146 lines
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146 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.468.8506" ID-GBIF-Dataset="df3f2d10-3b5e-480f-936c-368e01ffd7b1" ID-PMC="PMC4296520" ID-Pensoft-Pub="1313-2970-468-1" ID-PubMed="25610341" ID-ZBK="AE5AB793FDC74DCD8A47AE96A141E2AD" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-468-1" ModsDocOrigin="ZooKeys 468" ModsDocTitle="Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1451244902156" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo, Panjaitan, Rawati, Menufandu, Herlina & Balke, Michael" docDate="2014" docId="24BD0B47690840719470103A68D9FB70" docLanguage="en" docName="ZooKeys 468: 1-83" docOrigin="ZooKeys 468" docSource="http://dx.doi.org/10.3897/zookeys.468.8506" docTitle="Exocelina pseudoedeltraudae Shaverdo & Balke, sp. n." docType="treatment" docUuid="C7536736-ABCE-42EC-89D0-678762B728AE" docUuidSource="ZooBank" docVersion="4" lastPageNumber="23" masterDocId="FFE9FFA5117AFFD17452FFB5FF94FFE3" masterDocTitle="Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="83" masterPageNumber="1" pageNumber="21" updateTime="1668159774671" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sagata, Katayo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panjaitan, Rawati</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Menufandu, Herlina</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>468</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.468.8506</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.468.8506</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-468-1</mods:identifier>
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<mods:identifier type="ZBK">AE5AB793FDC74DCD8A47AE96A141E2AD</mods:identifier>
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<mods:identifier type="ZooBank">AE5AB793FDC74DCD8A47AE96A141E2AD</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152057388" LSID="urn:lsid:zoobank.org:act:C7536736-ABCE-42EC-89D0-678762B728AE" httpUri="http://treatment.plazi.org/id/24BD0B47690840719470103A68D9FB70" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
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<subSubSection pageId="20" pageNumber="21" type="multiple">
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<paragraph pageId="20" pageNumber="21">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="21" type="nomenclature">
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<paragraph pageId="20" pageNumber="21">
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15.
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<taxonomicName LSID="http://zoobank.org/C7536736-ABCE-42EC-89D0-678762B728AE" authority="Shaverdo & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina pseudoedeltraudae" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pseudoedeltraudae">Exocelina pseudoedeltraudae Shaverdo & Balke</taxonomicName>
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<taxonomicNameLabel pageId="20" pageNumber="21">sp. n.</taxonomicNameLabel>
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Figs 9, 38
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<bibRefCitation author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="45" pageNumber="46" pagination="1 - 76" title="Introduction of the Exocelinaekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="10.3897/zookeys.250.3715" volume="250" year="2012">
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; 4
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<normalizedToken originalValue="A–F">A-F</normalizedToken>
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, 30 in Shaverdo et al. (2012)
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</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="21" type="reference_group">
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<paragraph pageId="20" pageNumber="21">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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undescribed sp. MB1288:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="Nature Communications" pageId="45" pageNumber="46" pagination="1 - 10" title="The towering orogeny of New Guinea as a trigger for arthropod megadiversity." url="10.1038/ncomms5001" volume="1" year="2014">Toussaint et al. 2014</bibRefCitation>
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: Supplementary figs 1-4, Tab. 2.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="21" type="type locality">
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<paragraph pageId="20" pageNumber="21">Type locality.</paragraph>
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<paragraph pageId="20" pageNumber="21">
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Papua New Guinea: Hela Province, Koroba,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-5.6975">05°41.85'S</geoCoordinate>
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;
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="142.7306">142°43.836'E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="type material">
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<paragraph pageId="20" pageNumber="21">Type material.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<pageBreakToken pageId="21" pageNumber="22" start="start">Holotype</pageBreakToken>
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: male "Papua New Guinea: Southern Highlands, Koroba, 1600m, 15.v.1994, 05.41.854S 142.43.836E, Balke (PNG 66)", "Paratypus
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina edeltraudae" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="edeltraudae">Exocelina edeltraudae</taxonomicName>
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sp. n. des. H.Shaverdo, L.Hendrich & M.Balke, 2012" (ZSM). Paratypes: 5 males, 1 female with the same labels as the holotype (NHMW, ZSM). 3 males "PAPUA N.G.: 6.-9.5.1998 Southern Highl. Prov. Tari-Koroba, Hedemari [Hedamali] 1700-1900 m, leg. Riedel", "Paratypus
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina edeltraudae" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="edeltraudae">Exocelina edeltraudae</taxonomicName>
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sp. n. des. H.Shaverdo, L.Hendrich & M.Balke, 2012" (NHMW). 1 male, 3 females "Papua New Guinea: Southern Highlands, Tari Komo road, 10km N Hides Gas, 1700m, 13.v.1994, Balke (PNG 61)", "Paratypus
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina edeltraudae" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="edeltraudae">Exocelina edeltraudae</taxonomicName>
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sp. n. des. H.Shaverdo, L.Hendrich & M.Balke, 2012", the male additionally with a green label "DNA M.Balke 1288" (ZSM). 5 males, 8 females "Papua New Guinea: Southern Highlands, Tari to Koroba, 1600m, 15.v.1994, 05.46.500S 142.50.000E, Balke (PNG 65)", "Paratypus
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina edeltraudae" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="edeltraudae">Exocelina edeltraudae</taxonomicName>
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sp. n. des. H.Shaverdo, L.Hendrich & M.Balke, 2012" (NARI, NHMW, ZSM). 1 female "Papua New Guinea: Southern Highlands, Tari to Koroba, 1600m, 15.v.1994, 05.46.500S 142.50.000E, Balke (PNG 65)" (ZSM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="diagnosis">
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<paragraph pageId="21" pageNumber="22">Diagnosis.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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Beetle medium-sized, piceous, submatt; pronotum with distinct lateral bead; male ventrite 6 slightly to distinctly concave apically; male antennomeres 3-5 distinctly enlarged, almost equal in size and shape, antennomeres 6-8 enlarged; male protarsomere 4 with large, slender, evidently curved anterolateral hook-like seta; median lobe with very strong submedian constriction and proximal part very broad in ventral view, apex of median lobe pointed and strongly curved downwards in lateral; paramere with distinct notch on dorsal side and subdistal part elongate, with numerous, dense, long, thin setae. The species is very similar to
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina edeltraudae" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="edeltraudae">Exocelina edeltraudae</taxonomicName>
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Shaverdo, Hendrich & Balke, 2012, from which differs with less shinier dorsal surface, due to stronger punctation and microreticulation, with larger and more rounded male antennomeres 3-5 (for male antennomeres 3 and 4, ratio width/length:> 1.0) and apex of median lobe narrower in lateral view.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
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<paragraph pageId="21" pageNumber="22">Description.</paragraph>
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<paragraph pageId="21" pageNumber="22">Size and shape: Beetle medium-sized (TL-H 3.45-4.0 mm, TL 3.85-4.45 mm, MW 1.8-2.2 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Dorsally piceous, with dark brown anterior margin of head and narrowly pronotal sides; head appendages and legs reddish to reddish brown, legs distally darker (Fig. 38). Teneral specimens paler.</paragraph>
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<paragraph pageId="21" pageNumber="22">Surface sculpture: Head with dense, coarse punctation (spaces between punctures 1-3 times size of punctures), especially on vertex. Pronotum with punctation finer, sparser, and more evenly distributed than on head. Elytra with punctation finer, sparser than on pronotum. Pronotum and elytra with strongly impressed microreticulation, dorsal surface submatt. Head with microreticulation stronger. Metaventrite and metacoxa distinctly microreticulate, metacoxal figs with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine sparse punctation, coarser and denser on two last abdominal ventrites.</paragraph>
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<paragraph pageId="21" pageNumber="22">Structures: Pronotum with distinct lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, rounded and smooth anteriorly, with small anterolateral extensions. Blade of prosternal process lanceolate, relatively narrow, convex, with distinct lateral bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal ventrite 6 broadly rounded, or slightly truncate, or concave apically.</paragraph>
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<paragraph pageId="22" pageNumber="23">
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<pageBreakToken pageId="22" pageNumber="23" start="start">Male</pageBreakToken>
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: Antennomeres 3-5 distinctly enlarged, almost equal in size, antennomeres 6-8 enlarged (Fig. 9A), antennomeres 3-7 rugose ventrally. Protarsomere 4 with large, slender, evidently curved anterolateral hook. Protarsomere 5 ventrally with anterior row of 15 and posterior row of 5 short setae (Fig. 9B). Abdominal ventrite 6 with 8-13 lateral striae on each side, slightly to distinctly concave apically (Fig. 9C). Median lobe with very strong submedian constriction and proximal part very broad in ventral view, apex of median lobe pointed and strongly curved downwards in lateral view (Fig. 9D, E). Paramere with distinct notch on dorsal side and subdistal part elongate, with numerous, dense, long, thin setae (Fig. 9F).
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</paragraph>
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<paragraph pageId="22" pageNumber="23">Holotype: TL-H 4.0 mm, TL 4.45 mm, MW 2.2 mm.</paragraph>
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<paragraph pageId="22" pageNumber="23">Female: Antennae simple, abdominal ventrite 6 broadly rounded or slightly truncate apically, without striae.</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="distribution">
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<paragraph pageId="22" pageNumber="23">Distribution.</paragraph>
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<paragraph pageId="22" pageNumber="23">Papua New Guinea. The species is known only from Hela Province (Fig. 53).</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="etymology">
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<paragraph pageId="22" pageNumber="23">Etymology.</paragraph>
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<paragraph pageId="22" pageNumber="23">
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In an earlier work (
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<bibRefCitation author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="45" pageNumber="46" pagination="1 - 76" title="Introduction of the Exocelinaekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al. 2012</bibRefCitation>
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), this species was mistaken for
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina edeltraudae" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="edeltraudae">Exocelina edeltraudae</taxonomicName>
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. The name is a noun in the nominative singular standing in apposition.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |