332 lines
25 KiB
XML
332 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit & Jochum, Adrienne" docDate="2023" docId="232000B627745E58857D6A60C113C991" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila apiostoma Pall-Gergely & Vermeulen 2023, sp. nov." docType="treatment" docUuid="7369BED7-8447-443D-8FC7-627FEB3B1CBB" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
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<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunyadi, Andras</mods:namePart>
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<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grego, Jozef</mods:namePart>
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<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reischuetz, Alexander</mods:namePart>
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<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
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<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Oerstan, Aydin</mods:namePart>
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<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fekete, Judit</mods:namePart>
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<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jochum, Adrienne</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1147</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>177</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
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<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:7369BED7-8447-443D-8FC7-627FEB3B1CBB" httpUri="http://treatment.plazi.org/id/232000B627745E58857D6A60C113C991" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="https://zoobank.org/7369BED7-8447-443D-8FC7-627FEB3B1CBB" authority="Páll-Gergely & Vermeulen" authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila apiostoma" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="apiostoma" status="sp. nov.">
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Angustopila apiostoma
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Vermeulen
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Angustopila apiostoma Pall-Gergely & Vermeulen, sp. nov. (holotype, RMNH 5006718). Apertural (A), ventral (B), apical (C), and lateral (D) sides of the shell; microstructure on the protoconch (E), continuous edentate aperture (F); spiral microstructure on ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure9" httpUri="https://binary.pensoft.net/fig/808818" pageId="0" pageNumber="1">Figs 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Conchological variability of Angustopila apiostoma Pall-Gergely & Vermeulen, sp. nov. from site WMVT. 0327 (A, C paratypes [RMNH 5006722] B holotype)." figureDoi="10.3897/zookeys.1147.93824.figure10" httpUri="https://binary.pensoft.net/fig/808819" pageId="0" pageNumber="1">, 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: Vietnam • 1 empty shell (H: 0.84 mm, D. 0.71 mm); Quang Ninh Province, Halong Bay area, Thay Cave on Congfo Island, collected inside cave (locality code: WMVT.0327);
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<geoCoordinate degrees="20" direction="north" minutes="52.07" orientation="latitude" precision="9" value="20.867834">20°52.07'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="12.06" orientation="longitude" precision="9" value="107.201">107°12.06'E</geoCoordinate>
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; 6 Sep. 2003; W.J.M. Maassen leg.; RMNH 5006718.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: Vietnam • 2 figured shells; same data as for holotype; RMNH 5006722 • 5 shells; same data as for holotype; HNHM 100441 • 5 shells; same data as for holotype; NMBE 550640 • 28 shells; same data as for holotype; coll. PGB (ex coll. W. Maassen) • 987 shells; same data as for holotype; RMNH 347758 • 10 shells; same data as for holotype; coll. HA • 53 shells; Quang Ninh Province, Halong Bay area, unnamed island in Dau Moi Temple area (locality code: WMVT.0328);
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<geoCoordinate degrees="20" direction="north" minutes="55.69" orientation="latitude" precision="9" value="20.928167">20°55.69'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="09.40" orientation="longitude" precision="9" value="107.15667">107°09.40'E</geoCoordinate>
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; 13 Sep. 2003; W.J.M. Maassen leg.; RMNH 347759 • 41 shells (one of them is figured: Fig.
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<figureCitation captionStart="Figure 99" captionStartId="F99" captionText="Figure 99. Six sympatric Angustopila species collected at Hang Lo Cao Khang Chi ến (Thanh Hoa Province, Vietnam) A synoptic photo of shells of all 6 species B A. fabella Pall-Gergely & Hunyadi, 2015 C A. tetradon Pall-Gergely & Hunyadi, sp. nov. D A. papaver Pall-Gergely & Hunyadi, sp. nov. E A. parallela Pall-Gergely & Hunyadi, sp. nov. F A. szekeresi Pall-Gergely & Hunyadi, 2015 G A. antidomedon Pall-Gergely & Hunyadi, sp. nov." figureDoi="10.3897/zookeys.1147.93824.figure99" httpUri="https://binary.pensoft.net/fig/808908" pageId="0" pageNumber="1">99D</figureCitation>
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); Haiphong Province, Cat Ba Island, Cave Qua Vang, inside cave, large, ecologically intact active cave with speleothems;
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<geoCoordinate degrees="20" direction="north" minutes="48.64" orientation="latitude" precision="9" value="20.810667">20°48.64'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="04.64" orientation="longitude" precision="9" value="107.07733">107°04.64'E</geoCoordinate>
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; 60 m a.s.l.; 6 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16614 • 1 shell; Haiphong Province, Cat Ba Island, Cave Qua Vang, around cave entrance, rocky limestone slope with low, rather mature forest; 6 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16604 • 397 shells; Haiphong Province, Cat Ba Island, large, somewhat ecologically-disturbed, active cave with speleothems;
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<geoCoordinate degrees="20" direction="north" minutes="50.06" orientation="latitude" precision="9" value="20.834333">20°50.06'N</geoCoordinate>
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,
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<geoCoordinate degrees="106" direction="east" minutes="55.91" orientation="longitude" precision="9" value="106.93183">106°55.91'E</geoCoordinate>
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; 0-50 m a.s.l.; 7 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16605 • 343 shells; Haiphong Province, Cat Ba Island, Cave Hoa Cuong, inside cave, polluted cave disturbed by tourism, with concrete paths;
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<geoCoordinate degrees="20" direction="north" minutes="50.41" orientation="latitude" precision="9" value="20.840166">20°50.41'N</geoCoordinate>
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,
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<geoCoordinate degrees="106" direction="east" minutes="59.15" orientation="longitude" precision="9" value="106.98583">106°59.15'E</geoCoordinate>
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; 50 m a.s.l.; 5 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16609 • 13 shells; Haiphong Province, Cat Ba Island, Cave Xa Bac, inside cave;
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<geoCoordinate degrees="20" direction="north" minutes="50.07" orientation="latitude" precision="9" value="20.8345">20°50.07'N</geoCoordinate>
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,
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<geoCoordinate degrees="106" direction="east" minutes="58.61" orientation="longitude" precision="9" value="106.97684">106°58.61'E</geoCoordinate>
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; 30 m a.s.l.; 9 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16615 • 29 shells; Quang Ninh Province, Halong Bay area, Dao Bo Hon, Song Sot Cave, drift material washed over sinkhole in cave;
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<geoCoordinate degrees="20" direction="north" minutes="50.83" orientation="latitude" precision="9" value="20.847166">20°50.83'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="05.67" orientation="longitude" precision="9" value="107.0945">107°05.67'E</geoCoordinate>
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; 2 Oct. 1998; J.J. Vermeulen & A.J. Whitten leg.; JJV 6268 • 248 shells; Quang Ninh Province, Halong Bay area, Dao Bo Hon, Song Sot Cave, guano-enriched sediments in cave;
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<geoCoordinate degrees="20" direction="north" minutes="50.83" orientation="latitude" precision="9" value="20.847166">20°50.83'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="05.67" orientation="longitude" precision="9" value="107.0945">107°05.67'E</geoCoordinate>
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; 2 Oct. 1998; J.J. Vermeulen & A.J. Whitten leg.; JJV 6269 • 4 shells; Haiphong Province, Cat Ba Island, Cave Xa Bac, around cave entrance;
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<geoCoordinate degrees="20" direction="north" minutes="50.07" orientation="latitude" precision="9" value="20.8345">20°50.07'N</geoCoordinate>
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,
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<geoCoordinate degrees="106" direction="east" minutes="58.61" orientation="longitude" precision="9" value="106.97684">106°58.61'E</geoCoordinate>
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; 30 m a.s.l.; 9 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16616 • 6 shells; Haiphong Province, Cat Ba Island, cave Trung Trang, inside cave, cave disturbed by tourism and concrete paths;
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<geoCoordinate degrees="20" direction="north" minutes="47.30" orientation="latitude" precision="9" value="20.788334">20°47.30'N</geoCoordinate>
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,
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<geoCoordinate degrees="106" direction="east" minutes="59.84" orientation="longitude" precision="9" value="106.99733">106°59.84'E</geoCoordinate>
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; 50 m a.s.l.; 5 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16602.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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A small to medium-sized, toothless
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<taxonomicName authorityName="Jochum, Slapnik & Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
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</taxonomicName>
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species with a very narrow umbilicus, conspicuously protruding aperture, and a characteristically shaped aperture (kidney-shaped with narrow sinulus).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell small to medium-sized for the genus, shell height variable, but always higher than wide; pale grey, conical, last whorl widest from standard apertural view; protoconch consists of 1.5 whorls, microstructure finely pitted and granular with a powdery superficial texture, spiral striation not discernible; teleoconch finely ornamented with irregularly spaced radial growth lines crossed by fine rows of equidistantly-spaced microscopic spiral threads (ca. 18-24 on last whorl in apertural view); on frontal and ventral surface of body whorl spiral and radial lines prominent; whorls 4.25-5, shouldered; aperture slightly oblique to shell axis from lateral view; umbilicus deep, very narrow; ventral side characteristic due to comparatively long and straight tuba; aperture kidney-shaped with narrow sinulus; peristome slightly expanded, not reflected; parietal callus slightly protruding, aperture detached from penultimate whorl; aperture toothless or with a vestigial, deeply-situated parietal tooth.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
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<paragraph pageId="0" pageNumber="1">
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H = 0.73-0.93, D = 0.67-0.85, H/D*100 = 101.2-128.2 (
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 14), RUD = 19.2-25.7 (
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 6).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila apiostoma" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="apiostoma">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila apiostoma</emphasis>
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</taxonomicName>
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sp. nov. can be distinguished from
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<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
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<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
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</taxonomicName>
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, by the very narrow umbilicus, the protruding aperture (visible in ventral view) and the kidney-shaped aperture. See also under
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<taxonomicName lsidName="A. fratermajor" pageId="0" pageNumber="1" rank="species" species="fratermajor">
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<emphasis italics="true" pageId="0" pageNumber="1">A. fratermajor</emphasis>
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</taxonomicName>
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sp. nov. and remarks.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The name
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<taxonomicName authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila apiostoma" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="apiostoma">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila apiostoma</emphasis>
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</taxonomicName>
|
|
derives from the Greek
|
|
<emphasis italics="true" pageId="0" pageNumber="1">
|
|
<normalizedToken originalValue="ἄπιον">ἄπιον</normalizedToken>
|
|
</emphasis>
|
|
(= a pear) and
|
|
<emphasis italics="true" pageId="0" pageNumber="1">
|
|
<normalizedToken originalValue="στόμα">στόμα</normalizedToken>
|
|
</emphasis>
|
|
(= mouth) referring to the pear-shaped aperture (to be used as a noun in apposition).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
This species is known from two nearby localities in the Halong Bay Area, Vietnam (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of Angustopila apiostoma Pall-Gergely & Vermeulen, sp. nov. and Angustopila babel Pall-Gergely & Vermeulen, sp. nov." figureDoi="10.3897/zookeys.1147.93824.figure11" httpUri="https://binary.pensoft.net/fig/808820" pageId="0" pageNumber="1">11</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1147.93824.figure9" httpUri="https://binary.pensoft.net/fig/808818" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
|
|
<taxonomicName authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila apiostoma" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="apiostoma">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila apiostoma</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
|
& Vermeulen, sp. nov. (holotype, RMNH 5006718). Apertural (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
), apical (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
), and lateral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
) sides of the shell; microstructure on the protoconch (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
), continuous edentate aperture (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
); spiral microstructure on ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
|
|
) and frontal (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
|
|
) surface of the body whorl.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.1147.93824.figure10" httpUri="https://binary.pensoft.net/fig/808819" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
|
|
Conchological variability of
|
|
<taxonomicName authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila apiostoma" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="apiostoma">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila apiostoma</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
|
& Vermeulen, sp. nov. from site WMVT.0327 (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A, C</emphasis>
|
|
paratypes [RMNH 5006722]
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
holotype).
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.1147.93824.figure11" httpUri="https://binary.pensoft.net/fig/808820" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
|
|
Distribution of
|
|
<taxonomicName authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila apiostoma" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="apiostoma">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila apiostoma</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
|
& Vermeulen, sp. nov. and
|
|
<taxonomicName authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila babel" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="babel">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila babel</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
|
& Vermeulen, sp. nov.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
The ventral side and the shape of the aperture of
|
|
<taxonomicName lsidName="A. apiostoma" pageId="0" pageNumber="1" rank="species" species="apiostoma">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">A. apiostoma</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. resembles those of
|
|
<taxonomicName authorityName="Pall-Gergely & Jochum" authorityYear="2017" class="Gastropoda" family="Gastrocoptidae" genus="Dentisphaera" kingdom="Animalia" lsidName="Dentisphaera maxema" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="maxema">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Dentisphaera maxema</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
|
& Jochum, 2017, which has a more protruding aperture (i.e., a longer tuba) and six apertural teeth. However, the two prominent teeth on the parietal side of
|
|
<taxonomicName authorityName="Pall-Gergely & Jochum" authorityYear="2017" class="Gastropoda" family="Gastrocoptidae" genus="Dentisphaera" kingdom="Animalia" lsidName="Dentisphaera" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Dentisphaera</emphasis>
|
|
</taxonomicName>
|
|
distinguishes this genus from
|
|
<taxonomicName authorityName="Jochum, Slapnik & Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |