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<document id="67F3C2113A8D794DDFA04B346E262F43" ID-CLB-Dataset="53387" ID-DOI="10.11646/zootaxa.5124.3.1" ID-GBIF-Dataset="de55f596-c349-4ca2-a826-0e2bad912aa3" ID-ISSN="1175-5326" ID-Zenodo-Dep="6410696" ID-ZooBank="67D4C618-C7C2-4FFC-925B-B260E11CD7B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649054363463" checkinUser="plazi" docAuthor="Jiang, Zhuo-Yin, Zhao, Shuang, Yang, Zhen-Yu, Jia, Feng-Long &amp; Hájek, Jiří" docDate="2022" docId="291CAA4FBF48FF8B1DC5FAD09ABBF843" docLanguage="en" docName="zootaxa.5124.3.1.pdf" docOrigin="Zootaxa 5124 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copelatus tengchongensis Hajek, Jiang &amp; Jia 2022, sp. nov." docType="treatment" docVersion="10" lastPageNumber="273" masterDocId="D525D237BF5DFF9D1D52FF849836FF9E" masterDocTitle="A review of Copelatus Erichson, 1832 of Mainland China, with description of ten new species from the japonicus complex (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="295" masterPageNumber="251" pageNumber="272" updateTime="1699319256616" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="BFCCDB6FE4B8B445376B666AFB51A207" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F7E193702BCA842114AD568618DFDE0C">
<mods:title id="70D53EB8571575E46E8958D5974D3928">A review of Copelatus Erichson, 1832 of Mainland China, with description of ten new species from the japonicus complex (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name id="5397D78BB3EF09420734DB7732A492F4" type="personal">
<mods:role id="1C402A6386DF2939EFA15820548E7EB8">
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</mods:role>
<mods:namePart id="10A4CC10F656BA4AE8057F7B8A574003">Jiang, Zhuo-Yin</mods:namePart>
<mods:nameIdentifier id="D9B504583DFABE3BCE60A56BBC29C54B" type="ORCID">0000-0001-6825-4283</mods:nameIdentifier>
<mods:affiliation id="52CE194C2E7DB1C301AAECF3F0E1013B">State Key Laboratory of Biocontrol and Institute of Entomology, Sun Yat-sen University, Guangzhou 510275, China. zhuoyin _ jiang @ 126. com; https: // orcid. org / 0000 - 0001 - 6825 - 4283</mods:affiliation>
</mods:name>
<mods:name id="C3532DA38B7CB244E884748BF7A6E2AC" type="personal">
<mods:role id="63494F6852FE46ABF989F04A0505B6F9">
<mods:roleTerm id="A2AB1C5770C33E6BE1A3DBA9C8C180BD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C392642EB8557F04CBE3E3E6CE5109DD">Zhao, Shuang</mods:namePart>
<mods:affiliation id="F73105AAEACD397AB9167B8B27F616F4">Guangzhou Customs Technology Center, Nansha District, Guangzhou, China.</mods:affiliation>
</mods:name>
<mods:name id="6BEF44659539046AB7635CD0B0B71972" type="personal">
<mods:role id="8116BED5651482ACB49DFDD0C5925CE3">
<mods:roleTerm id="A2E7FD28284F919D982E9ECD32B2F340">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27FC4AB00EC175924BB9A7FA35859344">Yang, Zhen-Yu</mods:namePart>
<mods:affiliation id="E7622D17EB1D1E025BD28188E0F53311">Brilliant Landscaping and Gardening Company Limited, Macau.</mods:affiliation>
</mods:name>
<mods:name id="24BF55AA21A4666FE473EDF5E52D68BA" type="personal">
<mods:role id="DE6C5A51AB452C1B8F1533CBB89C816C">
<mods:roleTerm id="7F6DDD1613EA2C09827B114D642DCF48">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75970A50A300DFEC46AF89D7C0E2E541">Jia, Feng-Long</mods:namePart>
<mods:nameIdentifier id="325D7ECBEDA1FBE9C9B519DCC9435773" type="ORCID">0000-0003-2391-5038</mods:nameIdentifier>
<mods:affiliation id="AB0AB88FE9DCC25EE3E8C3C5BD591DD1">State Key Laboratory of Biocontrol and Institute of Entomology, Sun Yat-sen University, Guangzhou 510275, China. fenglongjia @ aliyun. com; lssjfl @ mail. sysu. edu. cn; http: // orcid. org / 0000 - 0003 - 2391 - 5038</mods:affiliation>
<mods:nameIdentifier id="836A2E123AF91E15F523B5253C22E626" type="email">fenglongjia@aliyun.com</mods:nameIdentifier>
</mods:name>
<mods:name id="AC290903D6C7568C760EF90421AC79BE" type="personal">
<mods:role id="64564AEF54493D3AA339C931ED55DA39">
<mods:roleTerm id="3BEED3732C50E607B9E395E2AFDAE058">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB4B58BEF4D9584C9892C80B0520F6C1">Hájek, Jiří</mods:namePart>
<mods:nameIdentifier id="91DBE49628392917F7DC98346A00021D" type="ORCID">0000-0001-5779-1542</mods:nameIdentifier>
<mods:affiliation id="3484D11F59749A98DA0510C65269A6FD">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic. jiri. hajek @ nm. cz; https: // orcid. org / 0000 - 0001 - 5779 - 1542</mods:affiliation>
<mods:nameIdentifier id="22939212CF46AFDE42507804C0A4E568" type="email">jiri.hajek@nm.cz</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="3E1AD03A5FB99F4EBF28EB25C2A1710A">text</mods:typeOfResource>
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<mods:titleInfo id="8EF397555F2FE57ADE2BEAC8EBB6AC3F">
<mods:title id="D7BE28E389A4263F93A5E43D72C870F4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B9D622C1B5247973F223C6F73109D044">
<mods:date id="7F8A29733392516BF2949F58BB1FB461">2022</mods:date>
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<mods:number id="ED843090035C2080B9AB6B508DD15E80">2022-04-04</mods:number>
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<mods:number id="E0702D0C655303FF73569AB0B5BDFA94">5124</mods:number>
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<mods:number id="6A4831DB3D4227607E9CE427F0B85E31">3</mods:number>
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<mods:classification id="CCC4266E49DED2706D47E04EC5D67ED1">journal article</mods:classification>
<mods:identifier id="3B56AECC71C20A9EF9FD6F5CB4BAFA44" type="CLB-Dataset">53387</mods:identifier>
<mods:identifier id="1B5195E464E2A433C2BE59AE3B0AB3A4" type="DOI">10.11646/zootaxa.5124.3.1</mods:identifier>
<mods:identifier id="7BBA987E5124F6603214631D93D06C06" type="GBIF-Dataset">de55f596-c349-4ca2-a826-0e2bad912aa3</mods:identifier>
<mods:identifier id="39006EBDD73B22788C32E56C97D284DF" type="ISSN">1175-5326</mods:identifier>
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<treatment id="291CAA4FBF48FF8B1DC5FAD09ABBF843" ID-DOI="http://doi.org/10.5281/zenodo.6410944" ID-GBIF-Taxon="194444076" ID-Zenodo-Dep="6410944" LSID="urn:lsid:plazi:treatment:291CAA4FBF48FF8B1DC5FAD09ABBF843" httpUri="http://treatment.plazi.org/id/291CAA4FBF48FF8B1DC5FAD09ABBF843" lastPageId="22" lastPageNumber="273" pageId="21" pageNumber="272">
<subSubSection id="E9AF48D2BF48FF881DC5FAD09B1DFAF1" box="[151,811,1363,1391]" pageId="21" pageNumber="272" type="nomenclature">
<paragraph id="A10A1B59BF48FF881DC5FAD09B1DFAF1" blockId="21.[151,811,1363,1427]" box="[151,811,1363,1391]" pageId="21" pageNumber="272">
<heading id="FA42AC35BF48FF881DC5FAD09B1DFAF1" bold="true" box="[151,811,1363,1391]" fontSize="11" level="1" pageId="21" pageNumber="272" reason="1">
<emphasis id="93C1C74BBF48FF881DC5FAD09B1DFAF1" bold="true" box="[151,811,1363,1391]" pageId="21" pageNumber="272">
<taxonomicName id="66B560DABF48FF881DC5FAD09AF0FAF0" authority="Hajek, Jiang &amp; Jia" authorityName="Hajek, Jiang &amp; Jia" authorityYear="2022" box="[151,710,1363,1391]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="272" phylum="Arthropoda" rank="species" species="tengchongensis" status="sp. nov.">
<emphasis id="93C1C74BBF48FF881DC5FAD099E4FAF1" bold="true" box="[151,466,1364,1391]" italics="true" pageId="21" pageNumber="272">Copelatus tengchongensis</emphasis>
Hájek, Jiang &amp; Jia
</taxonomicName>
<taxonomicNameLabel id="88F27A30BF48FF881F9CFAD19B1DFAF1" box="[718,811,1365,1391]" pageId="21" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E9AF48D2BF48FF881DC5FAFC994DFA0D" box="[151,379,1400,1427]" pageId="21" pageNumber="272" type="description">
<paragraph id="A10A1B59BF48FF881DC5FAFC994DFA0D" blockId="21.[151,811,1363,1427]" box="[151,379,1400,1427]" pageId="21" pageNumber="272">
(
<figureCitation id="398E07DCBF48FF881DCDFAFC9916FA0D" box="[159,288,1400,1427]" captionStart="FIGURES 21–24" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[207,1375,184,1914]" captionTargetId="figure-23@36.[204,1382,181,1920]" captionTargetPageId="36" captionText="FIGURES 21–24. Habitus of Copelatus in China. 21. C. sociennus (♂; Hengqin Island, Guangdong); 22. C. tenebrosus (♂; Hengqin Island, Guangdong); 23. C. tengchongensis sp. nov. (holotype ♂; Beihai, Yunnan); 24. C. tengchongensis sp. nov. (paratype ♀; Beihai, Yunnan). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6411112" httpUri="https://zenodo.org/record/6411112/files/figure.png" pageId="21" pageNumber="272">Figs 23–24</figureCitation>
,
<figureCitation id="398E07DCBF48FF881C7EFAFC9942FA0D" box="[300,372,1400,1427]" captionStart="FIGURES 63–70" captionStartId="41.[151,264,1403,1427]" captionTargetBox="[216,1366,200,1362]" captionTargetId="figure-18@41.[205,1383,181,1379]" captionTargetPageId="41" captionText="FIGURES 63–70. Male genitalia of Copelatus. 63, 65, 67, 69. median lobe in lateral view; 64, 66, 68, 70. paramere. 63–64. C. tengchongensis sp. nov. (holotype, Beihai, Yunnan); 65–66. C. weymarni (Honshu, Japan); 67–68. C. xiniudong sp. nov. (holotype, Xiniudong, Guizhou); 69–70. C. zimmermanni (Danxiashan, Guangdong). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6410764" httpUri="https://zenodo.org/record/6410764/files/figure.png" pageId="21" pageNumber="272">63–64</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E9AF48D2BF48FF881DC5FA459D5AF964" pageId="21" pageNumber="272" type="materials_examined">
<paragraph id="A10A1B59BF48FF881DC5FA4599AEFA61" blockId="21.[151,1436,1472,1537]" pageId="21" pageNumber="272">
<materialsCitation id="11DD1104BF48FF881DC5FA4599A5FA61" ID-GBIF-Occurrence="3746435387" country="China" elevation="1730" latitude="25.125" location="Zing Hai lake" longLatPrecision="124" longitude="98.555" municipality="Beihai Wetlands" pageId="21" pageNumber="272" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="93C1C74BBF48FF881DC5FA459900FA44" bold="true" box="[151,310,1472,1499]" pageId="21" pageNumber="272">
<typeStatus id="7E0EA5FBBF48FF881DC5FA4598E4FA45" box="[151,210,1473,1499]" pageId="21" pageNumber="272">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="D9A25BC9BF48FF881C6FFA4499B5FA44" box="[317,387,1472,1498]" name="China" pageId="21" pageNumber="272">China</collectingCountry>
,
<collectingRegion id="6371D5BBBF48FF881CDFFA459A66FA44" box="[397,592,1472,1499]" country="China" name="Yunnan" pageId="21" pageNumber="272">Yunnan Province</collectingRegion>
,
<locationDeviation id="B0B8027BBF48FF881F09FA459B62FA44" box="[603,852,1472,1499]" location="Tengchong" pageId="21" pageNumber="272">
<quantity id="664DB6BCBF48FF881F09FA459A94FA44" box="[603,674,1472,1499]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="272" unit="km" value="12.0">12 km</quantity>
NE Tengchong
</locationDeviation>
,
<collectingMunicipality id="416E8123BF48FF881E32FA449C2FFA44" box="[864,1049,1472,1498]" pageId="21" pageNumber="272">Beihai Wetlands</collectingMunicipality>
,
<location id="A46A4D82BF48FF881977FA449CF6FA44" LSID="urn:lsid:plazi:treatment:291CAA4FBF48FF8B1DC5FAD09ABBF843:A46A4D82BF48FF881977FA449CF6FA44" box="[1061,1216,1472,1498]" country="China" latitude="25.125" longLatPrecision="124" longitude="98.555" municipality="Beihai Wetlands" name="Zing Hai lake" pageId="21" pageNumber="272" stateProvince="Yunnan">Zing Hai lake</location>
env., ca.
<geoCoordinate id="C4817D9EBF48FF881878FA469DAAFA43" box="[1322,1436,1474,1501]" degrees="25" direction="north" minutes="07.5" orientation="latitude" pageId="21" pageNumber="272" precision="92" value="25.125">25°07.5′N</geoCoordinate>
<geoCoordinate id="C4817D9EBF48FF881DC5FA62993FF99F" box="[151,265,1510,1537]" degrees="98" direction="east" minutes="33.3" orientation="longitude" pageId="21" pageNumber="272" precision="92" value="98.555">98°33.3′E</geoCoordinate>
, ca.
<quantity id="664DB6BCBF48FF881C69FA6199A5FA61" box="[315,403,1509,1535]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="21" pageNumber="272" unit="m" value="1730.0">
<elevation id="2A98FC6ABF48FF881C69FA6199A5FA61" box="[315,403,1509,1535]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="21" pageNumber="272" unit="m" value="1730.0">1730 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="A10A1B59BF48FF881D95F9A99D5AF964" blockId="21.[151,1437,1580,2039]" pageId="21" pageNumber="272">
<emphasis id="93C1C74BBF48FF881D95F9A9994EF9D8" bold="true" box="[199,376,1580,1607]" pageId="21" pageNumber="272">Type material.</emphasis>
<materialsCitation id="11DD1104BF48FF881CD1F9A89CD0F911" ID-GBIF-Occurrence="3746435351" collectingDate="1999-10-27" collectionCode="NMPC" collectorName="J. Stastny" county="China" determinerName="Jia" location="Sic" municipality="Behai" pageId="21" pageNumber="272" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="7E0EA5FBBF48FF881CD1F9A899DDF9D8" box="[387,491,1580,1606]" pageId="21" pageNumber="272" type="holotype">Holotype</typeStatus>
<specimenCount id="B7B3D0D0BF48FF881CA7F9A99A3CF9D9" box="[501,522,1581,1607]" pageId="21" pageNumber="272" type="male">♂</specimenCount>
(
<collectionCode id="C7A4839CBF48FF881F4EF9A99A47F9D9" box="[540,625,1581,1607]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="21" pageNumber="272">NMPC</collectionCode>
), labelled: “
<collectingCounty id="486B63D5BF48FF881FAEF9A89B72F9D8" box="[764,836,1580,1606]" pageId="21" pageNumber="272">China</collectingCounty>
,
<collectingRegion id="6371D5BBBF48FF881E00F9A99BD8F9D9" box="[850,1006,1581,1607]" country="China" name="Yunnan" pageId="21" pageNumber="272">Yünnan prov.</collectingRegion>
/
<locationDeviation id="B0B8027BBF48FF881958F9A99D38F9D8" box="[1034,1294,1580,1607]" location="Tengchong" pageId="21" pageNumber="272">
<quantity id="664DB6BCBF48FF881958F9A99C62F9D8" box="[1034,1108,1580,1607]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="272" unit="km" value="10.0">10 km</quantity>
NE Tengchong
</locationDeviation>
;
<collectingMunicipality id="416E8123BF48FF88184EF9A89D6BF9D8" box="[1308,1373,1580,1606]" pageId="21" pageNumber="272">Behai</collectingMunicipality>
[
<emphasis id="93C1C74BBF48FF881823F9A99DAAF9D9" box="[1393,1436,1581,1607]" italics="true" pageId="21" pageNumber="272">
<location id="A46A4D82BF48FF881823F9A99DA2F9D9" LSID="urn:lsid:plazi:treatment:291CAA4FBF48FF8B1DC5FAD09ABBF843:A46A4D82BF48FF881823F9A99DA2F9D9" box="[1393,1428,1581,1607]" county="China" municipality="Behai" name="Sic" pageId="21" pageNumber="272" stateProvince="Yunnan">Sic</location>
!
</emphasis>
<location id="A46A4D82BF48FF881DC5F9D59889F9F5" LSID="urn:lsid:plazi:treatment:291CAA4FBF48FF8B1DC5FAD09ABBF843:A46A4D82BF48FF881DC5F9D59889F9F5" box="[151,191,1617,1643]" county="China" municipality="Behai" name="See" pageId="21" pageNumber="272" stateProvince="Yunnan">See</location>
the type locality.] /
<location id="A46A4D82BF48FF881CFBF9D49A08F9F4" LSID="urn:lsid:plazi:treatment:291CAA4FBF48FF8B1DC5FAD09ABBF843:A46A4D82BF48FF881CFBF9D49A08F9F4" box="[425,574,1616,1642]" county="China" municipality="Behai" name="ZingHai lake" pageId="21" pageNumber="272" stateProvince="Yunnan">ZingHai lake</location>
env; / small flowing pools, dense veget. /
<date id="D50B3D99BF48FF88197BF9D59C9FF9F5" box="[1065,1193,1617,1643]" pageId="21" pageNumber="272" value="1999-10-27">
<collectingDate id="C54FC471BF48FF88197BF9D59C9FF9F5" box="[1065,1193,1617,1643]" pageId="21" pageNumber="272" value="1999-10-27">27.10.1999</collectingDate>
</date>
, leg.
<collectorName id="0C407E8FBF48FF8819BBF9D59D6CF9F3" box="[1257,1370,1617,1645]" pageId="21" pageNumber="272">J. Šťastný</collectorName>
[p] //
<typeStatus id="7E0EA5FBBF48FF881DC5F9F1991DF911" box="[151,299,1653,1679]" pageId="21" pageNumber="272" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="66B560DABF48FF881C6DF9F199ECF911" authorityName="Erichson" authorityYear="1832" box="[319,474,1653,1679]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="93C1C74BBF48FF881C6DF9F199ECF911" box="[319,474,1653,1679]" italics="true" pageId="21" pageNumber="272">COPELATUS</emphasis>
</taxonomicName>
/
<taxonomicName id="66B560DABF48FF881CBCF9F09AADF910" authority="Jiang &amp; Zhao &amp; Yang &amp; Jia &amp; Hájek, 2022" authorityName="Jiang &amp; Zhao &amp; Yang &amp; Jia &amp; Hájek" authorityYear="2022" box="[494,667,1652,1678]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="272" phylum="Arthropoda" rank="species" species="tengchongensis" status="sp. nov.">
<emphasis id="93C1C74BBF48FF881CBCF9F09AADF910" box="[494,667,1652,1678]" italics="true" pageId="21" pageNumber="272">tengchongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88F27A30BF48FF881FF2F9F19AC0F911" box="[672,758,1653,1679]" pageId="21" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
/ Hájek, Jiang &amp;
<determinerName id="54D6F79DBF48FF881EE5F9F09BE0F910" box="[951,982,1652,1678]" pageId="21" pageNumber="272">Jia</determinerName>
det. 2021 [red label, p]”
</materialsCitation>
.
<materialsCitation id="11DD1104BF48FF8819A3F9F19A81F92D" ID-GBIF-Occurrence="3746435356" collectingDate="1999-10-27" collectionCode="JSCL, NMPC" collectorName="J. Stastny" county="China" determinerName="Jia" location="Sic" municipality="Behai" pageId="21" pageNumber="272" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="7E0EA5FBBF48FF8819A3F9F19D54F911" box="[1265,1378,1653,1679]" pageId="21" pageNumber="272" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="B7B3D0D0BF48FF88183EF9F19DA1F911" box="[1388,1431,1653,1679]" pageId="21" pageNumber="272" type="male">2 ♂</specimenCount>
,
<specimenCount id="B7B3D0D0BF48FF881DC5F91D98F7F92D" box="[151,193,1689,1715]" pageId="21" pageNumber="272" type="female">4 ♀</specimenCount>
, same label data as holotype (
<collectionCode id="C7A4839CBF48FF881F5FF91D9A79F92D" box="[525,591,1689,1715]" pageId="21" pageNumber="272">JSCL</collectionCode>
,
<collectionCode id="C7A4839CBF48FF881F09F91D9A86F92D" box="[603,688,1689,1715]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="21" pageNumber="272">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="11DD1104BF48FF881F93F91D9D5EF964" ID-GBIF-Occurrence="3746435425" collectingDate="2009-08-27" collectionCode="NMPC, SYSU, ZSMG" collectorName="D. W. Wrase" country="China" elevation="1750" latitude="25.58889" location="Gaoligong Shan" longLatPrecision="21" longitude="98.6725" pageId="21" pageNumber="272" specimenCount="14" specimenCount-female="7" specimenCount-male="7" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="B7B3D0D0BF48FF881F93F91D9ADBF92D" box="[705,749,1689,1715]" pageId="21" pageNumber="272" type="male">7 ♂</specimenCount>
,
<specimenCount id="B7B3D0D0BF48FF881FAAF91D9B17F92D" box="[760,801,1689,1715]" pageId="21" pageNumber="272" type="female">7 ♀</specimenCount>
, labelled: “
<collectingCountry id="D9A25BC9BF48FF881ECDF91D9BCFF92D" box="[927,1017,1689,1715]" name="China" pageId="21" pageNumber="272">CHINA</collectingCountry>
(
<collectingRegion id="6371D5BBBF48FF881955F91D9C56F92D" box="[1031,1120,1689,1715]" country="China" name="Yunnan" pageId="21" pageNumber="272">Yunnan</collectingRegion>
) / Baoshan Pref.,
<location id="A46A4D82BF48FF881874F91C98F9F948" LSID="urn:lsid:plazi:treatment:291CAA4FBF48FF8B1DC5FAD09ABBF843:A46A4D82BF48FF881874F91C98F9F948" country="China" latitude="25.58889" longLatPrecision="21" longitude="98.6725" name="Gaoligong Shan" pageId="21" pageNumber="272" stateProvince="Yunnan">Gaoligong Shan</location>
/
<locationDeviation id="B0B8027BBF48FF881DBAF93899C1F948" box="[232,503,1724,1751]" location="Tengchong" pageId="21" pageNumber="272">
<quantity id="664DB6BCBF48FF881DBAF9389907F948" box="[232,305,1724,1751]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="21" pageNumber="272" unit="km" value="65.0">65 km</quantity>
NNE Tengchong
</locationDeviation>
<quantity id="664DB6BCBF48FF881CADF9389A60F949" box="[511,598,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="21" pageNumber="272" unit="m" value="1750.0">
<elevation id="2A98FC6ABF48FF881CADF9389A60F949" box="[511,598,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="21" pageNumber="272" unit="m" value="1750.0">1750 m</elevation>
</quantity>
/
<geoCoordinate id="C4817D9EBF48FF881F3CF93A9AC0F947" box="[622,758,1726,1753]" degrees="25" direction="north" minutes="35" orientation="latitude" pageId="21" pageNumber="272" precision="15" seconds="20" value="25.58889">25°35′20″N</geoCoordinate>
,
<geoCoordinate id="C4817D9EBF48FF881E51F93A9BB3F947" box="[771,901,1726,1753]" degrees="98" direction="east" minutes="40" orientation="longitude" pageId="21" pageNumber="272" precision="15" seconds="21" value="98.6725">98°40′21″E</geoCoordinate>
/ (small brook at edge of sec. / mixed forest) /
<date id="D50B3D99BF48FF881DC5F9659911F965" box="[151,295,1761,1787]" pageId="21" pageNumber="272" value="2009-08-27">
<collectingDate id="C54FC471BF48FF881DC5F9659911F965" box="[151,295,1761,1787]" pageId="21" pageNumber="272" value="2009-08-27">27.VIII.2009</collectingDate>
</date>
<collectorName id="0C407E8FBF48FF881C75F965999CF965" box="[295,426,1761,1787]" pageId="21" pageNumber="272">D.W.Wrase</collectorName>
(10G) [p]” (
<collectionCode id="C7A4839CBF48FF881F64F9659ABCF965" box="[566,650,1761,1787]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="21" pageNumber="272">NMPC</collectionCode>
,
<collectionCode id="C7A4839CBF48FF881FC7F9659AE9F965" box="[661,735,1761,1787]" pageId="21" pageNumber="272">SYSU</collectionCode>
,
<collectionCode id="C7A4839CBF48FF881FB9F9659B08F965" box="[747,830,1761,1787]" pageId="21" pageNumber="272">ZSMG</collectionCode>
). All
<typeStatus id="7E0EA5FBBF48FF881E2BF9659BD2F965" box="[889,996,1761,1787]" pageId="21" pageNumber="272" type="paratype">paratypes</typeStatus>
with the respective
<typeStatus id="7E0EA5FBBF48FF881994F9659D10F965" box="[1222,1318,1761,1787]" pageId="21" pageNumber="272" type="paratype">paratype</typeStatus>
label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E9AF48D2BF48FF8B1D95F8809D7EF9EB" lastPageId="22" lastPageNumber="273" pageId="21" pageNumber="272" type="description">
<paragraph id="A10A1B59BF48FF881D95F8809AC8F8DC" blockId="21.[151,1437,1580,2039]" pageId="21" pageNumber="272">
<emphasis id="93C1C74BBF48FF881D95F8809AB3F881" bold="true" box="[199,645,1796,1823]" pageId="21" pageNumber="272">
Description of male
<typeStatus id="7E0EA5FBBF48FF881CE7F8819A28F881" box="[437,542,1797,1823]" pageId="21" pageNumber="272" type="holotype">holotype</typeStatus>
.
<emphasis id="93C1C74BBF48FF881F7BF8809AB3F881" bold="true" box="[553,645,1796,1823]" italics="true" pageId="21" pageNumber="272">Habitus</emphasis>
</emphasis>
(
<figureCitation id="398E07DCBF48FF881FC1F8809AD5F881" box="[659,739,1796,1823]" captionStart="FIGURES 21–24" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[207,1375,184,1914]" captionTargetId="figure-23@36.[204,1382,181,1920]" captionTargetPageId="36" captionText="FIGURES 21–24. Habitus of Copelatus in China. 21. C. sociennus (♂; Hengqin Island, Guangdong); 22. C. tenebrosus (♂; Hengqin Island, Guangdong); 23. C. tengchongensis sp. nov. (holotype ♂; Beihai, Yunnan); 24. C. tengchongensis sp. nov. (paratype ♀; Beihai, Yunnan). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6411112" httpUri="https://zenodo.org/record/6411112/files/figure.png" pageId="21" pageNumber="272">Fig. 23</figureCitation>
) oblong-oval, with continuous outline, broadest in anterior 1/3 of elytral length, slightly convex. Dorsal surface shiny.
</paragraph>
<paragraph id="A10A1B59BF48FF881D95F8C899E2F84C" blockId="21.[151,1437,1580,2039]" pageId="21" pageNumber="272">
<emphasis id="93C1C74BBF48FF881D95F8C8996CF8F9" bold="true" box="[199,346,1868,1895]" italics="true" pageId="21" pageNumber="272">Colouration.</emphasis>
Head brown blackish, clypeus reddish brown; pronotum brown blackish, laterally broadly orangebrown, anterior and posterior margin orange translucent; elytra dark brown, paler laterally, base with distinct, orange, comb-like shaped transverse band reaching neither lateral margin nor suture; appendages orange-brown; ventral side brown blackish.
</paragraph>
<paragraph id="A10A1B59BF48FF8B1D95F8589B51FEDF" blockId="21.[151,1437,1580,2039]" lastBlockId="22.[151,1437,151,2013]" lastPageId="22" lastPageNumber="273" pageId="21" pageNumber="272">
<emphasis id="93C1C74BBF48FF881D95F858993AF869" bold="true" box="[199,268,2012,2039]" italics="true" pageId="21" pageNumber="272">Head.</emphasis>
Moderately broad, ca. 0.63× width of pronotum, semicircular. Anterior margin of clypeus indistinctly concave. Antenna with antennomeres long and slender. Reticulation consisting of moderately deeply impressed polygonal isodiametric meshes. Punctation double, consisting of coarse setigerous punctures, and fine punctures spread sparsely on surface; row of coarse punctures present alongside inner margin of eyes, several punctures present at frontal level of eyes, and antero-laterally to eyes in fronto-clypeal depressions; fine punctures smaller and sparser on clypeus, becoming larger and more dense posteriorly.
</paragraph>
<paragraph id="A10A1B59BF4BFF8B1D95FECF9A32FD87" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95FECF9975FEFB" bold="true" box="[199,323,331,357]" italics="true" pageId="22" pageNumber="273">Pronotum.</emphasis>
Transverse (width/length ratio = 2.69), broadest between posterior angles, lateral margins moderately curved. Lateral sides with beading very thin and indistinct. Reticulation similar to that of head. Punctation similar to that of head; rows of coarse setigerous punctures present along anterior margin, laterally close to sides, several punctures present also in shallow baso-lateral depressions along basal margin.Pronotum baso-laterally and especially laterally with irregular wrinkles and very few short longitudinal strioles.
<collectingRegion id="6371D5BBBF4BFF8B1E86FE5F9C29FE6B" box="[980,1055,475,501]" country="France" name="Centre" pageId="22" pageNumber="273">Centre</collectingRegion>
of disc with shallowly impressed medial longitudinal smooth line.
</paragraph>
<paragraph id="A10A1B59BF4BFF8B1D95FDA69BFAFC8B" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95FDA69923FDA3" bold="true" box="[199,277,546,573]" italics="true" pageId="22" pageNumber="273">Elytra.</emphasis>
Base of elytra as broad as pronotal base; lateral margins of elytra slightly diverging in basal third, then distinctly narrowing to apex. Six discal and one submarginal longitudinal striae present on each elytron: striae 1 and 5 beginning slightly posteriorly to base; striae 2–4, 6 beginning at base; stria 1 longest, ending close to apex; striae 2–5 ending subapically, even striae generally shorter than odd striae; stria 6 ending at apical fourth; submarginal stria long, beginning before elytral mid-length and ending at apical fourth. Reticulation similar to that of head and pronotum, but less impressed. Punctation consisting of coarse setigerous punctures and very fine sparse punctures; coarse punctures present along elytral striae and lateral margins of elytra.
</paragraph>
<paragraph id="A10A1B59BF4BFF8B1D95FC9B9C85FCC3" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95FC9B9933FCA7" bold="true" box="[199,261,799,825]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95FC9B98CBFCA7" bold="true" box="[199,253,799,825]" italics="true" pageId="22" pageNumber="273">Legs</emphasis>
.
</emphasis>
Protibia modified, angled near base, distinctly broadened anteriorly, club shaped. Pro- and mesotarsomeres 1–3 distinctly broadened, with four rows of adhesive setae on their ventral side; claws simple.
</paragraph>
<paragraph id="A10A1B59BF4BFF8B1D95FCE29A73FB5B" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95FCE29961FC1F" bold="true" box="[199,343,870,897]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95FCE29966FC1F" bold="true" box="[199,336,870,897]" italics="true" pageId="22" pageNumber="273">Ventral side</emphasis>
.
</emphasis>
Prosternum sinuate anteriorly, obtusely keeled medially. Prosternal process shortly lanceolate, in cross-section convex, apex obtuse; process distinctly bordered laterally; reticulation not perceptible. Metaventrite with microsculpture consisting of polygonal meshes, laterally with oblique strioles; lateral parts of metaventrite (“metasternal wings”) tongue-shaped, slender. Metacoxal lines nearly complete, absent only close to metaventrite. Metacoxal plates covered with long, oblique strioles; reticulation consisting of elongate, longitudinal or oblique polygonal meshes. Metacoxal processes rounded at posterior margin. Abdominal ventrites I–II with longitudinal strioles; ventrites III–IV with oblique strioles laterally. Tuft of setae present antero-medially on ventrites III–V; ventrite VI with setigerous punctures laterally on either side.Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I–II, oblique on ventrite III and transverse on ventrites IV–VI. Punctation consisting of fine, sparsely distributed punctures.
</paragraph>
<paragraph id="A10A1B59BF4BFF8B1D95FB4A9C85FAAC" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95FB4A994FFB77" bold="true" box="[199,377,1230,1257]" italics="true" pageId="22" pageNumber="273">Male genitalia.</emphasis>
Median lobe of aedeagus in lateral view simple, sickle-shaped, broadest at mid-length due to shallow swelling on ventral side; in apical third narrowing continuously to pointed apex (
<figureCitation id="398E07DCBF4BFF8B19E9FB779D27FA90" box="[1211,1297,1267,1294]" captionStart="FIGURES 63–70" captionStartId="41.[151,264,1403,1427]" captionTargetBox="[216,1366,200,1362]" captionTargetId="figure-18@41.[205,1383,181,1379]" captionTargetPageId="41" captionText="FIGURES 63–70. Male genitalia of Copelatus. 63, 65, 67, 69. median lobe in lateral view; 64, 66, 68, 70. paramere. 63–64. C. tengchongensis sp. nov. (holotype, Beihai, Yunnan); 65–66. C. weymarni (Honshu, Japan); 67–68. C. xiniudong sp. nov. (holotype, Xiniudong, Guizhou); 69–70. C. zimmermanni (Danxiashan, Guangdong). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6410764" httpUri="https://zenodo.org/record/6410764/files/figure.png" pageId="22" pageNumber="273">Fig. 63</figureCitation>
). Parameres moderately broad, ‘D’-shaped; apex short and broad; apical lobes long, club-shaped (
<figureCitation id="398E07DCBF4BFF8B1900FA939C90FAAC" box="[1106,1190,1303,1330]" captionStart="FIGURES 63–70" captionStartId="41.[151,264,1403,1427]" captionTargetBox="[216,1366,200,1362]" captionTargetId="figure-18@41.[205,1383,181,1379]" captionTargetPageId="41" captionText="FIGURES 63–70. Male genitalia of Copelatus. 63, 65, 67, 69. median lobe in lateral view; 64, 66, 68, 70. paramere. 63–64. C. tengchongensis sp. nov. (holotype, Beihai, Yunnan); 65–66. C. weymarni (Honshu, Japan); 67–68. C. xiniudong sp. nov. (holotype, Xiniudong, Guizhou); 69–70. C. zimmermanni (Danxiashan, Guangdong). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6410764" httpUri="https://zenodo.org/record/6410764/files/figure.png" pageId="22" pageNumber="273">Fig. 64</figureCitation>
).
</paragraph>
<paragraph id="A10A1B59BF4BFF8B1D95FABF9A59FA5F" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95FABF9913FACB" bold="true" box="[199,293,1339,1365]" pageId="22" pageNumber="273">Female.</emphasis>
Identical to male in habitus. Protibia simple, not angled basally and only slightly broadened distally; pro- and mesotarsomeres not broadened, without adhesive setae. Females dimorphic; smooth form differs from male in more developed striolation laterally on pronotum; striolate form submatt, with long strioles covering pronotum and elytra except for apical fifth (
<figureCitation id="398E07DCBF4BFF8B1F5CFA239A55FA5F" box="[526,611,1447,1473]" captionStart="FIGURES 21–24" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[207,1375,184,1914]" captionTargetId="figure-23@36.[204,1382,181,1920]" captionTargetPageId="36" captionText="FIGURES 21–24. Habitus of Copelatus in China. 21. C. sociennus (♂; Hengqin Island, Guangdong); 22. C. tenebrosus (♂; Hengqin Island, Guangdong); 23. C. tengchongensis sp. nov. (holotype ♂; Beihai, Yunnan); 24. C. tengchongensis sp. nov. (paratype ♀; Beihai, Yunnan). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6411112" httpUri="https://zenodo.org/record/6411112/files/figure.png" pageId="22" pageNumber="273">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="A10A1B59BF4BFF8B1D95FA4E9B6AF9B3" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95FA4E997BFA7A" bold="true" box="[199,333,1482,1508]" pageId="22" pageNumber="273">Variability.</emphasis>
All specimens of the
<typeStatus id="7E0EA5FBBF4BFF8B1F6EFA4F9A5AFA7B" box="[572,620,1483,1509]" pageId="22" pageNumber="273">type</typeStatus>
series are rather uniform. There is only slight variability in dorsal surface colouration, especially in extent and brightness of pale basal band on elytra. Minor variability can be seen also in number and length of strioles presenting laterally on pronotum.
</paragraph>
<paragraph id="A10A1B59BF4BFF8B1D95F9B39D7EF9EB" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95F9B39948F9CF" bold="true" box="[199,382,1591,1617]" pageId="22" pageNumber="273">Measurements.</emphasis>
TL:
<quantity id="664DB6BCBF4BFF8B1CEBF9B29A75F9CF" box="[441,579,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.5" metricValueMin="4.7" pageId="22" pageNumber="273" unit="mm" value="5.1" valueMax="5.5" valueMin="4.7">4.7–5.5 mm</quantity>
(mean value: 5.3 ±
<quantity id="664DB6BCBF4BFF8B1E7FF9B39BBFF9CF" box="[813,905,1591,1617]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="273" unit="mm" value="0.2">0.2 mm</quantity>
);
<typeStatus id="7E0EA5FBBF4BFF8B1ECDF9B39C33F9CF" box="[927,1029,1591,1617]" pageId="22" pageNumber="273" type="holotype">holotype</typeStatus>
: 5.0 mm. TL-h:
<quantity id="664DB6BCBF4BFF8B1996F9B29D7BF9CF" box="[1220,1357,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="5.1" metricValueMin="4.2" pageId="22" pageNumber="273" unit="mm" value="4.65" valueMax="5.1" valueMin="4.2">4.2–5.1 mm</quantity>
(mean value: 4.8 ±
<quantity id="664DB6BCBF4BFF8B1C71F9DF994BF9EB" box="[291,381,1627,1653]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="273" unit="mm" value="0.2">0.2 mm</quantity>
);
<typeStatus id="7E0EA5FBBF4BFF8B1CC3F9DF99C1F9EB" box="[401,503,1627,1653]" pageId="22" pageNumber="273" type="holotype">holotype</typeStatus>
:
<quantity id="664DB6BCBF4BFF8B1F53F9DE9A6CF9EB" box="[513,602,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="22" pageNumber="273" unit="mm" value="4.5">4.5 mm</quantity>
. MW:
<quantity id="664DB6BCBF4BFF8B1FF4F9DF9B1BF9EB" box="[678,813,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="22" pageNumber="273" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3–2.8 mm</quantity>
(mean value: 2.6 ±
<quantity id="664DB6BCBF4BFF8B195EF9DF9C50F9EB" box="[1036,1126,1627,1653]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="273" unit="mm" value="0.1">0.1 mm</quantity>
);
<typeStatus id="7E0EA5FBBF4BFF8B1928F9DF9CD6F9EB" box="[1146,1248,1627,1653]" pageId="22" pageNumber="273" type="holotype">holotype</typeStatus>
:
<quantity id="664DB6BCBF4BFF8B19B9F9DE9D72F9EB" box="[1259,1348,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="22" pageNumber="273" unit="mm" value="2.5">2.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="E9AF48D2BF4BFF8B1D95F9FA99E5F8D0" pageId="22" pageNumber="273" type="diagnosis">
<paragraph id="A10A1B59BF4BFF8B1D95F9FA99E5F8D0" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95F9FA99F8F906" bold="true" box="[199,462,1662,1688]" pageId="22" pageNumber="273">Differential diagnosis.</emphasis>
Based on the presence of six dorsal striae and a submarginal stria on each elytron, the new species can be classified within
<taxonomicName id="66B560DABF4BFF8B1F13F9279ACBF922" authorityName="Regimbart" authorityYear="1899" box="[577,765,1699,1725]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="irinus">
<emphasis id="93C1C74BBF4BFF8B1F13F9279ACBF922" box="[577,765,1699,1725]" italics="true" pageId="22" pageNumber="273">Copelatus irinus</emphasis>
</taxonomicName>
species group. Elongate habitus, complete elytral striation and male median lobe without distinct process place
<taxonomicName id="66B560DABF4BFF8B1E5AF9439BECF97F" authority="Jiang &amp; Zhao &amp; Yang &amp; Jia &amp; Hájek, 2022" authorityName="Jiang &amp; Zhao &amp; Yang &amp; Jia &amp; Hájek" authorityYear="2022" box="[776,986,1735,1761]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="tengchongensis" status="sp. nov.">
<emphasis id="93C1C74BBF4BFF8B1E5AF9439BECF97F" box="[776,986,1735,1761]" italics="true" pageId="22" pageNumber="273">C. tengchongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88F27A30BF4BFF8B1EB7F9439C76F97F" box="[997,1088,1735,1761]" pageId="22" pageNumber="273" rank="species">sp. nov.</taxonomicNameLabel>
in the
<taxonomicName id="66B560DABF4BFF8B19CAF9439D1CF97E" authorityName="Sharp" authorityYear="1884" box="[1176,1322,1735,1761]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="93C1C74BBF4BFF8B19CAF9439D1CF97E" box="[1176,1322,1735,1761]" italics="true" pageId="22" pageNumber="273">C. japonicus</emphasis>
</taxonomicName>
complex. Within the complex, the new species can be undoubtedly recognised only based on the shape of male genitalia:
<taxonomicName id="66B560DABF4BFF8B18D1F96F9972F8B7" authority="Jiang &amp; Zhao &amp; Yang &amp; Jia &amp; Hájek, 2022" authorityName="Jiang &amp; Zhao &amp; Yang &amp; Jia &amp; Hájek" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="tengchongensis" status="sp. nov.">
<emphasis id="93C1C74BBF4BFF8B18D1F96F9972F8B7" italics="true" pageId="22" pageNumber="273">C. tengchongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88F27A30BF4BFF8B1C19F88B9995F8B7" box="[331,419,1807,1833]" pageId="22" pageNumber="273" rank="species">sp. nov.</taxonomicNameLabel>
is characteristic with the median lobe broadest in midpart and the apical part only slightly narrowing to apex (
<figureCitation id="398E07DCBF4BFF8B1C20F8B799F0F8D0" box="[370,454,1843,1870]" captionStart="FIGURES 63–70" captionStartId="41.[151,264,1403,1427]" captionTargetBox="[216,1366,200,1362]" captionTargetId="figure-18@41.[205,1383,181,1379]" captionTargetPageId="41" captionText="FIGURES 63–70. Male genitalia of Copelatus. 63, 65, 67, 69. median lobe in lateral view; 64, 66, 68, 70. paramere. 63–64. C. tengchongensis sp. nov. (holotype, Beihai, Yunnan); 65–66. C. weymarni (Honshu, Japan); 67–68. C. xiniudong sp. nov. (holotype, Xiniudong, Guizhou); 69–70. C. zimmermanni (Danxiashan, Guangdong). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6410764" httpUri="https://zenodo.org/record/6410764/files/figure.png" pageId="22" pageNumber="273">Fig. 63</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E9AF48D2BF4BFF8B1D95F8D39CB4F80B" pageId="22" pageNumber="273" type="etymology">
<paragraph id="A10A1B59BF4BFF8B1D95F8D39CB4F80B" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95F8D39978F8EF" bold="true" box="[199,334,1879,1905]" pageId="22" pageNumber="273">Etymology.</emphasis>
The new species is named after its area of occurrence – Tengchong County, where both known localities of the species lie. The specific epithet is an adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="E9AF48D2BF4BFF8B1D95F81A9ABBF843" pageId="22" pageNumber="273" type="distribution">
<paragraph id="A10A1B59BF4BFF8B1D95F81A9ABBF843" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="273">
<emphasis id="93C1C74BBF4BFF8B1D95F81A9957F826" bold="true" box="[199,353,1950,1976]" pageId="22" pageNumber="273">Distribution.</emphasis>
The species is so far known only from two close localities on western side of the Gaoligong mountains, western
<collectingRegion id="6371D5BBBF4BFF8B1C25F84799E6F843" box="[375,464,1987,2013]" country="China" name="Yunnan" pageId="22" pageNumber="273">Yunnan</collectingRegion>
,
<collectingCountry id="D9A25BC9BF4BFF8B1C88F8479A2BF843" box="[474,541,1987,2013]" name="China" pageId="22" pageNumber="273">China</collectingCountry>
(
<figureCitation id="398E07DCBF4BFF8B1F7FF8479AB6F843" box="[557,640,1987,2013]" captionStart="FIGURE 83" captionStartId="44.[151,250,2008,2032]" captionTargetBox="[273,1315,1132,1980]" captionTargetId="figure-39@44.[267,1320,1127,1984]" captionTargetPageId="44" captionText="FIGURE 83. Map of distribution of Copelatus in China Mainland." figureDoi="http://doi.org/10.5281/zenodo.6410823" httpUri="https://zenodo.org/record/6410823/files/figure.png" pageId="22" pageNumber="273">Fig. 83</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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