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<document id="919F3C7BC85CB85CA09236F8F0696C2E" ID-DOI="http://dx.doi.org/10.3897/zookeys.263.4158" ID-GBIF-Dataset="5fe6c912-0997-4d09-800d-101a2a9c26e8" ID-PMC="PMC3591764" ID-Pensoft-Pub="1313-2970-263-59" ID-PubMed="23653517" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-263-59" ModsDocOrigin="ZooKeys 263" ModsDocTitle="Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species" checkinTime="1451247727841" checkinUser="pensoft" docAuthor="Taylor, Christopher K." docDate="2013" docId="29C2B1CEE225ABF01A9003E9F1C3F255" docLanguage="en" docName="ZooKeys 263: 59-73" docOrigin="ZooKeys 263" docSource="http://dx.doi.org/10.3897/zookeys.263.4158" docTitle="Pantopsalis cheliferoides Colenso 1882" docType="treatment" docVersion="4" lastPageNumber="70" masterDocId="776AFFECFFB1FFC40947FF959858FFB7" masterDocTitle="Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species" masterLastPageNumber="73" masterPageNumber="59" pageNumber="67" updateTime="1732707370824" updateUser="ExternalLinkService">
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<mods:title id="CB7926F86C03FD2B27C2F8C72C016652">Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species</mods:title>
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<mods:namePart id="34239DB45FCDD35112347BEB343D20EB">Taylor, Christopher K.</mods:namePart>
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<mods:identifier id="084B7B9CF6E7EF4BED1C0D14747D7D8E" type="DOI">http://dx.doi.org/10.3897/zookeys.263.4158</mods:identifier>
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<treatment id="29C2B1CEE225ABF01A9003E9F1C3F255" ID-GBIF-Taxon="152040054" LSID="urn:lsid:plazi:treatment:29C2B1CEE225ABF01A9003E9F1C3F255" httpUri="http://treatment.plazi.org/id/29C2B1CEE225ABF01A9003E9F1C3F255" lastPageId="12" lastPageNumber="70" pageId="8" pageNumber="67" scope_family="Neopilionidae" scope_order="Opiliones" scope_subFamily="Enantiobuninae">
<subSubSection id="8E01D7AD04642BDB11D4C8C7D7FFDF4F" pageId="8" pageNumber="67" type="nomenclature">
<paragraph id="07F45475EA38832740101657F707DCFF" pageId="8" pageNumber="67">
<taxonomicName id="100DB388CFD1FA88B59DA2533BC668D6" ID-CoL="7XVBJ" LSID="http://species-id.net/wiki/Pantopsalis_cheliferoides" authority="Colenso, 1882" authorityName="Colenso" authorityYear="1882" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis cheliferoides" order="Opiliones" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="cheliferoides">Pantopsalis cheliferoides (Colenso, 1882)</taxonomicName>
Figure 4
</paragraph>
</subSubSection>
<subSubSection id="63620C82C6465E649BDEFA81EDFDDEFA" pageId="8" pageNumber="67" type="reference_group">
<paragraph id="6A2A5C9572CE2DF2F8CE386896FC77C0" pageId="8" pageNumber="67">
<taxonomicName id="008758560CBBE02B6C95D8C37D3A5DEC" class="Arachnida" family="Phalangiidae" genus="Phalangium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangium (Phrynus) cheliferoides" order="Opiliones" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="cheliferoides" subGenus="Phrynus">Phalangium (Phrynus) cheliferoides</taxonomicName>
<bibRefCitation id="44100ADEB1F48BCE0985C8371C1DA641" author="Colenso, W" journalOrPublisher="Transactions and Proceedings of the New Zealand Institute" pageId="12" pageNumber="71" pagination="165 - 73" title="On some newly-discovered New Zealand arachnids." volume="15" year="1882">Colenso 1882</bibRefCitation>
: 166.
</paragraph>
<paragraph id="E16BA3B94968119697339DF2D08C526F" pageId="8" pageNumber="67">
<taxonomicName id="080123AC63265703915C2747FDF24D4B" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis cheliferoides" order="Opiliones" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="cheliferoides">Pantopsalis cheliferoides</taxonomicName>
(
<bibRefCitation id="E157165DA37A073DED024696A8A91B97" author="Colenso, W" journalOrPublisher="Transactions and Proceedings of the New Zealand Institute" pageId="12" pageNumber="71" pagination="165 - 73" title="On some newly-discovered New Zealand arachnids." volume="15" year="1882">Colenso 1882</bibRefCitation>
): Vink in
<bibRefCitation id="E12935C15DF1870098B6A28ACADE8E5A" author="Nicholls, DC" journalOrPublisher="Records of the Canterbury Museum" pageId="13" pageNumber="72" pagination="37 - 48" title="A list of arachnid primary types held in Canterbury Museum." volume="14" year="2000">Nicholls et al. 2000</bibRefCitation>
: 46,
<bibRefCitation id="832D146B455808A6ED1FE2AEE8E803D2" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
: 65 (as nomen dubium).
</paragraph>
</subSubSection>
<subSubSection id="4B932BA455B2ED4082AE7654AA1503F8" pageId="8" pageNumber="67" type="specimens examined">
<paragraph id="1F33EA5F5D13DFC1218822CC7AE84598" pageId="8" pageNumber="67">Specimens examined.</paragraph>
<paragraph id="4872A1FCC8845744C0A5D5F87DCB68FC" pageId="8" pageNumber="67">
1 ♂, New Zealand, GB. Lake Waikaremoana, 19 November 1975, G. W. Ransay, beating; 1 ♂, New Zealand, GB. Te Urewera National Park, Lake Waikaremoana, Kaitawa,
<geoCoordinate id="E9A0D08D431BDB6EA3DAE8F5F852852D" direction="south" orientation="latitude" precision="925" value="-38.766666">38°46'S</geoCoordinate>
,
<geoCoordinate id="5C8A4766622C88BC3DB5BE7F85622BD1" direction="east" orientation="longitude" precision="925" value="178.38333">177°83'E</geoCoordinate>
, 18 November 2004, D. King, on outside of house.
</paragraph>
</subSubSection>
<subSubSection id="7F6B72E9AB26358B12C4C862625C64C1" lastPageId="10" lastPageNumber="69" pageId="9" pageNumber="68" type="description">
<paragraph id="1F9113A2A61502A84EBE55CDFB114F50" pageId="9" pageNumber="68">
<pageBreakToken id="CCE1F66151EA478BD59059AF0A77FC62" pageId="9" pageNumber="68" start="start">Description</pageBreakToken>
:
</paragraph>
<paragraph id="7180FA301F76FFA20D33E89172E1ED06" pageId="9" pageNumber="68">
Male: Total body length 3.55-4.03 (former measurement refers to 2004 specimen), prosoma length 1.62-2.06, prosoma width 2.55-3.70. Dorsal prosomal plate medium brown, with some yellowish patches laterally; anterior propeltidium and ocularium heavily denticulate, remainder of prosoma unarmed. Dorsum of opisthosoma dark purplish brown with few white spots medially in one specimen, longitudinal purple medial stripe in other; larger white spots in present in central part of opisthosomal dorsum, comparable to lateral
<normalizedToken id="C58F53AD5355781F0EEACABD2CC7FF5B" originalValue="‘arms’">'arms'</normalizedToken>
of median stripe in females of other
<taxonomicName id="BD3C77851697CCB1DC6BA1C7784428F4" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="9" pageNumber="68" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
species. Coxae medium brown mottled with honey brown; venter of opisthosoma medium purplish brown mottled with lighter purple.
</paragraph>
<paragraph id="2F6069D68ABA369EE0570C4F62145C45" pageId="10" pageNumber="69">
<pageBreakToken id="75355EFE3B5BBA5359261FFB939612BE" pageId="10" pageNumber="69" start="start">Chelicerae</pageBreakToken>
: Segment I 4.73-6.23, segment II 6.36-8.47. Segment I medium brown with cream patches at distal end; segment II orange-brown; both segments heavily denticulate. Segment II inflated in larger specimen, slender in smaller. Cheliceral fingers (Fig. 4c) short, mobile finger crescent-shaped.
</paragraph>
<paragraph id="B23E4973C56994B60312B7F59FAC9DB3" pageId="10" pageNumber="69">Pedipalps: Femur 1.81-2.32, patella 0.97-1.15, tibia 1.01-1.18, tarsus 2.10-3.02. Femur light purple at base, remainder of pedipalp shining white. Patella and tibia (Fig. 4d) prolaterally hypersetose, patella bulging prodistally but without distinct apophysis. Microtrichia on distalmost end of tarsus only. Tarsal claw with single ventral tooth.</paragraph>
<paragraph id="872092578AB076AF5082EB991D0C7175" pageId="10" pageNumber="69">Legs: Leg I femur 7.91-8.56, patella 1.36-1.58, tibia 6.03-6.14; leg II femur 13.25-13.69, patella 1.82-1.80, tibia 11.18-11.47; leg III femur 6.91-7.38, patella 1.32-1.45, tibia 5.22-5.48; leg IV femur 9.88-10.81, patella 1.57-1.59, tibia7.53-7.38. Femora evenly but irregularly denticulate, except distal third of femur II unarmed; remaining segments unarmed. Legs medium brown mottled with yellowish, tibiae and tarsi tinged with purple, tibiae spotted with white; tarsi with white band at base of telotarsi. Tibia II with five pseudosegments; tibia IV undivided in larger specimen, with two pseudosegments in smaller.</paragraph>
<paragraph id="EC470855742A8F4ABA658A9C3FF20762" pageId="10" pageNumber="69">
Penis (Fig. 4
<normalizedToken id="FF0D2F9FB5E6D32122065D4ACBB9A733" originalValue="e–f">e-f</normalizedToken>
): Glans medium length, sides parabolic in ventral view; subtriangular in lateral view but not markedly flattened, slight dorsomedial bulge but keel essentially absent. Bristle groups short. Tendon long.
</paragraph>
<caption id="35358857344998E51BC3398B7E419176" pageId="10" pageNumber="69">
<paragraph id="D18F7D11EE34D6784D78EBA3C31E0C87" pageId="10" pageNumber="69">
Figure 4.
<taxonomicName id="A0D9925A66FA2DEE7734B6EFF6F57DFE" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis cheliferoides" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="cheliferoides">Pantopsalis cheliferoides</taxonomicName>
, specimen collected in 1975. A Dorsal view B lateral view C fingers of left chelicera, anterior view D patella and tibia of right pedipalp, dorsal view E glans, ventral view F glans, left lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="EDE1396025B4CC94CA9D3FCCE1016A37" lastPageId="12" lastPageNumber="71" pageId="10" pageNumber="69" type="comments">
<paragraph id="66ECDAAB27A202999DB338B1A735D067" pageId="10" pageNumber="69">Comments.</paragraph>
<paragraph id="A784D2F70D3742A61960B35471AE391E" pageId="10" pageNumber="69">
There is some variation in coloration between the two specimens available, most notably the presence of a medial stripe on the opisthosoma of one but not the other, with the former specimen also being overall lighter in coloration than the latter. It is possible that this difference may reflect differences in maturity between the two specimens, similar to what has been recorded for other
<taxonomicName id="290E97E0E5187284F13D4023F686A45E" genus="Opiliones" lsidName="Opiliones" pageId="10" pageNumber="69" rank="genus">Opiliones</taxonomicName>
species (
<bibRefCitation id="AF05A163FA15F9E0D98F190BF328643B" author="Shultz, JW" journalOrPublisher="Journal of Arachnology" pageId="13" pageNumber="72" pagination="184 - 186" title="Leiobunum nigripes is a junior synonym of Leiobunum verrucosum (Opiliones, Sclerosomatidae)." url="10.1636/H07-48SC.1" volume="36" year="2008">Shultz 2008</bibRefCitation>
). Such long-term darkening after moulting may also explain the anomalously pale specimens of
<taxonomicName id="83EEF53249103C15B073D49CD8103120" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis albipalpis" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="albipalpis">Pantopsalis albipalpis</taxonomicName>
recorded by
<bibRefCitation id="1C0470DF83DB6A1A8F435E742C060499" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor (2004)</bibRefCitation>
.
</paragraph>
<paragraph id="F2902FD7C60AD9753929320D6BC787B7" pageId="10" pageNumber="69">
Long regarded as something of a mystery after its initial description by
<bibRefCitation id="B6FCAA3260B54D4D9BEC14D66B242209" author="Colenso, W" journalOrPublisher="Transactions and Proceedings of the New Zealand Institute" pageId="12" pageNumber="71" pagination="165 - 73" title="On some newly-discovered New Zealand arachnids." volume="15" year="1882">Colenso (1882)</bibRefCitation>
,
<taxonomicName id="1E98695C6237DE8A57EDCCA07C95F62A" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis cheliferoides" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="cheliferoides">Pantopsalis cheliferoides</taxonomicName>
was referred to only in footnotes by
<bibRefCitation id="F00F6F89BED7700C845A8F60E44458DD" author="Forster, RR" journalOrPublisher="Records of the Dominion Museum" pageId="12" pageNumber="71" pagination="183 - 192" title="The genus Megalopsalis Roewer in New Zealand with keys to the New Zealand genera of Opiliones." volume="1" year="1944">Forster (1944)</bibRefCitation>
and
<bibRefCitation id="1D95AF023F69DBF22493294B5B848FC2" author="Marples, BJ" journalOrPublisher="Transactions of the Royal Society of New Zealand" pageId="13" pageNumber="72" pagination="313 - 14" title="A new species of harvestman of the genus Megalopsalis." volume="73" year="1944">Marples (1944)</bibRefCitation>
, and treated as a nomen dubium by
<bibRefCitation id="02CB2E4C9AE746FB03C5163AA9C2E4A1" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor (2004)</bibRefCitation>
due to the poor condition of the type specimen. This was unfortunate as, with the recognition of
<normalizedToken id="4C52FCD66FA9B6D72908558C0960EEAA" originalValue="‘Pantopsalis’">'Pantopsalis'</normalizedToken>
wattsi
<bibRefCitation id="1B4CB6145FF746F9EBBB0B7097E95A3F" pageId="10" pageNumber="69">Hogg 1920</bibRefCitation>
as a species of
<taxonomicName id="C6FB25FF936AC9506BDCAEA6C3378FD5" class="Arachnida" family="Neopilionidae" genus="Forsteropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Forsteropsalis" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="genus">Forsteropsalis</taxonomicName>
(
<bibRefCitation id="F0FDCD4328ABE698A5637777148DA00F" pageId="10" pageNumber="69">Taylor 2011</bibRefCitation>
),
<taxonomicName id="AE6B6418B9291046D1CAF816E12608A8" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis cheliferoides" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="cheliferoides">Pantopsalis cheliferoides</taxonomicName>
was the only species of
<taxonomicName id="29B382FE1FAB456D78670A63F9D5E790" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
described from the North Island. The two specimens examined here, though collected some distance from the type locality, are consistent with
<normalizedToken id="206812683191ED32F02CD12DC69D4C89" originalValue="Colenso’s">Colenso's</normalizedToken>
(1882) original description and discernable features of the type specimen (
<bibRefCitation id="6FB5B4D7DCA03B9988419E472DA41AB1" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
) and, in the absence of evidence to the contrary, can be identified as
<taxonomicName id="F38C80D06861A8B4CB13267004F30F41" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis cheliferoides" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="cheliferoides">Pantopsalis cheliferoides</taxonomicName>
. This species can, therefore, be confirmed as distinct from other identifiable species of
<taxonomicName id="DE0D0925D8641B046688E05ABA054DAA" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
.
</paragraph>
<paragraph id="7DFF1BB67C690C4160A874AEB4060F03" lastPageId="11" lastPageNumber="70" pageId="10" pageNumber="69">
The absence of a distinct dorsal keel on the glans of the penis clearly distinguishes
<taxonomicName id="6E04D16C58911CB65C7FACF209826927" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis cheliferoides" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="cheliferoides">Pantopsalis cheliferoides</taxonomicName>
from all other
<taxonomicName id="452E197BA176A37D03EB3B95CF4DDE50" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
species except
<taxonomicName id="E75ACD6F9F2E00CDD3AED34838E65D56" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis luna" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="luna">Pantopsalis luna</taxonomicName>
and possibly
<taxonomicName id="CA292B129C425F5C3EA80AA8CF4AFF75" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis pococki" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="pococki">Pantopsalis pococki</taxonomicName>
(for which the genital morphology remains unknown).
<taxonomicName id="C3C2D126C59ED1D35D4BC949433910B2" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis pococki" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="pococki">Pantopsalis pococki</taxonomicName>
has a very distinct colour pattern, with transverse light coloured stripes on the dorsum of the opisthosoma (
<bibRefCitation id="38399BEF2071F95A9E423B079FF32FCB" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
).
<taxonomicName id="BAB1FE98FB05A2342D4986B1E36792C2" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis luna" order="Opiliones" pageId="10" pageNumber="69" phylum="Arthropoda" rank="species" species="luna">Pantopsalis luna</taxonomicName>
lacks a ventral tooth on the pedipalpal tarsal claw, has the ocularium unarmed with denticulation on the prosoma restricted
<pageBreakToken id="772FB6627D830373053CC50C40D5BCF4" pageId="11" pageNumber="70" start="start">to</pageBreakToken>
the anterior propeltidial region, and has narrow transverse stripes on the dorsum of the opisthosoma (personal observations, male specimens from Waiho Gorge, South Island, cited by
<bibRefCitation id="FA24F4B33FB58EBBE25BF52192E933D0" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
).
</paragraph>
<paragraph id="2AA85CF008B57D1E89C0EAE6C242FF14" pageId="11" pageNumber="70">
The presence of dimorphic males as described for other
<taxonomicName id="8C6B732B1D9586091B05753B257501A8" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
species by
<bibRefCitation id="1E5A58AEC3F46D46612B38331F7C7808" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor (2004)</bibRefCitation>
in both
<taxonomicName id="6AF4F34EDD8F551357A3F8BD81A22949" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
and
<taxonomicName id="9F5DDC9AC1C001485899569675DA17A5" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis cheliferoides" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="cheliferoides">Pantopsalis cheliferoides</taxonomicName>
is of note. Previous species in which this phenomenon has been observed (
<taxonomicName id="BE15B5DFB9F57899A9703199F694BDA8" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis albipalpis" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="albipalpis">Pantopsalis albipalpis</taxonomicName>
,
<taxonomicName id="4276BDF8C619B0E65A6B375818F824EC" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis coronata" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="coronata">Pantopsalis coronata</taxonomicName>
,
<taxonomicName id="C9D4C5F78DF033A1AE955613AEA76366" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis johnsi" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="johnsi">Pantopsalis johnsi</taxonomicName>
and
<taxonomicName id="9761E1A5A469E74880D411FB03DD4C4A" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis phocator" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="phocator">Pantopsalis phocator</taxonomicName>
) all belong to the well-marked species group whose members possess a strong dorsal keel on the glans (
<bibRefCitation id="07B570D381E66AF05B32F9D3BF186188" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
) and it was previously unknown whether such male dimorphism occurred outside this species group. Its presence in the two species treated herein indicates that it does, and it may indeed be a synapomorphy for
<taxonomicName id="49195E7ADA6FB0EF3560B7D9272652CF" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
as a whole.
</paragraph>
<subSection id="ED7EB2BA34E7C5860F657C0A54B09736" lastPageId="12" lastPageNumber="71" pageId="11" pageNumber="70" type="key to males of pantopsalis">
<paragraph id="2B26D4A0E826EEB6DD84BCBA1431EE68" pageId="11" pageNumber="70">
Key to males of
<taxonomicName id="E10718CEE31592167A578A807D6BD671" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
</paragraph>
<paragraph id="B3E179EAE4B18DC9941FF47385464318" pageId="11" pageNumber="70">
The last author to provide a key to species of
<taxonomicName id="D90197EB090EF182CA4439BA0F2723CF" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
was
<bibRefCitation id="63C45B33D045449D700DC57A42C08536" pageId="11" pageNumber="70">Roewer (1923)</bibRefCitation>
. Since then, a number of species have been described, several of the species referred to by Roewer have been synonymised, and characters of the chelicerae referred to by Roewer have been shown to vary within species (
<bibRefCitation id="2AC0D881045AC651A98832092B495B33" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
). Therefore, the opportunity is taken to provide an updated key to
<taxonomicName id="99B052B5231454ECD7ABD085E53940A1" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
males.
<taxonomicName id="0506776930CDE92E88A471ADDF6AB98B" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis halli" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="halli">Pantopsalis halli</taxonomicName>
<bibRefCitation id="543A33611932D2B061636D21F4B5C501" pageId="11" pageNumber="70">Hogg 1920</bibRefCitation>
is omitted from the following key, as it is currently based only on a female specimen and not identifiable (
<bibRefCitation id="17CCBBC702F92262B54E23E86301CB84" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
).
<taxonomicName id="5FF3A9403B87F01CC1367B50E4C736FB" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis albipalpis" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="albipalpis">Pantopsalis albipalpis</taxonomicName>
and
<taxonomicName id="E90B41DB9F936847BE750D1E720A4149" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis johnsi" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="johnsi">Pantopsalis johnsi</taxonomicName>
are not currently distinguishable, but
<bibRefCitation id="72B04EC6A5B2CE9343470A1B6DC1C66D" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor (2004)</bibRefCitation>
refrained from synonymising them on the basis of their widely disjunct distributions. Their relationship requires further investigation.
</paragraph>
<paragraph id="568C1AE36FE0E2CAA5510BE944B5273C" lastPageId="12" lastPageNumber="71" pageId="11" pageNumber="70">
<table id="B3DB772E88F654A953625335C2D989AA" lastPageId="12" lastPageNumber="71" pageId="11" pageNumber="70">
<tr id="31B42DD66C268916918A8B9025577DB2" pageId="11" pageNumber="70">
<td id="7A1A45C40CE985050D97E1689B9C599C" colspan="1" pageId="11" pageNumber="70" rowspan="1">
<taxonomicName id="C3A2EA96A889F2B3F38108F854BB2B6B" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis phocator" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="phocator">Pantopsalis phocator</taxonomicName>
<bibRefCitation id="A8C787A410B2D9B0BC27A85948261065" author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
</td>
</tr>
<tr id="D97A7B1EA3CA7D75734F3CAC5BEC71DC" pageId="11" pageNumber="70">
<td id="A3886F3ECA1A26D09FC9FA92433F4088" colspan="1" pageId="11" pageNumber="70" rowspan="1">
<taxonomicName id="B7C9A14C46E06465CAD202FC3B5EF56B" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis pococki" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="pococki">Pantopsalis pococki</taxonomicName>
<bibRefCitation id="C0F033784265AF46DBA5A9502C03E55B" pageId="11" pageNumber="70">Hogg 1920</bibRefCitation>
</td>
</tr>
<tr id="A11FE68731F7A09BBC43C7DDF03BB40F" pageId="11" pageNumber="70">
<td id="6B1ED85102E22560FC6A5FA8CCB9F3C9" colspan="1" pageId="11" pageNumber="70" rowspan="1">
<taxonomicName id="6C58241A2D5A5D3867C5008758933E91" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis coronata" order="Opiliones" pageId="11" pageNumber="70" phylum="Arthropoda" rank="species" species="coronata">Pantopsalis coronata</taxonomicName>
<bibRefCitation id="DC7E81B894541151C44EB69C5D815606" pageId="11" pageNumber="70">Pocock 1903b</bibRefCitation>
</td>
</tr>
<tr id="07E74090575EF0602DE87A10748116C2" pageId="12" pageNumber="71">
<td id="099B3B66FB5EFCABC4215E3863FB4C4F" colspan="1" pageId="12" pageNumber="71" rowspan="1">
<taxonomicName id="6A1EA2078F4ED9329F39FD018453B2B7" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis albipalpis" order="Opiliones" pageId="12" pageNumber="71" phylum="Arthropoda" rank="species" species="albipalpis">
<pageBreakToken id="37E4B5BAB1EAE2DC414FDBADAFF2B2A0" pageId="12" pageNumber="71" start="start">Pantopsalis</pageBreakToken>
albipalpis
</taxonomicName>
<bibRefCitation id="6E7CD5939D4F42C7F8CAC5F483BCA502" author="Pocock, RI" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="13" pageNumber="72" pagination="392 - 413" title="On some new harvest-spiders of the order Opiliones from the southern continents." volume="1902" year="1903 a">Pocock 1903a</bibRefCitation>
<taxonomicName id="C17777909D9466A4E8F15C66F506AB6F" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis johnsi" order="Opiliones" pageId="12" pageNumber="71" phylum="Arthropoda" rank="species" species="johnsi">Pantopsalis johnsi</taxonomicName>
<bibRefCitation id="1FF3EB9097508C27ADBC532F81F669ED" author="Forster, RR" journalOrPublisher="Pacific Insects Monograph" pageId="12" pageNumber="71" pagination="58 - 115" title="The Araneae and Opiliones of the subantarctic islands of New Zealand." volume="7" year="1964">Forster 1964</bibRefCitation>
</td>
</tr>
<tr id="85EC051D83194C91CFD31E2781592F67" pageId="12" pageNumber="71">
<td id="F033090302F255457DC8073B56F159B1" colspan="1" pageId="12" pageNumber="71" rowspan="1">
<taxonomicName id="881F7BC876B7BA793F7EEC2491320221" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="12" pageNumber="71" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
<bibRefCitation id="5BF4ADED347090F898CB207D35793FB7" pageId="12" pageNumber="71">White 1849</bibRefCitation>
</td>
</tr>
<tr id="38F82307C9A1BBF5D045311DBFAF8BDF" pageId="12" pageNumber="71">
<td id="7C4230AC6A8C1BF74F4E85D59A6616C8" colspan="1" pageId="12" pageNumber="71" rowspan="1">
<taxonomicName id="EF397234F4648BD7504B1E7614681845" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis cheliferoides" order="Opiliones" pageId="12" pageNumber="71" phylum="Arthropoda" rank="species" species="cheliferoides">Pantopsalis cheliferoides</taxonomicName>
<bibRefCitation id="04C6EE4F277758D2D013E83418BD6A4F" author="Colenso, W" journalOrPublisher="Transactions and Proceedings of the New Zealand Institute" pageId="12" pageNumber="71" pagination="165 - 73" title="On some newly-discovered New Zealand arachnids." volume="15" year="1882">Colenso 1882</bibRefCitation>
</td>
</tr>
<tr id="0C420B6BFE4F22ABF79068532E1EFA13" pageId="12" pageNumber="71">
<td id="2F0DAFF446BD50EB729AB93682E4DC92" colspan="1" pageId="12" pageNumber="71" rowspan="1">
<taxonomicName id="4DE12B62E1E38A2968FA16B20A6413DE" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis rennelli" order="Opiliones" pageId="12" pageNumber="71" phylum="Arthropoda" rank="species" species="rennelli">Pantopsalis rennelli</taxonomicName>
<bibRefCitation id="12D5C520502CDA710EAF76BFF0C551E6" author="Forster, RR" journalOrPublisher="Pacific Insects Monograph" pageId="12" pageNumber="71" pagination="58 - 115" title="The Araneae and Opiliones of the subantarctic islands of New Zealand." volume="7" year="1964">Forster 1964</bibRefCitation>
</td>
</tr>
<tr id="CD180E436E3875B4005EEB80EA4545D0" pageId="12" pageNumber="71">
<td id="EB59531987C0435AE0F5CB9322F940F1" colspan="1" pageId="12" pageNumber="71" rowspan="1">
<taxonomicName id="29EBC98825699FD5ED8BDDBDD3F64057" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis snaresensis" order="Opiliones" pageId="12" pageNumber="71" phylum="Arthropoda" rank="species" species="snaresensis">Pantopsalis snaresensis</taxonomicName>
<bibRefCitation id="C99659423A8E015D7C6B04EDC2F7A7DF" author="Forster, RR" journalOrPublisher="Pacific Insects Monograph" pageId="12" pageNumber="71" pagination="58 - 115" title="The Araneae and Opiliones of the subantarctic islands of New Zealand." volume="7" year="1964">Forster 1964</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
</subSection>
</subSubSection>
</treatment>
</document>
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