treatments-xml/data/22/27/87/222787B2FFEAFF83FD0C839B4658FB5D.xml

<document id="3D589B58E9E3CBFF682459A71B2CA023" ID-CLB-Dataset="22220" ID-DOI="10.5852/ejt.2017.262" ID-GBIF-Dataset="bc93a100-df15-48bf-9453-34aafe93d9f0" ID-ISSN="2118-9773" ID-Zenodo-Dep="3823118" ID-ZooBank="10676882-F6AC-4BC8-A0F1-9C0882EA36C2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589314723106" checkinUser="carolina" docAuthor="Pavlek, Martina &amp; Ribera, Carles" docDate="2017" docId="222787B2FFEAFF83FD0C839B4658FB5D" docLanguage="en" docName="ejt-2017-262.pdf.imf" docOrigin="European Journal of Taxonomy 262" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Kryptonesticus Pavlek &amp; Ribera 2017, gen. nov." docType="treatment" docUuid="19EABFE7-08F7-4AB8-860B-0AB2E59EBC03" docUuidSource="ZooBank" docVersion="8" lastPageNumber="6" masterDocId="DE1EFFCAFFEEFF86FF8B825A4563FF9B" masterDocTitle="Kryptonesticus deelemanae gen. et sp. nov. (Araneae, Nesticidae), with notes on the Mediterranean cave species" masterLastPageNumber="27" masterPageNumber="1" pageNumber="5" updateTime="1698839289807" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BE65CF10C10C63B72AA9291FF5FCDEB3">Kryptonesticus deelemanae gen. et sp. nov. (Araneae, Nesticidae), with notes on the Mediterranean cave species</mods:title>
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<mods:namePart id="2804842BFDAD95032D323404EAA34D75">Pavlek, Martina</mods:namePart>
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<mods:affiliation id="4732F45BA292701E02B2DFE5C5B08D66">Ruđer Bošković Institute, Bijenička 54, 10000 Zagreb, Croatia; and Croatian Biospeleological Society, Demetrova 1, 10000 Zagreb, Croatia. &amp; urn: lsid: zoobank. org: author: 681 A 5613 - 37 C 0 - 4684 - B 3 B 5 - 8 BBE 6 BEAB 4 BB</mods:affiliation>
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<mods:namePart id="C9C75EA24BC960AE622A327D22CD4876">Ribera, Carles</mods:namePart>
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<mods:affiliation id="74EB11FBE0D14CF5B73858AC99E7F808">Institut de Recerca de la Biodiversitat, Departament de Biologia Animal, Universitat de Barcelona, Av. Diagonal, 643, Barcelona- 08028, Spain. &amp; Email: cribera @ ub. edu &amp; urn: lsid: zoobank. org: author: 509826 D 6 - 1 E 4 D- 4201 - B 08 C- 07 D 0 F 0044 CC 7</mods:affiliation>
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<taxonomicName id="6D8E4D27FFEAFF82FD0C839B4659FE40" ID-CoL="598Z" ID-ENA="1959300" authority="Pavlek &amp; Ribera, 2017" authorityName="Pavlek &amp; Ribera" authorityYear="2017" box="[647,826,449,475]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="98FAEAB6FFEAFF82FD0C839B4659FE40" bold="true" box="[647,826,449,475]" italics="true" pageId="4" pageNumber="5">Kryptonesticus</emphasis>
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<taxonomicNameLabel id="83C957CDFFEAFF82FCC9839846CEFE47" box="[834,941,450,476]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
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<figureCitation id="32B52A21FFEAFF82FD6C805D462EFDB9" box="[743,845,519,546]" captionStart-0="Fig 1" captionStart-1="Fig 2" captionStart-2="Fig 3" captionStart-3="Fig 4" captionStart-4="Fig 5" captionStartId-0="6.[191,232,1965,1991]" captionStartId-1="7.[189,230,1967,1993]" captionStartId-2="8.[189,230,1929,1955]" captionStartId-3="9.[189,230,1930,1956]" captionStartId-4="10.[190,231,1838,1864]" captionTargetBox-0="[191,1399,1531,1929]" captionTargetBox-1="[189,1398,1428,1942]" captionTargetBox-2="[189,1393,447,1881]" captionTargetBox-3="[189,1396,965,1873]" captionTargetBox-4="[190,1398,592,1797]" captionTargetId-0="figure@6.[191,1399,1531,1929]" captionTargetId-1="figure@7.[189,1398,1428,1942]" captionTargetId-2="figure@8.[189,1393,447,1881]" captionTargetId-3="figure@9.[189,1396,965,1873]" captionTargetId-4="figure@10.[190,1398,592,1797]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="Fig 1. Habitus of Kryptonesticus deelemanae sp. nov. A. Male, body length 3.6 mm. B. Female, body length 4.3 mm. (Photos by M. Pavlek.)" captionText-1="Fig 2. Male bulbus. A. Kryptonesticus deelemanae gen. et sp. nov. B. K. fagei (Kratochvíl, 1933). C. K. arenstorffi (Kulczyński, 1914)." captionText-2="Fig 3. Male bulbus. A–C. Krytponesticus deelemanae gen. et sp. nov. D–F. K. fagei (Kratochvíl, 1933). G–I. K. arenstorffi (Kulczyński, 1914). A, D, G. Ventral view. B, E, H. Dorsal view. C, F, I. Apical view. The scale (shown on I) is the same for all photos." captionText-3="Fig 4. Epigyne and vulva. A. Kryptonesticus deelemanae gen. et sp. nov., epigyne. B. K. fagei (Kratochvíl, 1933), epigyne. C. K. arenstorffi (Kulczyński, 1914), epigyne. D. K. deelemanae gen. et sp. nov. vulva. Broken lines mark the positions of the spermathecae." captionText-4="Fig 5. Epigyne and vulva. A–D. Kryptonesticus deelemanae gen. et sp. nov. E–H. K. fagei (Kratochvíl, 1933). I–L. K. arenstorffi (Kulczyński, 1914). A, E, I. Epigyne in ventral view. B, F, J. Epigyne in ventral view after detachment from the opisthosoma. C, G, K. Vulva after treatment with 30% KOH. D, H, L. Epigyne in lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.3823120" figureDoi-1="http://doi.org/10.5281/zenodo.3823122" figureDoi-2="http://doi.org/10.5281/zenodo.3823124" figureDoi-3="http://doi.org/10.5281/zenodo.3823126" figureDoi-4="http://doi.org/10.5281/zenodo.3823128" httpUri-0="https://zenodo.org/record/3823120/files/figure.png" httpUri-1="https://zenodo.org/record/3823122/files/figure.png" httpUri-2="https://zenodo.org/record/3823124/files/figure.png" httpUri-3="https://zenodo.org/record/3823126/files/figure.png" httpUri-4="https://zenodo.org/record/3823128/files/figure.png" pageId="4" pageNumber="5">Figs 1–5</figureCitation>
</paragraph>
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<paragraph id="AA3136A4FFEAFF82FF368013443BFDF8" blockId="4.[189,344,585,611]" box="[189,344,585,611]" pageId="4" pageNumber="5">
<heading id="F17981C8FFEAFF82FF368013443BFDF8" bold="true" box="[189,344,585,611]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="98FAEAB6FFEAFF82FF368013443BFDF8" bold="true" box="[189,344,585,611]" pageId="4" pageNumber="5">Type species</emphasis>
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<taxonomicName id="6D8E4D27FFEAFF82FF3680224763FD0A" ID-CoL="3RF2D" authority="Pavlek &amp; Ribera, 2017" authorityName="Pavlek &amp; Ribera" authorityYear="2017" box="[189,512,631,657]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="deelemanae" status="gen. et sp. nov.">
<emphasis id="98FAEAB6FFEAFF82FF3680224763FD0A" box="[189,512,631,657]" italics="true" pageId="4" pageNumber="5">Kryptonesticus deelemanae</emphasis>
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<taxonomicNameLabel id="83C957CDFFEAFF82FD83802347CBFD0B" box="[520,680,633,657]" pageId="4" pageNumber="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="E294652FFFEAFF82FF3680E34478F9B0" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="AA3136A4FFEAFF82FF3680E34455FD48" blockId="4.[189,310,697,723]" box="[189,310,697,723]" pageId="4" pageNumber="5">
<heading id="F17981C8FFEAFF82FF3680E34455FD48" bold="true" box="[189,310,697,723]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="98FAEAB6FFEAFF82FF3680E34455FD48" bold="true" box="[189,310,697,723]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="AA3136A4FFEAFF82FF3680B24478F9B0" blockId="4.[189,1399,743,1579]" pageId="4" pageNumber="5">
<taxonomicName id="6D8E4D27FFEAFF82FF3680B2440EFC9A" authority="Pavlek &amp; Ribera, 2017" authorityName="Pavlek &amp; Ribera" authorityYear="2017" box="[189,365,744,769]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="98FAEAB6FFEAFF82FF3680B2440EFC9A" box="[189,365,744,769]" italics="true" pageId="4" pageNumber="5">Kryptonesticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEAFF82FEFF80B244BCFC99" box="[372,479,744,770]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
differs from other nesticid genera by the morphology of both male and female copulatory organs. In males, the most conspicuous diagnostic character is the shape and size of the median apophysis (MA), which is large and conspicuous, highly developed and thickly sclerotized, emerging from the middle part of the tegulum and reaching its apical part. This characteristic is shared among a group of species including 
<emphasis id="98FAEAB6FFEAFF82FD89812F4125FC15" box="[514,1094,884,910]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="6D8E4D27FFEAFF82FD89812F47E2FC15" authorityName="Simon" authorityYear="1879" box="[514,641,885,910]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="eremita">K. eremita</taxonomicName>
, 
<taxonomicName id="6D8E4D27FFEAFF82FD04812F467EFC15" authorityName="Lopez-Pancorbo, Kunt &amp; Ribera" authorityYear="2013" box="[655,797,884,910]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dimensis">K. dimensis</taxonomicName>
, 
<taxonomicName id="6D8E4D27FFEAFF82FCA1812F46B9FC15" authorityName="Bosselaers" authorityYear="1998" box="[810,986,884,910]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="henderickxi">K. henderickxi</taxonomicName>
, 
<taxonomicName id="6D8E4D27FFEAFF82FC6C812F4125FC15" authorityName="Kratochvil" authorityYear="1933" box="[999,1094,884,910]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fagei">K. fagei</taxonomicName>
</emphasis>
and 
<taxonomicName id="6D8E4D27FFEAFF82FB09812F407CFC15" authorityName="Kulczynski" authorityYear="1914" box="[1154,1311,884,910]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arenstorffi">
<emphasis id="98FAEAB6FFEAFF82FB09812F407CFC15" box="[1154,1311,884,910]" italics="true" pageId="4" pageNumber="5">K. arenstorffi</emphasis>
</taxonomicName>
(males of 
<taxonomicName id="6D8E4D27FFEAFF82FF5681C24404FC2A" authorityName="Deltshev" authorityYear="1979" box="[221,359,919,945]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="beshkovi">
<emphasis id="98FAEAB6FFEAFF82FF5681C24404FC2A" box="[221,359,919,945]" italics="true" pageId="4" pageNumber="5">K. beshkovi</emphasis>
</taxonomicName>
and 
<taxonomicName id="6D8E4D27FFEAFF82FE2B81C24498FC2A" box="[416,507,919,945]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="biroi">
<emphasis id="98FAEAB6FFEAFF82FE2B81C24498FC2A" box="[416,507,919,945]" italics="true" pageId="4" pageNumber="5">K. biroi</emphasis>
</taxonomicName>
are unknown). In other nesticid genera, the MA is absent, inconspicuous or poorly developed (
<taxonomicName id="6D8E4D27FFEAFF82FE1081E14631FC4E" authority="Lehtinen &amp; Saaristo, 1980" authorityName="Lehtinen &amp; Saaristo" authorityYear="1980" box="[411,850,955,981]" class="Arachnida" family="Nesticidae" genus="Nesticella" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FE1081E14773FC4E" box="[411,528,955,981]" italics="true" pageId="4" pageNumber="5">Nesticella</emphasis>
<bibRefCitation id="CE1F4B55FFEAFF82FD9381E14631FC4E" author="Lehtinen P. T. &amp; Saaristo M. I." box="[536,850,955,981]" pageId="4" pageNumber="5" pagination="47 - 66" refId="ref10854" refString="Lehtinen P. T. &amp; Saaristo M. I. 1980. Spiders of the Oriental-Australian region. II. Nesticidae. Annales Zoologici Fennici 17: 47 - 66." type="journal article" year="1980">Lehtinen &amp; Saaristo, 1980</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="6D8E4D27FFEAFF82FCD481E141C6FC4E" authority="Ballarin &amp; Li, 2015" authorityName="Ballarin &amp; Li" authorityYear="2015" box="[863,1189,954,981]" class="Arachnida" family="Nesticidae" genus="Wraios" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FCD481E146D1FC4F" box="[863,946,955,980]" italics="true" pageId="4" pageNumber="5">Wraios</emphasis>
Ballarin &amp; Li, 2015
</taxonomicName>
; 
<taxonomicName id="6D8E4D27FFEAFF82FB3981E14427FC62" authority="Ballarin &amp; Li, 2015" authorityName="Ballarin &amp; Li" authorityYear="2015" class="Arachnida" family="Nesticidae" genus="Nescina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FB3981E14072FC4F" box="[1202,1297,955,980]" italics="true" pageId="4" pageNumber="5">Nescina</emphasis>
Ballarin &amp; Li, 2015
</taxonomicName>
; 
<taxonomicName id="6D8E4D27FFEAFF82FED9818547FFFC62" authority="Ballarin &amp; Li, 2015" authorityName="Ballarin &amp; Li" authorityYear="2015" box="[338,668,990,1017]" class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FED9818544C6FC63" box="[338,421,991,1016]" italics="true" pageId="4" pageNumber="5">Hamus</emphasis>
Ballarin &amp; Li, 2015
</taxonomicName>
; 
<taxonomicName id="6D8E4D27FFEAFF82FD2181844161FC63" authority="Keyserling, 1884" authorityName="Keyserling" authorityYear="1884" box="[682,1026,990,1016]" class="Arachnida" family="Nesticidae" genus="Gaucelmus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FD218184464DFC63" box="[682,814,990,1016]" italics="true" pageId="4" pageNumber="5">Gaucelmus</emphasis>
Keyserling, 1884
</taxonomicName>
; 
<taxonomicName id="6D8E4D27FFEAFF82FB9F81844011FC63" authority="Komatsu, 1942" authorityName="Komatsu" authorityYear="1942" box="[1044,1394,990,1016]" class="Arachnida" family="Nesticidae" genus="Cyclocarcina" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FB9F818441D2FC63" box="[1044,1201,990,1016]" italics="true" pageId="4" pageNumber="5">Cyclocarcina</emphasis>
Komatsu, 1942
</taxonomicName>
; 
<taxonomicName id="6D8E4D27FFEAFF82FF36865B4723FB80" authority="Kulczynski, 1914" authorityName="Kulczynski" authorityYear="1914" box="[189,576,1025,1051]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FF36865B4408FB80" box="[189,363,1025,1051]" italics="true" pageId="4" pageNumber="5">Typhlonesticus</emphasis>
<bibRefCitation id="CE1F4B55FFEAFF82FEFB865B4723FB80" author="Kulczynski W." box="[368,576,1025,1051]" pageId="4" pageNumber="5" pagination="353 - 387" refId="ref10687" refString="Kulczynski W. 1914. Aranearum species novae minusve cognate in montibus Kras dictis a Dre C. Absolon aliisque colectae. Bulletin international de l'Academie des Sciences de Cracovie 13: 353 - 387. Available from http: // biodiversitylibrary. org / page / 4553795 [accessed 17 Jan. 2017]." type="journal article" year="1914">Kulczyński, 1914</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="6D8E4D27FFEAFF82FDC1865B4686FB80" authority="Wunderlich, 1992" authorityName="Wunderlich" authorityYear="1992" box="[586,997,1025,1051]" class="Arachnida" family="Nesticidae" genus="Canarionesticus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FDC1865B4669FB80" box="[586,778,1025,1051]" italics="true" pageId="4" pageNumber="5">Canarionesticus</emphasis>
Wunderlich, 1992
</taxonomicName>
); in the form of a small, curved and pointed hook (
<taxonomicName id="6D8E4D27FFEAFF82FEEF867E4608FBA5" authority="Lehtinen &amp; Saaristo, 1980" authorityName="Lehtinen &amp; Saaristo" authorityYear="1980" box="[356,875,1060,1086]" class="Arachnida" family="Nesticidae" genus="Carpathonesticus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FEEF867E4750FBA5" box="[356,563,1060,1086]" italics="true" pageId="4" pageNumber="5">Carpathonesticus</emphasis>
<bibRefCitation id="CE1F4B55FFEAFF82FDB3867E4608FBA5" author="Lehtinen P. T. &amp; Saaristo M. I." box="[568,875,1060,1086]" pageId="4" pageNumber="5" pagination="47 - 66" refId="ref10854" refString="Lehtinen P. T. &amp; Saaristo M. I. 1980. Spiders of the Oriental-Australian region. II. Nesticidae. Annales Zoologici Fennici 17: 47 - 66." type="journal article" year="1980">Lehtinen &amp; Saaristo, 1980</bibRefCitation>
</taxonomicName>
); compact, transverse and not very developed (
<taxonomicName id="6D8E4D27FFEAFF82FF4C861D476DFBF9" authority="Roewer, 1935" authorityName="Roewer" authorityYear="1935" box="[199,526,1095,1122]" class="Arachnida" family="Nesticidae" genus="Eidmannella" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FF4C861D443EFBFA" box="[199,349,1095,1121]" italics="true" pageId="4" pageNumber="5">Eidmannella</emphasis>
Roewer, 1935
</taxonomicName>
); more conspicuous and transverse (
<taxonomicName id="6D8E4D27FFEAFF82FC348612402EFBF9" authority="Esyunin &amp; Efimik, 1998" authorityName="Esyunin &amp; Efimik" authorityYear="1998" box="[959,1357,1095,1122]" class="Arachnida" family="Nesticidae" genus="Aituaria" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FC3486124142FBF9" box="[959,1057,1096,1122]" italics="true" pageId="4" pageNumber="5">Aituaria</emphasis>
Esyunin &amp; Efimik, 1998
</taxonomicName>
) or it is well developed (
<taxonomicName id="6D8E4D27FFEAFF82FE3A86314626FB1E" authority="(Clerck, 1757)" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[433,837,1130,1157]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cellulanus">
<emphasis id="98FAEAB6FFEAFF82FE3A863147F6FB1E" box="[433,661,1131,1157]" italics="true" pageId="4" pageNumber="5">Nesticus cellulanus</emphasis>
(Clerck, 1757)
</taxonomicName>
and 
<taxonomicName id="6D8E4D27FFEAFF82FCF58631418DFB1E" authority="Liu &amp; Li, 2013" authorityName="Liu &amp; Li" authorityYear="2013" box="[894,1262,1131,1157]" class="Arachnida" family="Nesticidae" genus="Pseudonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FCF586314150FB1E" box="[894,1075,1131,1157]" italics="true" pageId="4" pageNumber="5">Pseudonesticus</emphasis>
Liu &amp; Li, 2013
</taxonomicName>
), but in this case the MA is long, thin and extended laterally. In 
<taxonomicName id="6D8E4D27FFEAFF82FC8486D546DCFB33" authority="Pavlek &amp; Ribera, 2017" authorityName="Pavlek &amp; Ribera" authorityYear="2017" box="[783,959,1167,1192]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="98FAEAB6FFEAFF82FC8486D546DCFB33" box="[783,959,1167,1192]" italics="true" pageId="4" pageNumber="5">Kryptonesticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEAFF82FC4D86D44151FB33" box="[966,1074,1166,1192]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
, the MA extends laterally at its base and then stretches towards the apex of the tegulum, parallel to the embolus. The theridioid tegular apophysis (TTA) is more compact than in 
<taxonomicName id="6D8E4D27FFEAFF82FD4A868F4603FB75" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[705,864,1236,1262]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cellulanus">
<emphasis id="98FAEAB6FFEAFF82FD4A868F4603FB75" box="[705,864,1236,1262]" italics="true" pageId="4" pageNumber="5">N. cellulanus</emphasis>
</taxonomicName>
or 
<taxonomicName id="6D8E4D27FFEAFF82FC07868E4122FB75" authorityName="Liu &amp; Li" authorityYear="2013" box="[908,1089,1236,1262]" class="Arachnida" family="Nesticidae" genus="Pseudonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEAFF82FC07868E4122FB75" box="[908,1089,1236,1262]" italics="true" pageId="4" pageNumber="5">Pseudonesticus</emphasis>
</taxonomicName>
. This is a very important character, since the TTA and MA are actively involved in sperm transfer (
<bibRefCitation id="CE1F4B55FFEAFF82FBAB86AD41D1FA89" author="Huber B. A." box="[1056,1202,1271,1298]" pageId="4" pageNumber="5" pagination="2437 - 2447" refId="ref10362" refString="Huber B. A. 1993. Genital mechanics and sexual selection in the spider Nesticus - cellulanus (Araneae, Nesticidae). Canadian Journal of Zoology 71: 2437 - 2447. http: // dx. doi. org / 10.1139 / z 93 - 340" type="journal article" year="1993">
Huber 
<date id="DE301064FFEAFF82FBF886A241D1FA89" box="[1139,1202,1272,1298]" pageId="4" pageNumber="5" value="1993">1993</date>
</bibRefCitation>
). 
<taxonomicName id="6D8E4D27FFEAFF82FB4D86A24015FA8A" authority="Pavlek &amp; Ribera, 2017" authorityName="Pavlek &amp; Ribera" authorityYear="2017" box="[1222,1398,1272,1297]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="98FAEAB6FFEAFF82FB4D86A24015FA8A" box="[1222,1398,1272,1297]" italics="true" pageId="4" pageNumber="5">Kryptonesticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEAFF82FF368741444BFAAE" box="[189,296,1307,1333]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
also differs from other nesticid genera in the shape, ramification and modifications associated with the paracymbium. 
<taxonomicName id="6D8E4D27FFEAFF82FE52876547EAFAC3" authority="Pavlek &amp; Ribera, 2017" authorityName="Pavlek &amp; Ribera" authorityYear="2017" box="[473,649,1343,1368]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="98FAEAB6FFEAFF82FE52876547EAFAC3" box="[473,649,1343,1368]" italics="true" pageId="4" pageNumber="5">Kryptonesticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEAFF82FD198764479CFAC3" box="[658,767,1342,1368]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
shows a well-developed dorsal process and a poorly differentiated ventral one. Distal and paradistal apophyses are absent and the dorsomedian apophysis is short and pointed, inconspicuous or absent. Females of this new genus show characteristic slender spermathecae, oval-shaped, with a shallow constriction in the middle, located above or just behind the apical part of the internal chitinous structures of the vulva. Insemination ducts depart from the basal part of the spermathecae, make a sharp turn-out and continue ventro-laterally, reaching the copulatory orifices.
</paragraph>
</subSubSection>
<subSubSection id="E294652FFFEAFF82FF368409472FF925" pageId="4" pageNumber="5" type="etymology">
<paragraph id="AA3136A4FFEAFF82FF3684094427F9F6" blockId="4.[189,324,1619,1645]" box="[189,324,1619,1645]" pageId="4" pageNumber="5">
<heading id="F17981C8FFEAFF82FF3684094427F9F6" bold="true" box="[189,324,1619,1645]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="98FAEAB6FFEAFF82FF3684094427F9F6" bold="true" box="[189,324,1619,1645]" pageId="4" pageNumber="5">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="AA3136A4FFEAFF82FF3684DB472FF925" blockId="4.[189,1398,1665,1727]" pageId="4" pageNumber="5">
The prefix “ 
<emphasis id="98FAEAB6FFEAFF82FEC684D844FDF900" box="[333,414,1666,1691]" italics="true" pageId="4" pageNumber="5">Krypto</emphasis>
”, from Ancient Greek κρυπτóς (kruptós “hidden”), alludes to the long time it took to diagnose this evolutionary line.
</paragraph>
</subSubSection>
<subSubSection id="E294652FFFEAFF82FF3684BF41ECF820" pageId="4" pageNumber="5" type="distribution">
<paragraph id="AA3136A4FFEAFF82FF3684BF4435F964" blockId="4.[189,342,1765,1791]" box="[189,342,1765,1791]" pageId="4" pageNumber="5">
<heading id="F17981C8FFEAFF82FF3684BF4435F964" bold="true" box="[189,342,1765,1791]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="98FAEAB6FFEAFF82FF3684BF4435F964" bold="true" box="[189,342,1765,1791]" pageId="4" pageNumber="5">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="AA3136A4FFEAFF82FF36854F41ECF820" blockId="4.[189,1399,1812,1979]" pageId="4" pageNumber="5">
From 
<collectingCountry id="D2997634FFEAFF82FE8D854E4408F8B5" box="[262,363,1812,1838]" name="Bulgaria" pageId="4" pageNumber="5">Bulgaria</collectingCountry>
and 
<collectingCountry id="D2997634FFEAFF82FE2C854E4498F8B5" box="[423,507,1812,1838]" name="Turkey" pageId="4" pageNumber="5">Turkey</collectingCountry>
to 
<collectingCountry id="D2997634FFEAFF82FDA8854E471FF8B5" box="[547,636,1812,1838]" name="Croatia" pageId="4" pageNumber="5">Croatia</collectingCountry>
, including 
<collectingCountry id="D2997634FFEAFF82FC89854F46F7F8B4" box="[770,916,1813,1839]" name="Serbia" pageId="4" pageNumber="5">Montenegro</collectingCountry>
, 
<collectingCountry id="D2997634FFEAFF82FC29854E41A5F8B5" box="[930,1222,1812,1838]" name="Bosnia and Herzegovina" pageId="4" pageNumber="5">Bosnia and Herzegovina</collectingCountry>
and Crete. All these species are known only from the 
<typeStatus id="75358806FFEAFF82FD01856247DEF8C9" box="[650,701,1848,1874]" pageId="4" pageNumber="5">type</typeStatus>
locality, or have small distribution ranges. 
<taxonomicName id="6D8E4D27FFEAFF82FB378562405BF8CA" authorityName="Simon" authorityYear="1879" box="[1212,1336,1848,1873]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="eremita">
<emphasis id="98FAEAB6FFEAFF82FB378562405BF8CA" box="[1212,1336,1848,1873]" italics="true" pageId="4" pageNumber="5">K. eremita</emphasis>
</taxonomicName>
is an exception; this species is linked to human activities and can be found in hangars, cellars and cottages. It has been cited from 
<collectingCountry id="D2997634FFEAFF82FE258524449DF803" box="[430,510,1918,1944]" name="France" pageId="4" pageNumber="5">France</collectingCountry>
and 
<collectingCountry id="D2997634FFEAFF82FDB38524470DF803" box="[568,622,1918,1944]" name="Italy" pageId="4" pageNumber="5">Italy</collectingCountry>
to 
<collectingCountry id="D2997634FFEAFF82FD1F8524479AF803" box="[660,761,1918,1944]" name="Bulgaria" pageId="4" pageNumber="5">Bulgaria</collectingCountry>
and 
<collectingCountry id="D2997634FFEAFF82FCB8852446EAF803" box="[819,905,1918,1944]" name="Turkey" pageId="4" pageNumber="5">Turkey</collectingCountry>
. It is a potentially invasive species, found in an abandoned air-raid tunnel in 
<collectingRegion id="684AF846FFEAFF82FDC485FB47A7F820" box="[591,708,1953,1979]" country="New Zealand" name="Auckland" pageId="4" pageNumber="5">Auckland</collectingRegion>
, 
<collectingCountry id="D2997634FFEAFF82FD4485FB460FF820" box="[719,876,1953,1979]" name="New Zealand" pageId="4" pageNumber="5">New Zealand</collectingCountry>
(
<bibRefCitation id="CE1F4B55FFEAFF82FCF785FB41E1F820" author="Vink C. J. &amp; Duperre N." box="[892,1154,1953,1979]" pageId="4" pageNumber="5" pagination="511 - 514" refId="ref11393" refString="Vink C. J. &amp; Duperre N. 2011. Nesticus eremita (Araneae: Nesticidae): redescription of a potentially invasive European spider found in New Zealand. Journal of Arachnology 39: 511 - 514. http: // dx. doi. org / 10.1636 / a 11 - 57.1" type="journal article" year="2011">
Vink &amp; Dupérré 
<date id="DE301064FFEAFF82FBCF85FB41E1F820" box="[1092,1154,1953,1979]" pageId="4" pageNumber="5" value="2011">2011</date>
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E294652FFFEBFF83FF3683514658FB5D" pageId="5" pageNumber="6" type="discussion">
<paragraph id="AA3136A4FFEBFF83FF36835144D4FEBE" blockId="5.[189,439,267,293]" box="[189,439,267,293]" pageId="5" pageNumber="6">
<heading id="F17981C8FFEBFF83FF36835144D4FEBE" bold="true" box="[189,439,267,293]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="98FAEAB6FFEBFF83FF36835144D4FEBE" bold="true" box="[189,439,267,293]" pageId="5" pageNumber="6">Taxonomic remarks</emphasis>
</heading>
</paragraph>
<paragraph id="AA3136A4FFEBFF83FF3683634032FDF6" blockId="5.[189,1399,313,622]" pageId="5" pageNumber="6">
The morphology of both male and female copulatory organs of this group of species shows important differences with respect to 
<taxonomicName id="6D8E4D27FFEBFF83FD85830747CCFEEC" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[526,687,349,375]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cellulanus">
<emphasis id="98FAEAB6FFEBFF83FD85830747CCFEEC" box="[526,687,349,375]" italics="true" pageId="5" pageNumber="6">N. cellulanus</emphasis>
</taxonomicName>
(the 
<typeStatus id="75358806FFEBFF83FD7E8307464BFEEC" box="[757,808,349,375]" pageId="5" pageNumber="6">type</typeStatus>
species of the genus 
<taxonomicName id="6D8E4D27FFEBFF83FBB0830741FDFEED" authorityName="Thorell" authorityYear="1869" box="[1083,1182,349,374]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEBFF83FBB0830741FDFEED" box="[1083,1182,349,374]" italics="true" pageId="5" pageNumber="6">Nesticus</emphasis>
</taxonomicName>
) and the western Mediterranean 
<taxonomicName id="6D8E4D27FFEBFF83FEE483DB44B1FE01" authorityName="Thorell" authorityYear="1869" box="[367,466,385,410]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEBFF83FEE483DB44B1FE01" box="[367,466,385,410]" italics="true" pageId="5" pageNumber="6">Nesticus</emphasis>
</taxonomicName>
species, as well as the representatives of 
<taxonomicName id="6D8E4D27FFEBFF83FC3283DA41EBFE01" authorityName="Lehtinen &amp; Saaristo" authorityYear="1980" box="[953,1160,384,410]" class="Arachnida" family="Nesticidae" genus="Carpathonesticus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEBFF83FC3283DA41EBFE01" box="[953,1160,384,410]" italics="true" pageId="5" pageNumber="6">Carpathonesticus</emphasis>
</taxonomicName>
and 
<taxonomicName id="6D8E4D27FFEBFF83FB4A83DA400CFE01" authorityName="Kulczynski" authorityYear="1914" box="[1217,1391,384,410]" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEBFF83FB4A83DA400CFE01" box="[1217,1391,384,410]" italics="true" pageId="5" pageNumber="6">Typhlonesticus</emphasis>
</taxonomicName>
. The asymmetrical development of the paracymbial ramification, the shape, size and arrangement of the median apophysis and some differences in the size and arrangement of the TTA p1 and p2 processes constitute the major differences in males. The number and position of spermathecae and the features of the insemination ducts are the most obvious characteristics in females. 
<taxonomicName id="6D8E4D27FFEBFF83FB99805741E1FDBC" authorityName="Deltshev" authorityYear="1977" box="[1042,1154,525,551]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beroni">
<emphasis id="98FAEAB6FFEBFF83FB99805741E1FDBC" box="[1042,1154,525,551]" italics="true" pageId="5" pageNumber="6">K. beroni</emphasis>
</taxonomicName>
and 
<taxonomicName id="6D8E4D27FFEBFF83FB3480574029FDBC" authorityName="Deltshev" authorityYear="1979" box="[1215,1354,525,551]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beshkovi">
<emphasis id="98FAEAB6FFEBFF83FB3480574029FDBC" box="[1215,1354,525,551]" italics="true" pageId="5" pageNumber="6">K. beshkovi</emphasis>
</taxonomicName>
are only known from females. Regardless, the shape and position of the spemathecae and the insemination ducts undoubtedly allow us to transfer these two species to the 
<taxonomicName id="6D8E4D27FFEBFF83FC2A800E4132FDF6" authorityName="Pavlek &amp; Ribera" authorityYear="2017" box="[929,1105,596,621]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEBFF83FC2A800E4132FDF6" box="[929,1105,596,621]" italics="true" pageId="5" pageNumber="6">Kryptonesticus</emphasis>
</taxonomicName>
evolutionary lineage.
</paragraph>
<paragraph id="AA3136A4FFEBFF83FF3680CE4143FC3E" blockId="5.[189,1399,660,933]" pageId="5" pageNumber="6">
Concerning species of 
<taxonomicName id="6D8E4D27FFEBFF83FE4D80CF474AFD35" authorityName="Thorell" authorityYear="1869" box="[454,553,661,686]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEBFF83FE4D80CF474AFD35" box="[454,553,661,686]" italics="true" pageId="5" pageNumber="6">Nesticus</emphasis>
</taxonomicName>
from other parts of the world (e.g., 
<collectingCountry id="D2997634FFEBFF83FC4380CF415BFD35" box="[968,1080,660,687]" name="China" pageId="5" pageNumber="6">East Asia</collectingCountry>
, North and South America) we can state that they do not show the diagnostic characters of the new genus described here. Surely this is a polyphyletic group that could be considered as a wastebasket group, and its inclusion within the genus 
<taxonomicName id="6D8E4D27FFEBFF83FE8280A5440FFC83" authorityName="Thorell" authorityYear="1869" box="[265,364,767,792]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEBFF83FE8280A5440FFC83" box="[265,364,767,792]" italics="true" pageId="5" pageNumber="6">Nesticus</emphasis>
</taxonomicName>
must be revised since the morphology of their genitalia is remarkably different from the 
<typeStatus id="75358806FFEBFF83FF3681784593FCA7" box="[189,240,802,828]" pageId="5" pageNumber="6">type</typeStatus>
species of the genus (
<taxonomicName id="6D8E4D27FFEBFF83FE70817847FBFCA0" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[507,664,801,827]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cellulanus">
<emphasis id="98FAEAB6FFEBFF83FE70817847FBFCA0" box="[507,664,801,827]" italics="true" pageId="5" pageNumber="6">N. cellulanus</emphasis>
</taxonomicName>
). Of course, this affirmation must be tested in a phylogenetic framework including all species of 
<taxonomicName id="6D8E4D27FFEBFF83FDEF811F47A4FCC5" authorityName="Thorell" authorityYear="1869" box="[612,711,837,862]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEBFF83FDEF811F47A4FCC5" box="[612,711,837,862]" italics="true" pageId="5" pageNumber="6">Nesticus</emphasis>
</taxonomicName>
described so far. Preliminary results on the phylogeny of European nesticids, including some American and Asian representatives (results not shown), indicate the polyphyly of 
<taxonomicName id="6D8E4D27FFEBFF83FE0D81D6448AFC3E" authorityName="Thorell" authorityYear="1869" box="[390,489,908,933]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="98FAEAB6FFEBFF83FE0D81D6448AFC3E" box="[390,489,908,933]" italics="true" pageId="5" pageNumber="6">Nesticus</emphasis>
</taxonomicName>
species described from other parts of the world.
</paragraph>
<paragraph id="AA3136A4FFEBFF83FF368196443EFC7D" blockId="5.[189,349,972,998]" box="[189,349,972,998]" pageId="5" pageNumber="6">
<heading id="F17981C8FFEBFF83FF368196443EFC7D" bold="true" box="[189,349,972,998]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="98FAEAB6FFEBFF83FF368196443EFC7D" bold="true" box="[189,349,972,998]" pageId="5" pageNumber="6">Composition</emphasis>
</heading>
</paragraph>
<paragraph id="AA3136A4FFEBFF83FF3681A64658FB5D" blockId="5.[189,1399,1019,1222]" pageId="5" pageNumber="6">
<taxonomicName id="6D8E4D27FFEBFF83FF3681A6449CFB8E" authority="Pavlek &amp; Ribera, 2017" authorityName="Pavlek &amp; Ribera" authorityYear="2017" box="[189,511,1019,1045]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="deelemanae" status="gen. et sp. nov.">
<emphasis id="98FAEAB6FFEBFF83FF3681A6449CFB8E" box="[189,511,1019,1045]" italics="true" pageId="5" pageNumber="6">Kryptonesticus deelemanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEBFF83FD8D81A147D6FB8E" box="[518,693,1019,1045]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="6D8E4D27FFEBFF83FD4B81A6468BFB8E" authority="(Simon, 1879)" authorityName="Pavlek &amp; Ribera" authorityYear="2017" baseAuthorityName="Simon" baseAuthorityYear="1879" box="[704,1000,1019,1045]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="eremita" status="comb. nov.">
<emphasis id="98FAEAB6FFEBFF83FD4B81A64659FB8E" box="[704,826,1020,1045]" italics="true" pageId="5" pageNumber="6">K. eremita</emphasis>
(Simon, 1879)
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEBFF83FC6481A14113FB8E" box="[1007,1136,1019,1045]" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
(Southern and Central Europe, introduced in 
<collectingCountry id="D2997634FFEBFF83FE4586454711FBA3" box="[462,626,1054,1081]" name="New Zealand" pageId="5" pageNumber="6">New Zealand</collectingCountry>
, potentially invasive species), 
<taxonomicName id="6D8E4D27FFEBFF83FC6C86454015FBA3" authority="(Kulczynski, 1914)" authorityName="Pavlek &amp; Ribera" authorityYear="2017" baseAuthorityName="Kulczynski" baseAuthorityYear="1914" box="[999,1398,1054,1080]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="arenstorffi" status="comb. nov.">
<emphasis id="98FAEAB6FFEBFF83FC6C864541E6FBA3" box="[999,1157,1054,1080]" italics="true" pageId="5" pageNumber="6">K. arenstorffi</emphasis>
(
<bibRefCitation id="CE1F4B55FFEBFF83FB138644400DFBA3" author="Kulczynski W." box="[1176,1390,1054,1080]" pageId="5" pageNumber="6" pagination="353 - 387" refId="ref10687" refString="Kulczynski W. 1914. Aranearum species novae minusve cognate in montibus Kras dictis a Dre C. Absolon aliisque colectae. Bulletin international de l'Academie des Sciences de Cracovie 13: 353 - 387. Available from http: // biodiversitylibrary. org / page / 4553795 [accessed 17 Jan. 2017]." type="journal article" year="1914">Kulczyński, 1914</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEBFF83FF36861B445DFBC7" box="[189,318,1089,1116]" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
(
<collectingCountry id="D2997634FFEBFF83FEC6861B470CFBC0" box="[333,623,1089,1115]" name="Bosnia and Herzegovina" pageId="5" pageNumber="6">Bosnia and Herzegovina</collectingCountry>
and 
<collectingCountry id="D2997634FFEBFF83FD2C86184658FBC7" box="[679,827,1090,1116]" name="Serbia" pageId="5" pageNumber="6">Montenegro</collectingCountry>
), 
<taxonomicName id="6D8E4D27FFEBFF83FCC5861841EFFBC0" authority="(Kratochvil, 1933)" authorityName="Pavlek &amp; Ribera" authorityYear="2017" baseAuthorityName="Kratochvil" baseAuthorityYear="1933" box="[846,1164,1089,1115]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fagei" status="comb. nov.">
<emphasis id="98FAEAB6FFEBFF83FCC5861846C8FBC0" box="[846,939,1089,1115]" italics="true" pageId="5" pageNumber="6">K. fagei</emphasis>
(
<bibRefCitation id="CE1F4B55FFEBFF83FC31861B41E7FBC0" author="Kratochvil J." box="[954,1156,1089,1115]" pageId="5" pageNumber="6" pagination="1 - 69" refId="ref10561" refString="Kratochvil J. 1933. Evropske druhy celedi Nesticidae Dahl. Prace Moravske Prirodovedecke Spolecnosti 8: 1 - 69." type="journal article" year="1933">Kratochvíl, 1933</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEBFF83FB18861B4076FBC7" box="[1171,1301,1089,1116]" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
(
<collectingCountry id="D2997634FFEBFF83FAA8861B44EDFBE4" name="Bosnia and Herzegovina" pageId="5" pageNumber="6">Bosnia and Herzegovina</collectingCountry>
), 
<taxonomicName id="6D8E4D27FFEBFF83FE2D863F478FFBE4" authority="(Deltshev, 1977)" authorityName="Pavlek &amp; Ribera" authorityYear="2017" baseAuthorityName="Deltshev" baseAuthorityYear="1977" box="[422,748,1124,1151]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beroni" status="comb. nov.">
<emphasis id="98FAEAB6FFEBFF83FE2D863F477AFBE5" box="[422,537,1124,1150]" italics="true" pageId="5" pageNumber="6">K. beroni</emphasis>
(
<bibRefCitation id="CE1F4B55FFEBFF83FDA7863E4787FBE4" author="Deltshev C." box="[556,740,1124,1151]" pageId="5" pageNumber="6" pagination="73 - 78" refId="ref10033" refString="Deltshev C. 1977. Genus Nesticus (Nesticidae, Araneae) from Bulgarian caves. Congres international de Speleologie 6 (Olomouc, 1973): 73 - 78." type="book chapter" year="1977">Deltshev, 1977</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEBFF83FD73863E461DFBE4" box="[760,894,1124,1151]" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
(
<collectingCountry id="D2997634FFEBFF83FC19863E469EFBE4" box="[914,1021,1124,1151]" name="Bulgaria" pageId="5" pageNumber="6">Bulgaria</collectingCountry>
), 
<taxonomicName id="6D8E4D27FFEBFF83FB9E863F4015FBE4" authority="(Deltshev, 1979)" authorityName="Pavlek &amp; Ribera" authorityYear="2017" baseAuthorityName="Deltshev" baseAuthorityYear="1979" box="[1045,1398,1124,1151]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beshkovi" status="comb. nov.">
<emphasis id="98FAEAB6FFEBFF83FB9E863F41C0FBE5" box="[1045,1187,1124,1150]" italics="true" pageId="5" pageNumber="6">K. beshkovi</emphasis>
(
<bibRefCitation id="CE1F4B55FFEBFF83FB3C863E400DFBE4" author="Deltshev C. D." box="[1207,1390,1124,1151]" pageId="5" pageNumber="6" pagination="53 - 63" refId="ref10064" refString="Deltshev C. D. 1979. A contribution to the study of cave spiders (Araneae) in Greece. Four new species (Araneae, Nesticidae, Linyphiidae) from the islands of Crete and Thera. Acta Zoologica Bulgarica 13: 53 - 63." type="journal article" year="1979">Deltshev, 1979</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEBFF83FF3686D24420FB39" box="[189,323,1160,1186]" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
(Crete), 
<taxonomicName id="6D8E4D27FFEBFF83FE3E86D2462DFB39" authority="(Bosselaers, 1998)" authorityName="Pavlek &amp; Ribera" authorityYear="2017" baseAuthorityName="Bosselaers" baseAuthorityYear="1998" box="[437,846,1160,1186]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="henderickxi" status="comb. nov.">
<emphasis id="98FAEAB6FFEBFF83FE3E86D24707FB39" box="[437,612,1160,1186]" italics="true" pageId="5" pageNumber="6">K. henderickxi</emphasis>
(
<bibRefCitation id="CE1F4B55FFEBFF83FDF386D24625FB39" author="Bosselaers J." box="[632,838,1160,1186]" pageId="5" pageNumber="6" pagination="9 - 14" refId="ref9919" refString="Bosselaers J. 1998. Nesticus henderickxi (Araneae, Nesticidae), a new blind troglobitic spider from Crete. Bulletin of the British Arachnological Society 11: 9 - 14." type="journal article" year="1998">Bosselaers, 1998</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="83C957CDFFEBFF83FCD186D24683FB39" box="[858,992,1160,1186]" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
(Crete) and 
<taxonomicName id="6D8E4D27FFEBFF83FB0A86D2406DFB39" authorityName="Lopez-Pancorbo, Kunt &amp; Ribera" authorityYear="2013" box="[1153,1294,1160,1186]" class="Arachnida" family="Nesticidae" genus="Kryptonesticus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dimensis">
<emphasis id="98FAEAB6FFEBFF83FB0A86D2406DFB39" box="[1153,1294,1160,1186]" italics="true" pageId="5" pageNumber="6">K. dimensis</emphasis>
</taxonomicName>
(López- Pancorbo, Kunt &amp; Ribera, 
<date id="DE301064FFEBFF83FE7C86F14750FB5E" box="[503,563,1195,1221]" pageId="5" pageNumber="6" value="2013">2013</date>
) comb. nov. (
<collectingCountry id="D2997634FFEBFF83FD5F86F1464CFB5D" box="[724,815,1195,1222]" name="Turkey" pageId="5" pageNumber="6">Turkey</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>