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<mods:title id="3D3947354DB362301A1626D26DB2A267">Amphi-Indian Ocean Disjunction in the Trans-Pacific Genus Archaeoglenes Brown (Coleoptera: Tenebrionidae: Phrenapatinae): New Taxonomic and Distributional Data</mods:title>
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<treatment id="2644879BFF93FFA2FF7A29DCCB80FDF4" ID-DOI="http://doi.org/10.5281/zenodo.6941410" ID-GBIF-Taxon="197870830" ID-Zenodo-Dep="6941410" LSID="urn:lsid:plazi:treatment:2644879BFF93FFA2FF7A29DCCB80FDF4" httpUri="http://treatment.plazi.org/id/2644879BFF93FFA2FF7A29DCCB80FDF4" lastPageNumber="160" pageId="8" pageNumber="160">
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<paragraph id="AE52368DFF93FFA2FF7A29DCCA05FB3B" blockId="8.[208,582,1171,1219]" box="[208,582,1171,1193]" pageId="8" pageNumber="160">
<heading id="F51A81E1FF93FFA2FF7A29DCCA05FB3B" box="[208,582,1171,1193]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="160" reason="2">
<taxonomicName id="69ED4D0EFF93FFA2FF7A29DCCA05FB3B" authority="Kaszab, 1982" authorityName="Kaszab" authorityYear="1982" box="[208,582,1171,1193]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="baloghi">
<emphasis id="9C99EA9FFF93FFA2FF7A29DCCA05FB3B" bold="true" box="[208,582,1171,1193]" pageId="8" pageNumber="160">
<emphasis id="9C99EA9FFF93FFA2FF7A29DCC9F0FB3B" bold="true" box="[208,435,1171,1193]" italics="true" pageId="8" pageNumber="160">Archaeoglenes baloghi</emphasis>
<bibRefCitation id="CA7C4B7CFF93FFA2FE1129DBCA05FB3B" author="Kaszab, Z." box="[443,582,1172,1193]" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="8" pageNumber="160" pagination="45 - 55" part="28" refId="ref7504" refString="Kaszab, Z. 1982 a. Die Papuanisch-Pazifischen Arten der Gattung Archeoglenes Brown, 1893 (Coleoptera: Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae 28: 45 - 55." title="Die Papuanisch-Pazifischen Arten der Gattung Archeoglenes Brown, 1893 (Coleoptera: Tenebrionidae)" type="journal article" year="1982">Kaszab, 1982</bibRefCitation>
</emphasis>
</taxonomicName>
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<paragraph id="AE52368DFF93FFA2FECF29E1C9ECFB51" blockId="8.[208,582,1171,1219]" box="[357,431,1198,1219]" pageId="8" pageNumber="160">
(
<figureCitation id="36D62A08FF93FFA2FEC729E1C9EBFB51" box="[365,424,1198,1219]" captionStart="Fig" captionStartId="9.[197,229,1662,1681]" captionTargetBox="[230,1132,193,1636]" captionTargetId="figure-1@9.[225,1141,192,1639]" captionTargetPageId="9" captionText="Fig. 6. Morphology and label data of the type specimens of Archaeoglenes species. A. baloghi: A) Habitus of holotype, B) Labels, C) Pronotal disc surface. A. novaeguineae: D) Habitus of holotype, E) Labels, F) Pronotal disc surface, G) Protarsus, H) Mesotarsus, I) Metatarsus." figureDoi="http://doi.org/10.5281/zenodo.4908296" httpUri="https://zenodo.org/record/4908296/files/figure.png" pageId="8" pageNumber="160">Fig. 6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E6F76506FF93FFA2FF3D2996C948FA9A" pageId="8" pageNumber="160" type="reference_group">
<paragraph id="AE52368DFF93FFA2FF3D2996C948FA9A" blockId="8.[151,638,1241,1528]" pageId="8" pageNumber="160">
<treatmentCitationGroup id="8EFD11A3FF93FFA2FF3D2996C948FA9A" pageId="8" pageNumber="160">
<treatmentCitation id="2F4C109CFF93FFA2FF3D2996CA6FFB7C" author="Kaszab" box="[151,556,1241,1262]" page="49" pageId="8" pageNumber="160" year="1982">
<taxonomicName id="69ED4D0EFF93FFA2FF3D2996CA6FFB7C" authority="Kaszab, 1982 a: 49" authorityName="Kaszab" authorityPageNumber="49" authorityYear="1982" box="[151,556,1241,1262]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="baloghi">
<emphasis id="9C99EA9FFF93FFA2FF3D2996C931FB7C" box="[151,370,1241,1262]" italics="true" pageId="8" pageNumber="160">Archaeoglenes baloghi</emphasis>
<bibRefCitation id="CA7C4B7CFF93FFA2FED22996CA6FFB7C" author="Kaszab" box="[376,556,1241,1262]" firstAuthor="Kaszab" page="49" pageId="8" pageNumber="160" pagination="45 - 55" refId="ref7504" refString="Kaszab, Z. 1982 a. Die Papuanisch-Pazifischen Arten der Gattung Archeoglenes Brown, 1893 (Coleoptera: Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae 28: 45 - 55." type="journal article" year="1982" yearSuffix="a">Kaszab, 1982a: 49</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Studied material:
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E6F76506FF93FFA2FEBF29BCCA55FA1F" pageId="8" pageNumber="160" type="reference_group">
<paragraph id="AE52368DFF93FFA2FEBF29BCCA3DFACA" blockId="8.[151,638,1241,1528]" pageId="8" pageNumber="160">
<materialsCitation id="1E853CD0FF93FFA2FEBF29BCCA3DFACA" ID-GBIF-Occurrence="3867547313" collectingDate="1968-09-27" collectingDateMax="1968-09-28" collectingDateMin="1968-09-27" collectionCode="HNHM" collectorName="Dr. J. Balogh" country="Guinea-Bissau" location="Mt. Kumbak" pageId="8" pageNumber="160" specimenCount="1" typeStatus="holotype">
<typeStatus id="7156882FFF93FFA2FEBF29BCC931FA9A" box="[277,370,1267,1288]" pageId="8" pageNumber="160" type="holotype">Holotype</typeStatus>
(
<collectionCode id="C8FCAE48FF93FFA2FE2929BCC996FA9A" box="[387,469,1267,1288]" pageId="8" pageNumber="160">HNHM</collectionCode>
): “/ No. NG-W- B.166./ / leg.
<collectorName id="0318535BFF93FFA2FE9C2841C9F6FAB1" box="[310,437,1294,1315]" pageId="8" pageNumber="160">Dr. J. Balogh</collectorName>
”, “NEW
<collectingCountry id="D6FA761DFF93FFA2FDBC2841CA33FAB1" box="[534,624,1294,1315]" name="Guinea-Bissau" pageId="8" pageNumber="160">GUINEA</collectingCountry>
/ NE/ / Wau,
<location id="AB326056FF93FFA2FE982867C9F6FAAF" LSID="urn:lsid:plazi:treatment:2644879BFF93FFA2FF7A29DCCB80FDF4:AB326056FF93FFA2FE982867C9F6FAAF" box="[306,437,1320,1341]" country="Guinea-Bissau" name="Mt. Kumbak" pageId="8" pageNumber="160">Mt. Kumbak</location>
/
<date id="DA53104DFF93FFA2FE7A2867CA2BFAAF" box="[464,616,1320,1341]" pageId="8" pageNumber="160" value="1968-09-27" valueMax="1968-09-28" valueMin="1968-09-27">
<collectingDate id="CA17E9A5FF93FFA2FE7A2867CA2BFAAF" box="[464,616,1320,1341]" pageId="8" pageNumber="160" value="1968-09-27" valueMax="1968-09-28" valueMin="1968-09-27">27-28.IX.1968</collectingDate>
</date>
.”, “Holotypus1981/Archeoglenes/baloghi/Kaszab”.
</materialsCitation>
</paragraph>
<paragraph id="AE52368DFF93FFA2FF3D2811CA55FA1F" blockId="8.[151,638,1241,1528]" pageId="8" pageNumber="160">
<treatmentCitationGroup id="8EFD11A3FF93FFA2FF3D2811CA55FA1F" pageId="8" pageNumber="160">
<treatmentCitation id="2F4C109CFF93FFA2FF3D2811CA3AFAE1" author="Kaszab" box="[151,633,1374,1395]" page="50" pageId="8" pageNumber="160" year="1982">
<taxonomicName id="69ED4D0EFF93FFA2FF3D2811CA3AFAE1" authority="Kaszab, 1982a: 50" authorityName="Kaszab" authorityPageNumber="50" authorityYear="1982" box="[151,633,1374,1395]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="novaeguineae">
<emphasis id="9C99EA9FFF93FFA2FF3D2811C9F6FAE1" box="[151,437,1374,1395]" italics="true" pageId="8" pageNumber="160">Archaeoglenes novaeguineae</emphasis>
<bibRefCitation id="CA7C4B7CFF93FFA2FE142811CA3AFAE1" author="Kaszab" box="[446,633,1374,1395]" firstAuthor="Kaszab" page="50" pageId="8" pageNumber="160" pagination="45 - 55" refId="ref7504" refString="Kaszab, Z. 1982 a. Die Papuanisch-Pazifischen Arten der Gattung Archeoglenes Brown, 1893 (Coleoptera: Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae 28: 45 - 55." type="journal article" year="1982" yearSuffix="a">Kaszab, 1982a: 50</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="9C99EA9FFF93FFA2FF1B2837C911FA1F" bold="true" box="[177,338,1400,1421]" pageId="8" pageNumber="160">
<taxonomicNameLabel id="87AA57E4FF93FFA2FF1B2837C911FA1F" box="[177,338,1400,1421]" pageId="8" pageNumber="160">new synonymy</taxonomicNameLabel>
</emphasis>
. Studied material:
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E6F76506FF93FFA2FDB52837CA7AFA65" pageId="8" pageNumber="160" type="reference_group">
<paragraph id="AE52368DFF93FFA2FDB52837CA7AFA65" blockId="8.[151,638,1241,1528]" pageId="8" pageNumber="160">
<materialsCitation id="1E853CD0FF93FFA2FDB52837CA7AFA65" ID-GBIF-Occurrence="3867547328" collectingDate="1968-09-03" collectionCode="HNHM" collectorName="Dr. J. Balogh" country="Guinea-Bissau" location="Golden ridge" municipality="Wau" originalDetermination="Archaeoglenes novaeguineae Kaszab, 1982" pageId="8" pageNumber="160" specimenCount="1" typeStatus="holotype">
<typeStatus id="7156882FFF93FFA2FDB52837CA3EFA1F" box="[543,637,1400,1421]" pageId="8" pageNumber="160" type="holotype">Holotype</typeStatus>
(
<collectionCode id="C8FCAE48FF93FFA2FF1228DCC948FA3A" box="[184,267,1427,1448]" pageId="8" pageNumber="160">HNHM</collectionCode>
): “/ No. NG-W-B.64./ / leg.
<collectorName id="0318535BFF93FFA2FDE928DCC943FA50" pageId="8" pageNumber="160">Dr. J. Balogh</collectorName>
”, “ NEW
<collectingCountry id="D6FA761DFF93FFA2FEDC28E2C998FA50" box="[374,475,1453,1474]" name="Guinea-Bissau" pageId="8" pageNumber="160">GUINEA</collectingCountry>
/ NE/ /
<collectingMunicipality id="4E36ACF7FF93FFA2FDEC28E2CA35FA50" box="[582,630,1453,1474]" pageId="8" pageNumber="160">Wau</collectingMunicipality>
,
<location id="AB326056FF93FFA2FF1B2887C972FA4F" LSID="urn:lsid:plazi:treatment:2644879BFF93FFA2FF7A29DCCB80FDF4:AB326056FF93FFA2FF1B2887C972FA4F" box="[177,305,1480,1501]" country="Guinea-Bissau" municipality="Wau" name="Golden ridge" pageId="8" pageNumber="160">Golden ridge</location>
/
<date id="DA53104DFF93FFA2FEE22887C9EFFA4F" box="[328,428,1480,1501]" pageId="8" pageNumber="160" value="1968-09-03">
<collectingDate id="CA17E9A5FF93FFA2FEE22887C9EFFA4F" box="[328,428,1480,1501]" pageId="8" pageNumber="160" value="1968-09-03">3.IX.1968</collectingDate>
</date>
.”, “Holotypus 1981 / Archeoglenes / novaeguineae / Kaszab”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E6F76506FF93FFA2FF052B42CB80FDF4" pageId="8" pageNumber="160" type="discussion">
<paragraph id="AE52368DFF93FFA2FF052B42CBEFFE3E" blockId="8.[151,638,1549,1729]" lastBlockId="8.[685,1173,194,614]" pageId="8" pageNumber="160">
<emphasis id="9C99EA9FFF93FFA2FF052B42C8ACF9B0" bold="true" box="[175,239,1549,1570]" pageId="8" pageNumber="160">Notes.</emphasis>
According to the original description,
<taxonomicName id="69ED4D0EFF93FFA2FF3D2B67C8BBF9AF" authorityName="Kaszab" authorityYear="1982" box="[151,248,1576,1597]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="baloghi">
<emphasis id="9C99EA9FFF93FFA2FF3D2B67C8BBF9AF" box="[151,248,1576,1597]" italics="true" pageId="8" pageNumber="160">A. baloghi</emphasis>
</taxonomicName>
and
<taxonomicName id="69ED4D0EFF93FFA2FE8E2B67C9FEF9AF" authorityName="Kaszab" authorityYear="1982" box="[292,445,1576,1597]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="novaeguineae">
<emphasis id="9C99EA9FFF93FFA2FE8E2B67C9FEF9AF" box="[292,445,1576,1597]" italics="true" pageId="8" pageNumber="160">A. novaeguineae</emphasis>
</taxonomicName>
can be distinguished from each other by different punctation on the pronotum (denser in
<taxonomicName id="69ED4D0EFF93FFA2FE902B12C997F9E0" authorityName="Kaszab" authorityYear="1982" box="[314,468,1629,1650]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="novaeguineae">
<emphasis id="9C99EA9FFF93FFA2FE902B12C997F9E0" box="[314,468,1629,1650]" italics="true" pageId="8" pageNumber="160">A. novaeguineae</emphasis>
</taxonomicName>
) and elytral shape (more elongated in
<taxonomicName id="69ED4D0EFF93FFA2FEFF2B38C9FBF91E" authorityName="Kaszab" authorityYear="1982" box="[341,440,1655,1676]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="baloghi">
<emphasis id="9C99EA9FFF93FFA2FEFF2B38C9FBF91E" box="[341,440,1655,1676]" italics="true" pageId="8" pageNumber="160">A. baloghi</emphasis>
</taxonomicName>
) (
<figureCitation id="36D62A08FF93FFA2FE652B38CA30F91E" box="[463,627,1655,1677]" captionStart="Fig" captionStartId="9.[197,229,1662,1681]" captionTargetBox="[230,1132,193,1636]" captionTargetId="figure-1@9.[225,1141,192,1639]" captionTargetPageId="9" captionText="Fig. 6. Morphology and label data of the type specimens of Archaeoglenes species. A. baloghi: A) Habitus of holotype, B) Labels, C) Pronotal disc surface. A. novaeguineae: D) Habitus of holotype, E) Labels, F) Pronotal disc surface, G) Protarsus, H) Mesotarsus, I) Metatarsus." figureDoi="http://doi.org/10.5281/zenodo.4908296" httpUri="https://zenodo.org/record/4908296/files/figure.png" pageId="8" pageNumber="160">Fig. 6AC, D, F</figureCitation>
). Unfortunately, both taxa are known only from
<typeStatus id="7156882FFF93FFA2FDE72BDDC882F953" pageId="8" pageNumber="160" type="holotype">holotype</typeStatus>
specimens. The examined material of other
<taxonomicName id="69ED4D0EFF93FFA2FD072D8DCBCEFF45" box="[685,909,194,215]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9C99EA9FFF93FFA2FD072D8DCB7EFF45" box="[685,829,194,215]" italics="true" pageId="8" pageNumber="160">Archaeoglenes</emphasis>
species
</taxonomicName>
(
<taxonomicName id="69ED4D0EFF93FFA2FC352D8DCBB9FF45" box="[927,1018,194,215]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="9C99EA9FFF93FFA2FC352D8DCBB9FF45" box="[927,1018,194,215]" italics="true" pageId="8" pageNumber="160">A. gomyi</emphasis>
</taxonomicName>
,
<taxonomicName id="69ED4D0EFF93FFA2FBA02D8DCC22FF45" box="[1034,1121,194,215]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="9C99EA9FFF93FFA2FBA02D8DCC22FF45" box="[1034,1121,194,215]" italics="true" pageId="8" pageNumber="160">A. loebli</emphasis>
</taxonomicName>
, and
<taxonomicName id="69ED4D0EFF93FFA2FD072D92CB6FFF60" box="[685,812,221,242]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="triplehorni">
<emphasis id="9C99EA9FFF93FFA2FD072D92CB6FFF60" box="[685,812,221,242]" italics="true" pageId="8" pageNumber="160">A. triplehorni</emphasis>
</taxonomicName>
) shows that the punctation and shape of the pronotum are extremely variable within the genus. In our opinion, the characters used by Kaszab to distinguish these
<specimenCount id="B8EBFD04FF93FFA2FCCE2C63CBAFFED3" box="[868,1004,300,321]" pageId="8" pageNumber="160" type="generic">two specimens</specimenCount>
should be considered as intraspecific variation of a single species. Moreover, the two individuals representing those creatures were collected in the same area (Wau, New
<collectingCountry id="D6FA761DFF93FFA2FD4B2CD8CB66FE3E" box="[737,805,407,428]" name="Guinea-Bissau" pageId="8" pageNumber="160">Guinea</collectingCountry>
) (
<figureCitation id="36D62A08FF93FFA2FC902CD8CBE3FE3E" box="[826,928,407,428]" captionStart="Fig" captionStartId="9.[197,229,1662,1681]" captionTargetBox="[230,1132,193,1636]" captionTargetId="figure-1@9.[225,1141,192,1639]" captionTargetPageId="9" captionText="Fig. 6. Morphology and label data of the type specimens of Archaeoglenes species. A. baloghi: A) Habitus of holotype, B) Labels, C) Pronotal disc surface. A. novaeguineae: D) Habitus of holotype, E) Labels, F) Pronotal disc surface, G) Protarsus, H) Mesotarsus, I) Metatarsus." figureDoi="http://doi.org/10.5281/zenodo.4908296" httpUri="https://zenodo.org/record/4908296/files/figure.png" pageId="8" pageNumber="160">Fig. 6B, E</figureCitation>
).
</paragraph>
<paragraph id="AE52368DFF93FFA2FD6F2CFECB80FDF4" blockId="8.[685,1173,194,614]" pageId="8" pageNumber="160">
During our investigation of the material of
<taxonomicName id="69ED4D0EFF93FFA2FD072C83CB5DFE73" authorityName="Kaszab" authorityYear="1982" box="[685,798,460,481]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="baloghi">
<emphasis id="9C99EA9FFF93FFA2FD072C83CB5DFE73" box="[685,798,460,481]" italics="true" pageId="8" pageNumber="160">A. baloghi</emphasis>
</taxonomicName>
and its newly proposed synonym (
<taxonomicName id="69ED4D0EFF93FFA2FD1F2CA9CB15FE69" authorityName="Kaszab" authorityYear="1982" box="[693,854,486,507]" class="Insecta" family="Tenebrionidae" genus="Archaeoglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="novaeguineae">
<emphasis id="9C99EA9FFF93FFA2FD1F2CA9CB15FE69" box="[693,854,486,507]" italics="true" pageId="8" pageNumber="160">A. novaeguineae</emphasis>
</taxonomicName>
), we noticed that this species is characterized by a 5-5-4 tarsal formula that is unique within the genus (
<figureCitation id="36D62A08FF93FFA2FCF32F54CBF4FDA2" box="[857,951,539,561]" captionStart="Fig" captionStartId="9.[197,229,1662,1681]" captionTargetBox="[230,1132,193,1636]" captionTargetId="figure-1@9.[225,1141,192,1639]" captionTargetPageId="9" captionText="Fig. 6. Morphology and label data of the type specimens of Archaeoglenes species. A. baloghi: A) Habitus of holotype, B) Labels, C) Pronotal disc surface. A. novaeguineae: D) Habitus of holotype, E) Labels, F) Pronotal disc surface, G) Protarsus, H) Mesotarsus, I) Metatarsus." figureDoi="http://doi.org/10.5281/zenodo.4908296" httpUri="https://zenodo.org/record/4908296/files/figure.png" pageId="8" pageNumber="160">Fig. 6GI</figureCitation>
). This information was omitted by
<bibRefCitation id="CA7C4B7CFF93FFA2FCB12F79CBF3FDD9" author="Kaszab" box="[795,944,566,587]" firstAuthor="Kaszab" pageId="8" pageNumber="160" pagination="45 - 55" refId="ref7504" refString="Kaszab, Z. 1982 a. Die Papuanisch-Pazifischen Arten der Gattung Archeoglenes Brown, 1893 (Coleoptera: Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae 28: 45 - 55." type="journal article" year="1982" yearSuffix="a">Kaszab (1982a)</bibRefCitation>
and subsequent authors (Watrous 1982;
<bibRefCitation id="CA7C4B7CFF93FFA2FCED2F1ECBF4FDF4" author="Sasaji" box="[839,951,593,614]" firstAuthor="Sasaji" pageId="8" pageNumber="160" pagination="17 - 52" refId="ref8173" refString="Sasaji, H. 1983. Contribution to the taxonomy of the superfamily Cucujoidea (Coleoptera) of Japan and her adjacent districts, 1. Memoir of the Faculty of Education Fukui University Series II Natural Science 33: 17 - 52." type="journal article" year="1983">Sasaji 1983</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>