathoracic flight wing, dorsal view A Paratachys terryli, macrophthalmic female; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B Tachys oahuensis Blackburn, female; Oahu: Honolulu, Public Health Dept. light trap, x- 1965, J. W. Beardsley (BPBM). Wing vein terminology follows Kukalova-Peck and Lawrence (1993) except for the oblongum cell of Forbes (1922). From front to back of wing, veins, and cells relevant to the text include the Radial Cell (RC). Radius Anterior (RA), Radius Posterior (RP), Oblongum cell (O), Media Posterior (MP), Cubitus Anterior (CuA), Anal Anterior (AA), and Anal Posterior (AP)." figureDoi="10.3897/zookeys.1044.59674.figure2" httpUri="https://binary.pensoft.net/fig/553678" pageId="0" pageNumber="229">2A</figureCitation>
|
||
), with length of alar surface 2.3
|
||
<normalizedToken id="A9000EA4930DD0602EF5D3DF210D5783" originalValue="×">x</normalizedToken>
|
||
breadth, and radial, medial, cubital and anal veins and an oblongum cell present.
|
||
<emphasis id="8819D2059FAC51FDFE825887C8A3FA82" bold="true" italics="true" pageId="0" pageNumber="229">Abdomen</emphasis>
|
||
with one seta each side of apical ventrite in males, two seta each side of ventrite in females.
|
||
<emphasis id="7A637E1D1D01B6AC9A52B5CE415290B3" bold="true" italics="true" pageId="0" pageNumber="229">Microsculpture</emphasis>
|
||
evident on all somites; frons covered with evident transverse mesh, sculpticells more isodiametric posteriorly on vertex; pronotum slightly iridescent due to elongate transverse sculpticells, median base and laterobasal depressions more opaque due to reticulated isodiametric microsculpture; elytra subiridescent due to a mix of elongate transverse-mesh and transverse-line microsculpture, the transverse lines denser laterally; elytral apex more opaque due to slightly raised isodiametric sculpticells between sutural interneur and apical recurrent groove; abdominal ventrites covered with elongate transverse sculpticells, the surface glossy to subiridescent.
|
||
<emphasis id="0AAA2852FD6CACB35BB385357CC753A9" bold="true" italics="true" pageId="0" pageNumber="229">Pelage</emphasis>
|
||
present on head, prothorax, elytra, pterothorax, abdominal ventrites and legs; pelage on head and pronotal disc, comprising microsetae separated by distances subequal to setal length (Fig.
|
||
<figureCitation id="2FA8AE7F146D5468F2F15582931842DB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1F, G</figureCitation>
|
||
), as well as along ommatidial margins of the eyes (Fig.
|
||
<figureCitation id="CA1AEB7D8FCCFA9E79B0B0AACE3D5A98" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1B</figureCitation>
|
||
); microsetae spaced slightly farther apart on elytra, with intersetal distances up to twice microsetal length; prosternum and mesosternum medially covered with microsetae as densely distributed as on frons and pronotum; pelage on abdomen present at middle of ventrite 2 between metalegs, and progressively more broadly on ventrites 3-6 (absent dorsad arc of metaleg movement); anterior surfaces of pro-, meso-, and metathoracic legs bearing pelage of elongate microsetae, the setal bases situated more closely than microsetal lengths, trochanters and coxae similarly covered with microsetae.
|
||
<emphasis id="C3025FEDBA53269E970D09F7C417D6C7" bold="true" italics="true" pageId="0" pageNumber="229">Coloration</emphasis>
|
||
moderately dark; vertex and frons dark brunneous to piceous, clypeus flavobrunneous, labrum flavous; basal two antennomeres flavous, outer antennomeres progressively darker, apical segments brunneous; maxillary and labial palps flavous; pronotal disc and elytra brunneous, elytral base near scutellum paler, rufobrunneous, elytral lateral marginal depression flavobrunneous, elytral epipleuron rufoflavous, contrasted to rufobrunneous thoracic ventrites; legs flavous from trochanters outward; pro- and mesocoxae concolorous with outer leg segments, metacoxae more brunneous laterally to match the dark brunneous ventrites.
|
||
<emphasis id="C2CB48F0D4010340217112CDE733D833" bold="true" italics="true" pageId="0" pageNumber="229">Legs</emphasis>
|
||
with basal male protarsomere alone bearing a blunt, antero-apical process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3C29DAD4A27E6669C10AF33ADB60D12B" pageId="0" pageNumber="229" type="variation">
|
||
<paragraph id="687061AD2827A6C173DFC5EA59574122" pageId="0" pageNumber="229">Variation.</paragraph>
|
||
<paragraph id="7024AD22D34C48EAE5C5A2140004ECA9" pageId="0" pageNumber="229">
|
||
The compound eyes vary dramatically in this species, from fully macrophthalmic (Fig.
|
||
<figureCitation id="B6EF5DD4014DA374A313A99989B7F227" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1C</figureCitation>
|
||
), to microphthalmic (Fig.
|
||
<figureCitation id="039655358A79714245709205FFE459F5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1I</figureCitation>
|
||
); OR ranging 1.15-1.35 for smallest- to largest-eyed individuals. That these differences reflect infraspecific variation is supported by presence of the extremes plus a variety of intermediate configurations (Fig.
|
||
<figureCitation id="9727B78A8DB75F9A0C197B2A0C3AB62A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1D-H</figureCitation>
|
||
) among specimens collected microsympatrically within the identical collecting series (Fig.
|
||
<figureCitation id="4B3E91245F7A3A131B006E52771988BE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1C-I</figureCitation>
|
||
).
|
||
<taxonomicName id="04F8430485D013000137D9383D0AD2DD" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys terryli" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="terryli">
|
||
<emphasis id="7B82FAD4AF17FF20E737C9B6C5847B48" italics="true" pageId="0" pageNumber="229">Paratachys terryli</emphasis>
|
||
</taxonomicName>
|
||
is also polymorphic for flight-wing development, with a single macrophthalmic female specimen (Fig.
|
||
<figureCitation id="8E6ABFDBDCEC1422BA949EBE4265EE54" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paratachys terryli A-I Kauai: Namolokama Mtn. A, B macrophthalmic, brachypterous female; 23 - v- 2005 lot 01 A dorsal habitus view B mentum and associated mouthparts, ventral view C-I compound eye, lateral view C macropterous female; 23 - v- 2005 lot 1 D brachypterous male; 21 - v- 2005 lot 8 E brachypterous male; 23 - v- 2005 lot 1 F brachypterous female; 23 - v- 2005 lot 1 G brachypterous female; 23 - v- 2005 lot 1 H brachypterous female; 23 - v- 2005 lot 1 I brachypterous female; 21 - v- 2005 lot 1 J Kauai: Upper Kawaikoi Stream at Alakai Swamp Trail, 16 - v- 1971; right metathoracic flight wing of male, vestigial morph (elytron removed); 14 - v- 1991 stop 91 - 26 B." figureDoi="10.3897/zookeys.1044.59674.figure1" httpUri="https://binary.pensoft.net/fig/553677" pageId="0" pageNumber="229">1C</figureCitation>
|
||
) bearing fully developed flight wings (Fig.
|
||
<figureCitation id="EEF7029ECEA7A34FF168DFC4FB1495FF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Right metathoracic flight wing, dorsal view A Paratachys terryli, macrophthalmic female; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B Tachys oahuensis Blackburn, female; Oahu: Honolulu, Public Health Dept. light trap, x- 1965, J. W. Beardsley (BPBM). Wing vein terminology follows Kukalova-Peck and Lawrence (1993) except for the oblongum cell of Forbes (1922). From front to back of wing, veins, and cells relevant to the text include the Radial Cell (RC). Radius Anterior (RA), Radius Posterior (RP), Oblongum cell (O), Media Posterior (MP), Cubitus Anterior (CuA), Anal Anterior (AA), and Anal Posterior (AP)." figureDoi="10.3897/zookeys.1044.59674.figure2" httpUri="https://binary.pensoft.net/fig/553678" pageId="0" pageNumber="229">2A</figureCitation>
|
||
). The wings of this female are of dimensions similar to those of fully flighted individuals of
|
||
<taxonomicName id="3779463C6A94815F935D1E09D05DB174" authorityName="Blackburn" authorityYear="1878" class="Insecta" family="Carabidae" genus="Tachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachys oahuensis" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="oahuensis">
|
||
<emphasis id="87FC3A66A7EACE631FDE496883BE84DD" italics="true" pageId="0" pageNumber="229">Tachys oahuensis</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="0D6F9E172F4846EF393D5E710622F794" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Right metathoracic flight wing, dorsal view A Paratachys terryli, macrophthalmic female; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B Tachys oahuensis Blackburn, female; Oahu: Honolulu, Public Health Dept. light trap, x- 1965, J. W. Beardsley (BPBM). Wing vein terminology follows Kukalova-Peck and Lawrence (1993) except for the oblongum cell of Forbes (1922). From front to back of wing, veins, and cells relevant to the text include the Radial Cell (RC). Radius Anterior (RA), Radius Posterior (RP), Oblongum cell (O), Media Posterior (MP), Cubitus Anterior (CuA), Anal Anterior (AA), and Anal Posterior (AP)." figureDoi="10.3897/zookeys.1044.59674.figure2" httpUri="https://binary.pensoft.net/fig/553678" pageId="0" pageNumber="229">2B</figureCitation>
|
||
) that were collected in ultraviolet light traps. Venation differs among the two compared species, with the wing of the female
|
||
<taxonomicName id="9F99762FC0457710C2946E2ABCA549CE" lsidName="P. terryli" pageId="0" pageNumber="229" rank="species" species="terryli">
|
||
<emphasis id="E30CD52BDC76589EB6B4FF082E50909A" italics="true" pageId="0" pageNumber="229">P. terryli</emphasis>
|
||
</taxonomicName>
|
||
exhibiting the oblongum cell but not the basal stem of the radius posterior (
|
||
<bibRefCitation id="FEC9B8CF744D7BEDED1B508223754BFC" DOI="https://doi.org/10.4039/Ent125181-2" author="Kukalova-Peck, J" journalOrPublisher="Canadian Entomologist" pageId="0" pageNumber="229" pagination="181 - 258" refId="B55" refString="Kukalova-Peck, J, Lawrence, JF, 1993. Evolution of the hind wing in Coleoptera. Canadian Entomologist 125: 181 - 258, DOI: https://doi.org/10.4039/Ent125181-2" title="Evolution of the hind wing in Coleoptera." url="https://doi.org/10.4039/Ent125181-2" volume="125" year="1993">
|
||
<normalizedToken id="CFFAD9CE6018F20247177D6A309CBBC2" originalValue="Kukalová-Peck">Kukalova-Peck</normalizedToken>
|
||
and Lawrence 1993
|
||
</bibRefCitation>
|
||
), whereas the wing of
|
||
<taxonomicName id="9F940A385D31E232D46D2CB08E62DD37" lsidName="T. oahuensis" pageId="0" pageNumber="229" rank="species" species="oahuensis">
|
||
<emphasis id="28719F1DD4B5CC7884D4EAD03A040816" italics="true" pageId="0" pageNumber="229">T. oahuensis</emphasis>
|
||
</taxonomicName>
|
||
lacks the oblongum but retains the radius posterior. Both wings are folded reflexively under the elytra, with the spring-like wing margin apicad the radial cell flipping the wing open when the wing is deployed. The various brachypterous individuals exhibit a variety of eye configurations. Wing configuration is not associated with morphological variation in elytral dimensions based on comparison of four simultaneously collected females (Namolokama Mountain, 23-v-2005, lot 1) with HuW/MEW for the macropterous female = 0.57, and HuW/MEW for the three brachypterous females = 0.57-0.58.
|
||
</paragraph>
|
||
<caption id="F86D936985F0D2C85AF3CD535DCC50B6" doi="10.3897/zookeys.1044.59674.figure2" httpUri="https://binary.pensoft.net/fig/553678" pageId="0" pageNumber="229" start="Figure 2" startId="F2">
|
||
<paragraph id="20C315083CB1790039CBFDDB445B8412" pageId="0" pageNumber="229">
|
||
<emphasis id="D2104FE437099FE2FC77BEF8FB460854" bold="true" pageId="0" pageNumber="229">Figure 2.</emphasis>
|
||
Right metathoracic flight wing, dorsal view
|
||
<emphasis id="44472DC401BC2E5DFD3C9473083235C4" bold="true" pageId="0" pageNumber="229">A</emphasis>
|
||
<taxonomicName id="4C32AB177086FC75A707A8F7706A6F6B" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys terryli" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="terryli">
|
||
<emphasis id="0B0486E031474D8DD2724149C188F3A7" italics="true" pageId="0" pageNumber="229">Paratachys terryli</emphasis>
|
||
</taxonomicName>
|
||
, macrophthalmic female; Kauai: Namolokama Mtn., 23-v-2005 (CUIC)
|
||
<emphasis id="EFA3BB76318B364F9F8D0AFE6B2E22AE" bold="true" pageId="0" pageNumber="229">B</emphasis>
|
||
<taxonomicName id="DC0F89EDDE4F335898E7DAFF1C9B1E3E" authorityName="Blackburn" authorityYear="1878" class="Insecta" family="Carabidae" genus="Tachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachys oahuensis" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="oahuensis">
|
||
<emphasis id="1742EC96F58A824F46AA75B6482A7F58" italics="true" pageId="0" pageNumber="229">Tachys oahuensis</emphasis>
|
||
</taxonomicName>
|
||
Blackburn, female; Oahu: Honolulu, Public Health Dept. light trap, x-1965, J. W. Beardsley (BPBM). Wing vein terminology follows
|
||
<bibRefCitation id="56438BD6C526A3676C1BA952C6B46B69" DOI="https://doi.org/10.4039/Ent125181-2" author="Kukalova-Peck, J" journalOrPublisher="Canadian Entomologist" pageId="0" pageNumber="229" pagination="181 - 258" refId="B55" refString="Kukalova-Peck, J, Lawrence, JF, 1993. Evolution of the hind wing in Coleoptera. Canadian Entomologist 125: 181 - 258, DOI: https://doi.org/10.4039/Ent125181-2" title="Evolution of the hind wing in Coleoptera." url="https://doi.org/10.4039/Ent125181-2" volume="125" year="1993">
|
||
<normalizedToken id="EC44F42ED990B7C9C34EBC6C5FBE648C" originalValue="Kukalová-Peck">Kukalova-Peck</normalizedToken>
|
||
and Lawrence (1993)
|
||
</bibRefCitation>
|
||
except for the oblongum cell of Forbes (1922). From front to back of wing, veins, and cells relevant to the text include the Radial Cell (RC). Radius Anterior (RA), Radius Posterior (RP), Oblongum cell (O), Media Posterior (MP), Cubitus Anterior (CuA), Anal Anterior (AA), and Anal Posterior (AP).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="BF852CA53424DF0939FDA940DA51D90E" pageId="0" pageNumber="229" type="male genitalia">
|
||
<paragraph id="124A70B298B0BE2C35ADE17A44C654A1" pageId="0" pageNumber="229">Male genitalia.</paragraph>
|
||
<paragraph id="91ABAEA3666E68353806392B06CB727F" pageId="0" pageNumber="229">
|
||
Aedeagal median lobe porrect, parallel-sided with an evenly rounded apex (Fig.
|
||
<figureCitation id="32E42FFCD6A967F364BD2370C417D93E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paratachys spp., male aedeagus, i. e., median lobe and associated smaller right paramere and larger left paramere (right lateral view) A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. arcanicola lectotype slide; " Tachys arcanicola Blackburn Type ♂ E. C. Z. (permanent slide, Euparal?, left paramere missing; NHMUK). " C P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) D-F P. aaa; Hawai'i I D Puna, Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC) E Kau, Ocean View, Kipuka Kanohina System, 21 - xi- 2017, lot HI 00179 (CUIC) F South Hilo, Kaumana Cave, 19 - iii- 2019, lot HI 00942 (CUIC); disassociated left paramere below, ectal view." figureDoi="10.3897/zookeys.1044.59674.figure3" httpUri="https://binary.pensoft.net/fig/553679" pageId="0" pageNumber="229">3A</figureCitation>
|
||
); flagellar complex elongate, with a sinuously scooped apical margin; right paramere narrow, strap-like with three apical setae; left paramere broad, elongate, with a broadly rounded apex and three apical setae.
|
||
</paragraph>
|
||
<caption id="82C22B88C810D5FD3A7057D3302F7733" doi="10.3897/zookeys.1044.59674.figure3" httpUri="https://binary.pensoft.net/fig/553679" pageId="0" pageNumber="229" start="Figure 3" startId="F3">
|
||
<paragraph id="ECA67F1EE97A413BCE35353A0FA90197" pageId="0" pageNumber="229">
|
||
<emphasis id="EC6D326E852B877989A54DB8D8C5679F" bold="true" pageId="0" pageNumber="229">Figure 3.</emphasis>
|
||
<taxonomicName id="2A40E6D18ECA40B0EE50961B0F56C0FB" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="399C22F709182B7628F77CCDE67DEBF2" italics="true" pageId="0" pageNumber="229">Paratachys</emphasis>
|
||
</taxonomicName>
|
||
spp., male aedeagus, i.e., median lobe and associated smaller right paramere and larger left paramere (right lateral view)
|
||
<emphasis id="744335126576D206107E61A96F2188F1" bold="true" pageId="0" pageNumber="229">A</emphasis>
|
||
<taxonomicName id="E8C6E61134E079F7A817DE3BEE2C7B9F" lsidName="P. terryli" pageId="0" pageNumber="229" rank="species" species="terryli">
|
||
<emphasis id="9870562374C183EB85157B7F3ADC33EB" italics="true" pageId="0" pageNumber="229">P. terryli</emphasis>
|
||
</taxonomicName>
|
||
; Kauai: Namolokama Mtn., 23-v-2005 (CUIC)
|
||
<emphasis id="E6B0BBDBEDCC05F15B2C2C2A0EAB4B9E" bold="true" pageId="0" pageNumber="229">B</emphasis>
|
||
<taxonomicName id="7C0E71F8A780934C11BA6DF8F440634F" lsidName="P. arcanicola" pageId="0" pageNumber="229" rank="species" species="arcanicola">
|
||
<emphasis id="80514EE0EFB09ED35A79619A21117D31" italics="true" pageId="0" pageNumber="229">P. arcanicola</emphasis>
|
||
</taxonomicName>
|
||
lectotype slide; "
|
||
<taxonomicName id="5159189D95B00934D58234402798EC45" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Tachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachys arcanicola" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="arcanicola">
|
||
<emphasis id="97A4EACB2A26D2A54AB1C155FBF271B8" italics="true" pageId="0" pageNumber="229">Tachys arcanicola</emphasis>
|
||
</taxonomicName>
|
||
Blackburn Type ♂ E.C.Z. (permanent slide, Euparal?, left paramere missing; NHMUK)."
|
||
<emphasis id="342A17C44AFB83D772673C2491B32B98" bold="true" pageId="0" pageNumber="229">C</emphasis>
|
||
<taxonomicName id="7A1500039461DCBE7D91AE94C69411B3" lsidName="P. haleakalae" pageId="0" pageNumber="229" rank="species" species="haleakalae">
|
||
<emphasis id="35B1A69DCA07B33ED73456B716D61AA4" italics="true" pageId="0" pageNumber="229">P. haleakalae</emphasis>
|
||
</taxonomicName>
|
||
; East Maui: Haleakala N. P., Kipahulu Vy., 30-iv-1991 (CUIC)
|
||
<emphasis id="3B01CDA794D827B9CFE4CCCF80F52785" bold="true" pageId="0" pageNumber="229">D-F</emphasis>
|
||
<taxonomicName id="5868DC573DC28755B8AC2D9429B24971" lsidName="P. aaa" pageId="0" pageNumber="229" rank="species" species="aaa">
|
||
<emphasis id="50EA366A4569F9772B3F9211A7DFA9B8" italics="true" pageId="0" pageNumber="229">P. aaa</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<normalizedToken id="35CD63F728F55A85FC9F118D45D48825" originalValue="Hawai‘i">Hawai'i</normalizedToken>
|
||
I
|
||
<emphasis id="C22E0D88D13B8E15C07DDB2FA12624E7" bold="true" pageId="0" pageNumber="229">D</emphasis>
|
||
Puna, Mountain View, Kazumura Cave, 21-xi-2018, lot HI00713 (CUIC)
|
||
<emphasis id="A443EB90E8E0C8117C01D7215A91E249" bold="true" pageId="0" pageNumber="229">E</emphasis>
|
||
Kau, Ocean View, Kipuka Kanohina System, 21-xi-2017, lot HI00179 (CUIC)
|
||
<emphasis id="B7520C3638FABF0AAE2A24459C6E1BBB" bold="true" pageId="0" pageNumber="229">F</emphasis>
|
||
South Hilo, Kaumana Cave, 19-iii-2019, lot HI00942 (CUIC); disassociated left paramere below, ectal view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="DB60E32A343DC62C901DC82AD37397F3" pageId="0" pageNumber="229" type="female reproductive tract">
|
||
<paragraph id="1FA19E12763B9374A7AB3D3170B0D236" pageId="0" pageNumber="229">Female reproductive tract.</paragraph>
|
||
<paragraph id="2F496058B12E408D86A502EEC2F72F3D" pageId="0" pageNumber="229">
|
||
Bursa copulatrix short, broad (as in Fig.
|
||
<figureCitation id="2C77CA7EE307A4103382E74B9BF90DD7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Paratachys aaa female reproductive tract, ventral view; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC). Abbreviations: bc, bursa copulatrix; co, common oviduct; sd, spermathecal duct; sg, spermathecal gland; sp, spermatheca. Schematic drawing breaks spermathecal duct at " A " to accommodate drawing to page size. Estimated length of spermathecal duct 1.7 mm." figureDoi="10.3897/zookeys.1044.59674.figure10" httpUri="https://binary.pensoft.net/fig/553686" pageId="0" pageNumber="229">10</figureCitation>
|
||
), spermathecal duct narrow and extremely elongate, at least 0.57 mm long in single dissection attempt; gonocoxa bipartite, basal gonocoxite 1 narrow, elongate, with a broad, lateral sclerotized apodeme and a single apical fringe seta situated near apex of apodeme (Fig.
|
||
<figureCitation id="4A3F16DD1FB8D1A75A7DD5FEDA0FBE54" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paratachys spp., left female gonocoxa, ventral view A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) C P. aaa; Hawai'i I.; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC). Abbreviations: afs, apical fringe seta; ans, apical nematiform setae; des, dorsal ensiform seta; gc 1, basal gonocoxite; gc 2, apical gonocoxite; les, lateral ensiform setae." figureDoi="10.3897/zookeys.1044.59674.figure4" httpUri="https://binary.pensoft.net/fig/553680" pageId="0" pageNumber="229">4A</figureCitation>
|
||
); apical gonocoxite falcate, broadly expanded laterally at base, apex finely acuminate; two peg-like lateral ensiform setae along mid-ventral line, and one like-sized dorsal ensiform seta dorsad the apical lateral seta; two apical nematiform setae in an elongate fossa situated at approximately 2/3 length of the apical gonocoxite.
|
||
</paragraph>
|
||
<caption id="3F24E4F0E9C6746FBBE191E29749FB52" doi="10.3897/zookeys.1044.59674.figure4" httpUri="https://binary.pensoft.net/fig/553680" pageId="0" pageNumber="229" start="Figure 4" startId="F4">
|
||
<paragraph id="D517B6D966A932B5688E70FA5BDD0304" pageId="0" pageNumber="229">
|
||
<emphasis id="342E113FE4A57B8A27A119B109694244" bold="true" pageId="0" pageNumber="229">Figure 4.</emphasis>
|
||
<taxonomicName id="89F9ED1F9AF3FE2A1FF3B1039496A8AB" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AF366746A2A0A8F1234F210AF9699871" italics="true" pageId="0" pageNumber="229">Paratachys</emphasis>
|
||
</taxonomicName>
|
||
spp., left female gonocoxa, ventral view
|
||
<emphasis id="540D2EA6C886A3C71F835DCA0847887C" bold="true" pageId="0" pageNumber="229">A</emphasis>
|
||
<taxonomicName id="2BC7E1F73508E73B7050BA5B3FC17F03" lsidName="P. terryli" pageId="0" pageNumber="229" rank="species" species="terryli">
|
||
<emphasis id="54B5538AFCF4523939A2C3CAB4F09EC0" italics="true" pageId="0" pageNumber="229">P. terryli</emphasis>
|
||
</taxonomicName>
|
||
; Kauai: Namolokama Mtn., 23-v-2005 (CUIC)
|
||
<emphasis id="210EF66CA6DB2041EB116C51E68B31A8" bold="true" pageId="0" pageNumber="229">B</emphasis>
|
||
<taxonomicName id="2C7030219CF9B8170AC9669DA7C12221" lsidName="P. haleakalae" pageId="0" pageNumber="229" rank="species" species="haleakalae">
|
||
<emphasis id="619C3C97F4A671596B66D29D9CB043EF" italics="true" pageId="0" pageNumber="229">P. haleakalae</emphasis>
|
||
</taxonomicName>
|
||
; East Maui: Haleakala N. P., Kipahulu Vy., 30-iv-1991 (CUIC)
|
||
<emphasis id="E8ACD9BC3CB3A1DEF4E752EBCD997BE9" bold="true" pageId="0" pageNumber="229">C</emphasis>
|
||
<taxonomicName id="163BAA0749BBDBB8CFBEF58B503B8F7B" lsidName="P. aaa" pageId="0" pageNumber="229" rank="species" species="aaa">
|
||
<emphasis id="9502D003919E44649D38FE0AAF49A982" italics="true" pageId="0" pageNumber="229">P. aaa</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<normalizedToken id="C10F3C2FB721A56F279CAB5579A51307" originalValue="Hawai‘i">Hawai'i</normalizedToken>
|
||
I.; Mountain View, Kazumura Cave, 21-xi-2018, lot HI00713 (CUIC). Abbreviations: afs, apical fringe seta; ans, apical nematiform setae; des, dorsal ensiform seta; gc1, basal gonocoxite; gc2, apical gonocoxite; les, lateral ensiform setae.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="9F68ED59AEB4D3CADFED063E9C7BAFAA" pageId="0" pageNumber="229" type="etymology">
|
||
<paragraph id="4309C5BA26B4EC08CDA547AAE904F50F" pageId="0" pageNumber="229">Etymology.</paragraph>
|
||
<paragraph id="62F92357EF250BDAE3309E4CA31D1B17" pageId="0" pageNumber="229">
|
||
This species honors Terry L. Erwin by combining his first name and middle initial to form the genitive patronym
|
||
<taxonomicName id="BCDE7458D3626F10AA7C5BC2EC8288C0" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys terryli" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="terryli">
|
||
<emphasis id="D7EAE91C38485FAFFA5236AABC35AE10" italics="true" pageId="0" pageNumber="229">Paratachys terryli</emphasis>
|
||
</taxonomicName>
|
||
. This construction follows that of
|
||
<taxonomicName id="E58B15A2D7A77CEEC59E2FDAEDC51B08" authorityName="Erwin & Kavanaugh" authorityYear="1981" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion carlhi" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="carlhi">
|
||
<emphasis id="279DB07FFF2F46926CF3D6CB06B430F0" italics="true" pageId="0" pageNumber="229">Bembidion carlhi</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="AF9B366576E491DE6FAC114A36A79C5B" author="Erwin, TL" journalOrPublisher="Entomologica Scandinavica Supplementum" pageId="0" pageNumber="229" pagination="33 - 72" refId="B35" refString="Erwin, TL, Kavanaugh, DH, 1981. Systematics and zoogeography of Bembidion Latreille: I. The carlhi and erasum groups of western North America (Coleoptera: Carabidae, Bembidiini). Entomologica Scandinavica Supplementum 15: 33 - 72" title="Systematics and zoogeography of Bembidion Latreille: I. The carlhi and erasum groups of western North America (Coleoptera: Carabidae, Bembidiini)." volume="15" year="1981">Erwin and Kavanaugh (1981)</bibRefCitation>
|
||
, a species that honors Carl H. Lindroth whose 'Carabid Beetles of Canada and
|
||
<normalizedToken id="26D68C94D6653AB5D33DBE3DB2E18F28" originalValue="Alaska’">Alaska'</normalizedToken>
|
||
monograph (Lindroth 1969 et. seq.) revolutionized the study of North American
|
||
<taxonomicName id="67419AE2EDE77931A411CCC068979EAD" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
||
. Analogous to
|
||
<taxonomicName id="E241685854CFB6485250F4909E5320C8" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. carlhi" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="carlhi">
|
||
<emphasis id="00776D8A03BE234C9176EDB92DA4891F" italics="true" pageId="0" pageNumber="229">B. carlhi</emphasis>
|
||
</taxonomicName>
|
||
, this epithet recognizes
|
||
<normalizedToken id="DBF60DE3A404E126FFAF0F0A9C491C25" originalValue="Terry’s">Terry's</normalizedToken>
|
||
immense impact on the study of Neotropical
|
||
<taxonomicName id="519A374A2FDEFED3DF8164889C1A7CF9" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
||
as well as tropical biodiversity writ large.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="70F6C3B5FE696B994C53A3048901AE00" pageId="0" pageNumber="229" type="distribution">
|
||
<paragraph id="873E0A831CE8FBCFA880F9622D25B893" pageId="0" pageNumber="229">Distribution and habitat.</paragraph>
|
||
<paragraph id="38B2855FE477534070F65C02C270831D" pageId="0" pageNumber="229">
|
||
This species is known from the Alakai Swamp west of the Wainiha River, and from Namolokama Mountain, a ridge to the east bordered by the Lumahei and Hanalei Rivers (Fig.
|
||
<figureCitation id="A3387DF15D7EA44DF1DF54FE820F9427" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distributional records for Paratachys terryli." figureDoi="10.3897/zookeys.1044.59674.figure5" httpUri="https://binary.pensoft.net/fig/553681" pageId="0" pageNumber="229">5</figureCitation>
|
||
). It has been most commonly collected in leaf and moss litter taken from low stature Ohia lehua (
|
||
<taxonomicName id="85E4D5E7D7E1701A0E1118CF69FE9F3E" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Metrosideros polymorpha" order="Myrtales" pageId="0" pageNumber="229" phylum="Tracheophyta" rank="species" species="polymorpha">
|
||
<emphasis id="03C717303DA4494E2BB1D9A89D2077A3" italics="true" pageId="0" pageNumber="229">Metrosideros polymorpha</emphasis>
|
||
</taxonomicName>
|
||
Gaud.;
|
||
<taxonomicName id="E1C82AA7B2EFDDDE9A415F1FA0426924" class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="0" pageNumber="229" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
|
||
) forest. At a site at 1340 m elevation on Namolokama, one sample (21-v-2005, lot 1) sifted and subsequently hand-picked contained five
|
||
<taxonomicName id="669D0F2E1A73F60C3D14A9B053FD99D1" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys terryli" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="terryli">
|
||
<emphasis id="2383AEF8856FB1FBBC61AB00E811038A" italics="true" pageId="0" pageNumber="229">Paratachys terryli</emphasis>
|
||
</taxonomicName>
|
||
in company with two
|
||
<taxonomicName id="C116E73276CB1233E61DEABA86F885BC" baseAuthorityName="Sharp" baseAuthorityYear="1903" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion admirandum" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="admirandum">
|
||
<emphasis id="0B84BAD503B13B0FB2479BE75FCA5252" italics="true" pageId="0" pageNumber="229">Bembidion admirandum</emphasis>
|
||
</taxonomicName>
|
||
(Sharp), two
|
||
<taxonomicName id="2B61ECAD15431B6AD75C414D7C572F79" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. corticarium" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="corticarium">
|
||
<emphasis id="7B8CF0EE5D53A4E9213B24B3FDA597F9" italics="true" pageId="0" pageNumber="229">B. corticarium</emphasis>
|
||
</taxonomicName>
|
||
(Sharp), and two
|
||
<taxonomicName id="6798C7D9496C837D6500E509778E4888" baseAuthorityName="Sharp" baseAuthorityYear="1903" class="Insecta" family="Carabidae" genus="Blackburnia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Blackburnia kauaiensis" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="kauaiensis">
|
||
<emphasis id="F9CF9A07EDC8CA181ABE2F34797EB969" italics="true" pageId="0" pageNumber="229">Blackburnia kauaiensis</emphasis>
|
||
</taxonomicName>
|
||
(Sharp). A second sample from the same site (21-v-2005, lot 8) was double-boiled on a stove, resulting in discovery of one
|
||
<taxonomicName id="538915035A241C073A60F6C787BAE301" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. admirandum" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="admirandum">
|
||
<emphasis id="D2FDE6EADAA5620967385FFF17664C87" italics="true" pageId="0" pageNumber="229">B. admirandum</emphasis>
|
||
</taxonomicName>
|
||
, eight
|
||
<taxonomicName id="833DF9E6F4EADC5FDA9CD1929E156C1C" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. corticarium" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="corticarium">
|
||
<emphasis id="2E84DC7FF6B2DBAE5C5A6A2669D93382" italics="true" pageId="0" pageNumber="229">B. corticarium</emphasis>
|
||
</taxonomicName>
|
||
, and one
|
||
<taxonomicName id="44737002B22EE47E1A4160A2C568B3BC" lsidName="P. terryli" pageId="0" pageNumber="229" rank="species" species="terryli">
|
||
<emphasis id="6BD454E895FFD52D8C80C7F014EAA516" italics="true" pageId="0" pageNumber="229">P. terryli</emphasis>
|
||
</taxonomicName>
|
||
. A third Namolokama sift sample from 1305 m elevation (23-v-2005, lot 1) contained seven
|
||
<taxonomicName id="8B27C3FC2AA43CEC11E8FA54FA7A80F9" lsidName="P. terryli" pageId="0" pageNumber="229" rank="species" species="terryli">
|
||
<emphasis id="489F959AD2F1B086CF145893127821DB" italics="true" pageId="0" pageNumber="229">P. terryli</emphasis>
|
||
</taxonomicName>
|
||
in company with specimens of three species of
|
||
<taxonomicName id="4711C9F21D97D3A709DC402A258633AA" class="Insecta" family="Carabidae" genus="Blackburnia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Blackburnia" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="501DFA75C456C53D21ED62F3F762D68F" italics="true" pageId="0" pageNumber="229">Blackburnia</emphasis>
|
||
</taxonomicName>
|
||
; two
|
||
<taxonomicName id="134D35E4247AAF06E3B57D55B9B3EB5B" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. posticata" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="posticata">
|
||
<emphasis id="6D45C6C917A7A310F14C6434CF231D45" italics="true" pageId="0" pageNumber="229">B. posticata</emphasis>
|
||
</taxonomicName>
|
||
(Sharp), and one each
|
||
<taxonomicName id="AA35A5EFE9343970E749DA1E77DC4023" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. bryophila" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="bryophila">
|
||
<emphasis id="C13542812F1B47DB47D93EFF98325694" italics="true" pageId="0" pageNumber="229">B. bryophila</emphasis>
|
||
</taxonomicName>
|
||
Liebherr and
|
||
<taxonomicName id="FE273E5C4CFD688AE01B798BC0DF15CC" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. kauaiensis" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="kauaiensis">
|
||
<emphasis id="05217B8C68764F8A4CB81ED2F424101C" italics="true" pageId="0" pageNumber="229">B. kauaiensis</emphasis>
|
||
</taxonomicName>
|
||
(Sharp). Most other collections of
|
||
<taxonomicName id="5D0387C4C8FE1E93A794C06BDD84248A" lsidName="P. terryli" pageId="0" pageNumber="229" rank="species" species="terryli">
|
||
<emphasis id="922A00AF0045EAA284A467204F16FB1F" italics="true" pageId="0" pageNumber="229">P. terryli</emphasis>
|
||
</taxonomicName>
|
||
represent singletons most often collected in company with
|
||
<taxonomicName id="D57633B28024DE9FDB76F3D99A8B9F24" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9555B9FA07D1C1BD9C2DC434376016B" italics="true" pageId="0" pageNumber="229">Bembidion</emphasis>
|
||
</taxonomicName>
|
||
spp., including: 16-V-1991 lot 5 from a rocky streambed also including five
|
||
<taxonomicName id="8A9158DAD44AB5CE50C14EF238FA5E64" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. ignicola" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="ignicola">
|
||
<emphasis id="F4F34AA5C0E1ACDBAE69439F9171739E" italics="true" pageId="0" pageNumber="229">B. ignicola</emphasis>
|
||
</taxonomicName>
|
||
(Blackburn); 17-v-2005 lot 4 from Berlese extraction of leaf litter siftate along with two
|
||
<taxonomicName id="84B64486CCE5D1DBF6FD8CC2648E050E" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. admirandum" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="admirandum">
|
||
<emphasis id="D59BE5776758A7FAF449E362B6CF2BED" italics="true" pageId="0" pageNumber="229">B. admirandum</emphasis>
|
||
</taxonomicName>
|
||
and three
|
||
<taxonomicName id="6E899C44F764491789A21597A947D318" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. munroi" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="munroi">
|
||
<emphasis id="80BD3FA292C73F29DBF6D19AA639AF05" italics="true" pageId="0" pageNumber="229">B. munroi</emphasis>
|
||
</taxonomicName>
|
||
; and 17-v-2005 lot 9, double boiled from leaf litter siftate along with six
|
||
<taxonomicName id="18BEB610E389A67055C91627DA5510E2" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="B. admirandum" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="admirandum">
|
||
<emphasis id="1F530E7365E54F18119F6BF48C0E4B52" italics="true" pageId="0" pageNumber="229">B. admirandum</emphasis>
|
||
</taxonomicName>
|
||
and two
|
||
<taxonomicName id="C8F9DC18FB9B175ED1A1276EDDF8BE8E" lsidName="Bl. posticata" pageId="0" pageNumber="229" rank="species" species="posticata">
|
||
<emphasis id="D9AFE676AD5D702D03E297144AE39587" italics="true" pageId="0" pageNumber="229">Bl. posticata</emphasis>
|
||
</taxonomicName>
|
||
. Thus, it would appear from presently available evidence that
|
||
<taxonomicName id="0DD15D968D32D9496E8C532A316C2E02" lsidName="P. terryli" pageId="0" pageNumber="229" rank="species" species="terryli">
|
||
<emphasis id="753BB31A286108AA5BB5D5C37B393363" italics="true" pageId="0" pageNumber="229">P. terryli</emphasis>
|
||
</taxonomicName>
|
||
occurs in microhabitats most often also occupied by
|
||
<taxonomicName id="BBCF14AAA30CA3F22444DD544DDF69F9" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FB400342823BBBDD0D4285342C2ED37B" italics="true" pageId="0" pageNumber="229">Bembidion</emphasis>
|
||
</taxonomicName>
|
||
beetles, either along streams, or in deep terrestrial leaf litter in montane wet ohia forest.
|
||
</paragraph>
|
||
<caption id="E8A3AAE77CE1B64181CABE38E52CD207" doi="10.3897/zookeys.1044.59674.figure5" httpUri="https://binary.pensoft.net/fig/553681" pageId="0" pageNumber="229" start="Figure 5" startId="F5">
|
||
<paragraph id="029BC6183DB51E42E09B81D33924EFF3" pageId="0" pageNumber="229">
|
||
<emphasis id="9374E1372FDB90E134FA529D6E63EA08" bold="true" pageId="0" pageNumber="229">Figure 5.</emphasis>
|
||
Distributional records for
|
||
<taxonomicName id="4ADE55645805DBDE4F71A676F94D8379" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys terryli" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="terryli">
|
||
<emphasis id="5EB55EA97326B0BB5A59261566CC43E9" italics="true" pageId="0" pageNumber="229">Paratachys terryli</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |