157 lines
19 KiB
XML
157 lines
19 KiB
XML
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<mods:title id="FD18D7D550C3CB530030F2DC2C8DE05C">Revision of the Ipsviciidae of the Late Triassic of Queensland (Hemiptera Cicadomorpha: Scytinopteroidea)</mods:title>
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<mods:namePart id="2C3FF948D209F180FF746D44BE4A40F6">Lambkin, Kevin J.</mods:namePart>
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<mods:title id="0E5F6FA3E6002CA75ED370E936D28E2E">Zootaxa</mods:title>
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<mods:date id="8E866866D7B8D8D2D4FAA020C43C3522">2020</mods:date>
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<mods:number id="ECE823E1774D57C7D7B8E3AB21237918">2020-10-14</mods:number>
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<treatment id="231AE47EFF85FFF863F2F95225CBFBE4" ID-DOI="http://doi.org/10.5281/zenodo.4414274" ID-GBIF-Taxon="177323951" ID-Zenodo-Dep="4414274" LSID="urn:lsid:plazi:treatment:231AE47EFF85FFF863F2F95225CBFBE4" httpUri="http://treatment.plazi.org/id/231AE47EFF85FFF863F2F95225CBFBE4" lastPageId="4" lastPageNumber="507" pageId="3" pageNumber="506" scope_family="Ipsviciidae" scope_order="Hemiptera" scope_subOrder="Cicadomorpha" scope_superFamily="Scytinopteroidea">
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<subSubSection id="E3A906E3FF85FFFF63F2F95220EEF8FC" box="[151,428,1775,1801]" pageId="3" pageNumber="506" type="nomenclature">
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<paragraph id="AB0C5568FF85FFFF63F2F95220EEF8FC" blockId="3.[151,428,1775,1801]" box="[151,428,1775,1801]" pageId="3" pageNumber="506">
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<heading id="F044E204FF85FFFF63F2F95220EEF8FC" bold="true" box="[151,428,1775,1801]" fontSize="11" level="1" pageId="3" pageNumber="506" reason="1">
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<taxonomicName id="6CB32EEBFF85FFFF63F2F95220EEF8FC" authority="Tillyard, 1919" authorityName="Tillyard" authorityYear="1919" box="[151,428,1775,1801]" class="Insecta" genus="Ipsvicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="genus">
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<emphasis id="99C7897AFF85FFFF63F2F95220EEF8FC" bold="true" box="[151,428,1775,1801]" pageId="3" pageNumber="506">
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<emphasis id="99C7897AFF85FFFF63F2F95221B4F8FC" bold="true" box="[151,246,1775,1801]" italics="true" pageId="3" pageNumber="506">Ipsvicia</emphasis>
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Tillyard, 1919
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E3A906E3FF85FFFF63F2F8892081F8B9" box="[151,451,1844,1868]" pageId="3" pageNumber="506" type="reference_group">
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<paragraph id="AB0C5568FF85FFFF63F2F8892081F8B9" blockId="3.[151,848,1844,1898]" box="[151,451,1844,1868]" pageId="3" pageNumber="506">
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<treatmentCitationGroup id="8BA37246FF85FFFF63F2F8892081F8B9" box="[151,451,1844,1868]" pageId="3" pageNumber="506">
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<treatmentCitation id="2A127379FF85FFFF63F2F88920FDF8B9" author="Tillyard, R. J." box="[151,447,1844,1868]" page="878" pageId="3" pageNumber="506" year="1919">
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<taxonomicName id="6CB32EEBFF85FFFF63F2F88920FDF8B9" authority="Tillyard, 1919 a: 878" authorityName="Tillyard" authorityPageNumber="878" authorityYear="1919" box="[151,447,1844,1868]" class="Insecta" genus="Ipsvicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="genus">
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<emphasis id="99C7897AFF85FFFF63F2F88921A5F8BE" box="[151,231,1844,1867]" italics="true" pageId="3" pageNumber="506">Ipsvicia</emphasis>
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<bibRefCitation id="CF222899FF85FFFF638BF88920FDF8B9" author="Tillyard, R. J." box="[238,447,1844,1868]" pageId="3" pageNumber="506" pagination="857 - 896" refId="ref9521" refString="Tillyard, R. J. (1919 a) Mesozoic insects of Queensland. No. 7. Hemiptera Homoptera; with a note on the phylogeny of the Suborder. Proceedings of the Linnean Society of New South Wales, 44, 857 - 896." type="journal article" year="1919">Tillyard, 1919a: 878</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="E3A906E3FF85FFF863F2F8EF244BFF20" lastPageId="4" lastPageNumber="507" pageId="3" pageNumber="506" type="type_taxon">
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<paragraph id="AB0C5568FF85FFFF63F2F8EF2212F89F" blockId="3.[151,848,1844,1898]" box="[151,848,1874,1898]" pageId="3" pageNumber="506">
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<emphasis id="99C7897AFF85FFFF63F2F8EF2061F89F" bold="true" box="[151,291,1874,1898]" inLineHeading="true" pageId="3" pageNumber="506" reason="1">
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<typeStatus id="7408EBCAFF85FFFF63F2F8EF218EF89F" box="[151,204,1874,1898]" pageId="3" pageNumber="506">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="6CB32EEBFF85FFFF624FF8EE2317F89F" authority="Tillyard, 1919" authorityName="Tillyard" authorityYear="1919" box="[298,597,1874,1898]" class="Insecta" genus="Ipsvicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="jonesi">
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<emphasis id="99C7897AFF85FFFF624FF8EE20FFF89F" box="[298,445,1875,1898]" italics="true" pageId="3" pageNumber="506">Ipsvicia jonesi</emphasis>
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Tillyard, 1919
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</taxonomicName>
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, by original designation.
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</paragraph>
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<paragraph id="AB0C5568FF85FFF863F2F824244BFF20" blockId="3.[151,1436,1944,2043]" lastBlockId="4.[151,1437,151,1041]" lastPageId="4" lastPageNumber="507" pageId="3" pageNumber="506">
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<emphasis id="99C7897AFF85FFFF63F2F82420C4F847" bold="true" box="[151,390,1944,1971]" pageId="3" pageNumber="506">Emended diagnosis.</emphasis>
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Tegmen
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<emphasis id="99C7897AFF85FFFF6295F82420BEF847" box="[496,508,1945,1970]" italics="true" pageId="3" pageNumber="506">c</emphasis>
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. 3 times longer than wide; surface sculpture complex, comprising numerous variably shaped patches of fine tubercles each encircled by a smooth, very narrow margin, the patches set in a coarser tuberculate/punctate groundmass, a little finer apically; costal margin distinctly convex; basal cell short and broad, not extending to midpoint of PCu; R evenly convex, parallel with costal margin, the basal R cell therefore of about equal width for entire length; two
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<emphasis id="99C7897AFF82FFF86173FF06237CFF21" box="[534,574,187,212]" italics="true" pageId="4" pageNumber="507">r-m</emphasis>
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; CuA strongly angulate at
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<emphasis id="99C7897AFF82FFF8600EFF0622F3FF21" box="[875,945,187,212]" italics="true" pageId="4" pageNumber="507">m-cua</emphasis>
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; base of 1A slightly upcurved.
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</paragraph>
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</subSubSection>
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<subSubSection id="E3A906E3FF82FFF863A2FF6325CBFBE4" pageId="4" pageNumber="507" type="description">
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<paragraph id="AB0C5568FF82FFF863A2FF632372FD5C" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="507">
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<emphasis id="99C7897AFF82FFF863A2FF63201BFF0D" bold="true" box="[199,345,222,248]" pageId="4" pageNumber="507">Description.</emphasis>
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Tegmen
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<emphasis id="99C7897AFF82FFF862A0FF622093FF0D" box="[453,465,223,248]" italics="true" pageId="4" pageNumber="507">c</emphasis>
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.
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<quantity id="6C4BF88DFF82FFF86284FF62231FFF0C" box="[481,605,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.2" metricValueMin="1.4" pageId="4" pageNumber="507" unit="mm" value="18.0" valueMax="22.0" valueMin="14.0">14–22 mm</quantity>
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long; marginal membrane only slightly widened postero-apically; costal fracture fairly upright; bSc variable convex; dSc very weakly developed, running along middle of costal space; R with 4–6 mostly simple post
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<emphasis id="99C7897AFF82FFF86167FE9A237AFEB4" box="[514,568,295,321]" italics="true" pageId="4" pageNumber="507">1r-m</emphasis>
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branches; M+CuA stem a little variable in length, extending just beyond ½ length of PCu; apical section of CuA directed posteriorly, straight or slightly convex; clavus extending to
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<emphasis id="99C7897AFF82FFF86648FEF6247BFE91" box="[1325,1337,331,356]" italics="true" pageId="4" pageNumber="507">c</emphasis>
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. 0.6 tegmen length; 1A about as long as PCu+1A. Hind wing: size variable in proportion to variable size of the individual, maximum length
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<emphasis id="99C7897AFF82FFF8623EFE2E2025FE59" box="[347,359,403,428]" italics="true" pageId="4" pageNumber="507">c</emphasis>
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.
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<quantity id="6C4BF88DFF82FFF86210FE2E2098FE58" box="[373,474,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="4" pageNumber="507" unit="mm" value="13.8">13.8 mm</quantity>
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, maximum width
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<emphasis id="99C7897AFF82FFF861C6FE2E23EDFE59" box="[675,687,403,428]" italics="true" pageId="4" pageNumber="507">c</emphasis>
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.
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<quantity id="6C4BF88DFF82FFF861D9FE2F2250FE58" box="[700,786,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="4" pageNumber="507" unit="mm" value="5.9">5.9 mm</quantity>
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in measurable specimen; R, M, CuA and CuP very closely approximated basally, CuA and CuP apparently without a common stem; R, M and CuA then more or less parallel, directed towards apical margin, gradually diverging apically, apex of CuA upcurved and forming an apical loop with R and M enclosing a small network of terminal short branches and crossveins; a long, backwardly oblique
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<emphasis id="99C7897AFF82FFF86633FE4224DEFDED" box="[1366,1436,511,536]" italics="true" pageId="4" pageNumber="507">m-cua</emphasis>
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(the single
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<emphasis id="99C7897AFF82FFF86270FD9E202DFDC9" box="[277,367,547,572]" italics="true" pageId="4" pageNumber="507">cua-cup</emphasis>
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illustrated by
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<bibRefCitation id="CF222899FF82FFF86177FD9E23FFFDC8" author="Tillyard, R. J." box="[530,701,547,573]" pageId="4" pageNumber="507" pagination="481 - 498" refId="ref9802" refString="Tillyard, R. J. (1923) Mesozoic insects of Queensland. No. 10. Summary of the Upper Triassic insect fauna of Ipswich, Q. (With an appendix describing new Hemiptera and Planipennia). Proceedings of the Linnean Society of New South Wales, 48, 481 - 498. [https: // www. biodiversitylibrary. org / page / 35067038]" type="journal article" year="1923">Tillyard (1923)</bibRefCitation>
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and
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<bibRefCitation id="CF222899FF82FFF86190FD9E22CEFDCB" author="Evans, J. W." box="[757,908,547,574]" pageId="4" pageNumber="507" pagination="17 - 23" refId="ref7521" refString="Evans, J. W. (1963) The systematic position of the Ipsviciidae (Upper Triassic Hemiptera) and some new Upper Permian and Middle Triassic Homoptera from Australia (Insecta). Journal of the Entomological Society of Queensland, 2, 17 - 23. https: // doi. org / 10.1111 / j. 1440 - 6055.1963. tb 00384. x" type="journal article" year="1963">Evans (1963)</bibRefCitation>
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not detected); CuP simple, very long, sinuous, its apex broadly separated from that of CuA and PCu; both PCu and 1A with thickened bases, both straight basally and sinuous apically, CuP and PCu approximating each other at their mid-points; anal area noticeably crinkled; marginal membrane well developed.
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</paragraph>
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<paragraph id="AB0C5568FF82FFF863A2FD0E25CBFBE4" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="507">
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<emphasis id="99C7897AFF82FFF863A2FD0E2053FD38" bold="true" box="[199,273,691,717]" pageId="4" pageNumber="507">Notes.</emphasis>
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The hind wing venation is noteworthy for the close alignment and apical looping of R, M and CuA, and the long, simple and sinuous CuP, PCu and 1A. The other known hind wings of the
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<taxonomicName id="6CB32EEBFF82FFF86737FD6A2452FD04" authorityName="Handlirsch" authorityYear="1906" box="[1106,1296,727,753]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="superFamily" superFamily="Scytinopteroidea">Scytinopteroidea</taxonomicName>
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are those of several species of
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<taxonomicName id="6CB32EEBFF82FFF86201FD4623FEFCE0" authority="Handlirsch, 1904" authorityName="Handlirsch" authorityYear="1904" box="[356,700,763,789]" class="Insecta" family="Scytinopteridae" genus="Scytinoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="genus">
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<emphasis id="99C7897AFF82FFF86201FD4620B0FCE0" box="[356,498,763,789]" italics="true" pageId="4" pageNumber="507">Scytinoptera</emphasis>
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Handlirsch, 1904
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</taxonomicName>
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, illustrated by
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<bibRefCitation id="CF222899FF82FFF86004FD46253EFCE0" author="Becker-Migdisova, E. E." box="[865,1148,763,789]" pageId="4" pageNumber="507" pagination="1 - 42" refId="ref6881" refString="Becker-Migdisova, E. E. (1948) Permian cicadas of the family Scytinopteridae from the Soyana River. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR, 15, 1 - 42. [in Russian]" type="journal article" year="1948">Becker-Migdisova (1948</bibRefCitation>
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, figs. 2, 33, 39–43). They apparently lack a marginal membrane, and their venation is much simpler than that of
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<taxonomicName id="6CB32EEBFF82FFF86711FCA2258FFCCD" authorityName="Tillyard" authorityYear="1919" box="[1140,1229,799,824]" class="Insecta" genus="Ipsvicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="genus">
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<emphasis id="99C7897AFF82FFF86711FCA2258FFCCD" box="[1140,1229,799,824]" italics="true" pageId="4" pageNumber="507">Ipsvicia</emphasis>
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</taxonomicName>
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, with M and CuA directed postero-apically rather than apically, R, M and CuA without apical looping, CuA and CuP with a common stem, CuP, PCu and 1A almost straight rather than sinuous, and without the long, backwardly oblique
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<emphasis id="99C7897AFF82FFF86793FCDA247EFC75" box="[1270,1340,871,896]" italics="true" pageId="4" pageNumber="507">m-cua</emphasis>
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. Indeed, as noted by
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<bibRefCitation id="CF222899FF82FFF8627BFC3620F5FC50" author="Evans, J. W." box="[286,439,906,933]" pageId="4" pageNumber="507" refId="ref7482" refString="Evans, J. W. (1956) Palaeozoic and Mesozoic Hemiptera (Insecta). Australian Journal of Zoology, 4, 165 - 158. https: // doi. org / 10.1071 / ZO 9560165" type="book" year="1956">Evans (1956)</bibRefCitation>
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, the venation of
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<taxonomicName id="6CB32EEBFF82FFF86116FC36238EFC51" authorityName="Tillyard" authorityYear="1919" box="[627,716,907,932]" class="Insecta" genus="Ipsvicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="genus">
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<emphasis id="99C7897AFF82FFF86116FC36238EFC51" box="[627,716,907,932]" italics="true" pageId="4" pageNumber="507">Ipsvicia</emphasis>
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</taxonomicName>
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was unlike any of the Palaeozoic or Mesozoic homopteran hind wings known at that time (see also those illustrated by
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<bibRefCitation id="CF222899FF82FFF86194FC122540FC3F" author="Becker-Migdisova, E. E." box="[753,1026,943,970]" pageId="4" pageNumber="507" pagination="286 - 393" refId="ref6918" refString="Becker-Migdisova, E. E. (1961) Suborder Rhynchota, In: Rohdendorf, B. B., Becker-Migdisova, E. E., Martynova, O. M. & Sharov, A. G. (Eds.), Paleozoic insects of the Kuznetsk Basin, Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR, 85, pp 286 - 393. [in Russian]" type="journal article" year="1961">Becker-Migdisova 1961</bibRefCitation>
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), and it differs from that of the extant families of
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<taxonomicName id="6CB32EEBFF82FFF86272FC6E20FEFC18" authorityName="Evans" authorityYear="1946" box="[279,444,979,1005]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="subOrder" subOrder="Cicadomorpha">Cicadomorpha</taxonomicName>
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(see figures in
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<bibRefCitation id="CF222899FF82FFF8610FFC6E2241FC18" author="Kramer, S." box="[618,771,978,1005]" pageId="4" pageNumber="507" pagination="1 - 111" refId="ref8211" refString="Kramer, S. (1950) The morphology and phylogeny of auchenorrhynchous Homoptera (Insecta). Illinois Biological Monographs, 20 (4), i-vii + 1 - 111. https: // doi. org / 10.5962 / bhl. title. 50340" type="journal article" year="1950">Kramer 1950</bibRefCitation>
|
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). It might be observed, however, that the form of R, M and CuA bears some resemblance to that of Heteroptera:
|
||
<taxonomicName id="6CB32EEBFF82FFF8618DFC4A2238FBE4" box="[744,890,1015,1041]" infraOrder="Nepomorpha" pageId="4" pageNumber="519" rank="infraOrder">Nepomorpha</taxonomicName>
|
||
(
|
||
<bibRefCitation id="CF222899FF82FFF860EDFC4A2551FBE4" author="Popov, Y. A." box="[904,1043,1015,1041]" pageId="4" pageNumber="507" pagination="1 - 230" refId="ref8833" refString="Popov, Y. A. (1971) Historical development of the infraorder Nepomorpha (Heteroptera). Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR, 129, 1 - 230. [in Russian]" type="journal article" year="1971">Popov 1971</bibRefCitation>
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, plate 52).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
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||
</document> |