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<document ID-DOI="http://dx.doi.org/10.3897/jhr.95.96784" ID-GBIF-Dataset="aa0785c0-f168-4124-a156-b90bf73f5523" ID-Pensoft-Pub="1314-2607-95-59" ID-Pensoft-UUID="93329D9520C759EE8AF1663C83BB9768" ID-ZooBank="DEAA4D16C2484EA8B26C83133240F854" ModsDocID="1314-2607-95-59" checkinTime="1676724327596" checkinUser="pensoft" docAuthor="Belokobylskij, Sergey A., Simutnik, Serguei A., Vasilenko, Dmitry V. &amp; Perkovsky, Evgeny E." docDate="2023" docId="2452DC0955845003853D89F3D43BBF3D" docLanguage="en" docName="JourHymenoptRes 95: 59-72" docOrigin="Journal of Hymenoptera Research 95" docPubDate="2023-02-17" docSource="http://dx.doi.org/10.3897/jhr.95.96784" docTitle="Eocenhecabolus Belokobylskij 2023, gen. nov." docType="treatment" docUuid="3E9686FC-03E0-4910-B954-E7FEBAC094F0" docUuidSource="ZooBank" docVersion="4" id="93329D9520C759EE8AF1663C83BB9768" lastPageNumber="59" masterDocId="93329D9520C759EE8AF1663C83BB9768" masterDocTitle="First record of the parasitoid subfamily Doryctinae (Hymenoptera, Braconidae) in Rovno amber: description of a new genus and species with stigma-like enlargement on the hind wing of the male" masterLastPageNumber="72" masterPageNumber="59" pageNumber="59" updateTime="1732566880503" updateUser="ExternalLinkService">
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<mods:titleInfo id="39FBA9C7679842602F527CE7219230D1">
<mods:title id="8A8FE4E14E83940D7DE83BA0958FA2C1">First record of the parasitoid subfamily Doryctinae (Hymenoptera, Braconidae) in Rovno amber: description of a new genus and species with stigma-like enlargement on the hind wing of the male</mods:title>
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<mods:namePart id="08EEB323DF0F577240CD31FB348CC667">Belokobylskij, Sergey A.</mods:namePart>
<mods:nameIdentifier id="CD3AFCFAF1E7412A3CA00B00BB3FD183" type="ORCID">https://orcid.org/0000-0002-3646-3459</mods:nameIdentifier>
<mods:affiliation id="A87ACDC0CCA707CB8D071EA32B4AEE60">Zoological Institute of the Russian Academy of Sciences, St Petersburg 199034, Russia</mods:affiliation>
<mods:nameIdentifier id="55E483CEF5409B4E89A0496C6EA0ADC8" type="email">doryctes@gmail.com</mods:nameIdentifier>
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<mods:namePart id="39F948AA2B6E510F80F642954A6FCA8C">Simutnik, Serguei A.</mods:namePart>
<mods:nameIdentifier id="4402E2AEFF4F5A0509ADEAD3D9722915" type="ORCID">https://orcid.org/0000-0002-2538-6216</mods:nameIdentifier>
<mods:affiliation id="675F58C454CFB3CE86C24FED5D6D6BF1">I. I. Schmalhausen Institute of Zoology (SIZK), National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, Kiev 01030, Ukraine</mods:affiliation>
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<mods:name id="30D088D2930C0385BCCF819D67E7A0E6" type="personal">
<mods:role id="ACAD0EB1E52365B349EBE3B23B407C8B">
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<mods:namePart id="6DCC0569F10F06EF77D252E7DCD9AFEB">Vasilenko, Dmitry V.</mods:namePart>
<mods:affiliation id="68F88EA36F1ADAE7A1673394D8171FF4">A. A. Borissiak Paleontological Institute of the Russian Academy of Sciences, Profsoyuznaya Str. 123, Moscow 117647, Russia &amp; Cherepovets State University, Cherepovets 162600, Russia</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="28B4F3F8877BA4A7AE947B6E1868F285">Perkovsky, Evgeny E.</mods:namePart>
<mods:nameIdentifier id="EBADCB808926181A933FF948C10BDA28" type="ORCID">https://orcid.org/0000-0002-7959-4379</mods:nameIdentifier>
<mods:affiliation id="7084C7CABE101E966D400BCB03E69E57">I. I. Schmalhausen Institute of Zoology (SIZK), National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, Kiev 01030, Ukraine</mods:affiliation>
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<mods:title id="DBB756EFEB4944E1D262626F7FC6FE1B">Journal of Hymenoptera Research</mods:title>
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<mods:part id="7549A83D087ADF8FD02CCC29C0DA8664">
<mods:date id="46329C25ACC63D705842EF10C62C3AC1">2023</mods:date>
<mods:detail id="4A6F0507ED1E60FB19E19631FC324171" type="pubDate">
<mods:number id="5DA89EB5B9CF918C08CE2890CA3CDDA7">2023-02-17</mods:number>
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<mods:detail id="3F3BAE3B6C9AC60FB9D63E66F83D9DDB" type="volume">
<mods:number id="84A9C8EB6E8D87AE2C8322865A19AFF7">95</mods:number>
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<mods:start id="856BB8CDC87EA30100903AC1B21F69C0">59</mods:start>
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<mods:classification id="3CA12839F8D2D960B6FC80D2C8FA7E28">journal article</mods:classification>
<mods:identifier id="A5E9BFCA78109DD1FCBF993A11DE2FB3" type="DOI">http://dx.doi.org/10.3897/jhr.95.96784</mods:identifier>
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<treatment id="2452DC0955845003853D89F3D43BBF3D" ID-GBIF-Taxon="209298084" LSID="urn:lsid:zoobank.org:act:3E9686FC-03E0-4910-B954-E7FEBAC094F0" httpUri="http://treatment.plazi.org/id/2452DC0955845003853D89F3D43BBF3D" lastPageNumber="59" pageId="0" pageNumber="59" scope_class="Insecta" scope_family="Braconidae" scope_order="Hymenoptera" scope_subFamily="Doryctinae">
<subSubSection id="920F18F50EE2C2B9A81E2B49A834F590" pageId="0" pageNumber="59" type="nomenclature">
<paragraph id="70D1C68316BB14409DE08A08498FB4AA" pageId="0" pageNumber="59">
Genus
<taxonomicName id="2466EFB9E6AE2982707E9D8F85AFB657" LSID="https://zoobank.org/3E9686FC-03E0-4910-B954-E7FEBAC094F0" authority="Belokobylskij" authorityName="Belokobylskij" authorityYear="2023" class="Insecta" family="Braconidae" genus="Eocenhecabolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eocenhecabolus" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus" status="gen. nov.">Eocenhecabolus Belokobylskij</taxonomicName>
<taxonomicNameLabel id="DB470A64F3FF7AD695AA1B2AED0104A5" pageId="0" pageNumber="59">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C04BC262D90939672978FFE80C850A14" pageId="0" pageNumber="59" type="type species">
<paragraph id="28C5294F88711F2B560E3FD396C2AF45" pageId="0" pageNumber="59">Type species.</paragraph>
<paragraph id="F3DC11D99DCFF04559D6F11133EFCA74" pageId="0" pageNumber="59">
<taxonomicName id="EEC0C779E6BF31DB175DE2A218F39A30" authorityName="Belokobylskij" authorityYear="2023" class="Insecta" family="Braconidae" genus="Eocenhecabolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eocenhecabolus kotenkoi" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="kotenkoi">
<emphasis id="19C18758E34E70A81DBDAAA1801AC31B" italics="true" pageId="0" pageNumber="59">Eocenhecabolus kotenkoi</emphasis>
</taxonomicName>
Belokobylskij, gen. et sp. nov., by present designation and monotypy.
</paragraph>
</subSubSection>
<subSubSection id="9F9A235985EEC895B9092B7A43AC902E" pageId="0" pageNumber="59" type="etymology">
<paragraph id="2867D3571A47C3659613D0B9CDC2862C" pageId="0" pageNumber="59">Etymology.</paragraph>
<paragraph id="E8D981237D77E0A7EDEA218EEFC97D11" pageId="0" pageNumber="59">
Named after
<normalizedToken id="06EEBA808182B34775145FC3FC0128DA" originalValue="“Eocene”">&quot;Eocene&quot;</normalizedToken>
from the geological epoch dated to the Rovno amber and the generic name of its extant type genus
<taxonomicName id="7DF9C4B2C60D28AF665A9DED83059663" class="Insecta" family="Braconidae" genus="Hecabolus" kingdom="Animalia" lsidName="Hecabolus" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="A6F534077130B410A84A036851E7049F" italics="true" pageId="0" pageNumber="59">Hecabolus</emphasis>
</taxonomicName>
of the tribe
<taxonomicName id="B4012B85CEB02D733C4AA924D3801899" lsidName="" pageId="0" pageNumber="59" rank="tribe" tribe="Hecabolini">Hecabolini</taxonomicName>
from subfamily
<taxonomicName id="2D915234D80D12BB0BBBA0AC1FD75943" lsidName="" pageId="0" pageNumber="59" rank="subFamily" subFamily="Doryctinae">Doryctinae</taxonomicName>
. Gender: masculine.
</paragraph>
</subSubSection>
<subSubSection id="56B6FA7F04816B5485FCE460FCD7A31C" pageId="0" pageNumber="59" type="description">
<paragraph id="4D47A54554F424A2989B88EA2A52DD9D" pageId="0" pageNumber="59">Description.</paragraph>
<paragraph id="BD55B463A8CB55B496C5133A5B267D01" pageId="0" pageNumber="59">
<emphasis id="3A769431E9015D453095EA782B8D6ACD" bold="true" italics="true" pageId="0" pageNumber="59">Head</emphasis>
(Fig.
<figureCitation id="BD3D1D77D94965A8A8D1D8CE43377E85" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A habitus, right dorso-lateral view B habitus, left ventro-lateral view C body, lateral view D body, ventro-lateral view E head and antenna, fronto-lateral view F head, dorsal view G propodeum and metasoma, dorsal view H head and antenna, lateral view." figureDoi="10.3897/jhr.95.96784.figure1" httpUri="https://binary.pensoft.net/fig/809232" pageId="0" pageNumber="59">1E, F, H</figureCitation>
) not depressed, weakly transverse. Ocelli medium-sized, weakly convex, arranged in triangle with base 1.3 times its sides. Frons almost not convex, without lateral protuberances. Eyes large, oval, glabrous. Face distinctly convex. Malar suture present, but weak. Clypeus relatively high, with distinct lower visor. Clypeal suture fine laterally, absent on wide distance dorsally. Anterior tentorial pits small. Occipital carina present and distinct at least laterally and dorsally. Mandibles robust. Maxillary palpus medial length.
<emphasis id="E63557AD38EC684B520349365B300717" bold="true" italics="true" pageId="0" pageNumber="59">Antenna</emphasis>
(Fig.
<figureCitation id="5D2838238E5ADE8C432F7648CFC21CD9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A habitus, right dorso-lateral view B habitus, left ventro-lateral view C body, lateral view D body, ventro-lateral view E head and antenna, fronto-lateral view F head, dorsal view G propodeum and metasoma, dorsal view H head and antenna, lateral view." figureDoi="10.3897/jhr.95.96784.figure1" httpUri="https://binary.pensoft.net/fig/809232" pageId="0" pageNumber="59">1C, E, H</figureCitation>
) mostly missing, only four segments present. Scape short and wide, approximately as long as maximum width. Pedicel relatively short and thick, about as long as scape. First flagellomere long, subcylindrical, weakly curved and without any modifications.
<emphasis id="1278386CF82C2050903DAA49933B5D6E" bold="true" italics="true" pageId="0" pageNumber="59">Mesosoma</emphasis>
(Fig.
<figureCitation id="64FD2FC71009CF57E19A2560E2903073" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A habitus, right dorso-lateral view B habitus, left ventro-lateral view C body, lateral view D body, ventro-lateral view E head and antenna, fronto-lateral view F head, dorsal view G propodeum and metasoma, dorsal view H head and antenna, lateral view." figureDoi="10.3897/jhr.95.96784.figure1" httpUri="https://binary.pensoft.net/fig/809232" pageId="0" pageNumber="59">1C, G</figureCitation>
) not depressed. Pronotum convex in posterior half, with distinct short longitudinal lateral carinae. Sides of pronotum mainly smooth with short rugae on oblique furrow. Mesoscutum distinctly (but not highly) roundly convex above pronotum, densely and rather distinctly granulate-punctate. Notauli present, deep and complete, reaching prescutellar furrow. Scutellum convex. Prepectal carina present, distinct. Mesopleuron mainly smooth. Precoxal sulcus present, but short (not more than half of mesopleuron length below), rather deep, almost straight, finely crenulate. Metascutum without dorsal tooth (lateral view). Propodeum evenly curved in lateral view, with areas delineated but distinct carinae, with wide, sub-round and smooth basolateral areas, with narrow and long areola, distinctly separated petiolate area and relatively short basomedial carina; without lateral tubercles; propodeal spiracle subcircular.
<emphasis id="F5A13CE95F1D20D870292872AC11378A" bold="true" italics="true" pageId="0" pageNumber="59">Wings</emphasis>
(Figs
<figureCitation id="F2513AF89B809BBDC6FFBD8F0AFF1D7C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A habitus, right dorso-lateral view B habitus, left ventro-lateral view C body, lateral view D body, ventro-lateral view E head and antenna, fronto-lateral view F head, dorsal view G propodeum and metasoma, dorsal view H head and antenna, lateral view." figureDoi="10.3897/jhr.95.96784.figure1" httpUri="https://binary.pensoft.net/fig/809232" pageId="0" pageNumber="59">1A, B</figureCitation>
,
<figureCitation id="8F2ABD3A8EBEF8074E8B8E0F18455237" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A wings B metasoma and hind leg, lateral view." figureDoi="10.3897/jhr.95.96784.figure2" httpUri="https://binary.pensoft.net/fig/809233" pageId="0" pageNumber="59">2A</figureCitation>
). Fore wing relatively wide, evenly faintly infuscate; pterostigma rather long and wide. Radial (marginal) cell not shortened, closed distally, wide, about 3.5 times longer than its maximum width. Metacarp (1-R1) 1.2 times longer than pterostigma. Radial vein (r) arising weakly before middle of pterostigma. First medial abscissa (1-SR+M) present and weakly sinuate. Both radiomedial veins (2-SR and r-m) present. Second radiomedial (submarginal) cell relatively long, pentagonal. Discoidal (first discal) cell petiolate anteriorly; petiole (1-SR) short. Recurrent vein (m-cu) distinctly postfurcal, weakly convergent posteriorly with basal vein (1-M). First mediocubital vein (M+CU1) well sclerotised and straight. Nervulus (cu-a) distinctly postfurcal. Brachial (first subdiscal) cell open posteriorly; brachial vein (CU1b) absent. Transverse anal veins (2A and a) absent. Hind wing. Second abscissa of costal vein (1-SC+R) with elementary elliptic stigma-like enlargement. Radial vein (SR) unsclerotised and transparent. Nervellus (cu-a) present. Submedial (subbasal) cell large. First abscissa of mediocubital vein (M+CU) more than twice longer than second abscissa (1-M).
<emphasis id="2A8FBADD1884E72C2A180FDAD34C1D69" bold="true" italics="true" pageId="0" pageNumber="59">Legs</emphasis>
(Figs
<figureCitation id="D0E6891BFDB6DD22CC9EA3B08C9A7EE2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A habitus, right dorso-lateral view B habitus, left ventro-lateral view C body, lateral view D body, ventro-lateral view E head and antenna, fronto-lateral view F head, dorsal view G propodeum and metasoma, dorsal view H head and antenna, lateral view." figureDoi="10.3897/jhr.95.96784.figure1" httpUri="https://binary.pensoft.net/fig/809232" pageId="0" pageNumber="59">1C, D</figureCitation>
,
<figureCitation id="C8BC178B75BC9136A7FE009C1F87CA1C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A wings B metasoma and hind leg, lateral view." figureDoi="10.3897/jhr.95.96784.figure2" httpUri="https://binary.pensoft.net/fig/809233" pageId="0" pageNumber="59">2B</figureCitation>
) rather robust and short. Fore tibia with distinct spines arranged almost in single line. Hind coxa elongate, without ventro-basal tubercle and corner, weakly shorter than propodeum. Hind femur short and wide, 0.7 times as long as hind tibia. Hind tibia weakly thickened distally, with at least two distinctly visible spines on its dorsal margin in distal quarter. Hind tibial spur glabrous, relatively short, about 0.3 times as long as hind basitarsus. Hind basitarsus short, about half as long as second to fifth segments combined. Tarsal claw medium size, simple and evenly curved.
<emphasis id="3C6026D464004AEC3D1D6B289C3CB35E" bold="true" italics="true" pageId="0" pageNumber="59">Metasoma</emphasis>
(Figs
<figureCitation id="A9B9C0047067433EB7E5FA6EAD6AC119" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A habitus, right dorso-lateral view B habitus, left ventro-lateral view C body, lateral view D body, ventro-lateral view E head and antenna, fronto-lateral view F head, dorsal view G propodeum and metasoma, dorsal view H head and antenna, lateral view." figureDoi="10.3897/jhr.95.96784.figure1" httpUri="https://binary.pensoft.net/fig/809232" pageId="0" pageNumber="59">1C, D, G</figureCitation>
,
<figureCitation id="540BC829207B88BA96E8738AEE7994A4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A wings B metasoma and hind leg, lateral view." figureDoi="10.3897/jhr.95.96784.figure2" httpUri="https://binary.pensoft.net/fig/809233" pageId="0" pageNumber="59">2B</figureCitation>
) elongate, oval in dissection, not pressed, segments behind third one distinctly exposed posteriorly. First metasomal tergite not wide, weakly widened distally, with deep dorsope, with distinct dorsomedial carinae situated closed to each other, with distinct lateral carinae, striate medially and smooth laterally, with spiracles situated on basal third of tergite, spiracular tubercles small, weakly shorter that second and third tergites combined. Suture between second and third tergites absent. Second tergite mainly smooth, with shallow and short sublateral depression. Laterotergites (epipleura) of segments behind first one perhaps not separated; spiracles placed on the lateral part of tergites. Genitalia distinctly visible from below.
</paragraph>
<caption id="544C86CBD055EAE440585B63F352E341" doi="10.3897/jhr.95.96784.figure1" httpUri="https://binary.pensoft.net/fig/809232" pageId="0" pageNumber="59" start="Figure 1" startId="F1">
<paragraph id="E81F67C6D6774FDC0A36DE74C86263D6" pageId="0" pageNumber="59">
<emphasis id="FE8E2393176DB3F6E3472ED8BD832489" bold="true" pageId="0" pageNumber="59">Figure 1.</emphasis>
<taxonomicName id="16B78E8A130A3021B020B419C95EE8E0" authorityName="Belokobylskij" authorityYear="2023" class="Insecta" family="Braconidae" genus="Eocenhecabolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eocenhecabolus kotenkoi" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="kotenkoi">
<emphasis id="D905BF5EE4C3D8F12DCCE4D1D34ADF15" italics="true" pageId="0" pageNumber="59">Eocenhecabolus kotenkoi</emphasis>
</taxonomicName>
gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT-607)
<emphasis id="142970BA9B3DA7884D568B72BB65105A" bold="true" pageId="0" pageNumber="59">A</emphasis>
habitus, right dorso-lateral view
<emphasis id="B1244E831AE4D146A92FEE88035CCF54" bold="true" pageId="0" pageNumber="59">B</emphasis>
habitus, left ventro-lateral view
<emphasis id="3230C95CEE4177BD528D8B3E8AF67FBD" bold="true" pageId="0" pageNumber="59">C</emphasis>
body, lateral view
<emphasis id="08C347C1B2B87921BAB6D07D4D56FACC" bold="true" pageId="0" pageNumber="59">D</emphasis>
body, ventro-lateral view
<emphasis id="8A7B108834F5ED385DF43A0A5794EAE9" bold="true" pageId="0" pageNumber="59">E</emphasis>
head and antenna, fronto-lateral view
<emphasis id="19B4B7514A00E23C5E26AB188C7C32F1" bold="true" pageId="0" pageNumber="59">F</emphasis>
head, dorsal view
<emphasis id="A57838052D1534A9AA54264FB93166C1" bold="true" pageId="0" pageNumber="59">G</emphasis>
propodeum and metasoma, dorsal view
<emphasis id="904AF44F7F66DB350041135A9E6AC067" bold="true" pageId="0" pageNumber="59">H</emphasis>
head and antenna, lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C284723FDDEC2689DE6D72609D30479E" pageId="0" pageNumber="59" type="comparative diagnosis">
<paragraph id="CA3DAF8580F3225D2E1455D9E93E3382" pageId="0" pageNumber="59">Comparative diagnosis.</paragraph>
<paragraph id="6FF9190C801A7C1B060DBBB67E83E880" pageId="0" pageNumber="59">
This new genus belongs to the tribe
<taxonomicName id="149ABAC524CF112F689866CE9A52570F" lsidName="" pageId="0" pageNumber="59" rank="tribe" tribe="Hecabolini">Hecabolini</taxonomicName>
based on the fore wing with a distally open brachial (subdiscal) cell and the hind wing of male with an elementary stigma-like enlargement. The latter character is similar to that found in the extant doryctine genera
<taxonomicName id="574452F187EB2A71125C9EE06BB58790" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="9758B056FBBB516C6B2C2E6645E9A2FD" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
Belokobylskij, 1992,
<taxonomicName id="189E424B55834899CCA09DB7ACF86CA8" authorityName="Wesmael" authorityYear="1838" class="Insecta" family="Braconidae" genus="Dendrosoter" kingdom="Animalia" lsidName="Dendrosoter" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="924A82CBDE7E5CCFC4823FB3E2C0EB59" italics="true" pageId="0" pageNumber="59">Dendrosoter</emphasis>
</taxonomicName>
Wesmael, 1838,
<taxonomicName id="8570946EA1A69AB940E7A241C49B4420" authorityName="Kocak &amp; Kemal" authorityYear="2008" class="Insecta" family="Braconidae" genus="Bracocesa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bracocesa" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="3790945A50F05D458C12481B21F3166E" italics="true" pageId="0" pageNumber="59">Bracocesa</emphasis>
</taxonomicName>
<normalizedToken id="E86989C4981C4EE49436AA3478955B96" originalValue="Koçak">Kocak</normalizedToken>
&amp; Kemal, 2008, and
<taxonomicName id="18632045BB65F55FCA41F438C7F59CDA" authorityName="Enderlein" authorityYear="1912" class="Insecta" family="Braconidae" genus="Doryctophasmus" kingdom="Animalia" lsidName="Doryctophasmus" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="621E2F910336AEE33917F3A6B72DF554" italics="true" pageId="0" pageNumber="59">Doryctophasmus</emphasis>
</taxonomicName>
Enderlein, 1912.
</paragraph>
<paragraph id="D953BB9F40692839B0756D454CA2B3FA" pageId="0" pageNumber="59">
<taxonomicName id="7EC7D51621A25B5D8905FD154E308202" authorityName="Belokobylskij" authorityYear="2023" class="Insecta" family="Braconidae" genus="Eocenhecabolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eocenhecabolus" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="D4DB4A600FE2C92903E521C16E57B6B7" italics="true" pageId="0" pageNumber="59">Eocenhecabolus</emphasis>
</taxonomicName>
gen. nov. is most similar to the Pantropical
<taxonomicName id="46B4D5196ACCAE2BEF4147C7437A0A8B" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="C6EDF0FA7BE5613B00DDE0A6B0EB44E7" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
Belokobylskij from the subtribe
<taxonomicName id="3F2AC4A42BF1D5F6914D5E738822AE7E" class="Insecta" family="Braconidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="subTribe" subTribe="Stenocorsina">Stenocorsina</taxonomicName>
(
<taxonomicName id="0D21D5FF41C658402C846237CBAD1037" lsidName="" pageId="0" pageNumber="59" rank="subFamily" subFamily="Doryctinae">Doryctinae</taxonomicName>
:
<taxonomicName id="BC2CD6C531BBD50AB55D83458DB418D4" lsidName="" pageId="0" pageNumber="59" rank="tribe" tribe="Hecabolini">Hecabolini</taxonomicName>
) by the wing venation and analogous enlargement on the hind wing. However, the new genus differs from
<taxonomicName id="80CDCB5EA453A24BA6A0D147A1820BF1" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="71D11AB686FDEA218E483166B5ADFC07" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
by the very short antennal scape, approximately as long as its maximum width (elongated, about 1.5 times longer than the maximum width of that in
<taxonomicName id="46239F3732045C26997DDD8CD18B1695" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="688A0D1676E51FFC1197928ABF8CDC01" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the enlarged pedicel, about as long as the scape (not enlarged and only about 0.5 times as long as the scape in
<taxonomicName id="AC3125EA6FB42A762956C32BB64D9E4A" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="81FC73F5F25DD368FF742F054D2A52BD" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the mostly smooth temple with additional sparse punctuation (densely granulate-striate in
<taxonomicName id="3B397BC2EFFC28F9D50A8FDA598D9735" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="2271993EDC12CF8D37951B67B895D2C4" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the mostly smooth side of the mesosoma (basically densely granulate in
<taxonomicName id="CC2F0048F7BFC341E285A3D083ADB5CF" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="0170CFD9D403FCB4355CE48DFEA84B15" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the propodeum with areas delineated by distinct carinae (without areas delineated by carinae in
<taxonomicName id="26E24819547A5D650B9289F831963424" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="8E48EAC42BD4772A4EF948B462CE0FF2" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the fore wing not maculate, but only faintly infuscate (distinctly maculate in
<taxonomicName id="305CC1AF5DE9BC99B84E587C69ADF8FB" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="A36351040E04FE0CD42462ACABC47087" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the distinctly postfurcal recurrent vein (m-cu) of the fore wing (usually distinctly antefurcal in
<taxonomicName id="2132BCFBAC0C51003CE1A1F5D9B341AE" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="7E619F80A93E1EA1A44E64A0EF61516F" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the relatively short discoidal (discal) cell of the fore wing (distinctly elongate in
<taxonomicName id="90C051FE8194C847F9484FB4528F59A0" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="E03E4E3A00253F0C5565884666B18642" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the weakly postfurcal nervulus (cu-a) in the fore wing (strongly postfurcal in
<taxonomicName id="AF9C46C15A4A2EE75B6266015A616C87" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="5DF32A8CC55901703D87BA70BEFF1EAC" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the first abscissa of the mediocubital vein (M+CU) of the hind wing distinctly longer than the second abscissa (1-M) (distinctly shorter in
<taxonomicName id="D0AC915A74075C209C95B344A8551B00" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="4760AC3E6A8E66DEA345C058CF8C6DB2" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the smooth and less thick hind femur, 3.0 times longer than its maximum width (densely granulate-reticulate and thicker, 2.5 times longer in
<taxonomicName id="5DA966D588337307A5A5C36771202B25" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="1F7337B640FC9FA30152251387043C30" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the hind tibia with relatively long setae and at least two distinct spines on its dorsal margin (with very short setae and without spines on the dorsal margin in
<taxonomicName id="890DF43A42FBE938CCA820F5F6197920" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="6DCA438F388094BFABBC2608470D59BA" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), the shortened hind tarsus with the segment not narrowed toward its distal margin (elongate and segments distinctly narrowed distally in
<taxonomicName id="7C9FE30654A6C2002221F5B6CC542DDB" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B75B144E722DECB83E131561BF689955" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
), and the smooth metasoma behind the first tergite (the second and part of third tergites heavily sculptured in
<taxonomicName id="692DA6CA40BB8322BBBB7CDFCC3D3C69" class="Insecta" family="Braconidae" genus="Hemidoryctes" kingdom="Animalia" lsidName="Hemidoryctes" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="A8C6550E7A2C9BFC0B0F41CCAD8539C3" italics="true" pageId="0" pageNumber="59">Hemidoryctes</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="F26E405A9424F7844FA062FCB08EC356" pageId="0" pageNumber="59">
Apart from several individual differences, the new genus differs from other three extant genera exhibiting stigma like enlargement on hind wing (
<taxonomicName id="AE242FF5A0249D35A75A0118121E1DFD" authorityName="Wesmael" authorityYear="1838" class="Insecta" family="Braconidae" genus="Dendrosoter" kingdom="Animalia" lsidName="Dendrosoter" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B91E45C64DEAD679DC2471048D8238B3" italics="true" pageId="0" pageNumber="59">Dendrosoter</emphasis>
</taxonomicName>
Wesmael,
<taxonomicName id="7E152CCAEBA07DDDD5C2216C586A84C5" authorityName="Kocak &amp; Kemal" authorityYear="2008" class="Insecta" family="Braconidae" genus="Bracocesa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bracocesa" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="BCEADABC30D9D2240101B065BE6DC486" italics="true" pageId="0" pageNumber="59">Bracocesa</emphasis>
</taxonomicName>
<normalizedToken id="080BC1D6D6F712FBDB73CB70789ECFE1" originalValue="Koçak">Kocak</normalizedToken>
&amp; Kemal and
<taxonomicName id="A259A5543A1153C3BB1BA1B956ACAFAE" authorityName="Enderlein" authorityYear="1912" class="Insecta" family="Braconidae" genus="Doryctophasmus" kingdom="Animalia" lsidName="Doryctophasmus" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="75AE72493B507EFEC100BA5788DA1215" italics="true" pageId="0" pageNumber="59">Doryctophasmus</emphasis>
</taxonomicName>
Enderlein) in having the open distally brachial (first subdiscal) cell and no brachial vein (CU1b) in the fore wing (this cell closed distally and the brachial vein present in all latter genera), and large submedial (subbasal) cell in the hind wing with the first abscissa of the mediocubital vein (M+CU) distinctly longer than the second abscissa (1-M) (this cell small and the first abscissa short in all three latter genera).
</paragraph>
<paragraph id="A75D227B00706B81FD8368CB927A5587" pageId="0" pageNumber="59">
Among known fossil
<taxonomicName id="DEA4ABF74A4F98A7D619705FD21F5517" lsidName="" pageId="0" pageNumber="59" rank="subFamily" subFamily="Doryctinae">Doryctinae</taxonomicName>
genera,
<taxonomicName id="E8067EF221866ABECBD999831F438824" authorityName="Belokobylskij" authorityYear="2023" class="Insecta" family="Braconidae" genus="Eocenhecabolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eocenhecabolus" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="19484F887DDEE76B3C616139F9187129" italics="true" pageId="0" pageNumber="59">Eocenhecabolus</emphasis>
</taxonomicName>
gen. nov. is superficially similar to the extinct
<taxonomicName id="712E3A8A19E91315F560D3B581FC24E3" authorityName="Brues" authorityYear="1933" class="Insecta" family="Braconidae" genus="Doryctomorpha" kingdom="Animalia" lsidName="Doryctomorpha tertiaria" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="tertiaria">
<emphasis id="E0FFFCBFF10AEAB807BA287EF6FFB24D" italics="true" pageId="0" pageNumber="59">Doryctomorpha tertiaria</emphasis>
</taxonomicName>
Brues, 1933 described based on a female from Baltic amber (Brues, 1933). However, the assignment of this species to the peculiar endemic New Zealand genus
<taxonomicName id="5DFDC64CF7048728A71333AF715BD951" authorityName="Ashmead" authorityYear="1900" class="Insecta" family="Braconidae" genus="Doryctomorpha" kingdom="Animalia" lsidName="Doryctomorpha" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="D92E507495E96B94E5C8F1134CBF7DDA" italics="true" pageId="0" pageNumber="59">Doryctomorpha</emphasis>
</taxonomicName>
Ashmead, 1900 from the subfamily
<taxonomicName id="FD0112F85D6648736F3C43A4ED06163E" authorityName="van Achterberg" authorityYear="1975" lsidName="" pageId="0" pageNumber="59" rank="subFamily" subFamily="Mesostoinae">Mesostoinae</taxonomicName>
is very doubtful and unsupported by known morphological characters. The female of
<taxonomicName id="B1475E69518BBF37C61243EBEEA20E43" lsidName="D. tertiaria" pageId="0" pageNumber="59" rank="species" species="tertiaria">
<emphasis id="0265C55164E75AC639C4D3A5818E7D99" italics="true" pageId="0" pageNumber="59">D. tertiaria</emphasis>
</taxonomicName>
Brues perhaps may belong to the new genus described here, but absence of important information in this species description (especially regarding wing venation and legs) and uninformative figure together with the loss of the type specimen prevent us to form a reliable opinion about its placement. Anyway,
<taxonomicName id="D89F9C7BAE50C1930EBABBBDE83AED82" authorityName="Belokobylskij" authorityYear="2023" class="Insecta" family="Braconidae" genus="Eocenhecabolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eocenhecabolus kotenkoi" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="kotenkoi">
<emphasis id="A700FFDEC340739332BC611152C0D93F" italics="true" pageId="0" pageNumber="59">Eocenhecabolus kotenkoi</emphasis>
</taxonomicName>
gen. et sp. nov. differs from
<taxonomicName id="B330B083D2957736A59A891150AB840C" lsidName="D. tertiaria" pageId="0" pageNumber="59" rank="species" species="tertiaria">
<emphasis id="FA450D68833C2978B9E0A302BB176A08" italics="true" pageId="0" pageNumber="59">D. tertiaria</emphasis>
</taxonomicName>
Brues by having the head transverse in dorsal view, with a transverse diameter of eye 1.5 times longer than the temple (head subquadrate and with a transverse diameter of eye 2.0 times longer than the temple in
<taxonomicName id="89D2F5B5924EC41059A4FF417AEBA2C2" lsidName="D. tertiaria" pageId="0" pageNumber="59" rank="species" species="tertiaria">
<emphasis id="D58ACF09B60FF3697446E9AB73DCDAD9" italics="true" pageId="0" pageNumber="59">D. tertiaria</emphasis>
</taxonomicName>
), the vertex transversely and sinuately striate (smooth in
<taxonomicName id="0879F75EF8EBD2B322C84227E7513E68" lsidName="D. tertiaria" pageId="0" pageNumber="59" rank="species" species="tertiaria">
<emphasis id="06F60AF3A8839D1FB3D4AEF3A3A5939D" italics="true" pageId="0" pageNumber="59">D. tertiaria</emphasis>
</taxonomicName>
), the propodeum with areas distinctly delineated by carinae (without areolation in
<taxonomicName id="B26C099B3926A9565FCC77F6260012CA" lsidName="D. tertiaria" pageId="0" pageNumber="59" rank="species" species="tertiaria">
<emphasis id="A082A8DF0B7C85E57BD7E80FC582E226" italics="true" pageId="0" pageNumber="59">D. tertiaria</emphasis>
</taxonomicName>
), and the hind coxa suboval and without a prominent lower corner (subtriangular and with a prominent lower corner in
<taxonomicName id="FB84230349C667D709254C03364E7EDC" lsidName="D. tertiaria" pageId="0" pageNumber="59" rank="species" species="tertiaria">
<emphasis id="4BED0F7F2988684EDAA188096D263C0B" italics="true" pageId="0" pageNumber="59">D. tertiaria</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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