402 lines
50 KiB
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402 lines
50 KiB
XML
<document id="D3CB29C9AEB9A0090642219399567909" ID-CLB-Dataset="132550" ID-DOI="10.1649/0010-065X-72.2.279" ID-GBIF-Dataset="3c3afcfd-63cc-4cec-89f3-efc5fa42a87e" ID-ISSN="1938-4394" ID-Zenodo-Dep="5381960" ID-ZooBank="urn:lsid:zoobank.org:pub:38A07005-9DF6-4468-BBC3-BD16FE6851B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630294548391" checkinUser="diego" docAuthor="Chatzimanolis, Stylianos" docDate="2018" docId="296D87B5FFED2406FF38F981FBD6FB4E" docLanguage="en" docName="TheColeopBull.72.2.279-291.pdf.imf" docOrigin="The Coleopterists Bulletin 72 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-72.2.279" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Dysanellus BERNHAUER AND 1911" docType="treatment" docVersion="7" lastPageNumber="282" masterDocId="D554FFCDFFEF2402FFC4FFCCFFEDFFC3" masterDocTitle="A Review of the Genera Dysanellus Bernhauer and Torobus Herman (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)" masterLastPageNumber="291" masterPageNumber="279" pageNumber="280" updateTime="1699240502839" updateUser="ExternalLinkService">
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<mods:title id="20CC4CD95C39F0B4847DEE3DAB62FF84">A Review of the Genera Dysanellus Bernhauer and Torobus Herman (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)</mods:title>
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<mods:namePart id="2891068179BF61EDE9468B04DA4634F6">Chatzimanolis, Stylianos</mods:namePart>
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<mods:title id="C262555923322AF546442B99CAF6C3CD">The Coleopterists Bulletin</mods:title>
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<mods:part id="BFAE6E8A0766C8A5F93E1C3755CC9DCD">
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<mods:date id="7E8E1D04AB3A0BF92CA38A3046A9F840">2018</mods:date>
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<mods:number id="1CA03248A64FFE14361FA6CB360EEB32">2018-06-20</mods:number>
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<mods:number id="F704BD27725FD9D3DBCD78CBAE364B95">72</mods:number>
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<mods:identifier id="EDA9DD1882328E3092C2918BC38B4FD6" type="DOI">10.1649/0010-065X-72.2.279</mods:identifier>
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<treatment id="296D87B5FFED2406FF38F981FBD6FB4E" ID-DOI="http://doi.org/10.5281/zenodo.7037275" ID-GBIF-Taxon="199882119" ID-Zenodo-Dep="7037275" LSID="urn:lsid:plazi:treatment:296D87B5FFED2406FF38F981FBD6FB4E" httpUri="http://treatment.plazi.org/id/296D87B5FFED2406FF38F981FBD6FB4E" lastPageId="4" lastPageNumber="282" pageId="2" pageNumber="280">
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<subSubSection id="E9DE6528FFED2400FF38F981FDF4F9A1" box="[252,537,1613,1634]" pageId="2" pageNumber="280" type="nomenclature">
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<paragraph id="A17B36A3FFED2400FF38F981FDF4F9A1" blockId="2.[213,576,1613,1661]" box="[252,537,1613,1634]" pageId="2" pageNumber="280">
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<heading id="FA3381CFFFED2400FF38F981FDF4F9A1" box="[252,537,1613,1634]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="280" reason="2">
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<taxonomicName id="66C44D20FFED2400FF38F981FDF4F9A1" ID-CoL="8XL36" authority="Bernhauer, 1911" authorityName="BERNHAUER AND" authorityYear="1911" box="[252,537,1613,1634]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="280" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0EAB1FFED2400FF38F981FDF4F9A1" bold="true" box="[252,537,1613,1634]" pageId="2" pageNumber="280">
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<emphasis id="93B0EAB1FFED2400FF38F981FE84F9A1" bold="true" box="[252,361,1613,1634]" italics="true" pageId="2" pageNumber="280">Dysanellus</emphasis>
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<bibRefCitation id="C5554B52FFED2400FEB5F981FDF4F9A1" author="Bernhauer" box="[369,537,1613,1634]" firstAuthor="Bernhauer" pageId="2" pageNumber="280" pagination="403 - 422" refId="ref8254" refString="Bernhauer, M. 1911. Zur Staphylinidenfauna von Sud- Amerika (Col.). Deutsche Entomologische Zeitschrift 1911: 403 - 422." type="journal article" year="1911">Bernhauer, 1911</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9DE6528FFED2400FF11F9A4FDADF9BE" box="[213,576,1639,1661]" pageId="2" pageNumber="280" type="description">
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<paragraph id="A17B36A3FFED2400FF11F9A4FDADF9BE" blockId="2.[213,576,1613,1661]" box="[213,576,1639,1661]" pageId="2" pageNumber="280">
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(
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<figureCitation id="39FF2A26FFED2400FF18F9A4FED4F9BE" box="[220,313,1639,1661]" captionStart="Figs" captionStartId="2.[709,748,854,873]" captionTargetBox="[693,1163,195,833]" captionTargetId="figure-504@2.[688,1170,194,838]" captionTargetPageId="2" captionText="Figs. 1–2. Dysanellus species, habitus. 1) D. bruchi, total length = 15.2 mm; 2) D. transverserugosus, total length = 17.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5381962" httpUri="https://zenodo.org/record/5381962/files/figure.png" pageId="2" pageNumber="280">Figs. 1–2</figureCitation>
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,
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<figureCitation id="39FF2A26FFED2400FE80F9A4FE87F9BE" box="[324,362,1639,1661]" captionStart="Figs" captionStartId="4.[175,214,992,1011]" captionTargetBox="[154,1169,194,976]" captionTargetId="figure-298@4.[154,1169,194,976]" captionTargetPageId="4" captionText="Figs. 5–6. Heads and pronota of Dysanellus species. 5) D. bruchi; 6) D. transverserugosus. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5381968" httpUri="https://zenodo.org/record/5381968/files/figure.png" pageId="2" pageNumber="280">5–6</figureCitation>
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,
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<figureCitation id="39FF2A26FFED2400FEB2F9A4FE69F9BE" box="[374,388,1640,1661]" captionStart="Figs" captionStartId="6.[175,214,861,880]" captionTargetBox="[154,1169,194,845]" captionTargetId="figure-440@6.[154,1169,194,845]" captionTargetPageId="6" captionText="Figs. 9–10. Head, ventral view. 9. Dysanellus transverserugosus; 10) Torobus laetipes. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5381972" httpUri="https://zenodo.org/record/5381972/files/figure.png" pageId="2" pageNumber="280">9</figureCitation>
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,
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<figureCitation id="39FF2A26FFED2400FE4BF9A4FE21F9BE" box="[399,460,1639,1661]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="7.[197,236,423,442]" captionStartId-1="7.[197,236,1663,1682]" captionTargetBox-0="[177,655,196,405]" captionTargetBox-1="[183,1188,1276,1644]" captionTargetId-0="figure-446@7.[175,657,194,407]" captionTargetId-1="figure-474@7.[175,1190,1274,1646]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="Figs. 11–12. Distal end of labial palpomere 3 in Dysanellus species. 11) D. bruchi; 12) D. transverserugosus. Scale bar = 50 µm." captionText-1="Figs. 13–15. Abdominal sterna VII and VIII. Arrows indicate the position of porose structure. 13) Dysanellus transverserugosus (While there is a broad area with no setae on VII, it is not clear if there is a porose structure. See text for more details); 14) Torobus laetipes; 15) Torobus purpurascens. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.5381976" figureDoi-1="http://doi.org/10.5281/zenodo.5381978" httpUri-0="https://zenodo.org/record/5381976/files/figure.png" httpUri-1="https://zenodo.org/record/5381978/files/figure.png" pageId="2" pageNumber="280">11–13</figureCitation>
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,
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<figureCitation id="39FF2A26FFED2400FE13F9A4FDF8F9BE" box="[471,533,1639,1661]" captionStart="Figs" captionStartId="8.[175,214,594,613]" captionTargetBox="[155,637,197,570]" captionTargetId="figure-266@8.[151,638,194,579]" captionTargetPageId="8" captionText="Figs. 16–18. Aedeagus of Dysanellus transverserugosus. 16) Dorsal view; 17) Lateral view; 18) Paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5381980" httpUri="https://zenodo.org/record/5381980/files/figure.png" pageId="2" pageNumber="280">16–18</figureCitation>
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,
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<figureCitation id="39FF2A26FFED2400FDE5F9A4FDD7F9BE" box="[545,570,1640,1661]" captionStart="Fig" captionStartId="8.[709,740,1687,1706]" captionTargetBox="[689,1168,1309,1669]" captionTargetId="figure-328@8.[688,1170,1307,1670]" captionTargetPageId="8" captionText="Fig. 25. Distribution map of Dysanellus bruchi and Dysanellus transverserugosus." figureDoi="http://doi.org/10.5281/zenodo.7033285" httpUri="https://zenodo.org/record/7033285/files/figure.png" pageId="2" pageNumber="280">25</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="E9DE6528FFED2400FF6BF95EFE42F901" pageId="2" pageNumber="280" type="type_taxon">
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<paragraph id="A17B36A3FFED2400FF6BF95EFE42F901" blockId="2.[151,638,1682,1730]" pageId="2" pageNumber="280">
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<emphasis id="93B0EAB1FFED2400FF6BF95EFEAFF964" bold="true" box="[175,322,1682,1703]" pageId="2" pageNumber="280">
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<typeStatus id="7E7F8801FFED2400FF6BF95EFF0CF964" box="[175,225,1682,1703]" pageId="2" pageNumber="280">Type</typeStatus>
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Species.
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</emphasis>
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<taxonomicName id="66C44D20FFED2400FE90F95EFDE1F964" ID-CoL="6DPL3" authorityName="Bernhauer" authorityYear="1911" box="[340,524,1682,1703]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="280" phylum="Arthropoda" rank="species" species="bruchi">
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<emphasis id="93B0EAB1FFED2400FE90F95EFDE1F964" box="[340,524,1682,1703]" italics="true" pageId="2" pageNumber="280">Dysanellus bruchi</emphasis>
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</taxonomicName>
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, fixed by monotypy (
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<bibRefCitation id="C5554B52FFED2400FEC2F961FE49F901" author="Bernhauer" box="[262,420,1709,1730]" firstAuthor="Bernhauer" pageId="2" pageNumber="280" pagination="403 - 422" refId="ref8254" refString="Bernhauer, M. 1911. Zur Staphylinidenfauna von Sud- Amerika (Col.). Deutsche Entomologische Zeitschrift 1911: 403 - 422." type="journal article" year="1911">Bernhauer 1911</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="E9DE6528FFED2400FD01FC2CFBB6FAF0" pageId="2" pageNumber="280" type="diagnosis">
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||
<paragraph id="A17B36A3FFED2400FD01FC2CFBB6FAF0" blockId="2.[685,1173,992,1730]" pageId="2" pageNumber="280">
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<emphasis id="93B0EAB1FFED2400FD01FC2CFCC0FC36" bold="true" box="[709,813,992,1013]" pageId="2" pageNumber="280">Diagnosis.</emphasis>
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The following character statess in combination can distinguish
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<taxonomicName id="66C44D20FFED2400FC0BFC36FBD5FBCC" authorityName="BERNHAUER AND" authorityYear="1911" box="[975,1080,1018,1039]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="280" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0EAB1FFED2400FC0BFC36FBD5FBCC" box="[975,1080,1018,1039]" italics="true" pageId="2" pageNumber="280">Dysanellus</emphasis>
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</taxonomicName>
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from all other genera in
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<taxonomicName id="66C44D20FFED2400FC9CFBD9FC0FFBE9" authorityName="Sharp" authorityYear="1884" box="[856,994,1045,1066]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="280" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
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: head transverse; antennomeres 1–7, 11 longer than wide; antennomeres 8–10 not transverse; mandibles long; labial palpomere 3 securiform; pronotum with several rows of punctures; and postcoxal process present. Species in
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<taxonomicName id="66C44D20FFED2400FCD7FB56FC5DFB6C" authority="Sharp" authorityName="Sharp" authorityYear="1884" box="[787,944,1178,1199]" class="Insecta" family="Staphylinidae" genus="Ocyolinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="280" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0EAB1FFED2400FCD7FB56FC9EFB6C" box="[787,883,1178,1199]" italics="true" pageId="2" pageNumber="280">Ocyolinus</emphasis>
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Sharp
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</taxonomicName>
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might be confused with
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<taxonomicName id="66C44D20FFED2400FD69FB78FCFBFB0A" authorityName="BERNHAUER AND" authorityYear="1911" box="[685,790,1204,1225]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="280" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0EAB1FFED2400FD69FB78FCFBFB0A" box="[685,790,1204,1225]" italics="true" pageId="2" pageNumber="280">Dysanellus</emphasis>
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</taxonomicName>
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, but the two genera can be easily distinguished by the shape of labial palpomere 3 (securiform in
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<taxonomicName id="66C44D20FFED2400FC85FB25FC47FB3D" authorityName="BERNHAUER AND" authorityYear="1911" box="[833,938,1257,1278]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="280" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0EAB1FFED2400FC85FB25FC47FB3D" box="[833,938,1257,1278]" italics="true" pageId="2" pageNumber="280">Dysanellus</emphasis>
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</taxonomicName>
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, dilated but not securiform in
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<taxonomicName id="66C44D20FFED2400FCC1FAC8FC88FADA" authorityName="Sharp" authorityYear="1884" box="[773,869,1284,1305]" class="Insecta" family="Staphylinidae" genus="Ocyolinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="280" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0EAB1FFED2400FCC1FAC8FC88FADA" box="[773,869,1284,1305]" italics="true" pageId="2" pageNumber="280">Ocyolinus</emphasis>
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||
</taxonomicName>
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) and presence of a postcoxal process in
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<taxonomicName id="66C44D20FFED2400FCD2FAD2FC92FAF0" authorityName="BERNHAUER AND" authorityYear="1911" box="[790,895,1310,1331]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="280" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0EAB1FFED2400FCD2FAD2FC92FAF0" box="[790,895,1310,1331]" italics="true" pageId="2" pageNumber="280">Dysanellus</emphasis>
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</taxonomicName>
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(absent in
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<taxonomicName id="66C44D20FFED2400FC29FAD2FBA0FAF0" authorityName="Sharp" authorityYear="1884" box="[1005,1101,1310,1331]" class="Insecta" family="Staphylinidae" genus="Ocyolinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="280" phylum="Arthropoda" rank="genus">
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||
<emphasis id="93B0EAB1FFED2400FC29FAD2FBA0FAF0" box="[1005,1101,1310,1331]" italics="true" pageId="2" pageNumber="280">Ocyolinus</emphasis>
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</taxonomicName>
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).
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9DE6528FFED2406FD01FAF5FE04FB4D" lastPageId="4" lastPageNumber="282" pageId="2" pageNumber="280" type="description">
|
||
<paragraph id="A17B36A3FFED2400FD01FAF5FC07F9FE" blockId="2.[685,1173,992,1730]" pageId="2" pageNumber="280">
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||
<emphasis id="93B0EAB1FFED2400FD01FAF5FCADFA8D" bold="true" box="[709,832,1337,1358]" pageId="2" pageNumber="280">Description.</emphasis>
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Habitus as in
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<figureCitation id="39FF2A26FFED2400FC0BFAF5FBC1FA8D" box="[975,1068,1336,1358]" captionStart="Figs" captionStartId="2.[709,748,854,873]" captionTargetBox="[693,1163,195,833]" captionTargetId="figure-504@2.[688,1170,194,838]" captionTargetPageId="2" captionText="Figs. 1–2. Dysanellus species, habitus. 1) D. bruchi, total length = 15.2 mm; 2) D. transverserugosus, total length = 17.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5381962" httpUri="https://zenodo.org/record/5381962/files/figure.png" pageId="2" pageNumber="280">Figs. 1–2</figureCitation>
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, body medium to large size,
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<quantity id="663C9B46FFED2400FCAAFA98FC16FAAA" box="[878,1019,1363,1385]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" metricValueMax="1.76" metricValueMin="1.3599999999999999" pageId="2" pageNumber="280" unit="mm" value="15.6" valueMax="17.6" valueMin="13.6">13.6–17.6 mm</quantity>
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||
in total length. Color of head and pronotum dark metallic brown with purple overtones or metallic green-blue; elytra metallic golden brown or metallic green-blue; mouthparts, mesoscutellum, ventral surface of body, and legs brown; proximal antennomeres brown, distal antennomeres dark orange; abdomen brown with segment VII orange-brown or brown and segment VIII orange-brown.
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</paragraph>
|
||
<paragraph id="A17B36A3FFED2401FD01F98FFEAAFA8D" blockId="2.[685,1173,992,1730]" lastBlockId="3.[173,660,1071,1730]" lastPageId="3" lastPageNumber="281" pageId="2" pageNumber="280">Head transverse, with medium-sized to large, setose punctures and microsculpture around margin of head; with various punctures and sculpture on epicranium. Clypeus slightly emarginate; anteclypeus not expanded. Eyes small to medium size, occupying 1/2 to 1/3 of lateral margins of head. Ventral surface of head with transverse microsculpture and scattered medium-sized to large punctures; postoccipital suture and ventral basal ridge present; infraorbital ridge short but well delineated; postmandibular ridge present, prominent, extending from near mandible to lateral side of head; gular sutures separated anteriorly, converging medially; nuchal depression prominent, forming well-defined neck; neck with microsculpture and micropunctures.</paragraph>
|
||
<caption id="F5BB662BFFEC2401FF01FC7EFEB9FC36" ID-DOI="http://doi.org/10.5281/zenodo.5381966" ID-Zenodo-Dep="5381966" httpUri="https://zenodo.org/record/5381966/files/figure.png" pageId="3" pageNumber="281" startId="3.[197,236,946,965]" targetBox="[178,635,196,928]" targetPageId="3">
|
||
<paragraph id="A17B36A3FFEC2401FF01FC7EFEB9FC36" blockId="3.[173,659,946,1013]" pageId="3" pageNumber="281">
|
||
<emphasis id="93B0EAB1FFEC2401FF01FC7EFEF3FC06" bold="true" box="[197,286,946,965]" pageId="3" pageNumber="281">Figs. 3–4.</emphasis>
|
||
<taxonomicName id="66C44D20FFEC2401FEF7FC7EFE2DFC06" box="[307,448,946,965]" class="Insecta" family="Staphylinidae" genus="Torobus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="281" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="93B0EAB1FFEC2401FEF7FC7EFE9AFC06" box="[307,375,946,965]" italics="true" pageId="3" pageNumber="281">Torobus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, habitus.
|
||
<emphasis id="93B0EAB1FFEC2401FDDCFC7EFDC4FC06" bold="true" box="[536,553,946,965]" pageId="3" pageNumber="281">3)</emphasis>
|
||
<taxonomicName id="66C44D20FFEC2401FDF7FC7EFD60FC06" baseAuthorityName="Bernhauer" baseAuthorityYear="1911" box="[563,653,946,965]" class="Insecta" family="Staphylinidae" genus="Torobus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="281" phylum="Arthropoda" rank="species" species="laetipes">
|
||
<emphasis id="93B0EAB1FFEC2401FDF7FC7EFD60FC06" box="[563,653,946,965]" italics="true" pageId="3" pageNumber="281">T. laetipes</emphasis>
|
||
</taxonomicName>
|
||
, total length = 17.2 mm;
|
||
<emphasis id="93B0EAB1FFEC2401FE6EFC06FE56FC1E" bold="true" box="[426,443,970,989]" pageId="3" pageNumber="281">4)</emphasis>
|
||
<taxonomicName id="66C44D20FFEC2401FE0EFC06FDB6FC1E" authorityName="Nordmann" authorityYear="1837" box="[458,603,970,989]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="281" phylum="Arthropoda" rank="species" species="purpurascens">
|
||
<emphasis id="93B0EAB1FFEC2401FE0EFC06FDB6FC1E" box="[458,603,970,989]" italics="true" pageId="3" pageNumber="281">T. purpurascens</emphasis>
|
||
</taxonomicName>
|
||
, total length = 18.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A17B36A3FFEC2401FF01FA98FDF7F9FE" blockId="3.[173,660,1071,1730]" pageId="3" pageNumber="281">Antennomeres 1–3 with multiple rows of macrosetae; antennomeres 4 (distally) and 5–11 with macrosetae but also with microtrichiae; antennomeres 1–7 and 11 longer than wide; antennomeres 8–10 longer than wide or subquadrate; antennomere 1 twice as long as antennomere 2; antennomere 3 longer than 2; antennomeres 4–7 subequal in size; antennomeres 8–10 becoming gradually shorter in length, shorter than antennomeres 4–7.</paragraph>
|
||
<paragraph id="A17B36A3FFEC2401FF01F98FFC78FE52" blockId="3.[173,660,1071,1730]" lastBlockId="3.[707,1194,194,1730]" pageId="3" pageNumber="281">
|
||
Mouthparts with labrum medially emarginate to its base. Mandibles large, curved, blunt; with or without long medial tooth; mandibles with dorsolateral groove extending from condyle to just above tooth; prostheca setose. Maxilla with galea and lacinia densely setose; maxillary palpi 4-segmented; P
|
||
<subScript id="3D4034E6FFEC2401FD14FF29FD34FF30" attach="left" box="[720,729,229,243]" fontSize="6" pageId="3" pageNumber="281">1</subScript>
|
||
small, about 1/3 as long as P
|
||
<subScript id="3D4034E6FFEC2401FBE6FF29FBC6FF30" attach="left" box="[1058,1067,229,243]" fontSize="6" pageId="3" pageNumber="281">2</subScript>
|
||
; P
|
||
<subScript id="3D4034E6FFEC2401FB88FF29FBB8FF30" attach="left" box="[1100,1109,229,243]" fontSize="6" pageId="3" pageNumber="281">2</subScript>
|
||
curved, elongate, longer than P
|
||
<subScript id="3D4034E6FFEC2401FC69FECCFC5BFECD" attach="right" box="[941,950,256,270]" fontSize="6" pageId="3" pageNumber="281">3</subScript>
|
||
; P
|
||
<subScript id="3D4034E6FFEC2401FC10FECCFC30FECD" attach="left" box="[980,989,256,270]" fontSize="6" pageId="3" pageNumber="281">2</subScript>
|
||
–P
|
||
<subScript id="3D4034E6FFEC2401FC33FECCFBEDFECD" attach="left" box="[1015,1024,256,270]" fontSize="6" pageId="3" pageNumber="281">3</subScript>
|
||
with large setae apically; P
|
||
<subScript id="3D4034E6FFEC2401FCEEFED7FCDEFEEA" attach="left" box="[810,819,283,297]" fontSize="6" pageId="3" pageNumber="281">4</subScript>
|
||
elongate, longer than P
|
||
<subScript id="3D4034E6FFEC2401FBDAFED7FBCAFEEA" attach="left" box="[1054,1063,283,297]" fontSize="6" pageId="3" pageNumber="281">3</subScript>
|
||
. Submentum with 1 long and 1 shorter anterolateral seta in each end; labial palpi 3-segmented; P
|
||
<subScript id="3D4034E6FFEC2401FC2BFE9CFC15FE9D" attach="left" box="[1007,1016,336,350]" fontSize="6" pageId="3" pageNumber="281">1</subScript>
|
||
subequal in length to P
|
||
<subScript id="3D4034E6FFEC2401FD23FEA6FD1DFEBB" attach="left" box="[743,752,362,376]" fontSize="6" pageId="3" pageNumber="281">2</subScript>
|
||
; both P
|
||
<subScript id="3D4034E6FFEC2401FCF3FEA6FCADFEBB" attach="left" box="[823,832,362,376]" fontSize="6" pageId="3" pageNumber="281">1</subScript>
|
||
and P
|
||
<subScript id="3D4034E6FFEC2401FCBCFEA6FC6CFEBB" attach="left" box="[888,897,362,376]" fontSize="6" pageId="3" pageNumber="281">2</subScript>
|
||
with several long setae; P
|
||
<subScript id="3D4034E6FFEC2401FBB5FEA6FB97FEBB" attach="left" box="[1137,1146,362,376]" fontSize="6" pageId="3" pageNumber="281">3</subScript>
|
||
with distal end securiform.
|
||
</paragraph>
|
||
<paragraph id="A17B36A3FFEC2401FD1FFE5AFBE5FCAC" blockId="3.[707,1194,194,1730]" pageId="3" pageNumber="281">Pronotum quadrate; lateral margins of pronotum concave in dorsal aspect; pronotum broadest in apical 1/3 and narrower at basal angles. Hypomeron expanded, with transverse microsculpture and few micropunctures; superior and inferior marginal lines of hypomeron separate throughout their lengths; superior line fully visible from above, extending around anterolateral margin of pronotum and contacting inferior line at neck fossa; no portion of dorsum of pronotum visible from below. Pronotum with multiple rows of punctures. Postcoxal process present,elongate.Mesoscutellum prominent, long, with dense polygonal microsculpture and micropunctures; with multiple rows of small punctures. Basisternum with dense polygonal microsculpture and carina; anterior marginal depression present; furcasternum with medial carina pointed vertically; furcasternum without polygonal microsculpture.</paragraph>
|
||
<paragraph id="A17B36A3FFEC2401FD1FFCB9FC38FBBA" blockId="3.[707,1194,194,1730]" pageId="3" pageNumber="281">Elytral width subequal to pronotal width; with multiple rows of punctures and large setae and sparse microsculpture; elytra shiny due to lack of extensive microsculpture. Hind wings fully developed. Mesoventrite with anterior margin forming “lip”; with dense polygonal microsculpture and few punctures along edges; without median carina. Metaventrite with dense, uniform, medium-sized punctures; metaventral process small, rounded, with v-shaped emargination.</paragraph>
|
||
<paragraph id="A17B36A3FFEC2401FD1FFBB3FD14FB3D" blockId="3.[707,1194,194,1730]" pageId="3" pageNumber="281">Tarsal segmentation 5-5-5; meso- and metatibiae with multiple rows of spurs. Protarsus enlarged in both sexes, with spatulate setae ventrally; meso- and metatarsi not enlarged. Empodium with 2 small setae.</paragraph>
|
||
<paragraph id="A17B36A3FFEC2401FD1FFAC8FC07F94E" blockId="3.[707,1194,194,1730]" pageId="3" pageNumber="281">
|
||
Abdomen with paired protergal glands; abdomen expanding from segment III to segment V (widest) and then becoming narrower towards segment VIII. Abdominal terga III–V with basal carina;
|
||
<taxonomicName id="66C44D20FFEC2401FB8AFA9FFB44FAAB" authorityName="Bernhauer" authorityYear="1911" box="[1102,1193,1363,1384]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="281" phylum="Arthropoda" rank="species" species="bruchi">
|
||
<emphasis id="93B0EAB1FFEC2401FB8AFA9FFB44FAAB" box="[1102,1193,1363,1384]" italics="true" pageId="3" pageNumber="281">D. bruchi</emphasis>
|
||
</taxonomicName>
|
||
with faint subbasal (arch-like) carina on III–V;
|
||
<taxonomicName id="66C44D20FFEC2401FB56FAA2FC05FA5E" authority="Bernhauer" authorityName="Bernhauer" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="281" phylum="Arthropoda" rank="species" species="transverserugosus">
|
||
<emphasis id="93B0EAB1FFEC2401FB56FAA2FC9CFA5E" italics="true" pageId="3" pageNumber="281">D. transverserugosus</emphasis>
|
||
Bernhauer
|
||
</taxonomicName>
|
||
without subbasal (arch-like) carina on III–V. Segments with dense, uniform, small punctures and microsculpture. Males with secondary sexual structures of sterna VIII–IX (only known for
|
||
<emphasis id="93B0EAB1FFEC2401FCA4FA3FFBDFF9CB" box="[864,1074,1523,1544]" italics="true" pageId="3" pageNumber="281">
|
||
<taxonomicName id="66C44D20FFEC2401FCA4FA3FFBC6F9CB" box="[864,1067,1523,1544]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="281" phylum="Arthropoda" rank="species" species="transverserugosus">D. transverserugosus</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
; presence of porose structure on abdominal sternum VII unclear (see Remarks on the description of
|
||
<taxonomicName id="66C44D20FFEC2401FB8EF9E4FCADF99B" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="281" phylum="Arthropoda" rank="species" species="transverserugosus">
|
||
<emphasis id="93B0EAB1FFEC2401FB8EF9E4FCADF99B" italics="true" pageId="3" pageNumber="281">D. transverserugosus</emphasis>
|
||
</taxonomicName>
|
||
for more details). Lateral tergal sclerites of abdominal segment IX long and straight, covered with long macrosetae.
|
||
</paragraph>
|
||
<paragraph id="A17B36A3FFEC2406FD1FF95EFE04FB4D" blockId="3.[707,1194,194,1730]" lastBlockId="4.[151,638,1065,1326]" lastPageId="4" lastPageNumber="282" pageId="3" pageNumber="281">
|
||
Male genitalia with aedeagus typical of
|
||
<taxonomicName id="66C44D20FFEC2401FBBDF95EFCCFF901" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="281" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
|
||
(
|
||
<figureCitation id="39FF2A26FFEC2401FCF5F961FC4AF901" box="[817,935,1708,1730]" captionStart="Figs" captionStartId="8.[175,214,594,613]" captionTargetBox="[155,637,197,570]" captionTargetId="figure-266@8.[151,638,194,579]" captionTargetPageId="8" captionText="Figs. 16–18. Aedeagus of Dysanellus transverserugosus. 16) Dorsal view; 17) Lateral view; 18) Paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5381980" httpUri="https://zenodo.org/record/5381980/files/figure.png" pageId="3" pageNumber="281">Figs. 16–18</figureCitation>
|
||
); with median lobe longer and wider than paramere; median lobe with single dorsal tooth; paramere wide and long, not divided into lobes. Paramere with peg setae and short apical setae. Spermatheca not sclerotized.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F5BB662BFFEB2406FF6BFC2CFBC3FC30" ID-DOI="http://doi.org/10.5281/zenodo.5381968" ID-Zenodo-Dep="5381968" box="[175,1070,992,1011]" httpUri="https://zenodo.org/record/5381968/files/figure.png" pageId="4" pageNumber="282" startId="4.[175,214,992,1011]" targetBox="[154,1169,194,976]" targetPageId="4">
|
||
<paragraph id="A17B36A3FFEB2406FF6BFC2CFBC3FC30" blockId="4.[175,1070,992,1011]" box="[175,1070,992,1011]" pageId="4" pageNumber="282">
|
||
<emphasis id="93B0EAB1FFEB2406FF6BFC2CFEEBFC30" bold="true" box="[175,262,992,1011]" pageId="4" pageNumber="282">Figs. 5–6.</emphasis>
|
||
Heads and pronota of
|
||
<taxonomicName id="66C44D20FFEB2406FE12FC2CFD6AFC30" box="[470,647,992,1011]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="undefined-5">
|
||
<emphasis id="93B0EAB1FFEB2406FE12FC2CFDDDFC30" box="[470,560,992,1011]" italics="true" pageId="4" pageNumber="282">Dysanellus</emphasis>
|
||
species.
|
||
<emphasis id="93B0EAB1FFEB2406FDB8FC2CFD6AFC30" bold="true" box="[636,647,992,1011]" pageId="4" pageNumber="282">5</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="93B0EAB1FFEB2406FD43FC2CFD60FC30" bold="true" box="[647,653,992,1011]" pageId="4" pageNumber="282">)</emphasis>
|
||
<taxonomicName id="66C44D20FFEB2406FD50FC2CFD09FC30" authorityName="Bernhauer" authorityYear="1911" box="[660,740,992,1011]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="bruchi">
|
||
<emphasis id="93B0EAB1FFEB2406FD50FC2CFD09FC30" box="[660,740,992,1011]" italics="true" pageId="4" pageNumber="282">D. bruchi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="93B0EAB1FFEB2406FD34FC2CFCECFC30" bold="true" box="[752,769,992,1011]" pageId="4" pageNumber="282">6)</emphasis>
|
||
<taxonomicName id="66C44D20FFEB2406FCCCFC2CFC5AFC30" box="[776,951,992,1011]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="transverserugosus">
|
||
<emphasis id="93B0EAB1FFEB2406FCCCFC2CFC5AFC30" box="[776,951,992,1011]" italics="true" pageId="4" pageNumber="282">D. transverserugosus</emphasis>
|
||
</taxonomicName>
|
||
. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E9DE6528FFEB2406FF6BFB58FBD6FB4E" pageId="4" pageNumber="282" type="discussion">
|
||
<paragraph id="A17B36A3FFEB2406FF6BFB58FED0FAED" blockId="4.[151,638,1065,1326]" pageId="4" pageNumber="282">
|
||
<emphasis id="93B0EAB1FFEB2406FF6BFB58FE6DFB6A" bold="true" box="[175,384,1172,1193]" pageId="4" pageNumber="282">Synoptic Catalogue.</emphasis>
|
||
The catalogue presented below follows the format of the catalogue presented in
|
||
<bibRefCitation id="C5554B52FFEB2406FF6BFB05FEA0FB1D" author="Herman" box="[175,333,1225,1246]" firstAuthor="Herman" pageId="4" pageNumber="282" pagination="1 - 4218" refId="ref8893" refString="Herman, L. H. 2001 b. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Bulletin of the American Museum of Natural History 265: 1 - 4218." type="journal article" year="2001" yearSuffix="b">Herman (2001b)</bibRefCitation>
|
||
, and its purpose here is to clarify the status of all taxa assigned to
|
||
<taxonomicName id="66C44D20FFEB2406FE12FB28FDD2FB3A" authorityName="BERNHAUER AND" authorityYear="1911" box="[470,575,1252,1273]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FE12FB28FDD2FB3A" box="[470,575,1252,1273]" italics="true" pageId="4" pageNumber="282">Dysanellus</emphasis>
|
||
</taxonomicName>
|
||
in the past. For the complete catalogue entry details, see
|
||
<bibRefCitation id="C5554B52FFEB2406FF53FAD5FED4FAED" author="Herman" box="[151,313,1305,1326]" firstAuthor="Herman" pageId="4" pageNumber="282" pagination="1 - 4218" refId="ref8893" refString="Herman, L. H. 2001 b. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Bulletin of the American Museum of Natural History 265: 1 - 4218." type="journal article" year="2001" yearSuffix="b">Herman (2001b)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<treatmentCitationGroup id="81D4118DFFEB2406FF53FA88FE9CFA3B" pageId="4" pageNumber="282">
|
||
<paragraph id="A17B36A3FFEB2406FF53FA88FEF1FAB0" blockId="4.[151,639,1348,1528]" pageId="4" pageNumber="282">
|
||
<treatmentCitation id="206510B2FFEB2406FF53FA88FE3DFA9A" author="Bernhauer, M." box="[151,464,1348,1369]" page="419" pageId="4" pageNumber="282" year="1911">
|
||
<taxonomicName id="66C44D20FFEB2406FF53FA88FE3DFA9A" ID-CoL="8XL36" authority="Bernhauer 1911: 419" authorityName="BERNHAUER AND" authorityPageNumber="419" authorityYear="1911" box="[151,464,1348,1369]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FF53FA88FEE9FA9A" bold="true" box="[151,260,1348,1369]" italics="true" pageId="4" pageNumber="282">Dysanellus</emphasis>
|
||
<bibRefCitation id="C5554B52FFEB2406FECDFA88FE3DFA9A" author="Bernhauer" box="[265,464,1348,1369]" firstAuthor="Bernhauer" page="419" pageId="4" pageNumber="282" pagination="403 - 422" refId="ref8254" refString="Bernhauer, M. 1911. Zur Staphylinidenfauna von Sud- Amerika (Col.). Deutsche Entomologische Zeitschrift 1911: 403 - 422." type="journal article" year="1911">Bernhauer 1911: 419</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(species included:
|
||
<taxonomicName id="66C44D20FFEB2406FF76FA92FEE2FAB0" authorityName="Bernhauer" authorityYear="1911" box="[178,271,1374,1395]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="bruchi">
|
||
<emphasis id="93B0EAB1FFEB2406FF76FA92FF2EFAB0" box="[178,195,1374,1395]" italics="true" pageId="4" pageNumber="282">D</emphasis>
|
||
.
|
||
<emphasis id="93B0EAB1FFEB2406FF15FA92FEE2FAB0" box="[209,271,1374,1395]" italics="true" pageId="4" pageNumber="282">bruchi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A17B36A3FFEB2406FF6BFAB5FE9CFA3B" blockId="4.[151,639,1348,1528]" pageId="4" pageNumber="282">
|
||
<treatmentCitation id="206510B2FFEB2406FF6BFAB5FDFAFA4D" author="Bernhauer, M." box="[175,535,1401,1422]" page="214" pageId="4" pageNumber="282" year="1934">
|
||
<taxonomicName id="66C44D20FFEB2406FF6BFAB5FDFAFA4D" authority="Bernhauer 1934: 214" authorityName="Bernhauer" authorityPageNumber="214" authorityYear="1934" box="[175,535,1401,1422]" class="Insecta" family="Staphylinidae" genus="Leptodiastemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FF6BFAB5FEA9FA4D" box="[175,324,1401,1422]" italics="true" pageId="4" pageNumber="282">Leptodiastemus</emphasis>
|
||
<bibRefCitation id="C5554B52FFEB2406FE8FFAB5FDFAFA4D" author="Bernhauer" box="[331,535,1401,1422]" firstAuthor="Bernhauer" page="214" pageId="4" pageNumber="282" pagination="212 - 221" refId="ref8348" refString="Bernhauer, M. 1934. Neue Staphyliniden aus Argentinien. Revista de Entomolog'ia 4: 112 - 119, 212 - 221." type="journal article" year="1934">Bernhauer 1934: 214</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, originally proposed as a subgenus of
|
||
<taxonomicName id="66C44D20FFEB2406FE07FA5FFDC1FA6B" authorityName="BERNHAUER AND" authorityYear="1911" box="[451,556,1427,1448]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FE07FA5FFDC1FA6B" box="[451,556,1427,1448]" italics="true" pageId="4" pageNumber="282">Dysanellus</emphasis>
|
||
</taxonomicName>
|
||
(species included:
|
||
<taxonomicName id="66C44D20FFEB2406FEE0FA62FE7BFA00" authorityName="Bernhauer" authorityYear="1934" box="[292,406,1454,1475]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="excellens">
|
||
<emphasis id="93B0EAB1FFEB2406FEE0FA62FED8FA00" box="[292,309,1454,1475]" italics="true" pageId="4" pageNumber="282">D</emphasis>
|
||
.
|
||
<emphasis id="93B0EAB1FFEB2406FE84FA62FE7BFA00" box="[320,406,1454,1475]" italics="true" pageId="4" pageNumber="282">excellens</emphasis>
|
||
</taxonomicName>
|
||
), currently in synonymy with
|
||
<taxonomicName id="66C44D20FFEB2406FEC6FA04FE30FA1E" authority="Bernhauer" authorityName="Bernhauer" authorityYear="1906" box="[258,477,1480,1501]" class="Insecta" family="Staphylinidae" genus="Xenopygus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FEC6FA04FE87FA1E" box="[258,362,1480,1501]" italics="true" pageId="4" pageNumber="282">Xenopygus</emphasis>
|
||
Bernhauer
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="C5554B52FFEB2406FE37FA04FE8AFA3B" author="Chatzimanolis and Caron" firstAuthor="Chatzimanolis" pageId="4" pageNumber="282" pagination="131 - 142" refId="ref8755" refString="Chatzimanolis, S., and E. Caron. 2016. New species and synonymies in Xenopygus Bernhauer (Staphylinidae: Staphylinini). Zootaxa 4200 (1): 131 - 142." type="journal article" year="2016">Chatzimanolis and Caron 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</treatmentCitationGroup>
|
||
<paragraph id="A17B36A3FFEB2406FF53F9C1FDD5F9B1" blockId="4.[151,638,1549,1730]" pageId="4" pageNumber="282">
|
||
<treatmentCitationGroup id="81D4118DFFEB2406FF53F9C1FDD5F9B1" pageId="4" pageNumber="282">
|
||
<heading id="FA3381CFFFEB2406FF53F9C1FD90F9E0" box="[151,637,1549,1571]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="282" reason="2">
|
||
<taxonomicName id="66C44D20FFEB2406FF53F9C1FDB0F9E0" ID-CoL="BRR7" authority="Cameron 1933: 347" authorityName="Cameron" authorityPageNumber="347" authorityYear="1933" box="[151,605,1549,1571]" class="Insecta" family="Staphylinidae" genus="Algon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="brevipennis">
|
||
<treatmentCitation id="206510B2FFEB2406FF53F9C1FE34F9E0" author="Cameron, M." box="[151,473,1549,1571]" page="347" pageId="4" pageNumber="282" year="1933">
|
||
<emphasis id="93B0EAB1FFEB2406FF53F9C1FEE4F9E1" bold="true" box="[151,265,1549,1570]" italics="true" pageId="4" pageNumber="282">brevipennis</emphasis>
|
||
<bibRefCitation id="C5554B52FFEB2406FED0F9C2FE34F9E0" author="Cameron" box="[276,473,1550,1571]" firstAuthor="Cameron" page="347" pageId="4" pageNumber="282" pagination="338 - 360" refId="ref8395" refString="Cameron, M. 1933. Staphylinidae (Col.) from Mount Kinabalu. Journal of the Federated Malay States Museums 17 (2): 338 - 360." type="journal article" year="1933">Cameron 1933: 347</bibRefCitation>
|
||
</treatmentCitation>
|
||
(
|
||
<emphasis id="93B0EAB1FFEB2406FE2EF9C2FDBEF9E0" box="[490,595,1550,1571]" italics="true" pageId="4" pageNumber="282">Dysanellus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
, in
|
||
</heading>
|
||
<taxonomicName id="66C44D20FFEB2406FF76F9E4FECEF9FE" authority="Sharp" authorityName="Sharp" authorityYear="1874" box="[178,291,1576,1597]" class="Insecta" family="Staphylinidae" genus="Algon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FF76F9E4FF04F9FE" box="[178,233,1576,1597]" italics="true" pageId="4" pageNumber="282">Algon</emphasis>
|
||
Sharp
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName id="66C44D20FFEB2406FEFAF9E4FF0FF99B" authority="Schillhammer" authorityName="Schillhammer" authorityYear="1999" class="Insecta" family="Staphylinidae" genus="Algon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="dysanelloides">
|
||
<emphasis id="93B0EAB1FFEB2406FEFAF9E4FE18F9FE" box="[318,501,1576,1597]" italics="true" pageId="4" pageNumber="282">Algon dysanelloides</emphasis>
|
||
Schillhammer, 1999
|
||
</taxonomicName>
|
||
(a replacement name for
|
||
<taxonomicName id="66C44D20FFEB2406FE15F98FFEBBF9B1" authorityName="Cameron" authorityYear="1933" class="Insecta" family="Staphylinidae" genus="Algon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="brevipennis">
|
||
<emphasis id="93B0EAB1FFEB2406FE15F98FFD93F99B" box="[465,638,1603,1624]" italics="true" pageId="4" pageNumber="282">Algon brevipennis</emphasis>
|
||
(
|
||
<bibRefCitation id="C5554B52FFEB2406FF7CF991FEA2F9B1" author="Cameron" box="[184,335,1629,1650]" firstAuthor="Cameron" pageId="4" pageNumber="282" pagination="338 - 360" refId="ref8395" refString="Cameron, M. 1933. Staphylinidae (Col.) from Mount Kinabalu. Journal of the Federated Malay States Museums 17 (2): 338 - 360." type="journal article" year="1933">Cameron, 1933</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
that was preoccupied).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="A17B36A3FFEB2406FF53F9B4FDDFF94E" blockId="4.[151,638,1549,1730]" box="[151,562,1656,1677]" pageId="4" pageNumber="282">
|
||
<treatmentCitationGroup id="81D4118DFFEB2406FF53F9B4FDDFF94E" box="[151,562,1656,1677]" pageId="4" pageNumber="282">
|
||
<taxonomicName id="66C44D20FFEB2406FF53F9B4FDC3F94E" ID-CoL="6DPL3" authority="Bernhauer 1911: 419" authorityName="Bernhauer" authorityPageNumber="419" authorityYear="1911" box="[151,558,1656,1677]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="bruchi">
|
||
<treatmentCitation id="206510B2FFEB2406FF53F9B4FE41F94E" author="Bernhauer, M." box="[151,428,1656,1677]" page="419" pageId="4" pageNumber="282" year="1911">
|
||
<emphasis id="93B0EAB1FFEB2406FF53F9B4FF35F94E" bold="true" box="[151,216,1656,1677]" italics="true" pageId="4" pageNumber="282">bruchi</emphasis>
|
||
<bibRefCitation id="C5554B52FFEB2406FF24F9B4FE41F94E" author="Bernhauer" box="[224,428,1656,1677]" firstAuthor="Bernhauer" page="419" pageId="4" pageNumber="282" pagination="403 - 422" refId="ref8254" refString="Bernhauer, M. 1911. Zur Staphylinidenfauna von Sud- Amerika (Col.). Deutsche Entomologische Zeitschrift 1911: 403 - 422." type="journal article" year="1911">Bernhauer 1911: 419</bibRefCitation>
|
||
</treatmentCitation>
|
||
(
|
||
<emphasis id="93B0EAB1FFEB2406FE7FF9B4FDC9F94E" box="[443,548,1656,1677]" italics="true" pageId="4" pageNumber="282">Dysanellus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="A17B36A3FFEB2406FF53F95EFDD4F901" blockId="4.[151,638,1549,1730]" pageId="4" pageNumber="282">
|
||
<treatmentCitationGroup id="81D4118DFFEB2406FF53F95EFDD4F901" pageId="4" pageNumber="282">
|
||
<treatmentCitation id="206510B2FFEB2406FF53F95EFE5FF964" author="Bernhauer, M." box="[151,434,1682,1703]" page="145" pageId="4" pageNumber="282" year="1915">
|
||
<emphasis id="93B0EAB1FFEB2406FF53F95EFF0CF964" bold="true" box="[151,225,1682,1703]" italics="true" pageId="4" pageNumber="282">elegans</emphasis>
|
||
<bibRefCitation id="C5554B52FFEB2406FF2CF95EFE5FF964" author="Bernhauer" box="[232,434,1682,1703]" firstAuthor="Bernhauer" page="145" pageId="4" pageNumber="282" pagination="95 - 189" refId="ref8279" refString="Bernhauer, M. 1915. Zur Staphyliniden-fauna des tropischen Afrika. Annales Musei Nationalis Hungarici 13: 95 - 189." type="journal article" year="1915">Bernhauer 1915: 145</bibRefCitation>
|
||
</treatmentCitation>
|
||
(
|
||
<taxonomicName id="66C44D20FFEB2406FE04F95EFDC4F964" ID-CoL="8XL36" authorityName="BERNHAUER AND" authorityYear="1911" box="[448,553,1682,1703]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FE04F95EFDC4F964" box="[448,553,1682,1703]" italics="true" pageId="4" pageNumber="282">Dysanellus</emphasis>
|
||
</taxonomicName>
|
||
, cited as
|
||
<emphasis id="93B0EAB1FFEB2406FF76F961FEFAF901" box="[178,279,1709,1730]" italics="true" pageId="4" pageNumber="282">Disanellus</emphasis>
|
||
), in
|
||
<taxonomicName id="66C44D20FFEB2406FE83F961FDD8F901" authority="(Hammond 1984)" baseAuthorityName="Hammond" baseAuthorityYear="1984" box="[327,565,1709,1730]" class="Insecta" family="Staphylinidae" genus="Algon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FE83F961FE6DF901" box="[327,384,1709,1730]" italics="true" pageId="4" pageNumber="282">Algon</emphasis>
|
||
(
|
||
<bibRefCitation id="C5554B52FFEB2406FE4AF961FDC3F901" author="Hammond" box="[398,558,1709,1730]" firstAuthor="Hammond" pageId="4" pageNumber="282" pagination="187 - 218" refId="ref8824" refString="Hammond, P. M. 1984. An annotated check-list of Staphylinidae (Insecta: Coleoptera) recorded from Borneo. The Sarawak Museum Journal 33 (54): 187 - 218." type="journal article" year="1984">Hammond 1984</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="A17B36A3FFEB2406FD69FBE5FC81FBB0" blockId="4.[685,1171,1065,1165]" pageId="4" pageNumber="282">
|
||
<treatmentCitationGroup id="81D4118DFFEB2406FD69FBE5FC81FBB0" pageId="4" pageNumber="282">
|
||
<heading id="FA3381CFFFEB2406FD69FBE5FB7EFBFD" box="[685,1171,1065,1086]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="282" reason="2">
|
||
<taxonomicName id="66C44D20FFEB2406FD69FBE5FC76FBFD" ID-CoL="6DPKY" authority="Bernhauer 1934" authorityName="Bernhauer" authorityYear="1934" box="[685,923,1065,1086]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="excellens">
|
||
<emphasis id="93B0EAB1FFEB2406FD69FBE5FCE9FBFD" bold="true" box="[685,772,1065,1086]" italics="true" pageId="4" pageNumber="282">excellens</emphasis>
|
||
<bibRefCitation id="C5554B52FFEB2406FCCCFBE5FC76FBFD" author="Bernhauer" box="[776,923,1065,1086]" firstAuthor="Bernhauer" pageId="4" pageNumber="282" pagination="212 - 221" refId="ref8348" refString="Bernhauer, M. 1934. Neue Staphyliniden aus Argentinien. Revista de Entomolog'ia 4: 112 - 119, 212 - 221." type="journal article" year="1934">Bernhauer 1934</bibRefCitation>
|
||
</taxonomicName>
|
||
:(215
|
||
<taxonomicName id="66C44D20FFEB2406FC14FBE5FBD8FBFD" authorityName="BERNHAUER AND" authorityYear="1911" box="[976,1077,1065,1086]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FC14FBE5FBD8FBFD" box="[976,1077,1065,1086]" italics="true" pageId="4" pageNumber="282">Dysanellus</emphasis>
|
||
</taxonomicName>
|
||
, subgenus
|
||
</heading>
|
||
<taxonomicName id="66C44D20FFEB2406FD0CFB8FFCBBFB9B" authorityName="Bernhauer" authorityYear="1934" box="[712,854,1091,1112]" class="Insecta" family="Staphylinidae" genus="Leptodiastemus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FD0CFB8FFCBBFB9B" box="[712,854,1091,1112]" italics="true" pageId="4" pageNumber="282">Leptodiastemus</emphasis>
|
||
</taxonomicName>
|
||
), in
|
||
<taxonomicName id="66C44D20FFEB2406FC51FB8FFC8FFBB0" authority="(Chatzimanolis and Caron 2016" authorityYear="2016" baseAuthorityName="Chatzimanolis and Caron" baseAuthorityYear="2016" class="Insecta" family="Staphylinidae" genus="Xenopygus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="93B0EAB1FFEB2406FC51FB8FFC17FB9B" box="[917,1018,1091,1112]" italics="true" pageId="4" pageNumber="282">Xenopygus</emphasis>
|
||
(
|
||
<bibRefCitation id="C5554B52FFEB2406FBCBFB8FFC8FFBB0" author="Chatzimanolis, S. & E. Caron" firstAuthor="Chatzimanolis" pageId="4" pageNumber="282" pagination="131 - 142" refId="ref8755" refString="Chatzimanolis, S., and E. Caron. 2016. New species and synonymies in Xenopygus Bernhauer (Staphylinidae: Staphylinini). Zootaxa 4200 (1): 131 - 142." type="journal article" year="2016">Chatzimanolis and Caron 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="A17B36A3FFEB2406FD69FBB4FBD6FB4E" blockId="4.[685,1171,1065,1165]" box="[685,1083,1144,1165]" pageId="4" pageNumber="282">
|
||
<treatmentCitationGroup id="81D4118DFFEB2406FD69FBB4FBD6FB4E" box="[685,1083,1144,1165]" pageId="4" pageNumber="282">
|
||
<treatmentCitation id="206510B2FFEB2406FD69FBB4FBD6FB4E" author="Bernhauer, M." box="[685,1083,1144,1165]" page="175" pageId="4" pageNumber="282" year="1921">
|
||
<taxonomicName id="66C44D20FFEB2406FD69FBB4FBDAFB4E" ID-CoL="6DPKQ" authority="Bernhauer 1921: 175" authorityName="Bernhauer" authorityPageNumber="175" authorityYear="1921" box="[685,1079,1144,1165]" class="Insecta" family="Staphylinidae" genus="Dysanellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="transverserugosus">
|
||
<emphasis id="93B0EAB1FFEB2406FD69FBB4FC8DFB4E" bold="true" box="[685,864,1144,1165]" italics="true" pageId="4" pageNumber="282">transverserugosus</emphasis>
|
||
<bibRefCitation id="C5554B52FFEB2406FCACFBB4FBDAFB4E" author="Bernhauer" box="[872,1079,1144,1165]" firstAuthor="Bernhauer" page="175" pageId="4" pageNumber="282" pagination="170 - 183" refId="ref8320" refString="Bernhauer, M. 1921. Zur Staphylinidenfauna Sudamerikas, insbesondere Argentiniens. Archiv fur Naturgeschichte (A) 86 (8): 170 - 183." type="journal article" year="1921">Bernhauer 1921: 175</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |