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<document id="6191A3AA7E131EDA9992894834792803" ID-CLB-Dataset="54817" ID-DOI="10.11646/zootaxa.5325.4.2" ID-GBIF-Dataset="83f870a7-384b-4bf7-926b-424795815e17" ID-ISSN="1175-5326" ID-Zenodo-Dep="8243618" ID-ZooBank="8FD2B222-ACC1-4C8B-B18C-799B1A2F9F1D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1691942856599" checkinUser="plazi" docAuthor="Liu, Haoyu &amp; Wang, Shuxia" docDate="2023" docId="223987C00F1CFFD4FF4C6853FD05FE1B" docLanguage="en" docName="zootaxa.5325.4.2.pdf" docOrigin="Zootaxa 5325 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5325.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thecobathra microsignata Liu 1980" docType="treatment" docVersion="6" lastPageNumber="496" masterDocId="DE00FFB80F10FFD9FFDB6D0CFFC2FF92" masterDocTitle="Taxonomic study of the genus Thecobathra Meyrick (Lepidoptera: Yponomeutidae) in China, with descriptions of eight new species" masterLastPageNumber="508" masterPageNumber="483" pageNumber="495" updateTime="1699557074084" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C90730BFDB6CDB02DC8A5D43C2EA7F0F">Taxonomic study of the genus Thecobathra Meyrick (Lepidoptera: Yponomeutidae) in China, with descriptions of eight new species</mods:title>
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<mods:namePart id="5B74633C05481BAC570DC9EBE335F20B">Liu, Haoyu</mods:namePart>
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<mods:affiliation id="975F1A9FDE799CDF11548633E336CA5B">College of Life Sciences, Nankai University, Tianjin 300071, China. &amp; liuhaoyulisa @ 163. com; https: // orcid. org / 0009 - 0001 - 9556 - 1526</mods:affiliation>
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<mods:namePart id="6F3EE75FBD138938BABCC2DFF17F4F99">Wang, Shuxia</mods:namePart>
<mods:affiliation id="FB23102C17E9C1B670496930517F1054">College of Life Sciences, Nankai University, Tianjin 300071, China.</mods:affiliation>
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<treatment id="223987C00F1CFFD4FF4C6853FD05FE1B" ID-DOI="http://doi.org/10.5281/zenodo.8243668" ID-GBIF-Taxon="212429965" ID-Zenodo-Dep="8243668" LSID="urn:lsid:plazi:treatment:223987C00F1CFFD4FF4C6853FD05FE1B" httpUri="http://treatment.plazi.org/id/223987C00F1CFFD4FF4C6853FD05FE1B" lastPageId="13" lastPageNumber="496" pageId="12" pageNumber="495">
<subSubSection id="E28A655D0F1CFFD5FF4C6853FD96FAEB" box="[151,596,1375,1402]" pageId="12" pageNumber="495" type="nomenclature">
<paragraph id="AA2F36D60F1CFFD5FF4C6853FD96FAEB" blockId="12.[151,596,1375,1437]" box="[151,596,1375,1402]" pageId="12" pageNumber="495">
<heading id="F16781BA0F1CFFD5FF4C6853FD96FAEB" bold="true" box="[151,596,1375,1402]" fontSize="11" level="1" pageId="12" pageNumber="495" reason="1">
<taxonomicName id="6D904D550F1CFFD5FF4C6853FD96FAEB" ID-CoL="BHYRT" authority="Liu, 1980" authorityName="Liu" authorityYear="1980" box="[151,596,1375,1402]" class="Insecta" family="Yponomeutidae" genus="Thecobathra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="495" phylum="Arthropoda" rank="species" species="microsignata">
<emphasis id="98E4EAC40F1CFFD5FF4C6853FD96FAEB" bold="true" box="[151,596,1375,1402]" pageId="12" pageNumber="495">
<emphasis id="98E4EAC40F1CFFD5FF4C6853FE15FAEB" bold="true" box="[151,471,1375,1402]" italics="true" pageId="12" pageNumber="495">Thecobathra microsignata</emphasis>
<bibRefCitation id="CE014B270F1CFFD5FE046853FD96FAEB" author="Liu, Y. Q." box="[479,596,1375,1401]" pageId="12" pageNumber="495" pagination="33 - 40" refId="ref20613" refString="Liu, Y. Q. (1980) A study of Chinese Thecobathra Meyrick (Lepidoptera: Yponomeutidae). Entomotaxonomia, 2 (1), 33 - 40." type="journal article" year="1980">Liu, 1980</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E28A655D0F1CFFD5FF4C688FFF31FA0F" box="[151,243,1411,1437]" pageId="12" pageNumber="495" type="description">
<paragraph id="AA2F36D60F1CFFD5FF4C688FFF31FA0F" blockId="12.[151,596,1375,1437]" box="[151,243,1411,1437]" pageId="12" pageNumber="495">
(
<figureCitation id="32AB2A530F1CFFD5FF44688FFF28FA0F" box="[159,234,1411,1437]" captionStart="FIGURES 15–18" captionStartId="11.[151,264,1803,1827]" captionTargetBox="[157,1420,194,1766]" captionTargetId="figure-18@11.[155,1433,181,1779]" captionTargetPageId="11" captionText="FIGURES 15–18. Male genitalia of Thecobathra spp. 15, T. meniscata sp. nov., holotype, slide No. LHY21528; 16, T. microsignata, slide No. LHY21734; 17, T. minutiprocessa sp. nov., holotype, slide No. LHY21667; 18, T. odontoidea sp. nov., holotype, slide No. LHY21068. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8243632" httpUri="https://zenodo.org/record/8243632/files/figure.png" pageId="12" pageNumber="495">Fig 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E28A655D0F1CFFD5FF4C68CEFC5BFA48" box="[151,921,1474,1498]" pageId="12" pageNumber="495" type="reference_group">
<paragraph id="AA2F36D60F1CFFD5FF4C68CEFC5BFA48" blockId="12.[151,921,1474,1498]" box="[151,921,1474,1498]" pageId="12" pageNumber="495">
<treatmentCitationGroup id="8A8011F80F1CFFD5FF4C68CEFC5BFA48" box="[151,921,1474,1498]" pageId="12" pageNumber="495">
<treatmentCitation id="2B3110C70F1CFFD5FF4C68CEFDF4FA48" author="Liu, Y. Q." box="[151,566,1474,1498]" page="37" pageId="12" pageNumber="495" year="1980">
<taxonomicName id="6D904D550F1CFFD5FF4C68CEFDF4FA48" ID-CoL="BHYRT" authority="Liu, 1980: 37" authorityName="Liu" authorityPageNumber="37" authorityYear="1980" box="[151,566,1474,1498]" class="Insecta" family="Yponomeutidae" genus="Thecobathra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="495" phylum="Arthropoda" rank="species" species="microsignata">
<emphasis id="98E4EAC40F1CFFD5FF4C68CEFE66FA48" box="[151,420,1474,1498]" italics="true" pageId="12" pageNumber="495">Thecobathra microsignata</emphasis>
<materialsCitation id="1AF83C8B0F1CFFD5FE7168CEFDF4FA48" ID-GBIF-Occurrence="4173775423" box="[426,566,1474,1498]" collectionCode="IZCAS" country="China" location="Yunnan" pageId="12" pageNumber="495" specimenCount="1" stateProvince="Yunnan">
<bibRefCitation id="CE014B270F1CFFD5FE7168CEFDF4FA48" author="Liu, Y. Q." box="[426,566,1474,1498]" pageId="12" pageNumber="495" pagination="33 - 40" refId="ref20613" refString="Liu, Y. Q. (1980) A study of Chinese Thecobathra Meyrick (Lepidoptera: Yponomeutidae). Entomotaxonomia, 2 (1), 33 - 40." type="journal article" year="1980">Liu, 1980: 37</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
. TL:
<collectingCountry id="D28776460F1CFFD5FDB768CEFD6BFA48" box="[620,681,1474,1498]" name="China" pageId="12" pageNumber="495">China</collectingCountry>
(
<collectingRegion id="6854F8340F1CFFD5FD6D68CEFCC8FA48" box="[694,778,1474,1498]" country="China" name="Yunnan" pageId="12" pageNumber="495">Yunnan</collectingRegion>
). TD: IZCAS.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E28A655D0F1CFFD5FF4C6B04FDBAF91C" pageId="12" pageNumber="495" type="materials_examined">
<paragraph id="AA2F36D60F1CFFD5FF4C6B04FDBAF91C" blockId="12.[151,1437,1544,2039]" pageId="12" pageNumber="495">
<emphasis id="98E4EAC40F1CFFD5FF4C6B04FE42F9B0" bold="true" box="[151,384,1544,1570]" pageId="12" pageNumber="495">Material examined.</emphasis>
<materialsCitation id="1AF83C8B0F1CFFD5FE5C6B04FB85F9DB" ID-GBIF-Occurrence="4173775427" collectingDate="2014-08-17" collectorName="KJ Teng" country="China" elevation="2009" latitude="24.82" location="Nankang" longLatPrecision="750" longitude="98.78" pageId="12" pageNumber="495" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan">
<emphasis id="98E4EAC40F1CFFD5FE5C6B04FD9AF9B1" bold="true" box="[391,600,1544,1571]" pageId="12" pageNumber="495">
<collectingCountry id="D28776460F1CFFD5FE5C6B04FE24F9B0" box="[391,486,1544,1570]" name="China" pageId="12" pageNumber="495">CHINA</collectingCountry>
,
<collectingRegion id="6854F8340F1CFFD5FE2A6B05FD91F9B1" box="[497,595,1545,1571]" country="China" name="Yunnan" pageId="12" pageNumber="495">Yunnan</collectingRegion>
:
</emphasis>
13♁,
<location id="AF4F600D0F1CFFD5FD456B04FCC1F9B0" LSID="urn:lsid:plazi:treatment:223987C00F1CFFD4FF4C6853FD05FE1B:AF4F600D0F1CFFD5FD456B04FCC1F9B0" box="[670,771,1544,1570]" country="China" latitude="24.82" longLatPrecision="750" longitude="98.78" name="Nankang" pageId="12" pageNumber="495" stateProvince="Yunnan">Nankang</location>
(
<geoCoordinate id="CFA450110F1CFFD5FCC96B05FCB7F9B1" box="[786,885,1545,1571]" degrees="24.82" direction="north" orientation="latitude" pageId="12" pageNumber="495" precision="555" value="24.82">24.82°N</geoCoordinate>
,
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),
<quantity id="6D689B330F1CFFD5FC296B05FB8BF9B1" box="[1010,1097,1545,1571]" metricMagnitude="3" metricUnit="m" metricValue="2.009" pageId="12" pageNumber="495" unit="m" value="2009.0">
<elevation id="21BDD1E50F1CFFD5FC296B05FB8BF9B1" box="[1010,1097,1545,1571]" metricMagnitude="3" metricUnit="m" metricValue="2.009" pageId="12" pageNumber="495" unit="m" value="2009.0">2009 m</elevation>
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,
<date id="DE2E10160F1CFFD5FB8F6B05FB2BF9B1" box="[1108,1257,1545,1571]" pageId="12" pageNumber="495" value="2014-08-17">
<collectingDate id="CE6AE9FE0F1CFFD5FB8F6B05FB2BF9B1" box="[1108,1257,1545,1571]" pageId="12" pageNumber="495" value="2014-08-17">17.VIII.2014</collectingDate>
</date>
, leg.
<collectorName id="076553000F1CFFD5FAFF6B05FA40F9B1" box="[1316,1410,1545,1571]" pageId="12" pageNumber="495">KJ Teng</collectorName>
<emphasis id="98E4EAC40F1CFFD5FA536B05FF6FF9D4" italics="true" pageId="12" pageNumber="495">et al</emphasis>
., slide Nos. LHY21535, LHY21733, LHY21734, LHY21737, LHY21738; 37♁
<specimenCount id="BC96FD5F0F1CFFD5FBFD6B22FB85F9DB" box="[1062,1095,1582,1609]" pageId="12" pageNumber="495" type="female">3♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="1AF83C8B0F1CFFD5FB896B20FA55F9F8" ID-GBIF-Occurrence="4173775413" collectingDate="2015-07-16" collectingDateMax="2015-07-21" collectingDateMin="2015-07-16" collectorName="KJ Teng &amp; X Bai" country="China" elevation="2009" latitude="24.82" location="Nankang" longLatPrecision="750" longitude="98.78" pageId="12" pageNumber="495" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<location id="AF4F600D0F1CFFD5FB896B20FB75F9D4" LSID="urn:lsid:plazi:treatment:223987C00F1CFFD4FF4C6853FD05FE1B:AF4F600D0F1CFFD5FB896B20FB75F9D4" box="[1106,1207,1580,1606]" country="China" latitude="24.82" longLatPrecision="750" longitude="98.78" name="Nankang" pageId="12" pageNumber="495" stateProvince="Yunnan">Nankang</location>
(
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,
<geoCoordinate id="CFA450110F1CFFD5FAEA6B21FA52F9D5" box="[1329,1424,1581,1607]" degrees="98.78" direction="east" orientation="longitude" pageId="12" pageNumber="495" precision="555" value="98.78">98.78°E</geoCoordinate>
),
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<elevation id="21BDD1E50F1CFFD5FF4C6B5DFF2EF9F9" box="[151,236,1617,1643]" metricMagnitude="3" metricUnit="m" metricValue="2.009" pageId="12" pageNumber="495" unit="m" value="2009.0">2009 m</elevation>
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,
<date id="DE2E10160F1CFFD5FF2D6B5CFE6EF9F9" box="[246,428,1616,1643]" pageId="12" pageNumber="495" value="2015-07-16" valueMax="2015-07-21" valueMin="2015-07-16">
<collectingDate id="CE6AE9FE0F1CFFD5FF2D6B5CFE6EF9F9" box="[246,428,1616,1643]" pageId="12" pageNumber="495" value="2015-07-16" valueMax="2015-07-21" valueMin="2015-07-16">16‒21.VII.2015</collectingDate>
</date>
, leg.
<collectorName id="076553000F1CFFD5FE3F6B5DFDFDF9F9" box="[484,575,1617,1643]" pageId="12" pageNumber="495">KJ Teng</collectorName>
&amp;
<collectorName id="076553000F1CFFD5FD846B5DFD63F9F8" box="[607,673,1616,1643]" pageId="12" pageNumber="495">X Bai</collectorName>
, slide Nos. LHY21546♁, LHY21547♁, LHY21548♁, LHY21647
<specimenCount id="BC96FD5F0F1CFFD5FA516B5CFA55F9F8" box="[1418,1431,1616,1642]" pageId="12" pageNumber="495" type="female">♀</specimenCount>
</materialsCitation>
, LHY21686
<specimenCount id="BC96FD5F0F1CFFD5FEFA6B78FEECF91C" box="[289,302,1652,1678]" pageId="12" pageNumber="495" type="female">♀</specimenCount>
, LHY21735♁, LHY21736
<specimenCount id="BC96FD5F0F1CFFD5FDBC6B78FDB6F91C" box="[615,628,1652,1678]" pageId="12" pageNumber="495" type="female">♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="E28A655D0F1CFFD5FF1C6B94FC2DF968" pageId="12" pageNumber="495" type="diagnosis">
<paragraph id="AA2F36D60F1CFFD5FF1C6B94FC2DF968" blockId="12.[151,1437,1544,2039]" pageId="12" pageNumber="495">
<emphasis id="98E4EAC40F1CFFD5FF1C6B94FE80F920" bold="true" box="[199,322,1688,1714]" pageId="12" pageNumber="495">Diagnosis.</emphasis>
<taxonomicName id="6D904D550F1CFFD5FE9C6B94FDACF920" authorityName="Liu" authorityYear="1980" box="[327,622,1688,1714]" class="Insecta" family="Yponomeutidae" genus="Thecobathra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="495" phylum="Arthropoda" rank="species" species="microsignata">
<emphasis id="98E4EAC40F1CFFD5FE9C6B94FDACF920" box="[327,622,1688,1714]" italics="true" pageId="12" pageNumber="495">Thecobathra microsignata</emphasis>
</taxonomicName>
is characterized by the subtriangular sacculus with sclerotized band along dorsal and ventral margins, and the cornuti with spines along inner margins. It is similar to
<taxonomicName id="6D904D550F1CFFD5FB546BB1FAC4F944" authority="Liu &amp; Wang, 2023" authorityName="Liu &amp; Wang" authorityYear="2023" box="[1167,1286,1724,1750]" class="Insecta" family="Yponomeutidae" genus="Thecobathra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="495" phylum="Arthropoda" rank="species" species="setulosa" status="sp. nov.">
<emphasis id="98E4EAC40F1CFFD5FB546BB1FAC4F944" box="[1167,1286,1724,1750]" italics="true" pageId="12" pageNumber="495">T. setulosa</emphasis>
</taxonomicName>
<emphasis id="98E4EAC40F1CFFD5FAD66BB1FAA4F945" bold="true" box="[1293,1382,1725,1751]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="83D757BF0F1CFFD5FAD66BB1FAA4F945" box="[1293,1382,1725,1751]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and the differences between them are stated in the diagnosis of the latter species.
</paragraph>
</subSubSection>
<subSubSection id="E28A655D0F1CFFD4FF1C6A08FD66FED3" lastPageId="13" lastPageNumber="496" pageId="12" pageNumber="495" type="description">
<paragraph id="AA2F36D60F1CFFD5FF1C6A08FD0FF88D" blockId="12.[151,1437,1544,2039]" box="[199,717,1796,1823]" pageId="12" pageNumber="495">
<emphasis id="98E4EAC40F1CFFD5FF1C6A08FE9BF88C" bold="true" box="[199,345,1796,1822]" pageId="12" pageNumber="495">Description.</emphasis>
Adult. Wingspan
<quantity id="6D689B330F1CFFD5FDF86A08FD0BF88D" box="[547,713,1796,1823]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.55" metricValueMin="1.45" pageId="12" pageNumber="495" unit="mm" value="15.0" valueMax="15.5" valueMin="14.5">14.5‒15.5 mm</quantity>
.
</paragraph>
<paragraph id="AA2F36D60F1CFFD4FF1C6A24FD66FED3" blockId="12.[151,1437,1544,2039]" lastBlockId="13.[151,1436,151,393]" lastPageId="13" lastPageNumber="496" pageId="12" pageNumber="495">
<emphasis id="98E4EAC40F1CFFD5FF1C6A24FEACF8D0" box="[199,366,1832,1858]" italics="true" pageId="12" pageNumber="495">Male genitalia</emphasis>
(
<figureCitation id="32AB2A530F1CFFD5FE5B6A24FE16F8D0" box="[384,468,1832,1858]" captionStart="FIGURES 15–18" captionStartId="11.[151,264,1803,1827]" captionTargetBox="[157,1420,194,1766]" captionTargetId="figure-18@11.[155,1433,181,1779]" captionTargetPageId="11" captionText="FIGURES 15–18. Male genitalia of Thecobathra spp. 15, T. meniscata sp. nov., holotype, slide No. LHY21528; 16, T. microsignata, slide No. LHY21734; 17, T. minutiprocessa sp. nov., holotype, slide No. LHY21667; 18, T. odontoidea sp. nov., holotype, slide No. LHY21068. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8243632" httpUri="https://zenodo.org/record/8243632/files/figure.png" pageId="12" pageNumber="495">Fig. 16</figureCitation>
). Uncus mound-shaped, bifid. Socius narrow basally, inflated medially, narrowed to apex distally, with two thorns apically. Subscaphium narrow, parallel-sided. Ventral plate of gnathos long, narrowed anteriorly, concave in V shape posteriorly, densely setose. Valva narrower and subparallel-sided basally, widened at basal 1/4, thereafter narrowed to rounded apex; dorsoproximal process gradually narrowed, with a long, slender spine curved backward at base, nearly straight distally; inner process narrow, elongate triangular. Sacculus subtriangular, truncate obliquely inward apically; sclerotized, narrow band along dorsal and ventral margins; dorsal band arched, rectangularly produced and free distally; ventral band about 1/2 length of dorsal band. Saccus with posterior 1/4 V-shaped, anterior 3/4 narrow and subparallel sided to rounded apex. Aedeagus straight, 1.5 times length of saccus, with a row of teeth along distal half of one side, densely dentate from 1/2 to 5/6 along the other side; cornuti consisting of two horns, with narrow bands extending from its base to middle length of aedeagus, with several spines along inner margin, apical spine largest.
</paragraph>
</subSubSection>
<subSubSection id="E28A655D0F1DFFD4FF1C6C46FD05FE1B" pageId="13" pageNumber="496" type="distribution">
<paragraph id="AA2F36D60F1DFFD4FF1C6C46FDE3FEF7" blockId="13.[151,1436,151,393]" box="[199,545,330,357]" pageId="13" pageNumber="496">
<emphasis id="98E4EAC40F1DFFD4FF1C6C46FEA3FEF6" bold="true" box="[199,353,330,356]" pageId="13" pageNumber="496">Distribution.</emphasis>
<collectingCountry id="D28776460F1DFFD4FEBC6C47FE68FEF7" box="[359,426,331,357]" name="China" pageId="13" pageNumber="496">China</collectingCountry>
(
<collectingRegion id="6854F8340F1DFFD4FE626C47FDD7FEF7" box="[441,533,331,357]" country="China" name="Yunnan" pageId="13" pageNumber="496">Yunnan</collectingRegion>
).
</paragraph>
<paragraph id="AA2F36D60F1DFFD4FF1C6C63FD05FE1B" blockId="13.[151,1436,151,393]" box="[199,711,367,393]" pageId="13" pageNumber="496">
<emphasis id="98E4EAC40F1DFFD4FF1C6C63FEC4FE1B" bold="true" box="[199,262,367,393]" pageId="13" pageNumber="496">Note.</emphasis>
The male is described for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>
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