139 lines
17 KiB
XML
139 lines
17 KiB
XML
<document id="FF724592EC221E82404A4787989C32CB" ID-CLB-Dataset="34950" ID-DOI="http://dx.doi.org/10.3897/zookeys.600.8913" ID-GBIF-Dataset="2a0dd984-89ac-4139-9cb4-a9dbb7c889eb" ID-PMC="PMC4926682" ID-Pensoft-Pub="1313-2970-600-75" ID-PubMed="27408601" ID-ZooBank="93F40A39F3B54BADBD61D3C8AFC36AFC" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-600-75" ModsDocOrigin="ZooKeys 600" ModsDocTitle="A preliminary synopsis on amber scorpions with special reference to Burmite species: an extraordinary development of our knowledge in only 20 years" checkinTime="1466665100100" checkinUser="pensoft" docAuthor="Lourenco, Wilson R." docDate="2016" docId="25101D0486B803E7CDE32E1E4412E947" docLanguage="en" docName="ZooKeys 600: 75-87" docOrigin="ZooKeys 600" docSource="http://dx.doi.org/10.3897/zookeys.600.8913" docTitle="Betaburmesebuthus bellus Lourenco, 2016, sp. n." docType="treatment" docUuid="21267812-28A4-4D1A-BD96-A2D71ACC11F8" docUuidSource="ZooBank" docVersion="8" lastPageNumber="82" masterDocId="CE36FFB61035FFC3FFD7E111FFAEE806" masterDocTitle="A preliminary synopsis on amber scorpions with special reference to Burmite species: an extraordinary development of our knowledge in only 20 years" masterLastPageNumber="87" masterPageNumber="75" pageNumber="78" updateTime="1732795294725" updateUser="ExternalLinkService">
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<mods:title id="4CA28BCC5414D1A551675974A9B7D3E2">A preliminary synopsis on amber scorpions with special reference to Burmite species: an extraordinary development of our knowledge in only 20 years</mods:title>
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<mods:name id="DE21F4A7E0CA2BA64BD559FC82AF3A9F" type="personal">
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<mods:namePart id="9D49BE5081BA9F8B78C0355728694882">Lourenco, Wilson R.</mods:namePart>
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<mods:title id="0C1F3268D9CA676AA482A214B3B260D6">ZooKeys</mods:title>
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<mods:date id="8DD32782632C1AE969524F696805A52F">2016</mods:date>
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<mods:number id="4991B6385454CBDA682C901AC169ADAC">600</mods:number>
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<mods:start id="01289F5D2BDA909921A339F62FC55B86">75</mods:start>
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<mods:url id="FFD28C0007AE411D271EBF3DCAFDA2D7">http://dx.doi.org/10.3897/zookeys.600.8913</mods:url>
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<mods:classification id="489CDDD94DB9F961F44072D1C7C22954">journal article</mods:classification>
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<mods:identifier id="AFA5B77797A069EC254C6E8F6624FB94" type="DOI">http://dx.doi.org/10.3897/zookeys.600.8913</mods:identifier>
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<mods:identifier id="6A6C2F1CB0B951B2A78A5683FB9B6192" type="Pensoft-Pub">1313-2970-600-75</mods:identifier>
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<mods:identifier id="7614D425E8D87F2E4E2FDBC8D15018EF" type="ZooBank">93F40A39F3B54BADBD61D3C8AFC36AFC</mods:identifier>
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<treatment id="25101D0486B803E7CDE32E1E4412E947" ID-GBIF-Taxon="127876518" LSID="urn:lsid:zoobank.org:act:21267812-28A4-4D1A-BD96-A2D71ACC11F8" httpUri="http://treatment.plazi.org/id/25101D0486B803E7CDE32E1E4412E947" lastPageId="7" lastPageNumber="82" pageId="3" pageNumber="78" scope_family="Palaeoburmesebuthidae">
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<subSubSection id="14ABB33CCD954876F6C87AD230066882" pageId="3" pageNumber="78" type="multiple">
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<paragraph id="106D217B3055A54048E17A4C366D3EF5" pageId="3" pageNumber="78">Taxon classification Animalia Scorpiones Palaeoburmesebuthidae</paragraph>
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</subSubSection>
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<subSubSection id="492338B49AE68581E03792AD346D787F" pageId="3" pageNumber="78" type="nomenclature">
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<paragraph id="E59BE1550308F902FF893D31C080D4E0" pageId="3" pageNumber="78">
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<taxonomicName id="F017B29EEEA346A5B165F248B51600A7" LSID="http://zoobank.org/21267812-28A4-4D1A-BD96-A2D71ACC11F8" genus="Betaburmesebuthus" lsidName="Betaburmesebuthus bellus" pageId="3" pageNumber="78" rank="species" species="bellus">Betaburmesebuthus bellus</taxonomicName>
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<taxonomicNameLabel id="07DF4EF47C4A7FD791E2C0432222FD61" pageId="3" pageNumber="78">sp. n.</taxonomicNameLabel>
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Figures 1-4, 5-7, 8-9, 10
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</paragraph>
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</subSubSection>
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<subSubSection id="E7F1C80134A3EB6841FD598E52E2D112" lastPageId="4" lastPageNumber="79" pageId="3" pageNumber="78" type="holotype">
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<paragraph id="5F283A6E5A8A5FFBB7FCEE1A22EA6220" pageId="3" pageNumber="78">Holotype.</paragraph>
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<paragraph id="64A5F5FA7E8B9FFC6E72D0573422CC34" lastPageId="4" lastPageNumber="79" pageId="3" pageNumber="78">
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A juvenile, most certainly a male. Very clear block of pale yellow amber that measured 21.5
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<normalizedToken id="254188A36D58710083F5D2279F1A03CA" originalValue="×">x</normalizedToken>
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17.0
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<normalizedToken id="9F25E657262ED3E360972AD3C8E312D6" originalValue="×">x</normalizedToken>
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2 mm. Only a few inclusions and bubbles prevent a 100%
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<pageBreakToken id="311DF04F20F60580C5856014200F2AC2" pageId="4" pageNumber="79" start="start">observation</pageBreakToken>
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of all characters. Type locality and horizon: Myanmar (Burma), Kachin; precise locality unknown; Lower Cretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="2CE5DDFBA2DD106A18AD0C32210FA195" pageId="4" pageNumber="79" type="etymology">
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<paragraph id="B893BA19A6DD1A95ED77658ED1F4D5C8" pageId="4" pageNumber="79">Etymology.</paragraph>
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<paragraph id="9D2ED1B67EC396E6A863F81440ECDC36" pageId="4" pageNumber="79">
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The specific name is an epythet from Latin
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<normalizedToken id="5681AEC8616D58DC489420EFDFEB1ADC" originalValue="“bellus”">"bellus"</normalizedToken>
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meaning beautiful, elegant.
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</paragraph>
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</subSubSection>
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<subSubSection id="30FBA7E282FC805274DADF14D2C2E25E" pageId="4" pageNumber="79" type="repository">
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<paragraph id="7DD5477870E1DDE24D3249C3B37116B7" pageId="4" pageNumber="79">Repository.</paragraph>
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<paragraph id="F61BD79E2CD4809F4FC564EED1099582" pageId="4" pageNumber="79">
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The type specimen is deposited in the collection of the Museum of the Geological-Palaeontological Institut of the University of Hamburg (CeNak - Centrum of Natural History), Coll. N° 4586, Coll.
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<normalizedToken id="72C21AF4EF9DFCBBA2FC22FBD623637F" originalValue="Gröhn">Groehn</normalizedToken>
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N° 11086.
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</paragraph>
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</subSubSection>
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<subSubSection id="853C87F1852D6078CE235EE79DEB5F02" lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79" type="diagnosis">
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<paragraph id="08C6296BD3053D1FC3527AA417569C33" pageId="4" pageNumber="79">Diagnosis.</paragraph>
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<paragraph id="7F1708FC95CA67D29E21670ECD4DB6F2" lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79">
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General morphology is globally similar with other species of the genus
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<taxonomicName id="7898E0D866ACDB1E8C994F8BE8CD0A97" genus="Betaburmesebuthus" lsidName="Betaburmesebuthus" pageId="4" pageNumber="79" rank="genus">Betaburmesebuthus</taxonomicName>
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<normalizedToken id="245C535664E9EDD27782CFC0FE09E4E4" originalValue="Lourenço">Lourenco</normalizedToken>
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, 2015 and also with extant buthoid scorpions. The
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<pageBreakToken id="32AFB3315BB5EC740CAD8C5132C63825" pageId="5" pageNumber="80" start="start">following</pageBreakToken>
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combination of features can be used to diagnose the new species: Carapace weakly to moderately granulated; anterior margin with a moderately marked median concavity. Sternum subpentagonal. Tergites with three carinae, the lateral totally inconspicuous. Sternites with small oval to slit-like spiracle. Metasomal segments I and II with ten carinae; setation on all metasomal segments strongly marked. Fixed and movable fingers of pedipalp chela with a series of small rounded granules and no conspicuous spinoid accessory granules. Trichobothrial pattern with elements ressembling those of extant buthid type A (
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<bibRefCitation id="51B761E3937A8B47B0323F5E549A2FF3" author="Vachon, M" journalOrPublisher="Paris" pageId="11" pageNumber="86" pagination="857 - 958" title="Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions." volumeTitle="Bulletin du Museum national d'Histoire naturelle" year="1974">Vachon 1974</bibRefCitation>
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): at least 1-3 internal, five dorsal, and two external trichobothria on the femur; dorsal trichobothria disposed in beta (β) configuration (
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<bibRefCitation id="B564C2C99E3D9574F991AD821365F8D2" author="Vachon, M" journalOrPublisher="Comptes Rendus des Seances de l'Academie des Sciences, Paris" pageId="11" pageNumber="86" pagination="1597 - 1599" title="Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des Scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon." volume="281" year="1975">Vachon 1975</bibRefCitation>
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); one internal and five dorsal trichobothria on patella; 6-7 external trichobothria can be suggested on patella by the presence of fine setae; some are displaced on ventral aspect; 4-5 dorso-external and two ventral on chelal hand; six on fixed finger. Tibial spurs present on legs III and IV.
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</paragraph>
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</subSubSection>
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<subSubSection id="9361AE74C95CFEC3B1CC388665B23476" pageId="5" pageNumber="80" type="description">
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<paragraph id="290A07433CE8EFD614386B18B532B338" pageId="5" pageNumber="80">Description.</paragraph>
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<paragraph id="6AB8959D81DF0B998D76E5C08540DBD9" pageId="5" pageNumber="80">Coloration: the scorpion is yellow to slightly reddish-yellow; carapace and tergites yellow; metasomal segments yellow; telson reddish-yellow; pedipalps and legs yellow. Ventral aspect yellow; coxapophysis slightly darker than dorsal aspect.</paragraph>
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</subSubSection>
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<subSubSection id="5DA0A2DD48392A9F637AAD01FFB7D3C2" lastPageId="7" lastPageNumber="82" pageId="5" pageNumber="80" type="morphology">
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<paragraph id="E94A67F209C9F17CB3A98DB8A51A6CAF" pageId="5" pageNumber="80">Morphology.</paragraph>
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<paragraph id="2D94A4B766F2E70A740F8CA89D171CD5" lastPageId="7" lastPageNumber="82" pageId="5" pageNumber="80">
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Carapace weakly granular; anterior margin with a moderately marked median concavity. Carinae inconspicuous; furrows weak. Median ocular tubercle clearly anterior to the centre of carapace; median eyes moderate in size and separated by more than one ocular diameter. Three pairs of lateral eyes of moderate
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<pageBreakToken id="AA4FF4F55A2CEFAB649D963E64598679" pageId="6" pageNumber="81" start="start">size</pageBreakToken>
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. Sternum subpentagonal. Mesosomal tergites weakly granular, with one median carina; lateral carinae totally inconspicuous; VII with five weakly marked carinae. Pectines large, with 18-17 teeth; fulcra absent. Sternites weakly granular to smooth, with small oval to slit-like spiracles. Metasomal segment I to IV rounded with 10-10-8-8 carinae; segment V slender with five carinae; dorsal carinae of segments I-IV with minute spinoid granules; dorsal aspect of segments I to V weakly depressed; setation on all segments strongly marked. Telson with a very long pear-shaped vesicle; weakly granular to smooth; aculeus extremely long and moderately curved; setation strongly marked. Cheliceral dentition only partially visible; fixed and movable fingers with one basal tooth observable; distal teeth moderately long (Vachon 1963). Pedipalp femur pentacarinate; patella with 6-7 carinae; internal face of femur and patella with spinoid granules. Chela with weakly marked carinae; all faces not granular, smooth. Fixed and movable fingers each with one series of small rounded granules and no conspicuous spinoid accessory granules; extremity of fingers with stronger spinoid granules; setation of pedipalps weakly marked. Trichobothriotaxy recalling type A (
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<bibRefCitation id="C573B8239F6852761D8BE384D4AB53FA" author="Vachon, M" journalOrPublisher="Paris" pageId="11" pageNumber="86" pagination="857 - 958" title="Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions." volumeTitle="Bulletin du Museum national d'Histoire naturelle" year="1974">Vachon 1974</bibRefCitation>
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) of extant buthids: at least 1-3 internal, five dorsal, and two external trichobothria on the femur; dorsal trichobothria disposed in beta (β) configuration (
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<bibRefCitation id="7CDDA0D7EC3AAD5FA7C0C950C4786D53" author="Vachon, M" journalOrPublisher="Comptes Rendus des Seances de l'Academie des Sciences, Paris" pageId="11" pageNumber="86" pagination="1597 - 1599" title="Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des Scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon." volume="281" year="1975">Vachon 1975</bibRefCitation>
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); one internal, five dorsal, and no ventral trichobothria on patella; 6-7 external trichobothria can be suggested on patella by the presence of fine setae; some are displaced on the
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<pageBreakToken id="50AC40E9D5E27D4D4C58E69DBA96CC09" pageId="7" pageNumber="82" start="start">ventral</pageBreakToken>
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aspect 5-6 dorso-external and two ventral on chelal hand; six on fixed finger. Tibial spurs present on legs III and IV, moderately marked.
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</paragraph>
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<paragraph id="B329274B82221F2D8834E723736D7299" pageId="7" pageNumber="82">
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Morphometric values (in mm) of male juvenile holotype of
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<taxonomicName id="8B4764957FEE6EEE50AC4F1834694C50" genus="Betaburmesebuthus" lsidName="Betaburmesebuthus bellus" pageId="7" pageNumber="82" rank="species" species="bellus">Betaburmesebuthus bellus</taxonomicName>
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sp. n.
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</paragraph>
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<paragraph id="81A6B3D797483FD9ADE63A37A3BB9F33" pageId="7" pageNumber="82">Total length 11.80 (including telson). Carapace: length 1.40, anterior width 0.87, posterior width 1.40. Mesosoma length 3.60. Metasomal segments. I: length 0.74, width 0.60; II: length 0.84, width 0.60; III: length 0.94, depth 0.57; IV: length 1.14, depth 0.50; V: length 1.67, depth 0.47. Telson length 1.47. Vesicle: depth 0.37. Pedipalp: femur length 1.14, width 0.34; patella length 1.24, width 0.47; chela length 2.24, width 0.34; movable finger length 1.57.</paragraph>
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<caption id="7F0C8E53828739558AB097977F9BBF1B" pageId="7" pageNumber="82">
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<paragraph id="1CD135DF7D7E418202291756A050EF8C" pageId="7" pageNumber="82">
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Figures 1-4.
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<taxonomicName id="28C8F1E1D0CDFCE160A3A151028ABD11" genus="Betaburmesebuthus" lsidName="Betaburmesebuthus bellus" pageId="7" pageNumber="82" rank="species" species="bellus">Betaburmesebuthus bellus</taxonomicName>
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sp. n. Male holotype. 1-2 Habitus, dorsal and ventral aspects 3 Ventral aspect in detail, showing sternum, genital operculum and pectines 4 Tergites VI-VII, metasomal segments and telson, dorso-lateral aspect.
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</paragraph>
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</caption>
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<caption id="09AEF4F00043F31C0859E32621F6634D" pageId="7" pageNumber="82">
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<paragraph id="C906440524C32D5624775836E2A4726E" pageId="7" pageNumber="82">
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Figures 5-7.
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<taxonomicName id="6D1B4F2799B4914C0155CF12CEE4C23C" genus="Betaburmesebuthus" lsidName="Betaburmesebuthus bellus" pageId="7" pageNumber="82" rank="species" species="bellus">Betaburmesebuthus bellus</taxonomicName>
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sp. n. Male holotype. 5 Chelicera, carapace and tergites I-III, dorsal aspect 6 Sternites V-VII showing carinae and spiracles and metasomal segments I-V and telson, ventro-lateral aspect 7 Ventral aspect, showing Coxapophysis, sternum, genital operculum, pectines and sternites with spiracles. Scale bars: 0.5 mm.
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</paragraph>
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</caption>
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<caption id="2CCAC2238416E5A28EE27648EA267EF1" pageId="7" pageNumber="82">
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<paragraph id="149E1ECF7E380B0043384926E7ED52E0" pageId="7" pageNumber="82">
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Figures 8-9.
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<taxonomicName id="6C7A091A975E3DD3967A7A8F5A6694A8" genus="Betaburmesebuthus" lsidName="Betaburmesebuthus bellus" pageId="7" pageNumber="82" rank="species" species="bellus">Betaburmesebuthus bellus</taxonomicName>
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sp. n. Male holotype. Righ pedipalp, dorsal and ventral aspects, showing trichobothrial pattern. Dorsal aspect is partially covered by an inclusion. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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<caption id="855F9CF594EA8237EE04D889050BA2E5" pageId="7" pageNumber="82">
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<paragraph id="B394AC148F597FECEE98F53D2DE9A6C4" pageId="7" pageNumber="82">Figure 10. World map showing the sites where scorpions included in amber have been found.</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |