244 lines
28 KiB
XML
244 lines
28 KiB
XML
<document id="5880E4817DF1B290BE558039BFCAA052" ID-DOI="10.5281/zenodo.212975" ID-GBIF-Dataset="d10859be-023f-4fca-a164-6836331e5fa6" ID-ISSN="1175-5326" ID-Zenodo-Dep="212975" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460504029448" checkinUser="plazi" docAuthor="Schoville, Sean D." docDate="2012" docId="222787CCFFB3FF96838354DA8A9A36CB" docLanguage="en" docName="zt03412p052.pdf" docOrigin="Zootaxa 3412" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Grylloblatta marmoreus Schoville, 2012, new species" docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="DE1EFFB4FFB4FF9E831452078F063171" masterDocTitle="Three new species of Grylloblatta Walker (Insecta: Grylloblattodea: Grylloblattidae), from southern Oregon and northern California" masterLastPageNumber="52" masterPageNumber="42" pageNumber="49" updateTime="1698312492540" updateUser="plazi">
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<mods:titleInfo id="61E0FEB3F59DD2AD366EC2554D24BF18">
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<mods:title id="4CB940A4EF214644AA8C2A9C8AEE4F7A">Three new species of Grylloblatta Walker (Insecta: Grylloblattodea: Grylloblattidae), from southern Oregon and northern California</mods:title>
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</mods:titleInfo>
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<mods:name id="7C33D834D73A2048EF6FA5A47C6BD125" type="personal">
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<mods:role id="BBA8911A2879E33D7AC5D7AEB13C50EA">
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<mods:roleTerm id="7875333B70AA96365CD59BEC87AAA446">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8CC53010ABE2347512C1072C41B89C0D">Schoville, Sean D.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C05322BB298FDA582F3F93D62CD5C675">text</mods:typeOfResource>
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<mods:title id="46C83229176FFB654FB8DACF4F8E2D3A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C07C2CCD1927ED4B13664FDE93F4C544">
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<mods:date id="36CBC9A7E84C083ECF1ADCC5DAFA1260">2012</mods:date>
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<mods:detail id="240F049338FC803CEDEE33589AFC285E" type="volume">
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<mods:number id="384A029F9527978B879B55FFB63EF4A6">3412</mods:number>
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<mods:start id="120F45B122F148B48AB5D3011472902C">42</mods:start>
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<mods:end id="0F409CA8F05E354D1F4400E19F436DC2">52</mods:end>
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<mods:classification id="39CB112BD35B92C98987D993E3EC3A6D">journal article</mods:classification>
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<mods:identifier id="11971DA3C34DCC62D8EBF96A6FE9A637" type="DOI">10.5281/zenodo.212975</mods:identifier>
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<mods:identifier id="AD4740DA0D2849E0601D14F1E67A8A1F" type="GBIF-Dataset">d10859be-023f-4fca-a164-6836331e5fa6</mods:identifier>
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<mods:identifier id="E88ACDB08F9164A483362578A8AD4FDF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B4E1269C3FF66CCD4B4E1AEC8851E9AA" type="Zenodo-Dep">212975</mods:identifier>
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</mods:mods>
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<treatment id="222787CCFFB3FF96838354DA8A9A36CB" ID-DOI="http://doi.org/10.5281/zenodo.5673771" ID-GBIF-Taxon="119522302" ID-Zenodo-Dep="5673771" LSID="urn:lsid:plazi:treatment:222787CCFFB3FF96838354DA8A9A36CB" httpUri="http://treatment.plazi.org/id/222787CCFFB3FF96838354DA8A9A36CB" lastPageId="8" lastPageNumber="50" pageId="7" pageNumber="49">
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<subSubSection id="E2946551FFB3FF99838354DA8E5E3669" pageId="7" pageNumber="49" type="nomenclature">
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<paragraph id="AA3136DAFFB3FF99838354DA8D563786" blockId="7.[151,592,1757,1816]" box="[151,592,1757,1783]" pageId="7" pageNumber="49">
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<heading id="F17981B6FFB3FF99838354DA8D563786" bold="true" box="[151,592,1757,1783]" fontSize="11" level="1" pageId="7" pageNumber="49" reason="1">
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<emphasis id="98FAEAC8FFB3FF99838354DA8D563786" bold="true" box="[151,592,1757,1783]" pageId="7" pageNumber="49">
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<taxonomicName id="6D8E4D59FFB3FF99838354DA8EBF3786" ID-CoL="3HG2M" box="[151,441,1757,1783]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="7" pageNumber="49" phylum="Arthropoda" rank="species" species="marmoreus" status="sp. nov.">
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<emphasis id="98FAEAC8FFB3FF99838354DA8EBF3786" bold="true" box="[151,441,1757,1783]" italics="true" pageId="7" pageNumber="49">Grylloblatta marmoreus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="83C957B3FFB3FF9982D454DA8D563786" box="[448,592,1757,1783]" pageId="7" pageNumber="49" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="AA3136DAFFB3FF99838355078E5E3669" blockId="7.[151,592,1757,1816]" box="[151,344,1792,1816]" pageId="7" pageNumber="49">
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(
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<figureCitation id="32B52A5FFFB3FF99838B55078FEF3669" box="[159,233,1792,1816]" captionStart="FIGURE 1" captionStartId="2.[151,250,1409,1432]" captionTargetBox="[182,1410,627,1381]" captionTargetId="figure@2.[176,1411,625,1388]" captionTargetPageId="2" captionText="FIGURE 1. Differences in size and coloration observed in type specimens. A) Grylloblatta oregonensis n. sp. B) G. siskiyouensis n. sp. C) G. m a r m o re u s n. sp." httpUri="https://zenodo.org/record/212976/files/figure.png" pageId="7" pageNumber="49">Figs. 1</figureCitation>
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C, 2E–H)
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</paragraph>
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</subSubSection>
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<subSubSection id="E2946551FFB3FF968383554E8A18306E" lastPageId="8" lastPageNumber="50" pageId="7" pageNumber="49" type="materials_examined">
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<paragraph id="AA3136DAFFB3FF998383554E8EA63682" blockId="7.[151,1437,1864,2036]" pageId="7" pageNumber="49">
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<emphasis id="98FAEAC8FFB3FF998383554E8E403610" bold="true" box="[151,326,1864,1889]" pageId="7" pageNumber="49">
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<typeStatus id="75358878FFB3FF998383554E8FD43610" box="[151,210,1865,1889]" pageId="7" pageNumber="49">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="75358878FFB3FF99825A554F8EB03611" box="[334,438,1864,1888]" pageId="7" pageNumber="49" type="holotype">Holotype</typeStatus>
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male in ethanol, deposited in
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<collectionCode id="CC9FAE1FFFB3FF99801B554F8C4E3611" LSID="urn:lsid:biocol.org:col:34700" box="[783,840,1864,1888]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="7" pageNumber="49">CAS</collectionCode>
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, labeled: “Big Foot Cave, Marble Mountain Wilderness Area, Siskiyou Co., CA, G.
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<collectingCountry id="D299764AFFB3FF9982EB556B8D1336F5" box="[511,533,1900,1924]" name="Iceland" pageId="7" pageNumber="49">O</collectingCountry>
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. Graening & Guy Graening,
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<date id="DE30101AFFB3FF99804C556B8CE136F5" box="[856,999,1900,1924]" pageId="7" pageNumber="49" value="2010-09-03">3 Sept. 2010</date>
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” [white label]/ “
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<typeStatus id="75358878FFB3FF99878A556B8A3236F5" box="[1182,1332,1900,1924]" pageId="7" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="6D8E4D59FFB3FF998628556A8EC836D9" authority="Schoville" authorityName="Schoville" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="7" pageNumber="49" phylum="Arthropoda" rank="species" species="marmoreus">
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<emphasis id="98FAEAC8FFB3FF998628556A8E5036D8" italics="true" pageId="7" pageNumber="49">Grylloblatta marmoreus</emphasis>
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Schoville
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</taxonomicName>
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det. S.D. Schoville 2011” [red label]; Big Foot Cave, located in Marble Valley, approximately
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<geoCoordinate id="CFBA501DFFB3FF99825555B08EEA36BE" box="[321,492,1972,1999]" direction="north" orientation="latitude" pageId="7" pageNumber="49" precision="5" value="41.5628">41.5628o North</geoCoordinate>
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and -
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<geoCoordinate id="CFBA501DFFB3FF99813855B08DE736BE" box="[556,737,1972,1999]" direction="west" orientation="longitude" pageId="7" pageNumber="49" precision="5" value="-123.1899">123.1899o West</geoCoordinate>
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, elevation
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<quantity id="6D769B3FFFB3FF99804F55B08CB036BE" box="[859,950,1975,1999]" metricMagnitude="3" metricUnit="m" metricValue="1.808988" pageId="7" pageNumber="49" unit="ft" value="5935.0">5,935 ft</quantity>
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, Marble Mountain Wilderness Area, Siskiyou County, California.
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</paragraph>
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<paragraph id="AA3136DAFFBCFF9683D352908A18306E" blockId="8.[151,1437,151,1979]" pageId="8" pageNumber="50">
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<emphasis id="98FAEAC8FFBCFF9683D352908E9131C1" bold="true" box="[199,407,151,176]" pageId="8" pageNumber="50">Other specimens.</emphasis>
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Juvenile in ethanol, deposited in
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<collectionCode id="CC9FAE1FFFBCFF96800052908C4B31DE" LSID="urn:lsid:biocol.org:col:34700" box="[788,845,151,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="8" pageNumber="50">CAS</collectionCode>
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, labeled: “Planetary Dairy Cave, G.
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<collectingCountry id="D299764AFFBCFF9687F652908BF131DE" box="[1250,1271,151,175]" name="Iceland" pageId="8" pageNumber="50">O</collectingCountry>
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. Graening and David Weaver,
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<date id="DE30101AFFBCFF96825E52BB8ED231A5" box="[330,468,188,212]" pageId="8" pageNumber="50" value="2010-07-04">4 July 2010</date>
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” [white label]/ “
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<taxonomicName id="6D8E4D59FFBCFF96818252BB8B1A31A5" authority="Schoville" authorityName="Schoville" box="[662,1052,188,212]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="marmoreus">
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<emphasis id="98FAEAC8FFBCFF96818252BB8CAF31A5" box="[662,937,188,212]" italics="true" pageId="8" pageNumber="50">Grylloblatta marmoreus</emphasis>
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Schoville
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</taxonomicName>
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det. S.D. Schoville 2011” [white label]; Planetary Dairy Cave, approximately
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<geoCoordinate id="CFBA501DFFBCFF96818152E58C35318B" box="[661,819,223,250]" direction="north" orientation="latitude" pageId="8" pageNumber="50" precision="55" value="41.56">41.560o North</geoCoordinate>
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and -
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<geoCoordinate id="CFBA501DFFBCFF96806052E58B1D318B" box="[884,1051,223,250]" direction="west" orientation="longitude" pageId="8" pageNumber="50" precision="55" value="-123.197">123.197o West</geoCoordinate>
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, elevation
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<quantity id="6D769B3FFFBCFF96878352E58BEF318B" box="[1175,1257,226,250]" metricMagnitude="3" metricUnit="m" metricValue="1.7132808" pageId="8" pageNumber="50" unit="ft" value="5621.0">5,621ft</quantity>
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, Marble Mountain Wilderness Area, Siskiyou County, California near the headwaters of the Canyon Creek watershed.
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</paragraph>
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</subSubSection>
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<subSubSection id="E2946551FFBCFF9683D3532C8DF33016" pageId="8" pageNumber="50" type="etymology">
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<paragraph id="AA3136DAFFBCFF9683D3532C8DF33016" blockId="8.[151,1437,151,1979]" pageId="8" pageNumber="50">
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<emphasis id="98FAEAC8FFBCFF9683D3532C8E483032" bold="true" box="[199,334,299,323]" pageId="8" pageNumber="50">Etymology.</emphasis>
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The specific epithet,
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<taxonomicName id="6D8E4D59FFBCFF968155532B8DB93032" box="[577,703,300,323]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="marmoreus">
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<emphasis id="98FAEAC8FFBCFF968155532B8DB93032" box="[577,703,300,323]" italics="true" pageId="8" pageNumber="50">marmoreus</emphasis>
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</taxonomicName>
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, means “of marble” in Latin and is named for the Marble Mountains of California where this specimen was collected.
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</paragraph>
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</subSubSection>
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<subSubSection id="E2946551FFBCFF9683D353758B50334E" pageId="8" pageNumber="50" type="diagnosis">
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<paragraph id="AA3136DAFFBCFF9683D353758B50334E" blockId="8.[151,1437,151,1979]" pageId="8" pageNumber="50">
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<emphasis id="98FAEAC8FFBCFF9683D353758E4730FA" bold="true" box="[199,321,370,395]" pageId="8" pageNumber="50">Diagnosis.</emphasis>
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Adult males of this species can be distinguished from those of all other Nearctic
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<taxonomicName id="6D8E4D59FFBCFF9687CF53748A6030FA" box="[1243,1382,371,395]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="98FAEAC8FFBCFF9687CF53748A6030FA" box="[1243,1382,371,395]" italics="true" pageId="8" pageNumber="50">Grylloblatta</emphasis>
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</taxonomicName>
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species by the following combination of character states: color buff yellow; small body size for genus; relatively short limbs; fewer than 30 antennomeres; ratio of antennomere lengths 2 and 1 1.5; head wider than pronotum; compound eyes well-developed and medium size; pronotum trapezoidal with rounded proximal end and forming soft triangular point at posterior end; subgenital plate with slight rounded tooth; right coxopodite with sharply curved process; left coxopodite with short, stubby stylus; cercus short with short cercomeres.
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</paragraph>
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</subSubSection>
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<subSubSection id="E2946551FFBCFF9683D3504D8E65359B" pageId="8" pageNumber="50" type="description">
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<paragraph id="AA3136DAFFBCFF9683D3504D8B9A3546" blockId="8.[151,1437,151,1979]" pageId="8" pageNumber="50">
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<emphasis id="98FAEAC8FFBCFF9683D3504D8D213312" bold="true" box="[199,551,586,611]" pageId="8" pageNumber="50">
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Description of
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<typeStatus id="75358878FFBCFF96826C504C8ED83312" box="[376,478,587,611]" pageId="8" pageNumber="50" type="holotype">holotype</typeStatus>
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male.
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</emphasis>
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Antennomere count on right antennae 27 and broken left antennae 25. Antennomere 2 1.5X larger than antennomere 1. Compound eyes well developed, length greater along body axis than width, and pigmentation black. Head slightly broader than pronotum but narrower than interorbital distance. Pronotum gently curved in anterior end and triangular at posterior end, overall trapezoidal as width at anterior end greater than posterior end, posterior end forming soft triangular point, transverse sulcus at anterior end curving slightly towards anterior end, length greater than width. Leg coloration amber yellow, slightly lighter than dorsal body color, and small brown setae present. Legs relatively short. Portion of right mid leg below femur removed and destroyed during DNA extraction. Cercus length short with short cercomeres. Subgenital plate more than twice as wide as long, distal edge hardened, distal end forming very slight rounded tooth. Right coxopodite longer than wide with a strong curve towards the ventral side of the body, a short process strongly curved towards anterior on dorsal side near base, stylus borne directly on distal end approximately 30% of the length of coxopodite, and heavily pubescent. Left coxopodite width greater than length, stylus at distal end short and stubby (
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<quantity id="6D769B3FFFBCFF9687C151D08A3B329E" box="[1237,1341,983,1007]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="50" unit="cm" value="0.021">0.021cm</quantity>
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). Dorsal and ventral body color buff yellow (87), appendages lighter amber yellow (83), antennomeres and cercomeres amber yellow fading to white at distal ends, subgenital plate and coxopodites amber yellow.
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</paragraph>
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<paragraph id="AA3136DAFFBCFF9683D356448E65359B" blockId="8.[151,1437,151,1979]" pageId="8" pageNumber="50">
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<emphasis id="98FAEAC8FFBCFF9683D356448D93352A" bold="true" box="[199,661,1090,1115]" pageId="8" pageNumber="50">
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Measurements of
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<typeStatus id="75358878FFBCFF9682B756448D0C352A" box="[419,522,1091,1115]" pageId="8" pageNumber="50" type="holotype">holotype</typeStatus>
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male (cm).
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</emphasis>
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BL 1.256, HL 0.228, HW 0.255, EL 0.054, EW 0.034, ID 0.208, Ant1L 0.02, Ant2L 0.03, AntRatio 1.5, PL 0.235, PW 0.214, FL front 0.295, FL mid 0.268, FL rear 0.3886, TL front 0.235, TL mid 0.275, TL rear 0.402, TrL front 0.067, TrL mid 0.1005, TrL rear 0.154, PTrL front 0.201, PTrL mid 0.201, PTrL rear 0.214, SbgL 0.0536, SbgW 0.1273, RCxL 0.1005, RCxW 0.0804, LCxL 0.1139, LCxW 0.134, CL 0.4104.
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</paragraph>
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</subSubSection>
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<subSubSection id="E2946551FFBCFF9683D356F18A9A36CB" pageId="8" pageNumber="50" type="materials_examined">
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<paragraph id="AA3136DAFFBCFF9683D356F18D053443" blockId="8.[151,1437,151,1979]" pageId="8" pageNumber="50">
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<emphasis id="98FAEAC8FFBCFF9683D356F18D45347E" bold="true" box="[199,579,1270,1295]" pageId="8" pageNumber="50">Description of other specimens.</emphasis>
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Juvenile specimen is amber yellow (83) on the head and pronotum, with a white abdomen and appendages.
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</paragraph>
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<paragraph id="AA3136DAFFBCFF9683D357398E3C37EB" blockId="8.[151,1437,151,1979]" pageId="8" pageNumber="50">
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<emphasis id="98FAEAC8FFBCFF9683D357398E6A3426" bold="true" box="[199,364,1342,1367]" pageId="8" pageNumber="50">Comparisons.</emphasis>
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<taxonomicName id="6D8E4D59FFBCFF96826D57388D883426" box="[377,654,1343,1367]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="marmoreus" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFBCFF96826D57388D883426" box="[377,654,1343,1367]" italics="true" pageId="8" pageNumber="50">Grylloblatta marmoreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFBCFF96818E57388DD83426" box="[666,734,1343,1367]" pageId="8" pageNumber="50" rank="species">
|
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<emphasis id="98FAEAC8FFBCFF96818E57388DD83426" bold="true" box="[666,734,1343,1367]" pageId="8" pageNumber="50">n. sp.</emphasis>
|
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</taxonomicNameLabel>
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can be distinguished from
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<taxonomicName id="6D8E4D59FFBCFF96873357478BD73426" box="[1063,1233,1344,1367]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="oregonensis" status="sp. nov.">
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<emphasis id="98FAEAC8FFBCFF96873357478BD73426" box="[1063,1233,1344,1367]" italics="true" pageId="8" pageNumber="50">G. oregonensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="83C957B3FFBCFF9687C957388A273426" box="[1245,1313,1343,1367]" pageId="8" pageNumber="50" rank="species">
|
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<emphasis id="98FAEAC8FFBCFF9687C957388A273426" bold="true" box="[1245,1313,1343,1367]" pageId="8" pageNumber="50">n. sp.</emphasis>
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</taxonomicNameLabel>
|
||
in having smaller body size, pronotum narrowing at posterior end and forming soft triangular point, and smaller antennomere 2:1 ratio. It can be distinguished from
|
||
<taxonomicName id="6D8E4D59FFBCFF968150578F8DF234EE" box="[580,756,1415,1439]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="siskiyouensis" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFBCFF968150578F8DF234EE" box="[580,756,1415,1439]" italics="true" pageId="8" pageNumber="50">G. siskiyouensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFBCFF9681E857808C3D34EE" box="[764,827,1415,1439]" pageId="8" pageNumber="50" rank="species">
|
||
<emphasis id="98FAEAC8FFBCFF9681E857808C3D34EE" bold="true" box="[764,827,1415,1439]" pageId="8" pageNumber="50">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in having fewer antennomeres, buff yellow coloration and weaker point at posterior end of pronotum. It can be distinguished from
|
||
<emphasis id="98FAEAC8FFBCFF96871757AC8B5334B2" box="[1027,1109,1451,1475]" italics="true" pageId="8" pageNumber="50">G. ro t h i</emphasis>
|
||
in having a subgenital plate without a lobelike projection on distal end and right coxopodite with a strongly down-curved process. It differs from
|
||
<taxonomicName id="6D8E4D59FFBCFF9683C257F48E67377A" box="[214,353,1523,1547]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="chandleri">
|
||
<emphasis id="98FAEAC8FFBCFF9683C257F48E67377A" box="[214,353,1523,1547]" italics="true" pageId="8" pageNumber="50">G. chandleri</emphasis>
|
||
</taxonomicName>
|
||
in having a subgenital plate with a more rounded tooth at distal end and right coxopodite with strongly down-curved process. It can be distinguished from
|
||
<taxonomicName id="6D8E4D59FFBCFF968042541F8CD6375E" box="[854,976,1559,1583]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="sculleni">
|
||
<emphasis id="98FAEAC8FFBCFF968042541F8CD6375E" box="[854,976,1559,1583]" italics="true" pageId="8" pageNumber="50">G. sculleni</emphasis>
|
||
</taxonomicName>
|
||
in having smaller body size and fewer antennomeres; from
|
||
<taxonomicName id="6D8E4D59FFBCFF96826B543C8EF53722" box="[383,499,1595,1619]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="gurneyi">
|
||
<emphasis id="98FAEAC8FFBCFF96826B543C8EF53722" box="[383,499,1595,1619]" italics="true" pageId="8" pageNumber="50">G. gurneyi</emphasis>
|
||
</taxonomicName>
|
||
in having a subgenital plate with a single rounded tooth at distal end and right coxopodite with strongly down-curved process; and from
|
||
<emphasis id="98FAEAC8FFBCFF96800454678C853706" box="[784,899,1631,1655]" italics="true" pageId="8" pageNumber="50">G. b a r b e r i</emphasis>
|
||
in smaller body size and having less than 30 antennomeres.
|
||
</paragraph>
|
||
<paragraph id="AA3136DAFFBCFF9683D354A18B013793" blockId="8.[151,1437,151,1979]" pageId="8" pageNumber="50">
|
||
<emphasis id="98FAEAC8FFBCFF9683D354A18EB937CE" bold="true" box="[199,447,1702,1727]" pageId="8" pageNumber="50">Habitat distribution.</emphasis>
|
||
The
|
||
<typeStatus id="75358878FFBCFF9682EE54AF8D2C37B1" box="[506,554,1704,1728]" pageId="8" pageNumber="50">type</typeStatus>
|
||
specimen was found in the twilight zone of a cave habitat, on a rock near ice. The juvenile was found on a rock at the bottom of a pit near the cave entrance.
|
||
</paragraph>
|
||
<paragraph id="AA3136DAFFBCFF9683D354E98DB3365B" blockId="8.[151,1437,151,1979]" pageId="8" pageNumber="50">
|
||
<emphasis id="98FAEAC8FFBCFF9683D354E98D003676" bold="true" box="[199,518,1774,1799]" pageId="8" pageNumber="50">Geographical distribution.</emphasis>
|
||
<taxonomicName id="6D8E4D59FFBCFF96811B54E88C273676" box="[527,801,1775,1799]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="marmoreus" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFBCFF96811B54E88C273676" box="[527,801,1775,1799]" italics="true" pageId="8" pageNumber="50">Grylloblatta marmoreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFBCFF96803E54E88C6D3676" box="[810,875,1775,1799]" pageId="8" pageNumber="50" rank="species">
|
||
<emphasis id="98FAEAC8FFBCFF96803E54E88C6D3676" bold="true" box="[810,875,1775,1799]" pageId="8" pageNumber="50">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is known from multiple cave sites in the Marble Valley of Marble Mountains, California (
|
||
<figureCitation id="32B52A5FFFBCFF96817755158DA1365B" box="[611,679,1810,1834]" captionStart="FIGURE 3" captionStartId="4.[151,250,1659,1682]" captionTargetBox="[177,1435,193,1608]" captionTargetId="figure@4.[151,1436,193,1638]" captionTargetPageId="4" captionText="FIGURE 3. Geographical distribution of grylloblattid populations (dots) and type localities (crosses and asterisks) in California and Oregon, USA." httpUri="https://zenodo.org/record/212978/files/figure.png" pageId="8" pageNumber="50">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="AA3136DAFFBCFF9683D355318A9A36CB" blockId="8.[151,1437,151,1979]" pageId="8" pageNumber="50">
|
||
<emphasis id="98FAEAC8FFBCFF9683D355318D01363E" bold="true" box="[199,519,1846,1871]" pageId="8" pageNumber="50">Phylogenetic relationships.</emphasis>
|
||
Based on mitochondrial cytochrome oxidase subunit II data,
|
||
<taxonomicName id="6D8E4D59FFBCFF9687AC553F8A53363E" box="[1208,1365,1848,1871]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="marmoreus" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFBCFF9687AC553F8A53363E" box="[1208,1365,1848,1871]" italics="true" pageId="8" pageNumber="50">G. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFBCFF96864955308A9A363E" box="[1373,1436,1847,1871]" pageId="8" pageNumber="50" rank="species">
|
||
<emphasis id="98FAEAC8FFBCFF96864955308A93363E" bold="true" box="[1373,1429,1847,1871]" pageId="8" pageNumber="50">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(GenBank Accession
|
||
<accessionNumber id="B5DDAB39FFBCFF96829F555D8EF83603" box="[395,510,1882,1906]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN612963" pageId="8" pageNumber="50">JN612963</accessionNumber>
|
||
&
|
||
<accessionNumber id="B5DDAB39FFBCFF968136555D8D913603" box="[546,663,1882,1906]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN612964" pageId="8" pageNumber="50">JN612964</accessionNumber>
|
||
) is distinct from (minimum ~10%) other nearby grylloblattid species (
|
||
<tableCitation id="E70C0361FFBCFF96838A55788FF436E6" box="[158,242,1919,1943]" captionStart="TABLE 1" captionStartId="5.[151,235,1016,1037]" captionTargetBox="[151,1436,1126,1580]" captionTargetPageId="5" captionText="TABLE 1. Genetic divergence of Grylloblatta siskiyouensis, G. oregonensis, and G. m a r m o re u s from nearby grylloblattid populations. Values are Kimura two-parameter corrected pairwise genetic distance at the cytochrome oxidase II gene. Intraspecific genetic variability, if available, is shown for each species." httpUri="http://table.plazi.org/id/FEF16652FFB1FF9B838351FF8C63353A" pageId="8" pageNumber="50" tableUuid="FEF16652FFB1FF9B838351FF8C63353A">Table 1</tableCitation>
|
||
). The adult
|
||
<typeStatus id="75358878FFBCFF96826955878EAB36E9" box="[381,429,1920,1944]" pageId="8" pageNumber="50">type</typeStatus>
|
||
specimen and the juvenile from Planetary Dairy Cave differ by 0.4%. In a neighbor-joining analysis, it is grouped with
|
||
<taxonomicName id="6D8E4D59FFBCFF9682E055A48D9F36CA" box="[500,665,1955,1979]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="oregonensis" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFBCFF9682E055A48D9F36CA" box="[500,665,1955,1979]" italics="true" pageId="8" pageNumber="50">G. oregonensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFBCFF96818B55A48DD836CB" box="[671,734,1955,1979]" pageId="8" pageNumber="50" rank="species">
|
||
<emphasis id="98FAEAC8FFBCFF96818B55A48DD036CA" bold="true" box="[671,726,1955,1979]" pageId="8" pageNumber="50">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="6D8E4D59FFBCFF96800055A48CC236CA" box="[788,964,1955,1979]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="8" pageNumber="50" phylum="Arthropoda" rank="species" species="siskiyouensis" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFBCFF96800055A48CC236CA" box="[788,964,1955,1979]" italics="true" pageId="8" pageNumber="50">G. siskiyouensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFBCFF9680DF55A48B0F36CB" box="[971,1033,1955,1979]" pageId="8" pageNumber="50" rank="species">
|
||
<emphasis id="98FAEAC8FFBCFF9680DF55A48B0436CA" bold="true" box="[971,1026,1955,1979]" pageId="8" pageNumber="50">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
in an unresolved polytomy (
|
||
<figureCitation id="32B52A5FFFBCFF96865E55A58A8936CB" box="[1354,1423,1954,1978]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1674]" captionTargetBox="[242,1290,187,1620]" captionTargetId="figure@7.[230,1359,167,1634]" captionTargetPageId="7" captionText="FIGURE 4. Relationships of new Grylloblatta species based on mitochondrial cytochrome oxidase subunit II gene data." httpUri="https://zenodo.org/record/212979/files/figure.png" pageId="8" pageNumber="50">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |