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<mods:title id="F37C3F57E48D7EAF5B6F8A54301CC770">On the paraphyly of Homaloptera (Teleostei: Balitoridae) and description of a new genus of hillstream loaches from the Western Ghats of India</mods:title>
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<treatment id="2573D038A91E9705FF4EFF05A6F10900" ID-DOI="http://doi.org/10.5281/zenodo.5622844" ID-GBIF-Taxon="127671913" ID-Zenodo-Dep="5622844" LSID="urn:lsid:plazi:treatment:2573D038A91E9705FF4EFF05A6F10900" httpUri="http://treatment.plazi.org/id/2573D038A91E9705FF4EFF05A6F10900" lastPageId="22" lastPageNumber="79" pageId="20" pageNumber="77">
<subSubSection id="E5C032A5A91E9707FF4EFF05A6DB0DE7" pageId="20" pageNumber="77" type="nomenclature">
<paragraph id="AD65612EA91E9707FF4EFF05A57D0D86" blockId="20.[151,524,151,211]" box="[151,524,151,178]" pageId="20" pageNumber="77">
<heading id="F62DD642A91E9707FF4EFF05A57D0D86" bold="true" box="[151,524,151,178]" fontSize="11" level="1" pageId="20" pageNumber="77" reason="1">
<taxonomicName id="6ADA1AADA91E9707FF4EFF05A57D0D86" ID-CoL="729G" ID-ENA="1590169" authority="Silas 1953" authorityName="Silas" authorityYear="1953" box="[151,524,151,178]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FF4EFF05A57D0D86" bold="true" box="[151,524,151,178]" pageId="20" pageNumber="77">
<emphasis id="9FAEBD3CA91E9707FF4EFF05A6F90D85" bold="true" box="[151,392,151,177]" italics="true" pageId="20" pageNumber="77">Pseudohomaloptera</emphasis>
<bibRefCitation id="C94B1CDFA91E9707FE56FF0AA57D0D86" author="Silas" box="[399,524,152,178]" pageId="20" pageNumber="77" refString="Silas, E. G. (1953) Classification, zoogeography and evolution of the fishes of the cyprinoid families Homalopteridae and Gastromyzonidae. Records of the Indian Museum, 50, 173 - 263." type="journal article" year="1953">Silas 1953</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AD65612EA91E9707FF4EFF28A6DB0DE7" blockId="20.[151,524,151,211]" box="[151,426,186,211]" pageId="20" pageNumber="77">
(
<figureCitation id="35E17DABA91E9707FF47FF28A6790DE7" box="[158,264,186,211]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[211,1328,347,1828]" captionTargetId="figure@8.[151,1436,336,1848]" captionTargetPageId="8" captionText="FIGURE 3. Lateral views of living (A) Homaloptera orthogoniata, not cataloged, Kalimantan, Borneo, Indonesia. Strong reddish tints on body likely breeding colors; (B) Homalopteroides smithi, UF 235740, 45.0 mm SL, Khlong Tasae, Salui Subdistrict, Chumphon, Thailand; (C) Homalopterula cf. ripleyi, not cataloged, Sumatra, Indonesia; (D) Balitoropsis zollingeri, not cataloged, Kalimantan, Borneo, Indonesia; (E) Pseudohomaloptera leonardi, UF 235735, 34.2 mm SL, Ta Pi River, Nakhon Si Thammarat, Thailand. Photos (A) &amp; (D) by Renny Hadiaty. Photo (C) by Daniel Lumbantobing. Specimens not to scale." httpUri="https://zenodo.org/record/288013/files/figure.png" pageId="20" pageNumber="77">Figures 3</figureCitation>
E, 4B, 5E, 14)
</paragraph>
</subSubSection>
<subSubSection id="E5C032A5A91E9707FF4EFE90A6590C03" pageId="20" pageNumber="77" type="reference_group">
<paragraph id="AD65612EA91E9707FF4EFE90A6590C03" blockId="20.[151,1436,258,311]" pageId="20" pageNumber="77">
<treatmentCitationGroup id="8DCA4600A91E9707FF4EFE90A6590C03" pageId="20" pageNumber="77">
<taxonomicName id="6ADA1AADA91E9707FF4EFE90A57B0C2C" ID-CoL="729G" authority="Silas, 1953:204" authorityName="Silas" authorityPageNumber="204" authorityYear="1953" box="[151,522,258,280]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FF4EFE90A6120C2C" box="[151,355,258,280]" italics="true" pageId="20" pageNumber="77">Pseudohomaloptera</emphasis>
<treatmentCitation id="2C7B473FA91E9707FEB0FE90A57B0C2C" author="Silas" box="[361,522,258,280]" page="204" pageId="20" pageNumber="77" year="1953">
<bibRefCitation id="C94B1CDFA91E9707FEB0FE90A6AD0C2C" author="Silas" box="[361,476,258,280]" pageId="20" pageNumber="77" refString="Silas, E. G. (1953) Classification, zoogeography and evolution of the fishes of the cyprinoid families Homalopteridae and Gastromyzonidae. Records of the Indian Museum, 50, 173 - 263." type="journal article" year="1953">Silas, 1953</bibRefCitation>
:204
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</taxonomicName>
. (
<typeStatus id="7261DF8CA91E9707FDC3FE90A5360C2C" box="[538,583,258,280]" pageId="20" pageNumber="77">type</typeStatus>
species:
<taxonomicName id="6ADA1AADA91E9707FD7DFE90A3360C2C" ID-CoL="3MDX2" authority="Popta 1905:180" authorityName="Popta" authorityPageNumber="180" authorityYear="1905" box="[676,1095,258,280]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="species" species="tatereganii">
<emphasis id="9FAEBD3CA91E9707FD7DFE90A4EE0C2C" box="[676,927,258,280]" italics="true" pageId="20" pageNumber="77">Homaloptera tatereganii</emphasis>
Popta 1905:180
</taxonomicName>
, by original designation). Gender feminine.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E5C032A5A91E9707FF4EFEF6A3560F1B" pageId="20" pageNumber="77" type="discussion">
<paragraph id="AD65612EA91E9707FF4EFEF6A3560F1B" blockId="20.[151,1437,356,920]" pageId="20" pageNumber="77">
<emphasis id="9FAEBD3CA91E9707FF4EFEF6A67A0C49" bold="true" box="[151,267,356,381]" pageId="20" pageNumber="77">Remarks.</emphasis>
<taxonomicName id="6ADA1AADA91E9707FEF8FEF6A59C0C49" authority="Popta 1905" authorityName="Popta" authorityYear="1905" box="[289,749,356,381]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="species" species="tatereganii">
<emphasis id="9FAEBD3CA91E9707FEF8FEF6A5360C48" box="[289,583,356,380]" italics="true" pageId="20" pageNumber="77">Homaloptera tatereganii</emphasis>
Popta 1905
</taxonomicName>
was designated as the
<typeStatus id="7261DF8CA91E9707FBE5FEF7A31D0C49" box="[1084,1132,357,381]" pageId="20" pageNumber="77">type</typeStatus>
species for the genus
<taxonomicName id="6ADA1AADA91E9707FF4EFE1AA6080C94" box="[151,377,392,416]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FF4EFE1AA6080C94" box="[151,377,392,416]" italics="true" pageId="20" pageNumber="77">Pseudohomaloptera</emphasis>
</taxonomicName>
by
<bibRefCitation id="C94B1CDFA91E9707FE78FE15A55D0C94" author="Silas" box="[417,556,391,416]" pageId="20" pageNumber="77" refString="Silas, E. G. (1953) Classification, zoogeography and evolution of the fishes of the cyprinoid families Homalopteridae and Gastromyzonidae. Records of the Indian Museum, 50, 173 - 263." type="journal article" year="1953">Silas (1953)</bibRefCitation>
.
<taxonomicName id="6ADA1AADA91E9707FDEFFE1AA4690C94" box="[566,792,392,416]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FDEFFE1AA4690C94" box="[566,792,392,416]" italics="true" pageId="20" pageNumber="77">Pseudohomaloptera</emphasis>
</taxonomicName>
was distinguished from
<taxonomicName id="6ADA1AADA91E9707FBF0FE1AA3CF0C94" box="[1065,1214,392,416]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FBF0FE1AA3CF0C94" box="[1065,1214,392,416]" italics="true" pageId="20" pageNumber="77">Homaloptera</emphasis>
</taxonomicName>
by the “presence of a rostral groove and other structures associated with the mouth” (
<bibRefCitation id="C94B1CDFA91E9707FC19FE3EA3050CF1" author="Silas" box="[960,1140,428,453]" pageId="20" pageNumber="77" refString="Silas, E. G. (1953) Classification, zoogeography and evolution of the fishes of the cyprinoid families Homalopteridae and Gastromyzonidae. Records of the Indian Museum, 50, 173 - 263." type="journal article" year="1953">Silas 1953:205</bibRefCitation>
).
<bibRefCitation id="C94B1CDFA91E9707FB56FE3FA2620CF0" author="Tan" box="[1167,1299,428,452]" pageId="20" pageNumber="77" refString="Tan, H. H. (2009) A new species of hill stream loach (Teleostei: Balitoridae) from central Kalimantan, with redescriptions of Homaloptera tateregani Popta and Homaloptera stephensoni Hora. Zootaxa, 2171, 48 - 64." type="journal article" year="2009">Tan (2009)</bibRefCitation>
recognized
<taxonomicName id="6ADA1AADA91E9707FF4EFE42A6080CDC" box="[151,377,464,488]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FF4EFE42A6080CDC" box="[151,377,464,488]" italics="true" pageId="20" pageNumber="77">Pseudohomaloptera</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="6ADA1AADA91E9707FD41FE42A45C0CDC" box="[664,813,464,488]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FD41FE42A45C0CDC" box="[664,813,464,488]" italics="true" pageId="20" pageNumber="77">Homaloptera</emphasis>
</taxonomicName>
, since all species of
<taxonomicName id="6ADA1AADA91E9707FBFAFE42A3C90CDC" box="[1059,1208,464,488]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FBFAFE42A3C90CDC" box="[1059,1208,464,488]" italics="true" pageId="20" pageNumber="77">Homaloptera</emphasis>
</taxonomicName>
(sensu lato) have a rostral and postoral groove to varying degrees.
<bibRefCitation id="C94B1CDFA91E9707FD38FE66A4E70F38" author="Kottelat" box="[737,918,500,525]" pageId="20" pageNumber="77" refString="Kottelat, M. (2012) Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology, Supplement 26, 1 - 199." type="book chapter" year="2012">Kottelat (2012)</bibRefCitation>
recognized
<taxonomicName id="6ADA1AADA91E9707FBF6FE67A3A20F38" box="[1071,1235,501,524]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="species" species="tatereganii">
<emphasis id="9FAEBD3CA91E9707FBF6FE67A3A20F38" box="[1071,1235,501,524]" italics="true" pageId="20" pageNumber="77">H. tatereganii</emphasis>
</taxonomicName>
as a species of
<taxonomicName id="6ADA1AADA91E9707FF4EFD8AA66E0F04" box="[151,287,536,560]" family="Balitoridae" genus="Balitoropsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FF4EFD8AA66E0F04" box="[151,287,536,560]" italics="true" pageId="20" pageNumber="77">Balitoropsis</emphasis>
</taxonomicName>
and treated
<taxonomicName id="6ADA1AADA91E9707FE71FD8AA5FB0F04" box="[424,650,536,560]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FE71FD8AA5FB0F04" box="[424,650,536,560]" italics="true" pageId="20" pageNumber="77">Pseudohomaloptera</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="6ADA1AADA91E9707FC41FD8AA3510F04" box="[920,1056,536,560]" family="Balitoridae" genus="Balitoropsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FC41FD8AA3510F04" box="[920,1056,536,560]" italics="true" pageId="20" pageNumber="77">Balitoropsis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E5C032A5A91E9707FF1EFDAEA4BB0EA3" pageId="20" pageNumber="77" type="reference_group">
<paragraph id="AD65612EA91E9707FF1EFDAEA4BB0EA3" blockId="20.[151,1437,356,920]" pageId="20" pageNumber="77">
<taxonomicName id="6ADA1AADA91E9707FF1EFDAEA6D80F60" box="[199,425,572,596]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FF1EFDAEA6D80F60" box="[199,425,572,596]" italics="true" pageId="20" pageNumber="77">Pseudohomaloptera</emphasis>
</taxonomicName>
is morphologically very similar to
<emphasis id="9FAEBD3CA91E9707FCE4FDAEA4BA0F60" box="[829,971,572,596]" italics="true" pageId="20" pageNumber="77">
<taxonomicName id="6ADA1AADA91E9707FCE4FDAEA4B60F60" box="[829,967,572,596]" family="Balitoridae" genus="Balitoropsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">Balitoropsis</taxonomicName>
,
</emphasis>
and the mouth characters given by
<bibRefCitation id="C94B1CDFA91E9707FABFFDAEA7900F4C" author="Silas" pageId="20" pageNumber="77" refString="Silas, E. G. (1953) Classification, zoogeography and evolution of the fishes of the cyprinoid families Homalopteridae and Gastromyzonidae. Records of the Indian Museum, 50, 173 - 263." type="journal article" year="1953">Silas (1953)</bibRefCitation>
cannot differentiate the two genera.
<bibRefCitation id="C94B1CDFA91E9707FDA7FDF2A5880F4C" author="Tan" box="[638,761,608,632]" pageId="20" pageNumber="77" refString="Tan, H. H. (2009) A new species of hill stream loach (Teleostei: Balitoridae) from central Kalimantan, with redescriptions of Homaloptera tateregani Popta and Homaloptera stephensoni Hora. Zootaxa, 2171, 48 - 64." type="journal article" year="2009">Tan (2009)</bibRefCitation>
gave a simple pelvic-fin ray count of 3 for
<taxonomicName id="6ADA1AADA91E9707FB3AFDF3A20E0F43" ID-CoL="3MDX2" box="[1251,1407,608,632]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="species" species="tatereganii">
<emphasis id="9FAEBD3CA91E9707FB3AFDF3A20E0F43" box="[1251,1407,608,632]" italics="true" pageId="20" pageNumber="77">H. tatereganii</emphasis>
</taxonomicName>
to distinguish it from species of
<taxonomicName id="6ADA1AADA91E9707FE3EFD16A50D0FA8" ID-CoL="4Y8C" box="[487,636,644,668]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91E9707FE3EFD16A50D0FA8" box="[487,636,644,668]" italics="true" pageId="20" pageNumber="77">Homaloptera</emphasis>
</taxonomicName>
(s.l.), which have 2 simple pelvic-fin rays. However, the
<typeStatus id="7261DF8CA91E9707FAD0FD16A21F0FA9" box="[1289,1390,644,669]" pageId="20" pageNumber="77" type="holotype">holotype</typeStatus>
, the only known specimen of
<taxonomicName id="6ADA1AADA91E9707FE65FD3BA52B0F8B" ID-CoL="3MDX2" box="[444,602,680,704]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="species" species="tatereganii">
<emphasis id="9FAEBD3CA91E9707FE65FD3BA52B0F8B" box="[444,602,680,704]" italics="true" pageId="20" pageNumber="77">H. tatereganii</emphasis>
</taxonomicName>
(RMNH 7632), has only 2 simple pelvic-fin rays (on both sides). Other counts that differ from those given by
<bibRefCitation id="C94B1CDFA91E9707FD93FD5FA5B70FD0" author="Tan" box="[586,710,716,740]" pageId="20" pageNumber="77" refString="Tan, H. H. (2009) A new species of hill stream loach (Teleostei: Balitoridae) from central Kalimantan, with redescriptions of Homaloptera tateregani Popta and Homaloptera stephensoni Hora. Zootaxa, 2171, 48 - 64." type="journal article" year="2009">Tan (2009)</bibRefCitation>
are the following: iii, 8½ vs. ii, 8 dorsal-fin rays; ii, 5½ vs. ii, 5 anal-fin rays; vii, 12 vs. viii, 12 pectoral-fin rays; 18 vs. 14 circumpeduncular scale count; and 6/7 vs. 5/6 transverse scale count. The following measurements differ from
<bibRefCitation id="C94B1CDFA91E9707FCA5FC87A48B0E18" author="Tan" box="[892,1018,788,812]" pageId="20" pageNumber="77" refString="Tan, H. H. (2009) A new species of hill stream loach (Teleostei: Balitoridae) from central Kalimantan, with redescriptions of Homaloptera tateregani Popta and Homaloptera stephensoni Hora. Zootaxa, 2171, 48 - 64." type="journal article" year="2009">Tan (2009)</bibRefCitation>
(owing likely to different methods): predorsal length 44.1% vs. 45.3% SL; body depth 12.5% vs. 10.4% SL; dorsal-fin base 16.7% vs. 18.8% SL; pectoral-fin length 29.4% vs. 28.5% SL; head depth 48.2% vs. 42.8% HL; head width 81.3% vs. 78.3% HL; snout length 61.9% vs. 57.2% HL; 15.1% vs. 14.5% HL; 40.3% vs. 37.7% HL.
</paragraph>
</subSubSection>
<caption id="F9A531A6A91E9707FF4EF81AA48C0A89" httpUri="https://zenodo.org/record/288024/files/figure.png" pageId="20" pageNumber="77" targetBox="[210,1370,982,1890]" targetPageId="20">
<paragraph id="AD65612EA91E9707FF4EF81AA48C0A89" blockId="20.[151,1436,1928,1981]" pageId="20" pageNumber="77">
<emphasis id="9FAEBD3CA91E9707FF4EF81AA6530AA9" bold="true" box="[151,290,1928,1950]" pageId="20" pageNumber="77">FIGURE 14.</emphasis>
Dorsal, lateral, and ventral views of preserved
<taxonomicName id="6ADA1AADA91E9707FCCCF81AA3290AAA" box="[789,1112,1928,1950]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="77" phylum="Chordata" rank="species" species="tatereganii">
<emphasis id="9FAEBD3CA91E9707FCCCF81AA3290AAA" box="[789,1112,1928,1950]" italics="true" pageId="20" pageNumber="77">Pseudohomaloptera tatereganii</emphasis>
</taxonomicName>
, RMNH 7632 (holotype), 64.6 mm SL, Bo River, Upper Mahakam River basin, East Kalimantan, Borneo, Indonesia.
</paragraph>
</caption>
<caption id="F9A531A6A91F9706FF4EF92EA2690BE6" box="[151,1304,1724,1746]" httpUri="https://zenodo.org/record/288025/files/figure.png" pageId="21" pageNumber="78" targetBox="[230,1337,216,1686]" targetPageId="21">
<paragraph id="AD65612EA91F9706FF4EF92EA2690BE6" blockId="21.[151,1304,1724,1746]" box="[151,1304,1724,1746]" pageId="21" pageNumber="78">
<emphasis id="9FAEBD3CA91F9706FF4EF92EA66E0BE5" bold="true" box="[151,287,1724,1746]" pageId="21" pageNumber="78">FIGURE 15.</emphasis>
Type localities for species of
<taxonomicName id="6ADA1AADA91F9706FD97F92EA46B0BE6" box="[590,794,1724,1746]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91F9706FD97F92EA46B0BE6" box="[590,794,1724,1746]" italics="true" pageId="21" pageNumber="78">Pseudohomaloptera</emphasis>
</taxonomicName>
. Asterisk represents the type species of the genus.
</paragraph>
</caption>
<subSubSection id="E5C032A5A91F9705FF1EF96DA6570FC4" lastPageId="22" lastPageNumber="79" pageId="21" pageNumber="78" type="diagnosis">
<paragraph id="AD65612EA91F9706FF1EF96DA54D0AC4" blockId="21.[151,1436,1791,2032]" pageId="21" pageNumber="78">
<emphasis id="9FAEBD3CA91F9706FF1EF96DA64B0A2C" bold="true" box="[199,314,1791,1816]" pageId="21" pageNumber="78">Diagnosis</emphasis>
. Distinguishing characters are given in
<tableCitation id="E0585495A91F9706FCFCF96DA40E0A23" box="[805,895,1791,1816]" captionStart="TABLE 4" captionStartId="7.[125,207,600,621]" captionTargetBox="[125,1823,686,815]" captionTargetPageId="7" captionText="TABLE 4. Characters distinguishing Homaloptera, Homalopteroides, Homalopterula, Balitoropsis, Pseudohomaloptera and Ghatsa. Values are given as ranges; numbers in parentheses represent modal values. For Ghatsa, data from original descriptions are included; values in parentheses are for G. montana (CAS-SU 39871)." httpUri="http://table.plazi.org/id/F9A531A6A90D9714FFA4FDCAA1230FBC" pageId="21" pageNumber="78" tableUuid="F9A531A6A90D9714FFA4FDCAA1230FBC">Table 4</tableCitation>
and shown in
<figureCitation id="35E17DABA91F9706FB99F96DA3C10A23" box="[1088,1200,1791,1816]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[211,1328,347,1828]" captionTargetId="figure@8.[151,1436,336,1848]" captionTargetPageId="8" captionText="FIGURE 3. Lateral views of living (A) Homaloptera orthogoniata, not cataloged, Kalimantan, Borneo, Indonesia. Strong reddish tints on body likely breeding colors; (B) Homalopteroides smithi, UF 235740, 45.0 mm SL, Khlong Tasae, Salui Subdistrict, Chumphon, Thailand; (C) Homalopterula cf. ripleyi, not cataloged, Sumatra, Indonesia; (D) Balitoropsis zollingeri, not cataloged, Kalimantan, Borneo, Indonesia; (E) Pseudohomaloptera leonardi, UF 235735, 34.2 mm SL, Ta Pi River, Nakhon Si Thammarat, Thailand. Photos (A) &amp; (D) by Renny Hadiaty. Photo (C) by Daniel Lumbantobing. Specimens not to scale." httpUri="https://zenodo.org/record/288013/files/figure.png" pageId="21" pageNumber="78">Figures 3</figureCitation>
E, 4B, 5E, and 14.
<taxonomicName id="6ADA1AADA91F9706FF4EF8B1A6080A0F" box="[151,377,1827,1851]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91F9706FF4EF8B1A6080A0F" box="[151,377,1827,1851]" italics="true" pageId="21" pageNumber="78">Pseudohomaloptera</emphasis>
</taxonomicName>
is distinguished by the following combination of characters: without reddish tints on fins in life (
<figureCitation id="35E17DABA91F9706FF14F8D5A6630A6B" box="[205,274,1863,1888]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[211,1328,347,1828]" captionTargetId="figure@8.[151,1436,336,1848]" captionTargetPageId="8" captionText="FIGURE 3. Lateral views of living (A) Homaloptera orthogoniata, not cataloged, Kalimantan, Borneo, Indonesia. Strong reddish tints on body likely breeding colors; (B) Homalopteroides smithi, UF 235740, 45.0 mm SL, Khlong Tasae, Salui Subdistrict, Chumphon, Thailand; (C) Homalopterula cf. ripleyi, not cataloged, Sumatra, Indonesia; (D) Balitoropsis zollingeri, not cataloged, Kalimantan, Borneo, Indonesia; (E) Pseudohomaloptera leonardi, UF 235735, 34.2 mm SL, Ta Pi River, Nakhon Si Thammarat, Thailand. Photos (A) &amp; (D) by Renny Hadiaty. Photo (C) by Daniel Lumbantobing. Specimens not to scale." httpUri="https://zenodo.org/record/288013/files/figure.png" pageId="21" pageNumber="78">Fig. 3</figureCitation>
E); dorsal-fin origin anterior to or above pelvic-fin origin; 8½ branched dorsal-fin rays; 89 branched pelvic-fin rays; forked caudal fin; keeled scales (
<figureCitation id="35E17DABA91F9706FD1DF8F9A47B0AB7" box="[708,778,1899,1924]" captionStart="FIGURE 4" captionStartId="9.[151,250,1660,1682]" captionTargetBox="[155,1431,193,1638]" captionTargetId="figure@9.[155,1432,193,1638]" captionTargetPageId="9" captionText="FIGURE 4. Scale sizes and keel patterns of predorsal region: Large, and keeled scales, (A) Balitoropsis zollingeri, UF 166094, 52.1 mm SL; (B) Pseudohomaloptera tatereganii, RMNH 7632 (holotype), 64.6 mm SL. Large, and wart-like / keeled scales (C) Homalopteroides wassinkii, UF 183330, 43.6 mm SL. Medium, and keeled scales, (D) Homaloptera cf. ocellata, UF 161718, 64.1 mm SL. Small, and smooth scales, (E) Homalopterula ripleyi, ANSP 188908, 43.1 mm SL; (F) Ghatsa montana, CAS-SU 39871 (holotype), 46.6 mm SL. Scale bar represents 1 mm. Left side of photo is anterior." httpUri="https://zenodo.org/record/288014/files/figure.png" pageId="21" pageNumber="78">Fig. 4</figureCitation>
B); 5061 total lateral-line scales; 1319 predorsal scales; anus closer to anal-fin origin than to pelvic-fin insertion; no adipose keel on caudal peduncle; large rostral cap; 2 thick rostral barbels in close proximity to one another; thick and triangular/crescentic upper lip; fleshy pad between lateral portions of lower lip (
<figureCitation id="35E17DABA91F9706FE01F845A56A0AC4" box="[472,539,2007,2032]" captionStart="FIGURE 5" captionStartId="10.[151,250,1357,1379]" captionTargetBox="[150,1436,267,1335]" captionTargetId="figure@10.[150,1436,267,1335]" captionTargetPageId="10" captionText="FIGURE 5. Oral morphology of (A) Homaloptera cf. ocellata, UF 161718, 64.1 mm SL; (B) Homalopteroides wassinkii, UMMZ 155660, 46.2 mm SL; (C) Homalopterula ripleyi, ANSP 188908, 43.1 mm SL; (D) Balitoropsis zollingeri, UF 166094, 52.1 mm SL; (E) Pseudohomaloptera tatereganii, RMNH 7632 (holotype), 64.6 mm SL; (F) Ghatsa montana, (holotype) CAS- SU 39871, 46.4 mm SL. Abbreviations: CF, central furrow; FL, fleshy lobe; FP, fleshy pad; LJ, lower jaw; LL, lower lip; LRB, lateral-rostral barbel; MB, maxillary barbel; MRB, medial-rostral barbel; RC, rostral cap; UJ, upper jaw; UL, upper lip." httpUri="https://zenodo.org/record/288015/files/figure.png" pageId="21" pageNumber="78">Fig. 5</figureCitation>
E).
</paragraph>
<paragraph id="AD65612EA91C9705FF1EFF05A4E30DCC" blockId="22.[151,1436,151,1076]" pageId="22" pageNumber="79">
<emphasis id="9FAEBD3CA91C9705FF1EFF05A6E70D84" bold="true" box="[199,406,151,176]" pageId="22" pageNumber="79">Species included.</emphasis>
<taxonomicName id="6ADA1AADA91C9705FE79FF0AA4D70D84" authority="Popta 1905" authorityName="Popta" authorityYear="1905" box="[416,934,151,176]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="tatereganii">
<emphasis id="9FAEBD3CA91C9705FE79FF0AA4770D9B" box="[416,774,152,176]" italics="true" pageId="22" pageNumber="79">Pseudohomaloptera tatereganii</emphasis>
(Popta 1905)
</taxonomicName>
,
<taxonomicName id="6ADA1AADA91C9705FC6CFF0BA2610D84" authority="Fowler 1934" authorityName="Fowler" authorityYear="1934" box="[949,1296,151,176]" family="Balitoridae" genus="Balitoropsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="sexmaculata">
<emphasis id="9FAEBD3CA91C9705FC6CFF0BA3110D84" box="[949,1120,152,176]" italics="true" pageId="22" pageNumber="79">P. sexmaculata</emphasis>
(Fowler 1934)
</taxonomicName>
,
<taxonomicName id="6ADA1AADA91C9705FAC6FF0BA6590DE0" authority="Hora 1941" authorityName="Hora" authorityYear="1941" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="9FAEBD3CA91C9705FAC6FF0BA2ED0D84" box="[1311,1436,152,176]" italics="true" pageId="22" pageNumber="79">P. leonardi</emphasis>
(Hora 1941)
</taxonomicName>
,
<taxonomicName id="6ADA1AADA91C9705FEEEFF2FA5F00DE0" authority="Chen 1978" authorityName="Chen" authorityYear="1978" box="[311,641,188,213]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="9FAEBD3CA91C9705FEEEFF2FA6900DE0" box="[311,481,189,212]" italics="true" pageId="22" pageNumber="79">P. yunnanensis</emphasis>
(Chen 1978)
</taxonomicName>
,
<taxonomicName id="6ADA1AADA91C9705FD48FF2FA3570DE0" authority="Kottelat &amp; Chu 1988" authorityName="Kottelat &amp; Chu" authorityYear="1988" box="[657,1062,188,213]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="vulgaris">
<emphasis id="9FAEBD3CA91C9705FD48FF2FA47D0DE0" box="[657,780,188,212]" italics="true" pageId="22" pageNumber="79">P. vulgaris</emphasis>
(
<bibRefCitation id="C94B1CDFA91C9705FCC6FF2EA36F0DE0" author="Kottelat" box="[799,1054,188,213]" pageId="22" pageNumber="79" refString="Kottelat, M. &amp; Chu, X. - L. (1988) The genus Homaloptera (Osteichthyes, Cypriniformes, Homalopteridae) in Yunnan, China. Cybium, 12, 103 - 106." type="journal article" year="1988">Kottelat &amp; Chu 1988</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="6ADA1AADA91C9705FBB3FF2FA2240DE0" authority="Tan 2009" authorityName="Tan" authorityYear="2009" box="[1130,1365,188,212]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="batek">
<emphasis id="9FAEBD3CA91C9705FBB3FF2FA3B60DE0" box="[1130,1223,188,212]" italics="true" pageId="22" pageNumber="79">P. batek</emphasis>
(
<bibRefCitation id="C94B1CDFA91C9705FB03FF2EA23F0DE0" author="Tan" box="[1242,1358,188,212]" pageId="22" pageNumber="79" refString="Tan, H. H. (2009) A new species of hill stream loach (Teleostei: Balitoridae) from central Kalimantan, with redescriptions of Homaloptera tateregani Popta and Homaloptera stephensoni Hora. Zootaxa, 2171, 48 - 64." type="journal article" year="2009">Tan 2009</bibRefCitation>
)
</taxonomicName>
.
<typeStatus id="7261DF8CA91C9705FABDFF2FA2ED0DE1" box="[1380,1436,189,213]" pageId="22" pageNumber="79">Type</typeStatus>
localities for species of
<taxonomicName id="6ADA1AADA91C9705FE78FF72A5F20DCC" box="[417,643,224,248]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91C9705FE78FF72A5F20DCC" box="[417,643,224,248]" italics="true" pageId="22" pageNumber="79">Pseudohomaloptera</emphasis>
</taxonomicName>
are shown in
<figureCitation id="35E17DABA91C9705FCC6FF4DA4FF0DCC" box="[799,910,223,248]" captionStart="FIGURE 15" captionStartId="21.[151,250,1724,1746]" captionTargetBox="[230,1337,216,1686]" captionTargetId="figure@21.[210,1377,193,1703]" captionTargetPageId="21" captionText="FIGURE 15. Type localities for species of Pseudohomaloptera. Asterisk represents the type species of the genus." httpUri="https://zenodo.org/record/288025/files/figure.png" pageId="22" pageNumber="79">Figure 15</figureCitation>
.
</paragraph>
<paragraph id="AD65612EA91C9705FF1EFE96A3E00C74" blockId="22.[151,1436,151,1076]" pageId="22" pageNumber="79">
<emphasis id="9FAEBD3CA91C9705FF1EFE96A6130C29" bold="true" box="[199,354,260,285]" pageId="22" pageNumber="79">Comparison.</emphasis>
<taxonomicName id="6ADA1AADA91C9705FEB0FE96A53A0C28" box="[361,587,260,284]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91C9705FEB0FE96A53A0C28" box="[361,587,260,284]" italics="true" pageId="22" pageNumber="79">Pseudohomaloptera</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="6ADA1AADA91C9705FC9CFE96A4AB0C28" box="[837,986,260,284]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91C9705FC9CFE96A4AB0C28" box="[837,986,260,284]" italics="true" pageId="22" pageNumber="79">Homaloptera</emphasis>
</taxonomicName>
by absence vs. presence of reddish tints on fins in life; 89, 8 (M) vs. 7 branched pelvic fin-rays; 1319 vs. 2027 predorsal scales.
</paragraph>
<paragraph id="AD65612EA91C9705FF1EFEDEA3FB0CE4" blockId="22.[151,1436,151,1076]" pageId="22" pageNumber="79">
<taxonomicName id="6ADA1AADA91C9705FF1EFEDEA6D80C50" box="[199,425,332,356]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91C9705FF1EFEDEA6D80C50" box="[199,425,332,356]" italics="true" pageId="22" pageNumber="79">Pseudohomaloptera</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="6ADA1AADA91C9705FD78FEDEA4120C50" box="[673,867,332,356]" class="Actinopterygii" family="Balitoridae" genus="Homalopteroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91C9705FD78FEDEA4120C50" box="[673,867,332,356]" italics="true" pageId="22" pageNumber="79">Homalopteroides</emphasis>
</taxonomicName>
by having dorsal-fin origin anterior to or at pelvicfin origin vs. posterior to pelvic-fin origin; 8½ vs. 68½, 7½ (M) branched dorsal-fin rays; large vs. small rostral cap; medial- and lateral-rostral barbels in close proximity to one another vs. barbels widely separated at base; thick vs. thin upper lip; presence vs. absence of fleshy pad between lateral portions of lower lip.
</paragraph>
<paragraph id="AD65612EA91C9705FF1EFE4EA5440F9C" blockId="22.[151,1436,151,1076]" pageId="22" pageNumber="79">
<taxonomicName id="6ADA1AADA91C9705FF1EFE4EA6D80CC0" box="[199,425,476,500]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91C9705FF1EFE4EA6D80CC0" box="[199,425,476,500]" italics="true" pageId="22" pageNumber="79">Pseudohomaloptera</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="6ADA1AADA91C9705FD71FE4EA4220CC0" box="[680,851,476,500]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91C9705FD71FE4EA4220CC0" box="[680,851,476,500]" italics="true" pageId="22" pageNumber="79">Homalopterula</emphasis>
</taxonomicName>
by having dorsal-fin origin anterior to or at pelvicfin origin vs. posterior to pelvic-fin origin; 8½ vs. 5½ and 7½, 7½ (M) branched dorsal-fin rays; 89 vs. 7 branched pelvic fin-rays; keeled vs. smooth scales; 1319 vs. 2856 predorsal scales; forked vs. truncated or emarginated caudal fin; absence vs. presence of adipose keel on caudal peduncle; large vs. small rostral cap; medial- and lateralrostral barbels in close proximity to one another vs. widely separated at base; presence of fleshy pad vs. lobes between lateral portions of lower lip.
</paragraph>
<paragraph id="AD65612EA91C9705FF1EFD26A6570FC4" blockId="22.[151,1436,151,1076]" pageId="22" pageNumber="79">
<taxonomicName id="6ADA1AADA91C9705FF1EFD26A6D80FF8" box="[199,425,692,716]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91C9705FF1EFD26A6D80FF8" box="[199,425,692,716]" italics="true" pageId="22" pageNumber="79">Pseudohomaloptera</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="6ADA1AADA91C9705FD7CFD26A45C0FF8" box="[677,813,692,716]" family="Balitoridae" genus="Balitoropsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA91C9705FD7CFD26A45C0FF8" box="[677,813,692,716]" italics="true" pageId="22" pageNumber="79">Balitoropsis</emphasis>
</taxonomicName>
by having anus closer to anal-fin origin than to pelvicfin insertion.
</paragraph>
</subSubSection>
<subSubSection id="E5C032A5A91C9705FF1EFD6EA6F10900" pageId="22" pageNumber="79" type="materials_examined">
<paragraph id="AD65612EA91C9705FF1EFD6EA6F10900" blockId="22.[151,1436,151,1076]" pageId="22" pageNumber="79">
<emphasis id="9FAEBD3CA91C9705FF1EFD6EA4330E21" bold="true" box="[199,834,764,789]" pageId="22" pageNumber="79">
Material examined.
<taxonomicName id="6ADA1AADA91C9705FE1EFD6EA4330E21" authority="Borneo" authorityName="Borneo" box="[455,834,764,789]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="tatereganii">
<emphasis id="9FAEBD3CA91C9705FE1EFD6EA4480E21" bold="true" box="[455,825,764,789]" italics="true" pageId="22" pageNumber="79">Pseudohomaloptera tatereganii</emphasis>
:
</taxonomicName>
</emphasis>
Borneo:
<collectionCode id="CBCBF9EBA91C9705FC62FD6FA37E0E20" LSID="urn:lsid:biocol.org:col:34992" box="[955,1039,765,788]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="79">RMNH</collectionCode>
7632 (
<typeStatus id="7261DF8CA91C9705FBB4FD6EA3A00E21" box="[1133,1233,764,789]" pageId="22" pageNumber="79" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FADFFD6EA6600E03" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="tatereganii">
<emphasis id="9FAEBD3CA91C9705FADFFD6EA6600E03" italics="true" pageId="22" pageNumber="79">Homaloptera tatereganii</emphasis>
</taxonomicName>
).
<emphasis id="9FAEBD3CA91C9705FEFEFCB2A6AC0E0C" bold="true" box="[295,477,799,824]" pageId="22" pageNumber="79">
<taxonomicName id="6ADA1AADA91C9705FEFEFCB2A6A20E0C" box="[295,467,799,824]" family="Balitoridae" genus="Balitoropsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="sexmaculata">
<emphasis id="9FAEBD3CA91C9705FEFEFCB2A6A20E0C" bold="true" box="[295,467,799,824]" italics="true" pageId="22" pageNumber="79">P. sexmaculata</emphasis>
</taxonomicName>
:
</emphasis>
<collectingCountry id="D5CD21BEA91C9705FE3AFC8DA53A0E0C" box="[483,587,799,824]" name="Thailand" pageId="22" pageNumber="79">Thailand</collectingCountry>
:
<collectionCode id="CBCBF9EBA91C9705FD8DFCB2A5ED0E0C" LSID="urn:lsid:biocol.org:col:34784" box="[596,668,800,824]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="22" pageNumber="79">ANSP</collectionCode>
56374 (
<typeStatus id="7261DF8CA91C9705FD2FFC8DA42B0E0C" box="[758,858,799,824]" pageId="22" pageNumber="79" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FCA7FCB2A3D70E0C" box="[894,1190,800,824]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="sexmaculata">
<emphasis id="9FAEBD3CA91C9705FCA7FCB2A3D70E0C" box="[894,1190,800,824]" italics="true" pageId="22" pageNumber="79">Homaloptera sexmaculata</emphasis>
</taxonomicName>
), 56375 (
<typeStatus id="7261DF8CA91C9705FAC8FCB2A20F0E0C" box="[1297,1406,800,824]" pageId="22" pageNumber="79" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FF4EFCD6A6BD0E68" box="[151,460,836,860]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="sexmaculata">
<emphasis id="9FAEBD3CA91C9705FF4EFCD6A6BD0E68" box="[151,460,836,860]" italics="true" pageId="22" pageNumber="79">Homaloptera sexmaculata</emphasis>
</taxonomicName>
) (2), 56402 (
<typeStatus id="7261DF8CA91C9705FD5FFCD6A59B0E69" box="[646,746,836,861]" pageId="22" pageNumber="79" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FCF0FCD6A3F80E68" box="[809,1161,836,860]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="septemmaculata">
<emphasis id="9FAEBD3CA91C9705FCF0FCD6A3F80E68" box="[809,1161,836,860]" italics="true" pageId="22" pageNumber="79">Homaloptera septemmaculata</emphasis>
</taxonomicName>
), 56403 (
<typeStatus id="7261DF8CA91C9705FAD7FCD6A2010E68" box="[1294,1392,836,860]" pageId="22" pageNumber="79" type="paratype">paratype</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FF4EFCFAA69D0EB4" box="[151,492,872,896]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="septemmaculata">
<emphasis id="9FAEBD3CA91C9705FF4EFCFAA69D0EB4" box="[151,492,872,896]" italics="true" pageId="22" pageNumber="79">Homaloptera septemmaculata</emphasis>
</taxonomicName>
); UF 183358 (2), 177819 (3),
<date id="D96447EEA91C9705FC96FCFAA4D30EB4" box="[847,930,872,896]" pageId="22" pageNumber="79" value="70-11-18">181170</date>
(3).
<taxonomicName id="6ADA1AADA91C9705FC04FCFAA3950EB4" authority="Peninsular" authorityName="Peninsular" box="[989,1252,871,896]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="9FAEBD3CA91C9705FC04FCFAA3150EB4" bold="true" box="[989,1124,871,896]" pageId="22" pageNumber="79">
<emphasis id="9FAEBD3CA91C9705FC04FCFAA32A0EB4" bold="true" box="[989,1115,871,896]" italics="true" pageId="22" pageNumber="79">P. leonardi</emphasis>
:
</emphasis>
Peninsular
</taxonomicName>
<collectingCountry id="D5CD21BEA91C9705FB37FCF5A2280EB4" box="[1262,1369,871,896]" name="Malaysia" pageId="22" pageNumber="79">Malaysia</collectingCountry>
:
<collectionCode id="CBCBF9EBA91C9705FABCFCFAA2EA0EB4" box="[1381,1435,872,896]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="79">ZRC</collectionCode>
1753 (
<typeStatus id="7261DF8CA91C9705FF3FFC1EA6390E90" box="[230,328,908,932]" pageId="22" pageNumber="79" type="paratype">paratype</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FEA7FC1EA5F10E90" box="[382,640,908,932]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="9FAEBD3CA91C9705FEA7FC1EA5F10E90" box="[382,640,908,932]" italics="true" pageId="22" pageNumber="79">Homaloptera leonardi</emphasis>
</taxonomicName>
);
<collectionCode id="CBCBF9EBA91C9705FD7BFC1FA5870E90" LSID="urn:lsid:biocol.org:col:34992" box="[674,758,909,932]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="79">RMNH</collectionCode>
23264 (6), 25921 (4); UF 169909 (3), 235746 (3).
<emphasis id="9FAEBD3CA91C9705FA5EFC1EA6430EFC" bold="true" pageId="22" pageNumber="79">
<taxonomicName id="6ADA1AADA91C9705FA5EFC1EA6580EFC" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="9FAEBD3CA91C9705FA5EFC1EA6580EFC" bold="true" italics="true" pageId="22" pageNumber="79">P. yunnanensis</emphasis>
</taxonomicName>
:
</emphasis>
<collectingCountry id="D5CD21BEA91C9705FEE0FC3DA6F00EFC" box="[313,385,943,968]" name="China" pageId="22" pageNumber="79">China</collectingCountry>
:
<collectionCode id="CBCBF9EBA91C9705FE53FC22A6970EFC" box="[394,486,944,968]" httpUri="http://grbio.org/cool/n2yr-c98h" name="Institute of Hydrobiology, Academia Sinica" pageId="22" pageNumber="79">IHASW</collectionCode>
<date id="D96447EEA91C9705FE35FC3DA5310EFC" box="[492,576,943,968]" bridgedPair="-" pageId="22" pageNumber="79" value="1960-07">60-VII-</date>
012 (
<typeStatus id="7261DF8CA91C9705FDA0FC3DA5AC0EFC" box="[633,733,943,968]" pageId="22" pageNumber="79" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FCD8FC22A36B0EF3" box="[769,1050,944,968]" family="Balitoridae" genus="Balitoropsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="9FAEBD3CA91C9705FCD8FC22A36B0EF3" box="[769,1050,944,968]" italics="true" pageId="22" pageNumber="79">Balitoropsis yunnanensis</emphasis>
</taxonomicName>
).
<emphasis id="9FAEBD3CA91C9705FBE8FC22A3C30EFC" bold="true" box="[1073,1202,943,968]" pageId="22" pageNumber="79">
<taxonomicName id="6ADA1AADA91C9705FBE8FC22A3D80EFC" box="[1073,1193,943,968]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="vulgaris">
<emphasis id="9FAEBD3CA91C9705FBE8FC22A3D80EFC" bold="true" box="[1073,1193,943,968]" italics="true" pageId="22" pageNumber="79">P. vulgaris</emphasis>
</taxonomicName>
:
</emphasis>
<collectingCountry id="D5CD21BEA91C9705FB61FC3DA38E0EFC" box="[1208,1279,943,968]" name="China" pageId="22" pageNumber="79">China</collectingCountry>
: 788229 (
<collectionCode id="CBCBF9EBA91C9705FAB2FC22A2ED0EFC" box="[1387,1436,944,968]" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="22" pageNumber="79">KIZ</collectionCode>
1978001047) (
<typeStatus id="7261DF8CA91C9705FE91FC46A6DD0ED9" box="[328,428,980,1005]" pageId="22" pageNumber="79" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FE34FC46A5820ED8" box="[493,755,980,1004]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="vulgaris">
<emphasis id="9FAEBD3CA91C9705FE34FC46A5820ED8" box="[493,755,980,1004]" italics="true" pageId="22" pageNumber="79">Homaloptera vulgaris</emphasis>
</taxonomicName>
), 788225-788227 (
<collectionCode id="CBCBF9EBA91C9705FC3FFC46A3670ED8" box="[998,1046,980,1004]" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="22" pageNumber="79">KIZ</collectionCode>
1978001048-50) (
<typeStatus id="7261DF8CA91C9705FADAFC46A2010ED8" box="[1283,1392,980,1004]" pageId="22" pageNumber="79" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FF4EFC6AA6E10924" box="[151,400,1016,1040]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="vulgaris">
<emphasis id="9FAEBD3CA91C9705FF4EFC6AA6E10924" box="[151,400,1016,1040]" italics="true" pageId="22" pageNumber="79">Homaloptera vulgaris</emphasis>
</taxonomicName>
).
<taxonomicName id="6ADA1AADA91C9705FE7EFC6AA518093B" authority="Borneo" authorityName="Borneo" box="[423,617,1015,1040]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="batek">
<emphasis id="9FAEBD3CA91C9705FE7EFC6AA57A0924" bold="true" box="[423,523,1015,1040]" pageId="22" pageNumber="79">
<emphasis id="9FAEBD3CA91C9705FE7EFC6AA5700924" bold="true" box="[423,513,1015,1040]" italics="true" pageId="22" pageNumber="79">P. batek</emphasis>
:
</emphasis>
Borneo
</taxonomicName>
:
<collectionCode id="CBCBF9EBA91C9705FDADFC6AA5C1093B" box="[628,688,1016,1039]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="22" pageNumber="79">MZB</collectionCode>
10990 (
<typeStatus id="7261DF8CA91C9705FCD5FC65A4010924" box="[780,880,1015,1040]" pageId="22" pageNumber="79" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FC4FFC6AA3010924" box="[918,1136,1016,1040]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="batek">
<emphasis id="9FAEBD3CA91C9705FC4FFC6AA3010924" box="[918,1136,1016,1040]" italics="true" pageId="22" pageNumber="79">Homaloptera batek</emphasis>
</taxonomicName>
);
<collectionCode id="CBCBF9EBA91C9705FB51FC6AA3CF0924" box="[1160,1214,1016,1040]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="79">ZRC</collectionCode>
51743 (
<typeStatus id="7261DF8CA91C9705FAC2FC6AA20C0924" box="[1307,1405,1016,1040]" pageId="22" pageNumber="79" type="paratype">paratype</typeStatus>
of
<taxonomicName id="6ADA1AADA91C9705FF4EFB8EA61E0900" box="[151,367,1052,1076]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="79" phylum="Chordata" rank="species" species="batek">
<emphasis id="9FAEBD3CA91C9705FF4EFB8EA61E0900" box="[151,367,1052,1076]" italics="true" pageId="22" pageNumber="79">Homaloptera batek</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>