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<document ID-CLB-Dataset="229647" ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-GBIF-Dataset="6ba6d2ca-5dad-4270-914f-1042327d503c" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="22E8B7F71CCF59498611B6AEE552340B" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Coarctana glabra Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="5BF609DC-F952-4CE6-BE27-11A2BA6AEC20" docUuidSource="ZooBank" docVersion="5" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1732669275789" updateUser="ExternalLinkService">
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<mods:title id="FED1E9DF5097FBE85F1B040634B69684">Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:name id="E0B9C840AB95FEBAA029072AF212768F" type="personal">
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<mods:namePart id="827168EA135CEF377FB233B68F65A5DB">Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier id="7AF60ED05FB2E5FB3AE3898B2D47F843" type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation id="ED2EB92FF8BB63975E9636ADB3AC3AAB">Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier id="8C70FC32826446DD0030571A2792EE76" type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="83B6B272F732D7C47A2014B1846E71D8">Cavichioli, Rodney Ramiro</mods:namePart>
<mods:nameIdentifier id="3C6AE317794226FD4E406EE158359F76" type="ORCID">https://orcid.org/0000-0001-5996-0629</mods:nameIdentifier>
<mods:affiliation id="CCC9472D5DE91A8EDD0F6E7FD6F0A109">Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:title id="428565767979B77164F242C68EFC0775">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date id="632EC606B09AA9412C983669F08E9A76">2023</mods:date>
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<treatment id="22E8B7F71CCF59498611B6AEE552340B" ID-GBIF-Taxon="209013703" LSID="urn:lsid:zoobank.org:act:5BF609DC-F952-4CE6-BE27-11A2BA6AEC20" httpUri="http://treatment.plazi.org/id/22E8B7F71CCF59498611B6AEE552340B" lastPageNumber="79" pageId="0" pageNumber="79" scope_family="Cicadellidae" scope_order="Hemiptera" scope_tribe="Gyponini">
<subSubSection id="4D80DD569C80468F226823020FD12239" pageId="0" pageNumber="79" type="nomenclature">
<paragraph id="185D210477471E4E3F7BFF4F7BE6CCD8" pageId="0" pageNumber="79">
<taxonomicName id="92658765C87A1FAE3E7A19D57CC6D0F5" LSID="https://zoobank.org/5BF609DC-F952-4CE6-BE27-11A2BA6AEC20" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana glabra" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="glabra" status="sp. nov.">Coarctana glabra</taxonomicName>
<taxonomicNameLabel id="CD156723C99DECCA65B2843B322FDF47" pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="A3A2EAE8160A88F0D7347B32C07D88E1" pageId="0" pageNumber="79" type="description">
<paragraph id="230311CF74A1AC75122D377B80F84307" pageId="0" pageNumber="79">
<figureCitation id="76BDC100388D0F1981116BDBE008AF56" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">Figs 19</figureCitation>
<figureCitation id="188DF0C169D162F0561362330D416073" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">, 31E, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="390F16DD329EE2917C69638B66C0F56B" pageId="0" pageNumber="79" type="diagnosis">
<paragraph id="269CAB2DAA149C0FB7015C80E541C029" pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph id="921039D1B4D43DF29C72E9A4D89C4E0F" pageId="0" pageNumber="79">
Subgenital plate, in ventral view (Fig.
<figureCitation id="0EDA4FB16C2E917155B6628CEFDB0F79" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19H</figureCitation>
), ovoid, outer margin with minute setae. Connective (Fig.
<figureCitation id="0D7C03B9E49FB838C9770C49E6EE127A" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19I</figureCitation>
) with stem thin, about as long as arm length. Style, in lateral view (Fig.
<figureCitation id="9D311778633BFAF74AB73D78109C06B0" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19K</figureCitation>
), ventral margin with median third with small, rounded lobe; apex strongly tapered, curved dorsally. Aedeagus (Fig.
<figureCitation id="57C5A2E8B1D2B11A1C038B7457BAEF30" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19L, M</figureCitation>
) apodemal process elongated, broadly curved dorsally; apex tapered and truncated, finely serrated; shaft apex with two pairs of short processes: subapical pair directed ventrally, and apical pair curved laterally, as long as the subapical pair.
</paragraph>
</subSubSection>
<subSubSection id="78DC208BA9D8AD535A93CD4DFEC0BECC" pageId="0" pageNumber="79" type="description">
<paragraph id="482E782E3E73A5EE2EEEDE9A594D9A54" pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph id="AC7657E7462F430773069146F72EA382" pageId="0" pageNumber="79">Total length: holotype male 8.0 mm.</paragraph>
</subSubSection>
<subSubSection id="F316C438053ED81057D82AE33CEAFEAC" pageId="0" pageNumber="79" type="description">
<paragraph id="96B5495E01405162F7041F3632053C7F" pageId="0" pageNumber="79">Description.</paragraph>
<paragraph id="1559A137F7F989AC26FB76F661677053" pageId="0" pageNumber="79">
Head, in frontal view (Fig.
<figureCitation id="B4BCF24FEA770E715A9D76FC049E02D8" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19B</figureCitation>
), frons 1.1
<normalizedToken id="3434D974A5C98EF8C2016B35BCAB0240" originalValue="×">x</normalizedToken>
wider than long; frontogenal suture distant from eye margin by approximately half maximum width of clypeus; clypeus 1.5
<normalizedToken id="DC7C501EA9B208D56397CF2CDCAF1D6E" originalValue="×">x</normalizedToken>
wider than long; lateral margins parallel; apex straight. Head, in lateral view (Fig.
<figureCitation id="69E105EA0E86560DCF70A95856E1C030" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19C</figureCitation>
), crown-face transition thin, almost foliaceus, with 3 transverse carinae; clypeus not inflated. Profemur AV row with 5-6 setae; PV row with 2 setae. Protibia PD row with 3 setae and undifferentiated intercalary setae; PV row with 2-3 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 24, 13 and 14-15 macrosetae respectively. Metatarsomere I moderately elongated, 3
<normalizedToken id="DE6BC46F7A2CBC4747E412ECF6C24BDD" originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 7-8 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection id="F723F437207B8CD7CFD35F0DD91FB7B6" pageId="0" pageNumber="79" type="coloration">
<paragraph id="BA53CEF0FFB46A5658C91E628468430C" pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph id="9290C410B84AF2AD05473712648AF127" pageId="0" pageNumber="79">
Head and thorax (Figs
<figureCitation id="BBE70B741CE1A3B0BB1BE2C5D8B2216E" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19A-D</figureCitation>
,
<figureCitation id="7D02B383DB94504337D7ED99C455D236" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">31E, F</figureCitation>
) reddish-brown. Head (Fig.
<figureCitation id="095B0A2E8955D7EF4663DFAB4ED4851F" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19D</figureCitation>
) with pair of small circular black spots, near posterior margin, behind ocelli. Gena (Fig.
<figureCitation id="08BCF0F5775CFBE1181205DE6C802326" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19B</figureCitation>
) yellowish; antennal pit dark brown. Pronotum (Fig.
<figureCitation id="CB12E3B34665CDA8F65985604FFD7968" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19A</figureCitation>
) with black punctures and irregular black spots near anterior margin; proepimeron (Fig.
<figureCitation id="4DCEDB6113DE9777449BDA9E0F343EEE" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19C</figureCitation>
), with black band below pronotal carina, wider near anterior margin; lateral carina yellow. Mesonotum (Fig.
<figureCitation id="DE557F1CCD56A8BC9398C19E3B379301" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19A</figureCitation>
) with dark-brown lateral angles, pair of small median black spots near scutoscutellar suture, pair of small yellow spots on lateral angles. Forewing (Fig.
<figureCitation id="534C470BAEB425465A6C8B0CE6E68ADC" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19D</figureCitation>
) strongly marked by dark-brown mottling; veins brown, outlined by dark brown. Legs yellowish with black punctures and maculae. Metatibia (Fig.
<figureCitation id="8CB44E033010D05BC33D4E5E9C27B4B6" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection id="29229002B046D073DBFC3ED10BFC6D7D" pageId="0" pageNumber="79" type="male terminalia">
<paragraph id="24913CD1F985937165D9A645D9D985C1" pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph id="638E977B557A4E98704AAE1DBEF6F9A6" pageId="0" pageNumber="79">
Sternite VIII (Fig.
<figureCitation id="03C9945E46E51094E6D02BAA66886ED4" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19E</figureCitation>
) 1.5
<normalizedToken id="DCB47F7317BE83A92CE914D284D65258" originalValue="×">x</normalizedToken>
wider than long; lateral margins parallel; posterior margin slightly excavated each side of broad median lobe. Valve (Fig.
<figureCitation id="A93F08FE26A6FF483ACD1AD9D019C2C2" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19F</figureCitation>
) 2
<normalizedToken id="0727C56F9D7A2B0F4756CF8196412B1F" originalValue="×">x</normalizedToken>
wider than long; posterior margin notched medially. Pygofer, in lateral view (Fig.
<figureCitation id="E589C9ECD29826595D1D4F1FB96E32D9" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19G</figureCitation>
), 1.5
<normalizedToken id="0DC4016FF07B82B06FAD38A6DB383269" originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process present but reduced; anteroventral margin truncated; posterodorsal margin rounded; posteroventral margin straight; apex broadly rounded. Subgenital plate, in lateral view (Fig.
<figureCitation id="4515846554CC2E7F4D4727DB8DA78F45" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19G</figureCitation>
), almost reaching pygofer apex; apex curved dorsally; in ventral view (Fig.
<figureCitation id="843C7A71E9F5712A9E72C2A49B9B786A" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19H</figureCitation>
), ovoid, 2.5
<normalizedToken id="BDBEC760B23F79851A131AF5992B21CE" originalValue="×">x</normalizedToken>
longer than wide, maximum width near basal third; outer margin with minute setae; lateral margins rounded; apex weakly tapered and rounded. Connective (Fig.
<figureCitation id="EC8A89B316141F0C747A3B43E18FDE61" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19I</figureCitation>
) wider than long; anterior margin weakly excavated; dorsal keel reduced; stem thin, about as long as arm length. Style, in dorsal view (Fig.
<figureCitation id="E0A70E3286C19B82A5C5065990C9D8F3" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19J</figureCitation>
), with outer lobe reduced; in lateral view (Fig.
<figureCitation id="2BB943BA5A8FC1F4E0C658D787A775E3" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19K</figureCitation>
), with blade wider near base and tapered toward apex; dorsal margin slightly sinuate; ventral margin with apical half serrated, median third with small, rounded lobe; apex strongly tapered, curved dorsally. Aedeagus (Fig.
<figureCitation id="BF25C1B9057585ACEE3CDBD765374A95" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19L, M</figureCitation>
) preatrium weakly developed; dorsal apodeme with dorsal margin slightly excavated, lateral margins produced ventrolaterally; apodemal process elongated, strongly curved dorsally, slightly wider medially; apex tapered and truncated, finely serrated; shaft slightly asymmetrical, basal half weakly flattened dorsoventrally and cylindrical apically, curved dorsally near base; apex slightly twisted laterally, with two pairs of short processes: subapical pair directed ventrally and apical pair curved laterally, as long as the subapical pair. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection id="280F8E9C72393FBC6BA6A183856E6D4F" pageId="0" pageNumber="79" type="female">
<paragraph id="2C04111E313F09E7A37B6D0B298DF9B3" pageId="0" pageNumber="79">Female.</paragraph>
<paragraph id="AA9BD7167D42B8B27B14E4E24C486B24" pageId="0" pageNumber="79">Unknown.</paragraph>
</subSubSection>
<subSubSection id="180DC29C9ED1620AC86133B525BAE735" pageId="0" pageNumber="79" type="materials_examined">
<paragraph id="A305AFAABF9CE127C3D1F1E52F46095E" pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph id="5E290D868EED7397B516EE6F8BE0E085" pageId="0" pageNumber="79">
<materialsCitation id="3ACD5DD375BA275CDF075A97F5998DFF" ID-GBIF-Occurrence="4017211311" collectingDate="2014-11" collectorName="J. A. Rafael, F. F. Xavier" country="Brazil" county="Serra" location="Igarape" municipality="Navio" specimenCount="1" stateProvince="Amapa" typeStatus="Holotype">
<emphasis id="9426389730CC5E9B6B8E938515719628" bold="true" pageId="0" pageNumber="79">
<typeStatus id="4C3729AD3223740F39623D99745EE1F8">Holotype</typeStatus>
male:
<collectingCountry id="14987A756C5B28C862ED68DD9E5C7D90" name="Brazil">Brazil</collectingCountry>
:
<normalizedToken id="3C835D7C9ECF4D780AEB8FF02BDC14C3" originalValue="Amapá">
<collectingRegion id="510C2E31CB8AD750D879A1F772F85398" country="Brazil" name="Amapa">Amapa</collectingRegion>
</normalizedToken>
</emphasis>
: &quot;Brasil, AP,
<collectingCounty id="CF0274ABC8DE70B5625F44284070EABA">Serra</collectingCounty>
do\
<collectingMunicipality id="EADB7AF4CECAF2BBC57702160EA9060D">Navio</collectingMunicipality>
,
<normalizedToken id="813E74DE6D23F7A0D58DE5D523B15FAE" originalValue="Igarapé">
<location id="88737DD91EDBE90FAAF15A35108927B1" LSID="urn:lsid:plazi:treatment:22E8B7F71CCF59498611B6AEE552340B:88737DD91EDBE90FAAF15A35108927B1" country="Brazil" county="Serra" municipality="Navio" name="Igarape" stateProvince="Amapa">Igarape</location>
</normalizedToken>
na\ estrada para
<location id="D9BD7A95A3DCA60FF4EAB7B881AC111E" LSID="urn:lsid:plazi:treatment:22E8B7F71CCF59498611B6AEE552340B:D9BD7A95A3DCA60FF4EAB7B881AC111E" country="Brazil" county="Serra" municipality="Navio" name="Rio Amapari" stateProvince="Amapa">Rio Amapari</location>
(
<location id="82A73EDAE0661EF1732A8BABACABBC19" LSID="urn:lsid:plazi:treatment:22E8B7F71CCF59498611B6AEE552340B:82A73EDAE0661EF1732A8BABACABBC19" country="Brazil" county="Serra" municipality="Navio" name="Pedra Preta" stateProvince="Amapa">Pedra Preta</location>
)\
<collectingDate id="BB237F6101AEA121598617272C19D886" value="2014-11">XI.2014</collectingDate>
<collectorName id="35398B153B5EC66551D6E56CB755D577">J.A. Rafael</collectorName>
&amp;
<collectorName id="033DDF86702C6330520D2F2D27B338A4">F.F. Xavier</collectorName>
&quot; (INPA).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3319A83D1BA873F8D92304FF43EA634" pageId="0" pageNumber="79" type="etymology">
<paragraph id="9B4CB7A3E0FC0ADD4BFB668F9195A5C3" pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph id="C0D75A2D63D930E3917A581CEB1F5663" pageId="0" pageNumber="79">The new species name refers to the subgenital plates without filiform setae.</paragraph>
</subSubSection>
<subSubSection id="0314FCBD0C55822457BBF57FC8952A9C" pageId="0" pageNumber="79" type="remarks">
<paragraph id="2CC16EF5BE3E6CFD6661FC1319CC6FCF" pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph id="92F57074F59CA114623FC777CF3E08A2" pageId="0" pageNumber="79">
<taxonomicName id="0C69E93EE111D7BFCB774F3D68A258BF" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. glabra" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="0CC0A76311D23886BD7B1743D38A02A0" italics="true" pageId="0" pageNumber="79">C. glabra</emphasis>
</taxonomicName>
<emphasis id="2E52119ADFFDC4D09AC09AE418FCF8B4" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
is sister to
<taxonomicName id="6E44F1BB8570AF56E3BFE3E67DF8EE31" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. rana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="rana">
<emphasis id="9A7C00E3A2165A44820D485268F10A6F" italics="true" pageId="0" pageNumber="79">C. rana</emphasis>
</taxonomicName>
<emphasis id="F0F784649D2D285EE85FDFF4613EED31" bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
+
<taxonomicName id="61192648C42D69E271F675FB493A55DA" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. serpenta" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="serpenta">
<emphasis id="D5B39598429839D7C33CA11BBD0D47B5" italics="true" pageId="0" pageNumber="79">C. serpenta</emphasis>
</taxonomicName>
<emphasis id="E80C3A42F1713EFDCDF6683A557E64E0" bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
with moderate branch support (SR = 65) (Fig.
<figureCitation id="4F2BFD8DB619CD54B3696AF4135A8097" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationships of Acuera found in the implied weighting analysis for the two equally most parsimonious trees, K = 3.700 and 4.284, length = 514, consistency index = 0.362, retention index = 0.771. Symmetric resampling values (&lt;50) are marked next to branches; red stars represent the type species of the three subgenera of Acuera. Synapomorphies are marked with black circles, while homoplasies are marked with empty circles." figureDoi="10.3897/asp.81.e81961.figure7" httpUri="https://binary.pensoft.net/fig/798160" pageId="0" pageNumber="79">7</figureCitation>
).
<taxonomicName id="7B70BF89CA2170828B6E3BBA2D3F2586" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana glabra" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="F8E743787DFD7AC60F864CAA40983C48" italics="true" pageId="0" pageNumber="79">Coarctana glabra</emphasis>
</taxonomicName>
<emphasis id="963BD409FFE7950F43B0E27FB2F8C9C2" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
resembles
<taxonomicName id="DD61482CF1650FADF0F74243B6FCF626" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. rana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="rana">
<emphasis id="D76E9BE2CA7179ACCB0812BD87C638C6" italics="true" pageId="0" pageNumber="79">C. rana</emphasis>
</taxonomicName>
<emphasis id="F2D7E2354A5E127F535879FFF6D22786" bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
and
<taxonomicName id="BDD980A59CEAE85B5CFDC7BE76F4C11B" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. serpenta" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="serpenta">
<emphasis id="1FD0B9ED189587554875FC351DBE5330" italics="true" pageId="0" pageNumber="79">C. serpenta</emphasis>
</taxonomicName>
<emphasis id="E197FEAFD8C61833B3632E9622AC1D54" bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
by having the forewings (Figs
<figureCitation id="B178265B3D81967BFDE24351FA21F8D1" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">31E</figureCitation>
,
<figureCitation id="0A438E3885396C40520D6C24CC392D7F" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32C-F</figureCitation>
) with several small black maculae and by the similar shape of the apodemal processes of aedeagus (Fig.
<figureCitation id="1BD7113908041034E89992CED2965DF7" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19L</figureCitation>
) slender and strongly curved dorsally. The new species is easily separated by the subgenital plate (Fig.
<figureCitation id="3A1C59902753CA39746436B1F6F44FCA" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19H</figureCitation>
) ovate and lacking long filiform setae.
</paragraph>
</subSubSection>
</treatment>
</document>
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