247 lines
33 KiB
XML
247 lines
33 KiB
XML
<document id="CD8335E9058731A2020E254EF89E12C6" ID-DOI="10.5281/zenodo.5458831" ID-ISSN="2345-7600" ID-Zenodo-Dep="5458831" ID-ZooBank="CAABF079-BFA8-48C9-986C-2BD100B3CB7E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630211616052" checkinUser="tatiana" docAuthor="Sutcharit, Chirasak, Tongkerd, Piyoros & Panha, Somsak" docDate="2014" docId="294B25641070861CD662FBB3B65AFA79" docLanguage="en" docName="62rbz330-338.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Pterocyclos frednaggsi Sutcharit & Panha 2014, new species" docType="treatment" docUuid="8A1871F9-01C3-4319-8284-28C80AC434AB" docUuidSource="ZooBank" docVersion="4" lastPageNumber="337" masterDocId="D5725D1C1076861BD550FFEEB55DFF8C" masterDocTitle="The land snail genus Pterocyclos Benson, 1832 (Caenogastropoda: Cyclophoridae) from Thailand and Peninsular Malaysia, with descriptions of two new species" masterLastPageNumber="338" masterPageNumber="330" pageNumber="336" updateTime="1699187624015" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="B7ADA24A38C9B03EC9B3D6FF35877C2D" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="27BC56D531AFDAD21BE0E0F83BE5F94D">
|
||
<mods:title id="4AD6A3E0A280D8D2C1F8935DAFFB102D">The land snail genus Pterocyclos Benson, 1832 (Caenogastropoda: Cyclophoridae) from Thailand and Peninsular Malaysia, with descriptions of two new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="6FF78B8C5E901C2CDAE656FFE3A382CD" type="personal">
|
||
<mods:role id="7DB0E9CF4E11B360B9C16DD2FF89B8F3">
|
||
<mods:roleTerm id="D4750AE5091C335FBC81A0AA9896EF72">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D6CD29E4D6B717276DA1FD023102A91A">Sutcharit, Chirasak</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="7DCE1560DC01717204C38DEFE7B67F9F" type="personal">
|
||
<mods:role id="F5A3445AFEA252EC03B78B7D83E11D2A">
|
||
<mods:roleTerm id="3B14F9D95D3155B03E78D41C1DF3D96D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1854B1F9473E551D1600A3DFE2632F5F">Tongkerd, Piyoros</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="AC7DF7484E1C77320F0465B1CDE70305" type="personal">
|
||
<mods:role id="DE87EBB55EA8530736FF2AB8CE1378E0">
|
||
<mods:roleTerm id="B9A9447DD3A32E13FB13CC936479F2EB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B65834C9175A00462DB4FF35C5C6A14F">Panha, Somsak</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="6078C6E7BAB92D47F91B9F727177F71D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A0567062DF05A74165F05B916A133DC6" type="host">
|
||
<mods:titleInfo id="297B41586EC27E14EEBA6B7A89237422">
|
||
<mods:title id="3799EBC9E9A8B3DF51F56E45E536A032">Raffles Bulletin of Zoology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="CEED03D476958E3075E0AB4BE3B7BCF9">
|
||
<mods:date id="2AF3460D85D58153BA6BE4FF5B6D7814">2014</mods:date>
|
||
<mods:detail id="E34B655F8D47418686CEA42D1FD704EC" type="pubDate">
|
||
<mods:number id="9422DE354F2A7C1239B0626068C965A8">2014-05-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="847B90CEAB973895E3F04292CCFCAC45" type="volume">
|
||
<mods:number id="DA2D70F833FEE8E85F721774F16FE06B">62</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="68866DA3971CBC8164804ED2A2D068EC" unit="page">
|
||
<mods:start id="0BC3CEF7834AE23739A3F9FCC94F120B">330</mods:start>
|
||
<mods:end id="D0ECADE8CC1E01C07B0693318A820BE5">338</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="DEB7D47677B65C583FC2506EF85286B5">journal article</mods:classification>
|
||
<mods:identifier id="31834A0AD727B26794AACC27DF09E4B1" type="DOI">10.5281/zenodo.5458831</mods:identifier>
|
||
<mods:identifier id="C18D0BA2F4EA4219DD748CC49FF7D3CD" type="ISSN">2345-7600</mods:identifier>
|
||
<mods:identifier id="97F2F3C17348F1282E557BEF895C0399" type="Zenodo-Dep">5458831</mods:identifier>
|
||
<mods:identifier id="AC1EA2546C0CFA6D1971AF08C7689607" type="ZooBank">CAABF079-BFA8-48C9-986C-2BD100B3CB7E</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="294B25641070861CD662FBB3B65AFA79" ID-DOI="http://doi.org/10.5281/zenodo.5450989" ID-Zenodo-Dep="5450989" LSID="urn:lsid:zoobank.org:act:8A1871F9-01C3-4319-8284-28C80AC434AB" httpUri="http://treatment.plazi.org/id/294B25641070861CD662FBB3B65AFA79" lastPageId="7" lastPageNumber="337" pageId="6" pageNumber="336">
|
||
<subSubSection id="E9F8C7F91070861DD662FBB3B0FBFBF9" box="[818,1446,1117,1141]" pageId="6" pageNumber="336" type="nomenclature">
|
||
<paragraph id="A15D94721070861DD662FBB3B0FBFBF9" blockId="6.[818,1446,1117,1173]" box="[818,1446,1117,1141]" pageId="6" pageNumber="336">
|
||
<heading id="FA15231E1070861DD662FBB3B0FBFBF9" bold="true" box="[818,1446,1117,1141]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="336" reason="2">
|
||
<emphasis id="939648601070861DD662FBB3B0FBFBF9" bold="true" box="[818,1446,1117,1141]" pageId="6" pageNumber="336">
|
||
<taxonomicName id="66E2EFF11070861DD662FBB3B04EFBF9" ID-CoL="4PWDK" authority="Sutcharit & Panha" authorityName="Sutcharit & Panha" authorityYear="2014" box="[818,1299,1117,1141]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="336" phylum="Mollusca" rank="species" species="frednaggsi" status="sp. nov.">
|
||
<emphasis id="939648601070861DD662FBB3B173FBF9" bold="true" box="[818,1070,1117,1141]" italics="true" pageId="6" pageNumber="336">Pterocyclos frednaggsi</emphasis>
|
||
Sutcharit & Panha
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="88A5F51B1070861DD071FBB0B0FBFBF9" box="[1313,1446,1118,1141]" pageId="6" pageNumber="336" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9F8C7F91070861DD6E0FB90B075FB19" box="[944,1320,1150,1173]" pageId="6" pageNumber="336" type="description">
|
||
<paragraph id="A15D94721070861DD6E0FB90B075FB19" blockId="6.[818,1446,1117,1173]" box="[944,1320,1150,1173]" pageId="6" pageNumber="336">
|
||
(
|
||
<figureCitation id="39D988F71070861DD6E8FB90B15EFB19" box="[952,1027,1150,1173]" captionStart="Fig" captionStartId="1.[136,171,2006,2027]" captionTargetBox="[136,774,1032,1975]" captionTargetId="figure-176@1.[136,774,1032,1975]" captionTargetPageId="1" captionText="Fig. 1. Distribution map of Pterocyclos species examined in this study. The numbered locality names are detailed in Table 1. The approximate type locality of Pterocyclos subalatus, and Pterocyclos umbraticus, are indicated by locality numbers 3 and 5, respectively." figureDoi="http://doi.org/10.5281/zenodo.5458833" httpUri="https://zenodo.org/record/5458833/files/figure.png" pageId="6" pageNumber="336">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="39D988F71070861DD143FB90B128FB19" box="[1043,1141,1150,1173]" captionStart="Fig" captionStartId="2.[113,148,2006,2027]" captionTargetBox="[113,1450,778,1971]" captionTargetId="figure-151@2.[113,1451,776,1972]" captionTargetPageId="2" captionText="Fig. 2. A–C, General anatomy of Pterocyclos frednaggsi Sutcharit & Panha, new species, from Gua Musang, Kelantan, Malaysia CUMZ 4944, showing: A, right side of male with testis and external penis; B, right side of female with ovary and vaginal groove; C, left side of female with lung cavity and heart. D, E, Radula morphology of: D, Pterocyclos spaleotes, topotype specimen CUMZ 4585; E, Pterocyclos frednaggsi Sutcharit & Panha, new species, paratype CUMZ 4581." figureDoi="http://doi.org/10.5281/zenodo.5458835" httpUri="https://zenodo.org/record/5458835/files/figure.png" pageId="6" pageNumber="336">
|
||
2A
|
||
<emphasis id="939648601070861DD164FB90B11CFB19" bold="true" box="[1076,1089,1150,1173]" pageId="6" pageNumber="336">–</emphasis>
|
||
C, E
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="39D988F71070861DD1D2FB90B19CFB19" box="[1154,1217,1150,1173]" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit & Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit & Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="6" pageNumber="336">
|
||
3Q
|
||
<emphasis id="939648601070861DD1F2FB90B1F2FB19" bold="true" box="[1186,1199,1150,1173]" pageId="6" pageNumber="336">–</emphasis>
|
||
S
|
||
</figureCitation>
|
||
;
|
||
<tableCitation id="EC60A1C91070861DD19EFB90B043FB19" box="[1230,1310,1150,1173]" captionStart="Table 1" captionStartId="5.[136,189,189,210]" captionText="Table 1. Shell size variation in Pterocyclos spp. Specimen collections and catalogue numbers are indicated in parentheses." pageId="6" pageNumber="336">Table 1</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9F8C7F91070861DD67DFB50B6F4FA79" pageId="6" pageNumber="336" type="materials_examined">
|
||
<paragraph id="A15D94721070861DD67DFB50B1AEFAF9" blockId="6.[813,1453,1214,1397]" pageId="6" pageNumber="336">
|
||
<emphasis id="939648601070861DD67DFB50B6AAFB59" bold="true" box="[813,1015,1214,1237]" pageId="6" pageNumber="336">Type specimens.</emphasis>
|
||
<materialsCitation id="118A9E2F1070861DD154FB50B614FAB9" collectionCode="CUMZ" country="Malaysia" latitude="3.4466333" location="Bukit Chintamanis" longLatPrecision="1" longitude="102.01645" pageId="6" pageNumber="336" specimenCode="CUMZ 4594" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<typeStatus id="7E592AD01070861DD154FB50B132FB59" box="[1028,1135,1214,1237]" pageId="6" pageNumber="336" type="holotype">Holotype</typeStatus>
|
||
<specimenCode id="F1443C091070861DD12AFB50B04DFB59" box="[1146,1296,1214,1237]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="6" pageNumber="336">CUMZ 4594</specimenCode>
|
||
(height 12.1 × diameter
|
||
<quantity id="661A39971070861DD6ECFB30B177FB79" box="[956,1066,1246,1269]" metricMagnitude="-2" metricUnit="m" metricValue="2.78" pageId="6" pageNumber="336" unit="mm" value="27.8">27.8 mm</quantity>
|
||
;
|
||
<figureCitation id="39D988F71070861DD16BFB30B1C1FB79" box="[1083,1180,1246,1269]" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit & Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit & Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="6" pageNumber="336">Fig. 3Q</figureCitation>
|
||
). Type locality:
|
||
<location id="A43DC2A91070861DD03BFB30B6EAFA99" LSID="urn:lsid:plazi:treatment:294B25641070861CD662FBB3B65AFA79:A43DC2A91070861DD03BFB30B6EAFA99" country="Malaysia" latitude="3.4466333" longLatPrecision="1" longitude="102.01645" name="Bukit Chintamanis" pageId="6" pageNumber="336" stateProvince="Pahang">Bukit Chintamanis</location>
|
||
,
|
||
<collectingRegion id="63265A901070861DD691FB10B14FFA99" box="[961,1042,1278,1301]" country="Malaysia" name="Pahang" pageId="6" pageNumber="336">Pahang</collectingRegion>
|
||
,
|
||
<collectingCountry id="D9F5D4E21070861DD14DFB10B122FA99" box="[1053,1151,1278,1301]" name="Malaysia" pageId="6" pageNumber="336">Malaysia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="C4D6F2B51070861DD1DEFB10B07DFA99" box="[1166,1312,1278,1301]" degrees="03" direction="north" minutes="26.798" orientation="latitude" pageId="6" pageNumber="336" precision="1" value="3.4466333">03°26.798' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C4D6F2B51070861DD07BFB10B61CFAB9" degrees="102" direction="east" minutes="00.987" orientation="longitude" pageId="6" pageNumber="336" precision="1" value="102.01645">102°00.987' E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="118A9E2F1070861DD606FAF0B14CFAD9" collectionCode="CUMZ" country="Malaysia" pageId="6" pageNumber="336" specimenCode="CUMZ 4581" specimenCount="18" stateProvince="Pahang" typeStatus="paratype">
|
||
<typeStatus id="7E592AD01070861DD606FAF0B6E3FAB9" box="[854,958,1310,1333]" pageId="6" pageNumber="336" type="paratype">Paratypes</typeStatus>
|
||
<specimenCode id="F1443C091070861DD698FAF0B10EFAB9" box="[968,1107,1310,1333]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="6" pageNumber="336">CUMZ 4581</specimenCode>
|
||
(
|
||
<specimenCount id="B7E45FFB1070861DD136FAF0B1A7FAB9" box="[1126,1274,1310,1333]" pageId="6" pageNumber="336" type="generic">18 specimens</specimenCount>
|
||
in ethanol,
|
||
<figureCitation id="39D988F71070861DD0D1FAF0B613FAD9" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit & Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit & Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="6" pageNumber="336">Fig. 3R</figureCitation>
|
||
), 4571 (29 shells)
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="C7F30CB71070861DD14CFAD0B1DCFAD9" box="[1052,1153,1342,1365]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="336" type="Museum">NHMUK</collectionCode>
|
||
(2 shells),
|
||
<collectionCode id="C7F30CB71070861DD1A7FAD0B066FAD9" box="[1271,1339,1342,1365]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="6" pageNumber="336">NMW</collectionCode>
|
||
(2 shells),
|
||
<collectionCode id="C7F30CB71070861DD67DFAB0B63FFAF9" box="[813,866,1374,1397]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="336">SMF</collectionCode>
|
||
(2 shells) are from the type locality.
|
||
</paragraph>
|
||
<paragraph id="A15D94721070861DD67DFA70B6F4FA79" blockId="6.[813,1452,1438,1525]" pageId="6" pageNumber="336">
|
||
<emphasis id="939648601070861DD67DFA70B116FA39" bold="true" box="[813,1099,1438,1461]" pageId="6" pageNumber="336">Other material examined.</emphasis>
|
||
<materialsCitation id="118A9E2F1070861DD100FA70B6F9FA79" collectionCode="CUMZ" country="Malaysia" county="Kelanta" location="Gua Pulai" municipality="Gua Musang" pageId="6" pageNumber="336" specimenCode="CUMZ 4597" specimenCount="4">
|
||
<location id="A43DC2A91070861DD100FA70B1E7FA39" LSID="urn:lsid:plazi:treatment:294B25641070861CD662FBB3B65AFA79:A43DC2A91070861DD100FA70B1E7FA39" box="[1104,1210,1438,1461]" country="Malaysia" county="Kelanta" municipality="Gua Musang" name="Gua Pulai" pageId="6" pageNumber="336">Gua Pulai</location>
|
||
,
|
||
<collectingMunicipality id="41390E081070861DD192FA70B017FA39" box="[1218,1354,1438,1461]" pageId="6" pageNumber="336">Gua Musang</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="483CECFE1070861DD003FA70B0FAFA39" box="[1363,1447,1438,1461]" pageId="6" pageNumber="336">Kelanta</collectingCounty>
|
||
,
|
||
<collectingCountry id="D9F5D4E21070861DD67DFA50B6D2FA59" box="[813,911,1470,1493]" name="Malaysia" pageId="6" pageNumber="336">Malaysia</collectingCountry>
|
||
<specimenCode id="F1443C091070861DD6C4FA50B146FA59" box="[916,1051,1470,1493]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="6" pageNumber="336">CUMZ 4597</specimenCode>
|
||
(9 shells,
|
||
<figureCitation id="39D988F71070861DD1D5FA50B18EFA59" box="[1157,1235,1470,1493]" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit & Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit & Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="6" pageNumber="336">Fig. 3S</figureCitation>
|
||
), 4944 (
|
||
<specimenCount id="B7E45FFB1070861DD079FA50B0F6FA59" box="[1321,1451,1470,1493]" pageId="6" pageNumber="336" type="generic">4 specimens</specimenCount>
|
||
in ethanol)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9F8C7F91070861DD67DF9F0B1D6F939" pageId="6" pageNumber="336" type="etymology">
|
||
<paragraph id="A15D94721070861DD67DF9F0B1D6F939" blockId="6.[813,1452,1566,1717]" pageId="6" pageNumber="336">
|
||
<emphasis id="939648601070861DD67DF9F0B6E5F9B9" bold="true" box="[813,952,1566,1589]" pageId="6" pageNumber="336">Etymology.</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="66E2EFF11070861DD19DF9F0B014F9B9" authorityName="Sutcharit & Panha" authorityYear="2014" box="[1229,1353,1566,1589]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="336" phylum="Mollusca" rank="species" species="frednaggsi">
|
||
<emphasis id="939648601070861DD19DF9F0B014F9B9" box="[1229,1353,1566,1589]" italics="true" pageId="6" pageNumber="336">frednaggsi</emphasis>
|
||
</taxonomicName>
|
||
” comes from Fred Naggs, malacologist at the Natural History Museum, London, who has enthusiastically encouraged and continuously supported land snail research in
|
||
<collectingCountry id="D9F5D4E21070861DD04EF990B020F919" box="[1310,1405,1662,1685]" name="Thailand" pageId="6" pageNumber="336">Thailand</collectingCountry>
|
||
and other Southeast Asian countries.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9F8C7F91070861CD67DF930B730FE59" lastPageId="7" lastPageNumber="337" pageId="6" pageNumber="336" type="diagnosis">
|
||
<paragraph id="A15D94721070861DD67DF930B008F799" blockId="6.[813,1452,1758,2069]" pageId="6" pageNumber="336">
|
||
<emphasis id="939648601070861DD67DF930B6F4F979" bold="true" box="[813,937,1758,1781]" pageId="6" pageNumber="336">Diagnosis.</emphasis>
|
||
<taxonomicName id="66E2EFF11070861DD6E5F930B0FBF979" authority="Sutcharit & Panha" authorityName="Sutcharit & Panha" authorityYear="2014" box="[949,1446,1758,1781]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="336" phylum="Mollusca" rank="species" species="frednaggsi" status="sp. nov.">
|
||
<emphasis id="939648601070861DD6E5F930B1E2F979" box="[949,1215,1758,1781]" italics="true" pageId="6" pageNumber="336">Pterocyclos frednaggsi</emphasis>
|
||
Sutcharit & Panha
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="88A5F51B1070861DD67DF910B6EEF899" box="[813,947,1790,1813]" pageId="6" pageNumber="336" rank="species">new species</taxonomicNameLabel>
|
||
, differs from
|
||
<taxonomicName id="66E2EFF11070861DD11DF910B1F2F899" authorityName="Benson" authorityYear="1851" box="[1101,1199,1790,1813]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="336" phylum="Mollusca" rank="species" species="blandi">
|
||
<emphasis id="939648601070861DD11DF910B1F2F899" box="[1101,1199,1790,1813]" italics="true" pageId="6" pageNumber="336">P. blandi</emphasis>
|
||
</taxonomicName>
|
||
by having a thick and yellowish periostracum, a broad, dark brown peripheral band, a channel shaped accessory respiratory structure, and a slight expansion of the upper part of the apertural lip. It differs from
|
||
<taxonomicName id="66E2EFF11070861DD6E6F890B168F819" authorityName="Tomlin" authorityYear="1932" box="[950,1077,1918,1941]" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="336" phylum="Mollusca" rank="species" species="spaleotes">
|
||
<emphasis id="939648601070861DD6E6F890B168F819" box="[950,1077,1918,1941]" italics="true" pageId="6" pageNumber="336">P. spaleotes</emphasis>
|
||
</taxonomicName>
|
||
by having a larger shell, a thicker periostracum, and a broad, dark brown peripheral band. The differences from
|
||
<taxonomicName id="66E2EFF11070861DD14EF850B02CF859" authority="Sutcharit & Panha" authorityName="Sutcharit & Panha" authorityYear="2014" box="[1054,1393,1982,2005]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="336" phylum="Mollusca" rank="species" species="diluvium" status="sp. nov.">
|
||
<emphasis id="939648601070861DD14EF850B1C7F859" box="[1054,1178,1982,2005]" italics="true" pageId="6" pageNumber="336">P. diluvium</emphasis>
|
||
Sutcharit & Panha
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="88A5F51B1070861DD02FF850B620F879" pageId="6" pageNumber="336" rank="species">new species</taxonomicNameLabel>
|
||
, include the broad, dark brown peripheral band and the channel shaped accessory respiratory structure.
|
||
</paragraph>
|
||
<paragraph id="A15D94721071861CD5D8FF50B730FE59" blockId="7.[136,775,190,469]" pageId="7" pageNumber="337">
|
||
This new species is superficially similar to
|
||
<emphasis id="939648601071861CD719FF50B782FF59" box="[585,735,190,213]" italics="true" pageId="7" pageNumber="337">
|
||
<taxonomicName id="66E2EFF11071861CD719FF50B786FF59" authorityName="Benthem Jutting" authorityYear="1949" box="[585,731,190,213]" class="Gastropoda" family="Cyclophoridae" genus="Cyclotus" kingdom="Animalia" order="Architaenioglossa" pageId="7" pageNumber="337" phylum="Mollusca" rank="species" species="umbraticus">P. umbraticus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but differs from this latter species by its relatively smooth shell surface, the broad dark brown peripheral band, and the fact that its distribution is limited to the lowland limestone area (altitude less than
|
||
<quantity id="661A39971071861CD403FED0B4C4FED9" box="[339,409,318,341]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="337" unit="m" value="200.0">200 m</quantity>
|
||
amsl). In contrast,
|
||
<taxonomicName id="66E2EFF11071861CD722FED0B65AFED9" authorityName="Benthem Jutting" authorityYear="1949" box="[626,775,318,341]" class="Gastropoda" family="Cyclophoridae" genus="Cyclotus" kingdom="Animalia" order="Architaenioglossa" pageId="7" pageNumber="337" phylum="Mollusca" rank="species" species="umbraticus">
|
||
<emphasis id="939648601071861CD722FED0B65AFED9" box="[626,775,318,341]" italics="true" pageId="7" pageNumber="337">P. umbraticus</emphasis>
|
||
</taxonomicName>
|
||
has a chevron-shaped, zigzag shell sculpture and tends to live in non-limestone forests of highland areas (altitude about
|
||
<quantity id="661A39971071861CD59FFE70B406FE39" box="[207,347,414,437]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="16.0" metricValueMin="2.0" pageId="7" pageNumber="337" unit="m" value="900.0" valueMax="1600.0" valueMin="200.0">
|
||
200
|
||
<emphasis id="939648601071861CD5A8FE70B458FE39" bold="true" box="[248,261,414,437]" italics="true" pageId="7" pageNumber="337">–</emphasis>
|
||
1600 m
|
||
</quantity>
|
||
amsl) at Maxwell’s Hill and Gunung Brinchang,
|
||
<collectingRegion id="63265A901071861CD456FE50B408FE59" box="[262,341,446,469]" country="Malaysia" name="Pahang" pageId="7" pageNumber="337">Pahang</collectingRegion>
|
||
(
|
||
<bibRefCitation id="C573E9831071861CD435FE50B73DFE59" author="van Benthem Jutting WSS" box="[357,608,446,469]" pageId="7" pageNumber="337" pagination="50 - 77" refId="ref6872" refString="van Benthem Jutting WSS (1949) On collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells. Bulletin of the Raffles Museum, 19: 50 - 77." type="journal article" year="1949">Benthem Jutting, 1949</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9F8C7F91071861CD5D8FE10B4D8FB39" pageId="7" pageNumber="337" type="description">
|
||
<paragraph id="A15D94721071861CD5D8FE10B7F2FCD9" blockId="7.[136,775,510,853]" pageId="7" pageNumber="337">
|
||
<emphasis id="939648601071861CD5D8FE10B445FD99" bold="true" box="[136,280,510,533]" pageId="7" pageNumber="337">Description.</emphasis>
|
||
Shell medium sized, depressed, thickened, widely umbilicated. Apex acute; spire flattened to slightly elevated; suture wide; 4
|
||
<emphasis id="939648601071861CD4DAFDD0B4CAFDD9" bold="true" box="[394,407,574,597]" italics="true" pageId="7" pageNumber="337">–</emphasis>
|
||
5 convex and regularly increasing whorls. Shell surface with thin growth lines; periostracum brown and corneous. Last whorl rounded, stout, with a broad dark brown peripheral band. Shell colour brownish or with a variegated, dark brown zigzag pattern on early whorl. Aperture rounded. Apertural lip white, upper part slightly expanded near suture. Accessory respiratory structure channel-like to short tubular shaped. Operculum calcareous, slightly concave inside, and multilamellae outside.
|
||
</paragraph>
|
||
<paragraph id="A15D94721071861CD5D8FC90B4D8FB39" blockId="7.[136,775,894,1205]" pageId="7" pageNumber="337">
|
||
<emphasis id="939648601071861CD5D8FC90B5BFFC19" bold="true" box="[136,226,894,917]" pageId="7" pageNumber="337">Radula.</emphasis>
|
||
Central tooth with well developed central cusp and two smaller lateral cusps on each side; central cusp small with pointed tip; four smaller lateral cusps on both sides with dull to pointed head. Lateral teeth have three cusps; outer cusp large, elongate shape, and two smaller inner lateral cusps with curved tips. Inner marginal teeth have three cusps; central cusp large and convex head, and flanked with smaller and pointed head of one inner and one outer lateral cusps. Outer marginal cusp bicuspid, each cusp with pointed head (
|
||
<figureCitation id="39D988F71071861CD474FB70B424FB39" box="[292,377,1182,1205]" captionStart="Fig" captionStartId="2.[113,148,2006,2027]" captionTargetBox="[113,1450,778,1971]" captionTargetId="figure-151@2.[113,1451,776,1972]" captionTargetPageId="2" captionText="Fig. 2. A–C, General anatomy of Pterocyclos frednaggsi Sutcharit & Panha, new species, from Gua Musang, Kelantan, Malaysia CUMZ 4944, showing: A, right side of male with testis and external penis; B, right side of female with ovary and vaginal groove; C, left side of female with lung cavity and heart. D, E, Radula morphology of: D, Pterocyclos spaleotes, topotype specimen CUMZ 4585; E, Pterocyclos frednaggsi Sutcharit & Panha, new species, paratype CUMZ 4581." figureDoi="http://doi.org/10.5281/zenodo.5458835" httpUri="https://zenodo.org/record/5458835/files/figure.png" pageId="7" pageNumber="337">Fig. 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9F8C7F91071861CD5D8FB30B655FA99" pageId="7" pageNumber="337" type="distribution">
|
||
<paragraph id="A15D94721071861CD5D8FB30B655FA99" blockId="7.[136,776,1246,1301]" pageId="7" pageNumber="337">
|
||
<emphasis id="939648601071861CD5D8FB30B445FB79" bold="true" box="[136,280,1246,1269]" pageId="7" pageNumber="337">Distribution.</emphasis>
|
||
This limestone dwelling species is known from the
|
||
<typeStatus id="7E592AD01071861CD5FEFB10B581FA99" box="[174,220,1278,1301]" pageId="7" pageNumber="337">type</typeStatus>
|
||
locality and from Gua Musang,
|
||
<collectingRegion id="63265A901071861CD762FB10B7C8FA99" box="[562,661,1278,1301]" country="Malaysia" name="Kelantan" pageId="7" pageNumber="337">Kelantan</collectingRegion>
|
||
,
|
||
<collectingCountry id="D9F5D4E21071861CD7CEFB10B65EFA99" box="[670,771,1278,1301]" name="Malaysia" pageId="7" pageNumber="337">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9F8C7F91071861CD5D8FAD0B65AFA79" pageId="7" pageNumber="337" type="discussion">
|
||
<paragraph id="A15D94721071861CD5D8FAD0B65AFA79" blockId="7.[136,775,1342,1525]" pageId="7" pageNumber="337">
|
||
<emphasis id="939648601071861CD5D8FAD0B5A4FAD9" bold="true" box="[136,249,1342,1365]" pageId="7" pageNumber="337">Remarks.</emphasis>
|
||
Specimens of this species from
|
||
<collectingRegion id="63265A901071861CD720FAD0B7C0FAD9" box="[624,669,1342,1365]" country="India" name="Goa" pageId="7" pageNumber="337">Gua</collectingRegion>
|
||
Musang,
|
||
<collectingRegion id="63265A901071861CD5D8FAB0B5B5FAF9" box="[136,232,1374,1397]" country="Malaysia" name="Kelantan" pageId="7" pageNumber="337">Kelantan</collectingRegion>
|
||
(
|
||
<figureCitation id="39D988F71071861CD5A5FAB0B41FFAF9" box="[245,322,1374,1397]" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit & Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit & Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="7" pageNumber="337">Fig. 3S</figureCitation>
|
||
) have slightly larger and darker shells with a narrower peripheral band than typical shells. However, the unique shape of the short tubular accessory respiratory structure and the multilamellae operculum suggest that these atypical forms represent intraspecific geographic variation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |