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<document id="6488ED0848BFB06F76B9D761EFF398BC" ID-CLB-Dataset="57399" ID-DOI="10.5852/ejt.2023.878.2153" ID-GBIF-Dataset="0dfb60c4-1de3-4062-9443-15b55e7e0c88" ID-ISSN="2118-9773" ID-Zenodo-Dep="8110550" ID-ZooBank="07EFE4B6-4184-41C7-82A5-747BF67BA093" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1688395572616" checkinUser="felipe" docAuthor="Grichanov, Igor Ya." docDate="2023" docId="282DC677E919FFB9C028CB6CFEFB7F2A" docLanguage="en" docName="ejt-878_1-52.pdf" docOrigin="European Journal of Taxonomy 878" docSource="http://zoobank.org/07efe4b6-4184-41c7-82a5-747bf67ba093" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Thinophilus manambato Grichanov 2023, sp. nov." docType="treatment" docUuid="C2866EE5-976A-4775-859C-BBD06301A798" docUuidSource="ZooBank" docVersion="11" lastPageNumber="36" masterDocId="D414BE0FE938FF9AC27ACA64FFC27B3F" masterDocTitle="A review of the Afrotropical Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae), with the descriptions of ten new species" masterLastPageNumber="52" masterPageNumber="1" pageNumber="34" updateTime="1699533079317" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F96197946051E75C5328B1B200C272A3">A review of the Afrotropical Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae), with the descriptions of ten new species</mods:title>
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<mods:namePart id="C6895868711F63FA66D850BA25506DBE">Grichanov, Igor Ya.</mods:namePart>
<mods:affiliation id="67CB702BC141D40BC74F8AEB00E3165C">All-Russian Institute of Plant Protection, Podbelskogo 3, St Petersburg, Pushkin, 196608, Russia.</mods:affiliation>
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<mods:title id="8DB70A9CB762AFEDFF129AF1823B6DD6">European Journal of Taxonomy</mods:title>
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<mods:part id="974EC050AF0CEA468C6E3C8650C0E31A">
<mods:date id="F011FD1839F643EFA61BF1CED6D846B0">2023</mods:date>
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<mods:number id="63189802985BB8B834C4157278659F12">2023-07-03</mods:number>
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<treatment id="282DC677E919FFB9C028CB6CFEFB7F2A" ID-DOI="http://doi.org/10.5281/zenodo.8119195" ID-GBIF-Taxon="210883684" ID-Zenodo-Dep="8119195" LSID="urn:lsid:zoobank.org:act:C2866EE5-976A-4775-859C-BBD06301A798" httpUri="http://treatment.plazi.org/id/282DC677E919FFB9C028CB6CFEFB7F2A" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
<subSubSection id="E89E24EAE919FFBBC028CB6CFC207A1B" box="[594,994,264,292]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="A03B7761E919FFBBC028CB6CFC207A1B" blockId="33.[370,1217,264,362]" box="[594,994,264,292]" pageId="33" pageNumber="34">
<heading id="FB73C00DE919FFBBC028CB6CFC207A1B" box="[594,994,264,292]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="34" reason="2">
<taxonomicName id="67840CE2E919FFBBC028CB6CFC427A1C" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[594,896,264,291]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="manambato" status="sp. nov.">
<emphasis id="92F0AB73E919FFBBC028CB6CFC427A1C" bold="true" box="[594,896,264,291]" italics="true" pageId="33" pageNumber="34">Thinophilus manambato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E919FFBBC1FDCB6EFC207A1B" box="[903,994,266,292]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE919FFBBC308CB48FB037A78" box="[370,1217,300,327]" pageId="33" pageNumber="34" type="description">
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="E89E24EAE919FFBBC08BCB2BFC807A55" box="[753,834,335,362]" pageId="33" pageNumber="34" type="description">
<paragraph id="A03B7761E919FFBBC08BCB2BFC807A55" blockId="33.[370,1217,264,362]" box="[753,834,335,362]" pageId="33" pageNumber="34">
<figureCitation id="38BF6BE4E919FFBBC08BCB2BFC807A55" box="[753,834,335,362]" captionStart="Fig" captionStartId="34.[189,232,1892,1918]" captionTargetBox="[216,1371,544,1852]" captionTargetId="figure-84@34.[216,1371,544,1854]" captionTargetPageId="34" captionText="Fig. 11. Thinophilus manambato sp. nov. AD. Holotype, ♂ (ZMUM). EG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna. D. Wing. E. Hypopygium after maceration, lateral view. F. Distal appendages of hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110572" httpUri="https://zenodo.org/record/8110572/files/figure.png" pageId="33" pageNumber="34">Fig. 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE919FFBBC2C7CBF5FE1B7992" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="A03B7761E919FFBBC2C7CBF5FEF57A94" blockId="33.[189,311,401,427]" box="[189,311,401,427]" pageId="33" pageNumber="34">
<heading id="FB73C00DE919FFBBC2C7CBF5FEF57A94" bold="true" box="[189,311,401,427]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="92F0AB73E919FFBBC2C7CBF5FEF57A94" bold="true" box="[189,311,401,427]" pageId="33" pageNumber="34">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E919FFBBC2C7CBDBFE1B7992" blockId="33.[188,1399,447,685]" pageId="33" pageNumber="34">
<taxonomicName id="67840CE2E919FFBBC2C7CBDBFE1C7AE6" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[189,478,447,473]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="manambato" status="sp. nov.">
<emphasis id="92F0AB73E919FFBBC2C7CBDBFE1C7AE6" box="[189,478,447,473]" italics="true" pageId="33" pageNumber="34">Thinophilus manambato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E919FFBBC39ECBA4FDFC7AE5" box="[484,574,448,474]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
is close in habitus to
<taxonomicName id="67840CE2E919FFBBC14DCBA4FC787AE6" authorityName="Curran" authorityYear="1926" box="[823,954,448,473]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="92F0AB73E919FFBBC14DCBA4FC787AE6" box="[823,954,448,473]" italics="true" pageId="33" pageNumber="34">T. capensis</emphasis>
</taxonomicName>
,
<taxonomicName id="67840CE2E919FFBBC1B3CBA4FBA67AE6" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[969,1124,448,473]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cataractae" status="sp. nov.">
<emphasis id="92F0AB73E919FFBBC1B3CBA4FBA67AE6" box="[969,1124,448,473]" italics="true" pageId="33" pageNumber="34">T. cataractae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E919FFBBC610CBA4FB067AE5" box="[1130,1220,448,474]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="67840CE2E919FFBBC681CBA4FAB47AE6" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[1275,1398,447,473]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fluvialis" status="sp. nov.">
<emphasis id="92F0AB73E919FFBBC681CBA4FAB47AE6" box="[1275,1398,447,473]" italics="true" pageId="33" pageNumber="34">T. fluvialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E919FFBBC2C7CB87FEDB7AC2" box="[189,281,483,509]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
(see key above), differing in fore coxa with mainly white setae, with at most 23 black apical bristles; mid coxa with white setae and one black bristle. The last three species have fore coxa with mainly black bristles and setae; mid coxa with black bristle and setae. The hypopygium of
<taxonomicName id="67840CE2E919FFBBC6B7C84EFAB7797C" authorityName="Grichanov" authorityYear="2023" box="[1229,1397,553,579]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="manambato">
<emphasis id="92F0AB73E919FFBBC6B7C84EFAB7797C" box="[1229,1397,553,579]" italics="true" pageId="33" pageNumber="34">T. manambato</emphasis>
</taxonomicName>
is similar to that in
<taxonomicName id="67840CE2E919FFBBC3DFC829FDE27959" authorityName="Grichanov" authorityYear="2023" box="[421,544,588,614]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fluvialis">
<emphasis id="92F0AB73E919FFBBC3DFC829FDE27959" box="[421,544,588,614]" italics="true" pageId="33" pageNumber="34">T. fluvialis</emphasis>
</taxonomicName>
, but the surstylus of the latter is thin (lateral view), bearing about seven short spine-like apicoventral setae; the surstylus of
<taxonomicName id="67840CE2E919FFBBC15BC814FC0E79B6" authorityName="Grichanov" authorityYear="2023" box="[801,972,623,649]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="manambato">
<emphasis id="92F0AB73E919FFBBC15BC814FC0E79B6" box="[801,972,623,649]" italics="true" pageId="33" pageNumber="34">T. manambato</emphasis>
</taxonomicName>
is broad (lateral view), with simple ventral and apical setae.
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE919FFBBC2C7C8B0FE08787F" pageId="33" pageNumber="34" type="etymology">
<paragraph id="A03B7761E919FFBBC2C7C8B0FE8779D1" blockId="33.[189,325,724,750]" box="[189,325,724,750]" pageId="33" pageNumber="34">
<heading id="FB73C00DE919FFBBC2C7C8B0FE8779D1" bold="true" box="[189,325,724,750]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="92F0AB73E919FFBBC2C7C8B0FE8779D1" bold="true" box="[189,325,724,750]" pageId="33" pageNumber="34">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E919FFBBC2C7C966FE08787F" blockId="33.[189,1399,770,832]" pageId="33" pageNumber="34">
The specific epithet refers to the Manambato village at the Lake Rasoabe near
<collectingRegion id="6240B983E919FFBBC617C966FB2E7823" box="[1133,1260,770,796]" country="Madagascar" name="Toamasina" pageId="33" pageNumber="34">Toamasina</collectingRegion>
city, where the
<typeStatus id="7F3FC9C3E919FFBBC290C942FEDF787F" box="[234,285,806,832]" pageId="33" pageNumber="34">type</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE919FFBBC2C7C903FC397F64" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="A03B7761E919FFBBC2C7C903FE6F78BE" blockId="33.[189,429,871,897]" box="[189,429,871,897]" pageId="33" pageNumber="34">
<heading id="FB73C00DE919FFBBC2C7C903FE6F78BE" bold="true" box="[189,429,871,897]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="92F0AB73E919FFBBC2C7C903FE6F78BE" bold="true" box="[189,429,871,897]" pageId="33" pageNumber="34">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E919FFBBC290C9F2FDC578C8" blockId="33.[189,1398,918,1015]" pageId="33" pageNumber="34">
<materialsCitation id="10EC7D3CE919FFBBC290C9F2FDC578C8" ID-GBIF-Occurrence="4145165354" collectingDate="2012-11-24" collectingDateMax="2012-11-30" collectingDateMin="2012-11-24" collectionCode="ZMUM" collectorName="A. Medvedev" country="Madagascar" latitude="-18.75" location="Manambato" longLatPrecision="765" longitude="49.15" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<emphasis id="92F0AB73E919FFBBC290C9F2FE9F788F" bold="true" box="[234,349,918,944]" pageId="33" pageNumber="34">
<typeStatus id="7F3FC9C3E919FFBBC290C9F2FE9F788F" box="[234,349,918,944]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="D89337F1E919FFBBC2C7C9DDFE4C78EC" box="[189,398,953,979]" name="Madagascar" pageId="33" pageNumber="34">MADAGASCAR</collectingCountry>
<specimenCount id="B682BCE8E919FFBBC3CCC9DDFE0D78EC" box="[438,463,953,979]" pageId="33" pageNumber="34" type="male"></specimenCount>
; “
<collectingRegion id="6240B983E919FFBBC397C9DDFD1078EC" box="[493,722,953,979]" country="Madagascar" name="Toamasina" pageId="33" pageNumber="34">Toamasina Region</collectingRegion>
,
<location id="A55B21BAE919FFBBC092C9DDFCBA78EC" LSID="urn:lsid:plazi:treatment:282DC677E919FFB9C028CB6CFEFB7F2A:A55B21BAE919FFBBC092C9DDFCBA78EC" box="[744,888,953,979]" country="Madagascar" latitude="-18.75" longLatPrecision="765" longitude="49.15" name="Manambato" pageId="33" pageNumber="34" stateProvince="Toamasina">Manambato</location>
;
<geoCoordinate id="C5B011A6E919FFBBC1F6C9DDFC3878EC" box="[908,1018,953,980]" degrees="18.75" direction="south" orientation="latitude" pageId="33" pageNumber="34" precision="555" value="-18.75">18.75° S</geoCoordinate>
,
<geoCoordinate id="C5B011A6E919FFBBC675C9DDFB4278EC" box="[1039,1152,953,980]" degrees="49.15" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="555" value="49.15">49.15° E</geoCoordinate>
;
<date id="D43A51A1E919FFBBC6EEC9DDFAB078EC" box="[1172,1394,953,979]" pageId="33" pageNumber="34" value="2012-11-24" valueMax="2012-11-30" valueMin="2012-11-24">
<collectingDate id="C47EA849E919FFBBC6EEC9DDFAB078EC" box="[1172,1394,953,979]" pageId="33" pageNumber="34" value="2012-11-24" valueMax="2012-11-30" valueMin="2012-11-24">2430 Nov. 2012</collectingDate>
</date>
;
<collectorName id="0D7112B7E919FFBBC2C7C9B8FE9478C9" box="[189,342,988,1014]" pageId="33" pageNumber="34">A.Medvedev</collectorName>
leg.”;
<collectionCode id="C695EFA4E919FFBBC3DEC9B9FDC178C8" box="[420,515,989,1015]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="33" pageNumber="34">ZMUM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="A03B7761E919FFBBC290CE7AFC397F64" blockId="33.[189,1019,1054,1115]" pageId="33" pageNumber="34">
<materialsCitation id="10EC7D3CE919FFBBC290CE7AFC357F64" ID-GBIF-Occurrence="4145165313" collectingDate="2012-11-24" collectingDateMax="2012-11-30" collectingDateMin="2012-11-24" collectionCode="ZMUM" collectorName="A. Medvedev" country="Madagascar" latitude="-18.75" location="Manambato" longLatPrecision="765" longitude="49.15" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
<emphasis id="92F0AB73E919FFBBC290CE7AFE9F7F07" bold="true" box="[234,349,1054,1080]" pageId="33" pageNumber="34">
<typeStatus id="7F3FC9C3E919FFBBC290CE7AFE9F7F07" box="[234,349,1054,1080]" pageId="33" pageNumber="34" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="D89337F1E919FFBBC2C7CE25FE4C7F64" box="[189,398,1089,1115]" name="Madagascar" pageId="33" pageNumber="34">MADAGASCAR</collectingCountry>
<specimenCount id="B682BCE8E919FFBBC3DDCE25FE147F64" box="[423,470,1089,1115]" pageId="33" pageNumber="34" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="C695EFA4E919FFBBC1E2CE25FC357F64" box="[920,1015,1089,1115]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="33" pageNumber="34">ZMUM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="A03B7761E919FFBBC2C7CEE6FE8D7FA3" blockId="33.[189,335,1154,1180]" box="[189,335,1154,1180]" pageId="33" pageNumber="34">
<heading id="FB73C00DE919FFBBC2C7CEE6FE8D7FA3" bold="true" box="[189,335,1154,1180]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="92F0AB73E919FFBBC2C7CEE6FE8D7FA3" bold="true" box="[189,335,1154,1180]" pageId="33" pageNumber="34">Description</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E919FFBBC290CED5FE6F7FF4" blockId="33.[189,1399,1201,1262]" box="[234,429,1201,1227]" pageId="33" pageNumber="34">
<emphasis id="92F0AB73E919FFBBC290CED5FEE87FF4" bold="true" box="[234,298,1201,1227]" pageId="33" pageNumber="34">Male</emphasis>
(
<figureCitation id="38BF6BE4E919FFBBC340CED5FE667FF4" box="[314,420,1201,1227]" captionStart="Fig" captionStartId="34.[189,232,1892,1918]" captionTargetBox="[216,1371,544,1852]" captionTargetId="figure-84@34.[216,1371,544,1854]" captionTargetPageId="34" captionText="Fig. 11. Thinophilus manambato sp. nov. AD. Holotype, ♂ (ZMUM). EG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna. D. Wing. E. Hypopygium after maceration, lateral view. F. Distal appendages of hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110572" httpUri="https://zenodo.org/record/8110572/files/figure.png" pageId="33" pageNumber="34">Fig. 11A</figureCitation>
)
</paragraph>
<paragraph id="A03B7761E919FFBBC2C7CEB0FAB57FD1" blockId="33.[189,1399,1201,1262]" box="[189,1399,1236,1262]" pageId="33" pageNumber="34">
<smallCapsWord id="A6DDE1BDE919FFBBC2C7CEB0FEAC7FD3" baselines="1255,1255" box="[189,366,1236,1262]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="33" pageNumber="34">MEASUREMENTS</smallCapsWord>
. Body length
<quantity id="677CDA84E919FFBBC068CEB0FDAC7FD1" box="[530,622,1236,1262]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="33" pageNumber="34" unit="mm" value="4.3">4.3 mm</quantity>
; antenna length
<quantity id="677CDA84E919FFBBC151CEB0FC4A7FD1" box="[811,904,1236,1262]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="34" unit="mm" value="0.9">0.9 mm</quantity>
; wing length
<quantity id="677CDA84E919FFBBC65ECEB0FB427FD1" box="[1060,1152,1236,1262]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="33" pageNumber="34" unit="mm" value="3.7">3.7 mm</quantity>
; wing width
<quantity id="677CDA84E919FFBBC76CCEB0FAB07FD1" box="[1302,1394,1236,1262]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="mm" value="1.2">1.2 mm</quantity>
.
</paragraph>
<paragraph id="A03B7761E919FFBBC2C7CF72FB4D7D53" blockId="33.[188,1399,1301,1645]" pageId="33" pageNumber="34">
<smallCapsWord id="A6DDE1BDE919FFBBC2C7CF72FF3E7E11" baselines="1321,1321" box="[189,252,1302,1328]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="33" pageNumber="34">HEAD</smallCapsWord>
(
<figureCitation id="38BF6BE4E919FFBBC374CF71FEB57E10" box="[270,375,1301,1328]" captionStart="Fig" captionStartId="34.[189,232,1892,1918]" captionTargetBox="[216,1371,544,1852]" captionTargetId="figure-84@34.[216,1371,544,1854]" captionTargetPageId="34" captionText="Fig. 11. Thinophilus manambato sp. nov. AD. Holotype, ♂ (ZMUM). EG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna. D. Wing. E. Hypopygium after maceration, lateral view. F. Distal appendages of hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110572" httpUri="https://zenodo.org/record/8110572/files/figure.png" pageId="33" pageNumber="34">Fig. 11B</figureCitation>
). Postcranium blackish blue, white pollinose; frons copper green, with violet spot under ocellar tubercle; face greenish blue; clypeus blackish blue, grey pollinose; face under antennae 1.7 × as wide as height of postpedicel; clypeus about ⅓ as long as epistoma, about 2 × as wide as long; palp yellow, bearing black bristly hairs; proboscis dark brown; 2 diverging ocellars; 1 vertical, 1 postvertical, much stronger and longer than, and not in row with upper postoculars; upper postoculars uniseriate, black; middle and lower postoculars multiseriate, white, long; antennal scape, pedicel and postpedicel blackish dorsally, yellow ventrally (
<figureCitation id="38BF6BE4E919FFBBC016CF8CFD157D3C" box="[620,727,1512,1539]" captionStart="Fig" captionStartId="34.[189,232,1892,1918]" captionTargetBox="[216,1371,544,1852]" captionTargetId="figure-84@34.[216,1371,544,1854]" captionTargetPageId="34" captionText="Fig. 11. Thinophilus manambato sp. nov. AD. Holotype, ♂ (ZMUM). EG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna. D. Wing. E. Hypopygium after maceration, lateral view. F. Distal appendages of hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110572" httpUri="https://zenodo.org/record/8110572/files/figure.png" pageId="33" pageNumber="34">Fig. 11C</figureCitation>
); scape with scale-like inner projection; pedicel simple, convex on inner side; postpedicel apically browned, rounded, with short pubescence, slightly higher than long (15/11); arista-like stylus dorsal, black and thick basally, whitish and thin distally, shortly pubescent; length ratio of scape to pedicel to postpedicel to stylus, 0.1/0.1/0.11/0.7.
</paragraph>
<paragraph id="A03B7761E919FFBBC2C7CCF0FC307DCB" blockId="33.[189,1399,1683,1781]" pageId="33" pageNumber="34">
<smallCapsWord id="A6DDE1BDE919FFBBC2C7CCF0FEDB7D93" baselines="1703,1703" box="[189,281,1684,1710]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="33" pageNumber="34">THORAX</smallCapsWord>
. Metallic, grey dusted; mesonotum blue-green; pleura bluish black; no acrostichals; 6 dorsocentrals decreasing in length anteriorly; scutellum with 2 strong marginals and 2 minute laterals; 78 upper and 78 lower, white propleural bristles of different length.
</paragraph>
<paragraph id="A03B7761E919FFBBC2C7CD78FDF07C66" blockId="33.[189,1399,1819,1881]" pageId="33" pageNumber="34">
<smallCapsWord id="A6DDE1BDE919FFBBC2C7CD78FF347C0B" baselines="1839,1839" box="[189,246,1820,1846]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="33" pageNumber="34">LEGS</smallCapsWord>
. Fore coxa yellow; mid and hind coxae black, yellow at apex; legs mostly yellow; tarsi gradually darkened from tip of basitarsus.
</paragraph>
<paragraph id="A03B7761E919FFB8C2C7CDE5FC377A78" blockId="33.[188,1398,1920,2017]" lastBlockId="34.[189,1398,265,327]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<smallCapsWord id="A6DDE1BDE919FFBBC2C7CDE5FF347CA6" baselines="1940,1940" box="[189,246,1921,1947]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Fore" pageId="33" pageNumber="34">FORE</smallCapsWord>
<smallCapsWord id="A6DDE1BDE919FFBBC287CDE2FEE77CA6" baselines="1940" box="[253,293,1926,1945]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg" pageId="33" pageNumber="34">LEG</smallCapsWord>
. Coxa with mainly white setae, with at most 23 black apical bristles; femur simple, with some short fine ventral setae at base; tibia bearing 3 anterodorsal, 2 posterodorsal, 23 ventral and 34 apical short bristles, glabrous anteriorly on distal ⅓; tarsal segments 14 ventrally with short simple setulae; segments 35 with elongate apicodorsal setulae; segment 5 inconspicuously widened; length of femur, tibia and tarsal segments (in mm): 1.24/1.16/0.51/0.25/0.19/0.13/0.17.
</paragraph>
<paragraph id="A03B7761E91AFFB9C2C7CB0AFEAD7A79" blockId="34.[189,1400,366,463]" lastBlockId="35.[189,1398,265,326]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<smallCapsWord id="A6DDE1BDE91AFFB8C2C7CB0AFF2F7AB9" baselines="385,385" box="[189,237,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mid" pageId="34" pageNumber="35">MID</smallCapsWord>
<smallCapsWord id="A6DDE1BDE91AFFB8C28CCB17FEDC7AB9" baselines="385" box="[246,286,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg" pageId="34" pageNumber="35">LEG</smallCapsWord>
. Coxa with white setae and 1 black bristle; femur with anteroventral row of short white hairs on basal half; 1 preapical posteroventral; tibia bearing 2 anterodorsal; 2 posterodorsal, 2 ventral short bristles, 4 apicals; tarsal segments 14 ventrally with short setae; segment 5 inconspicuously flattened dorso-ventrally and widened; length of femur, tibia and tarsal segments (in mm): 1.44/1.52/0.88/0.35/ 0.24/0.16/0.17.
</paragraph>
<caption id="F4FB27E9E91AFFB8C2C7CD00FE257CD7" ID-DOI="http://doi.org/10.5281/zenodo.8110572" ID-Zenodo-Dep="8110572" httpUri="https://zenodo.org/record/8110572/files/figure.png" pageId="34" pageNumber="35" startId="34.[189,232,1892,1918]" targetBox="[216,1371,544,1852]" targetPageId="34">
<paragraph id="A03B7761E91AFFB8C2C7CD00FE257CD7" blockId="34.[189,1399,1892,2024]" pageId="34" pageNumber="35">
<emphasis id="92F0AB73E91AFFB8C2C7CD00FED97C40" bold="true" box="[189,283,1892,1919]" pageId="34" pageNumber="35">Fig. 11.</emphasis>
<taxonomicName id="67840CE2E91AFFB8C35DCD01FD887C40" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[295,586,1893,1919]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="manambato" status="sp. nov.">
<emphasis id="92F0AB73E91AFFB8C35DCD01FD887C40" box="[295,586,1893,1919]" italics="true" pageId="34" pageNumber="35">Thinophilus manambato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E91AFFB8C02ECD01FD737C40" box="[596,689,1893,1919]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="92F0AB73E91AFFB8C0C0CD00FD337C41" bold="true" box="[698,753,1892,1918]" pageId="34" pageNumber="35">AD</emphasis>
. Holotype, ♂ (ZMUM).
<emphasis id="92F0AB73E91AFFB8C665CD00FB9A7C41" bold="true" box="[1055,1112,1892,1918]" pageId="34" pageNumber="35">EG</emphasis>
. Paratype, ♂ (ZMUM).
<emphasis id="92F0AB73E91AFFB8C2C7CDECFF107C9D" bold="true" box="[189,210,1928,1954]" pageId="34" pageNumber="35">A</emphasis>
. Habitus.
<emphasis id="92F0AB73E91AFFB8C32ACDECFEA67C9D" bold="true" box="[336,356,1928,1954]" pageId="34" pageNumber="35">B</emphasis>
. Head.
<emphasis id="92F0AB73E91AFFB8C3BECDECFE1B7C9D" bold="true" box="[452,473,1928,1954]" pageId="34" pageNumber="35">C</emphasis>
. Antenna.
<emphasis id="92F0AB73E91AFFB8C025CDECFDB17C9D" bold="true" box="[607,627,1928,1954]" pageId="34" pageNumber="35">D</emphasis>
. Wing.
<emphasis id="92F0AB73E91AFFB8C0ADCDECFD2B7C9D" bold="true" box="[727,745,1928,1954]" pageId="34" pageNumber="35">E</emphasis>
. Hypopygium after maceration, lateral view.
<emphasis id="92F0AB73E91AFFB8C777CDECFADD7C9D" bold="true" box="[1293,1311,1928,1954]" pageId="34" pageNumber="35">F</emphasis>
. Distal appendages of hypopygium after maceration, lateral view.
<emphasis id="92F0AB73E91AFFB8C103CDCFFC527CFA" bold="true" box="[889,912,1963,1989]" pageId="34" pageNumber="35">G</emphasis>
. Distal appendages of hypopygium after maceration, ventral view.
</paragraph>
</caption>
<paragraph id="A03B7761E91BFFB9C2C7CB0AFACC7ACD" blockId="35.[188,1398,365,498]" pageId="35" pageNumber="36">
<smallCapsWord id="A6DDE1BDE91BFFB9C2C7CB0AFF357AB9" baselines="385,385" box="[189,247,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Hind" pageId="35" pageNumber="36">HIND</smallCapsWord>
<smallCapsWord id="A6DDE1BDE91BFFB9C284CB17FEE47AB9" baselines="385" box="[254,294,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg" pageId="35" pageNumber="36">LEG</smallCapsWord>
. Coxa with 1 black exterior bristle; femur ventrally without remarkable setae; 67 anterodorsal bristles; tibia bearing 4 anterodorsal, 4 posterodorsal bristles, 3 short ventrals, 4 apicals; tarsal segments 14 ventrally with short setae; segment 5 with elongate apicodorsal setulae; segment 5 inconspicuously widened; length of femur, tibia and tarsal segments (in mm): 1.68/1.9/0.61/0.45/0.28/0.21/0.2.
</paragraph>
<paragraph id="A03B7761E91BFFB9C2C7C87DFB4579A3" blockId="35.[189,1399,537,669]" pageId="35" pageNumber="36">
<smallCapsWord id="A6DDE1BDE91BFFB9C2C7C87DFF3C790E" baselines="556,556" box="[189,254,537,563]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Wing" pageId="35" pageNumber="36">WING</smallCapsWord>
(
<figureCitation id="38BF6BE4E91BFFB9C36BC87DFEBF790C" box="[273,381,537,563]" captionStart="Fig" captionStartId="34.[189,232,1892,1918]" captionTargetBox="[216,1371,544,1852]" captionTargetId="figure-84@34.[216,1371,544,1854]" captionTargetPageId="34" captionText="Fig. 11. Thinophilus manambato sp. nov. AD. Holotype, ♂ (ZMUM). EG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna. D. Wing. E. Hypopygium after maceration, lateral view. F. Distal appendages of hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110572" httpUri="https://zenodo.org/record/8110572/files/figure.png" pageId="35" pageNumber="36">Fig. 11D</figureCitation>
). Hyaline, without darker shades; veins yellow-brown, more yellowish at base; distal part of M 1+2 convex; tip of R 4+5 parallel with M 1+2; ratio of part of costa between R 2+3 and R 4+5 to that between R
<subScript id="3C007524E91BFFB9C340C816FE9779BF" attach="left" box="[314,341,626,640]" fontSize="6" pageId="35" pageNumber="36">4+5</subScript>
and M
<subScript id="3C007524E91BFFB9C3D3C816FE0679BF" attach="left" box="[425,452,626,640]" fontSize="6" pageId="35" pageNumber="36">1+2</subScript>
(in mm), 0.40/0.26; crossvein dm-m straight; ratio of dm-m to distal part of M
<subScript id="3C007524E91BFFB9C71DC816FAB279BF" attach="right" box="[1383,1392,626,640]" fontSize="6" pageId="35" pageNumber="36">4</subScript>
, 0.33/0.35; anal vein distinct; halter yellow; lower calypter yellow, with white cilia.
</paragraph>
<paragraph id="A03B7761E91BFFB9C2C7C8A0FDC8788E" blockId="35.[188,1399,708,945]" pageId="35" pageNumber="36">
<smallCapsWord id="A6DDE1BDE91BFFB9C2C7C8A0FEEE79E3" baselines="727,727" box="[189,300,708,734]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="35" pageNumber="36">ABDOMEN</smallCapsWord>
. Blackish violet, grey dusted; tergite 6 shining blue; setae and hind-marginal bristles on tergites black, short; sternites with short white setae. Hypopygium (
<figureCitation id="38BF6BE4E91BFFB9C67FC883FB57783E" box="[1029,1173,743,769]" captionStart="Fig" captionStartId="34.[189,232,1892,1918]" captionTargetBox="[216,1371,544,1852]" captionTargetId="figure-84@34.[216,1371,544,1854]" captionTargetPageId="34" captionText="Fig. 11. Thinophilus manambato sp. nov. AD. Holotype, ♂ (ZMUM). EG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna. D. Wing. E. Hypopygium after maceration, lateral view. F. Distal appendages of hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110572" httpUri="https://zenodo.org/record/8110572/files/figure.png" pageId="35" pageNumber="36">Fig. 11EG</figureCitation>
) black with black appendages; epandrial lobe reduced; hypandrium fused with epandrium, short and broad, apically concave; phallosoma broad, narrow at apex, not reaching apex of surstyli; phallus short, simple, apically broad, hidden under phallosoma; surstylus straight and broad (lateral view), with 3 long dorsal bristles and several short and long ventral and apical setae (
<figureCitation id="38BF6BE4E91BFFB9C169C910FCBA78B1" box="[787,888,884,910]" captionStart="Fig" captionStartId="34.[189,232,1892,1918]" captionTargetBox="[216,1371,544,1852]" captionTargetId="figure-84@34.[216,1371,544,1854]" captionTargetPageId="34" captionText="Fig. 11. Thinophilus manambato sp. nov. AD. Holotype, ♂ (ZMUM). EG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna. D. Wing. E. Hypopygium after maceration, lateral view. F. Distal appendages of hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110572" httpUri="https://zenodo.org/record/8110572/files/figure.png" pageId="35" pageNumber="36">Fig. 11F</figureCitation>
); cerci dorsally adjoined, leaflike, with long marginal bristles (
<figureCitation id="38BF6BE4E91BFFB9C3EBC9F3FE3F788E" box="[401,509,919,945]" captionStart="Fig" captionStartId="34.[189,232,1892,1918]" captionTargetBox="[216,1371,544,1852]" captionTargetId="figure-84@34.[216,1371,544,1854]" captionTargetPageId="34" captionText="Fig. 11. Thinophilus manambato sp. nov. AD. Holotype, ♂ (ZMUM). EG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna. D. Wing. E. Hypopygium after maceration, lateral view. F. Distal appendages of hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110572" httpUri="https://zenodo.org/record/8110572/files/figure.png" pageId="35" pageNumber="36">Fig. 11G</figureCitation>
).
</paragraph>
<paragraph id="A03B7761E91BFFB9C290C9BCFE8478CD" blockId="35.[189,326,984,1045]" box="[234,326,984,1010]" pageId="35" pageNumber="36">
<emphasis id="92F0AB73E91BFFB9C290C9BCFE8478CD" bold="true" box="[234,326,984,1010]" pageId="35" pageNumber="36">Female</emphasis>
</paragraph>
<paragraph id="A03B7761E91BFFB9C2C7C99FFEFB7F2A" blockId="35.[189,326,984,1045]" box="[189,313,1019,1045]" pageId="35" pageNumber="36">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>