156 lines
17 KiB
XML
156 lines
17 KiB
XML
<document id="711AA891E95E5AAEE2061CC6B8A4F293" ID-CLB-Dataset="34479" ID-DOI="http://dx.doi.org/10.3897/zookeys.145.1895" ID-GBIF-Dataset="9d00e643-a484-4a0f-9e4c-0486f131b545" ID-PMC="PMC3267458" ID-Pensoft-Pub="1313-2970-145-59" ID-PubMed="22287884" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-145-59" ModsDocOrigin="ZooKeys 145" ModsDocTitle="Synopsis of Falsocis Pic (Coleoptera, Ciidae), new species, new records and an identification key" checkinTime="1451249704874" checkinUser="pensoft" docAuthor="Lopes-Andrade, Cristiano & Lawrence, John F." docDate="2011" docId="22111DA02A83B75E5F48C458D243DD39" docLanguage="en" docName="ZooKeys 145: 59-78" docOrigin="ZooKeys 145" docSource="http://dx.doi.org/10.3897/zookeys.145.1895" docTitle="Falsocis egregius Lopes-Andrade & Lawrence, sp. n." docType="treatment" docVersion="6" lastPageNumber="67" masterDocId="A425F539795BFFF1FFD7FF81FF82F33E" masterDocTitle="Synopsis of Falsocis Pic (Coleoptera, Ciidae), new species, new records and an identification key" masterLastPageNumber="78" masterPageNumber="59" pageNumber="65" updateTime="1732761432594" updateUser="ExternalLinkService">
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<mods:mods id="29A5180407092D93074AB65074682C84" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C376DCAA924998E0847D09AA6796803B">
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<mods:title id="02E14A6B41A426C39E8B1B4F7C59E7E2">Synopsis of Falsocis Pic (Coleoptera, Ciidae), new species, new records and an identification key</mods:title>
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<mods:name id="7602BCEF0479B6D9370A1E5E0053CC8D" type="personal">
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<mods:roleTerm id="C814BCB28210E7042BFCD1F26B7481D0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D9AFDFEC2C2E714533A890A9BE2EC900">Lopes-Andrade, Cristiano</mods:namePart>
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<mods:name id="91A917EFEA41F441FFF52602805FE831" type="personal">
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<mods:roleTerm id="C1C4D70D73C67887D0F68072F70E6FEB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3EC973D6F48C1C1F9760D630451070CA">Lawrence, John F.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7CD98C5B30F9466271AFA61D99B84773">text</mods:typeOfResource>
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<mods:titleInfo id="E433A0350E95623B5215FB7377642ACA">
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<mods:title id="D9B81106C6FEB4F871A6B7873A9D3547">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="CF6D40EB3E122879F63B7010185AD8D8">
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<mods:date id="46B4546A83308F98C5F8304EBCC72725">2011</mods:date>
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<mods:detail id="00DB4B543CB37D7A146BBA45F579DBDD" type="volume">
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<mods:number id="CA462FA839973465AE0E7F1C5F48D378">145</mods:number>
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</mods:detail>
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<mods:extent id="9E0AC114E8292458B38CC797B37391F6" unit="page">
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<mods:start id="D4B79EECF0C69932B0A72128A12E5D16">59</mods:start>
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<mods:end id="F535A19751E69987E2C1872DE2EEA2BC">78</mods:end>
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<mods:location id="EAF36BBF0104407C33624B500931018E">
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<mods:url id="D8CC3AC9420D0E46E356733746844EEA">http://dx.doi.org/10.3897/zookeys.145.1895</mods:url>
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</mods:location>
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<mods:classification id="7C66A5709D71C422231F1CDA630462C7">journal article</mods:classification>
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<mods:identifier id="19EE46719B0661E4DDE97E47DFAB4726" type="DOI">http://dx.doi.org/10.3897/zookeys.145.1895</mods:identifier>
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<mods:identifier id="346548B22E4BC474CC3440A43C10A625" type="Pensoft-Pub">1313-2970-145-59</mods:identifier>
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</mods:mods>
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<treatment id="22111DA02A83B75E5F48C458D243DD39" ID-GBIF-Taxon="152031877" LSID="urn:lsid:zoobank.org:act:E6BFE936-A896-449F-8D6A-ADE8528F8728" httpUri="http://treatment.plazi.org/id/22111DA02A83B75E5F48C458D243DD39" lastPageId="8" lastPageNumber="67" pageId="6" pageNumber="65" scope_family="Ciidae" scope_order="Coleoptera">
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<subSubSection id="9D15C057E153F036E3D3955E3DF18FAF" pageId="6" pageNumber="65" type="nomenclature">
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<paragraph id="CE494B5D312F08C0A2EA71C44214C489" pageId="6" pageNumber="65">
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<taxonomicName id="6C9DF4E730ABAC574FAE0D3262A5626F" ID-CoL="4G54K" LSID="urn:lsid:zoobank.org:act:E6BFE936-A896-449F-8D6A-ADE8528F8728" authority="Lopes-Andrade & Lawrence" genus="Falsocis" lsidName="Falsocis egregius" pageId="6" pageNumber="65" rank="species" species="egregius">Falsocis egregius Lopes-Andrade & Lawrence</taxonomicName>
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<taxonomicNameLabel id="14C11BD22DCAB34EF06F306E0405DC1B" pageId="6" pageNumber="65">sp. n.</taxonomicNameLabel>
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Figs 10-1641
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</paragraph>
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</subSubSection>
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<subSubSection id="B2E719B237E842AB99A662B22B6B777D" pageId="6" pageNumber="65" type="type locality">
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<paragraph id="8546F63F71DAA57BAB55A21379A98B13" pageId="6" pageNumber="65">Type locality.</paragraph>
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<paragraph id="3D1E065381BFE18A6C351D76A7B8E420" pageId="6" pageNumber="65">
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<normalizedToken id="52328C3E392D626FE31697295B928046" originalValue="Santarém">Santarem</normalizedToken>
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, in the state of
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<normalizedToken id="C52714803470E4C1CD316033158F191C" originalValue="Pará">Para</normalizedToken>
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, northern Brazil (
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<geoCoordinate id="086C1D9C406AF4712BD9CFB41A3CFC02" direction="south" orientation="latitude" precision="925" value="-2.4333334">2°26'S</geoCoordinate>
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,
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<geoCoordinate id="84EE8E644A5BBDB4EC19895C35867E85" direction="west" orientation="longitude" precision="925" value="-54.7">54°42'W</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="D37962AD86F7AD790DCB30937D2C2CBE" pageId="6" pageNumber="65" type="etymology">
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<paragraph id="164B5143973DAB68EBF976FA6B0F1110" pageId="6" pageNumber="65">Etymology.</paragraph>
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<paragraph id="4824AC1F1096D12D28659331156D906B" pageId="6" pageNumber="65">
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The specific epithet is from the Latin
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<normalizedToken id="1DE6DD385012AE00A278394C2905326F" originalValue="“egregius”">"egregius"</normalizedToken>
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(adjective), which means
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<normalizedToken id="E3F97C5A81032FDC01DE3AF4D3EF409A" originalValue="“singular”">"singular"</normalizedToken>
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,
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<normalizedToken id="7962F8A70D0D1EEB72A0E4102B103C52" originalValue="“extraordinary”">"extraordinary"</normalizedToken>
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, in a reference to the outstanding head morphology of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="68DF3716A97F18796036945962D39074" pageId="6" pageNumber="65" type="diagnosis">
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<paragraph id="71E6D8EFEDC5D939B6DA95A19E31BE46" pageId="6" pageNumber="65">Diagnosis.</paragraph>
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<paragraph id="2F80C13A1D5B64FC6DAC7B035FA6055D" pageId="6" pageNumber="65">Head of male with sides of vertex produced laterally to form a pair of triangular plates, each with acute apex (Fig. 13, arrows); frontoclypeal horns wide at base and tapering to apex, slightly arcuate. Pronotum with anterior angles broadly rounded, barely produced forward; males with a row of long setae along the apex of the anterior projection but not forming distinct tufts (Figs 10, 12, arrows).</paragraph>
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</subSubSection>
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<subSubSection id="480EEF3DA45A80A7F40E9D9B8103DB57" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65" type="description">
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<paragraph id="D64686EFE141F98274FE748A72FC3D88" pageId="6" pageNumber="65">Description.</paragraph>
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<paragraph id="3B58A1072E60D2942713877688127D65" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65">
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Male holotype (Figs 10-13), measurements in mm: TL 2.85; PL 1.25; PW 1.50; EL 1.55; EW 1.55; GD 1.35. Ratios: PL/PW 0.83; EL/EW 1.00; EL/PL 1.24; GD/EW 0.87; TL/EW 1.84. Body oblong, strongly convex, mostly light yellowish brown; mouthparts and apices of femora reddish brown; ventral vestiture consisting of very fine decumbent setae. Head strongly developed, the anterior and lateral margins easily visible from above; dorsum concave, slightly tumid on disc; punctation relatively fine, shallow, sparse; in between punctures finely granulate; frontoclypeal ridge explanate, each side produced upward forming a conspicuous horn that is slightly arcuate, bearing two small tubercles between them; each side of vertex explanate and produced laterally forming a conspicuous triangular plate with an acute apex (Fig. 13, arrows). Eyes coarsely facetted; greatest width 0.18mm. Each antenna (left antenna measured; FL 0.22mm; CL 0.23mm; CL/FL 1.05) with length of antennomeres (in mm) as follows: 0.12; 0.07; 0.08; 0.04; 0.04; 0.03; 0.03; 0.06; 0.06; 0.11. Pronotum with shallow, coarse, single punctation; punctures separated by a distance about a puncture-width; in between punctures smooth at disc and finely granulate at the anterior projection; vestiture indistinctly dual, consisting of conspicuous yellowish stout erect bristles (~0.05mm) and very fine decumbent setae (<0.02mm); anterior angles broadly rounded, just slightly produced forward; anterior margin (beyond anterior angles) explanate, produced forward forming a plate that slightly curves downward and narrows toward a slightly arcuate apex with a row of very long slender setae (>0.35mm; Figs 10, 12, arrows). Scutellum subtriangular; punctation finer than those of pronotum and elytra; vestiture consisting of stout decumbent bristles (better seen in lateral view); basal width 0.21mm. Hindwings fully developed. Elytra subparallel at basal two-thirds, with posterolateral angles broadly rounded (as seen from above) and then converging to a blunt apex; punctation single and confused, a bit sparser than that of
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<pageBreakToken id="0E0E8329637CC75AE3F937980A2F38BF" pageId="7" pageNumber="66" start="start">pronotum</pageBreakToken>
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; in between punctures smooth; vestiture indistinctly dual, consisting of conspicuous stout erect bristles (~0.07mm) and very minute decumbent setae (<0.02mm); lateral and apical margins not visible from above; epipleura tapering from base to apex. Surface of the ventral thoracic and abdominal sclerites granulate, somewhat rugose. Prosternum flat; prosternal process laminate, almost half the length of procoxae. First abdominal ventrite more than twice as long as the second at midline; setose sex patch circular and margined, located at the middle of the ventrite and with a diameter of one-third the length of the ventrite at midline.
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</paragraph>
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<paragraph id="4BDC6D1944A520CACC5A2C612FF71AD3" pageId="7" pageNumber="66">Male terminalia in paratypes (Figs 15-16). Eighth sternite (Fig. 15) with posterior margin curved inward; angles not produced. Basal piece (Fig. 16) almost half the length of tegmen. Tegmen (Fig. 16) with anterior portion broadly rounded; lateral margins almost straight; posterior portion bearing a deep V-shaped emargination reaching the middle of the structure and forming two lateral lobes. Penis (Fig. 16) subcylindrical, lateral margins straight for most of their lengths and a bit expanded to a rounded apex; the penis is turned and lays laterally in the slide preparation.</paragraph>
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<paragraph id="30E5680E933F19AEFD3D70B146CA4332" pageId="7" pageNumber="66">Female (Fig. 14).Similar to males, but frontoclypeal ridge straight, barely sinuous, with rounded angles. Anterior pronotal margin broadly rounded. Abdomen devoid of sex patch.</paragraph>
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</subSubSection>
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<subSubSection id="5BF4C510B6424F579BD98DA1A6806F77" pageId="7" pageNumber="66" type="variation">
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<paragraph id="236D376F2EACFB9ABCE02FE743D244D5" pageId="7" pageNumber="66">Variation.</paragraph>
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<paragraph id="402AB84D4D25F52AEC3724C6F67C5676" pageId="7" pageNumber="66">
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Males, measurements in mm (n = 5, including the holotype): TL 2.75-2.95 (2.87
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<normalizedToken id="0EB8B33A1E6732665BCD016A612BA99B" originalValue="±">+/-</normalizedToken>
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0.08); PL 1.20-1.35 (1.26
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<normalizedToken id="C9ACA9B782E089FCED82BDA1E1A18363" originalValue="±">+/-</normalizedToken>
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0.05); PW 1.50-1.55 (1.52
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<normalizedToken id="520B117F3A3983A9FE7EFC4D43347DA3" originalValue="±">+/-</normalizedToken>
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0.03); EL 1.50-1.65 (1.56
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<normalizedToken id="74FDEE761A34A94FCDB6487930DB72B3" originalValue="±">+/-</normalizedToken>
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0.05); EW 1.55-1.65 (1.57
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<normalizedToken id="5EABC2CF7A6CB5EB4D02EC6576EE46CF" originalValue="±">+/-</normalizedToken>
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0.04); GD 1.30-1.40 (1.35
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<normalizedToken id="CCA8608BE35EC4B176C5B845C971207B" originalValue="±">+/-</normalizedToken>
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0.04). Ratios: PL/PW 0.80-0.87 (0.83
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<normalizedToken id="BD5CFFEF0A741521C09CADFA7761D5D5" originalValue="±">+/-</normalizedToken>
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0.03); EL/EW 0.97-1.00 (0.99
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<normalizedToken id="1C6E1B019B0BE4282B0411CA8AF90EF6" originalValue="±">+/-</normalizedToken>
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0.01); EL/PL 1.15-1.32 (1.24
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<normalizedToken id="EC2D8E004C84DA326E04601DD6FF9D45" originalValue="±">+/-</normalizedToken>
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0.06); GD/EW 0.84-0.87 (0.86
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<normalizedToken id="6BFB7CDFC5458E4CC2DBA833710E87D8" originalValue="±">+/-</normalizedToken>
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0.02); TL/EW 1.77-1.87 (1.83
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<normalizedToken id="BBB97DC0CACD9E37CAA5CBE318E04CC8" originalValue="±">+/-</normalizedToken>
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0.05).
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</paragraph>
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<paragraph id="3350A03DDDDDB3455A23BB97C810C5B4" pageId="7" pageNumber="66">Female, measurements in mm (n = 1): TL 2.50; PL 0.95; PW 1.45; EL 1.55; EW 1.50; GD 1.05. Ratios: PL/PW 0.66; EL/EW 1.03; EL/PL 1.63; GD/EW 0.70; TL/EW 1.67.</paragraph>
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</subSubSection>
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<subSubSection id="0C1CA3F3BF98D1AEF2F3D8E7E082991E" pageId="7" pageNumber="66" type="type series">
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<paragraph id="729FA10A9AF2D201E6124824E55E2756" pageId="7" pageNumber="66">Type series.</paragraph>
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<paragraph id="1C6D67DF2ED2175E619318DFE0BE32C0" pageId="7" pageNumber="66">
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Holotype. (CMNH) Brazil: \Santarem Brazil Acc. No.2966 \ CM \
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<taxonomicName id="1045EF7EE719595E873414AC04EB930B" genus="Falsocis" lsidName="Falsocis" pageId="7" pageNumber="66" rank="genus">Falsocis</taxonomicName>
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egregius Lopes-Andrade & Lawrence HOLOTYPUS [printed on red paper]\. Paratypes.Brazil: 4 males (1 ANIC; 2 CMNH; 1 LAPC, dissected) and 1 female (CMNH) \Santarem Brazil Acc. No.2966 \ CM\, same data as holotype. All paratypes distinguished labeled \
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<taxonomicName id="5DAAA0A5B27B0B67DFCA9A83E1B78988" genus="Falsocis" lsidName="Falsocis egregius" pageId="7" pageNumber="66" rank="species" species="egregius">Falsocis egregius</taxonomicName>
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Lopes-Andrade & Lawrence PARATYPUS [printed on yellow paper]\.
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</paragraph>
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</subSubSection>
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<subSubSection id="2D6FA4D54C977549D2326CC8E08A1D53" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66" type="comments and comparative notes">
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<paragraph id="5038668571387A3CE2A43EDB1C1E1C5A" pageId="7" pageNumber="66">Comments and comparative notes.</paragraph>
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<paragraph id="AFF1AF55ADD34B23BF850F42E8FA9E2D" pageId="7" pageNumber="66">
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The species is currently known from a single collection in
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<normalizedToken id="D3A253A90D5245196443B06391D1CC71" originalValue="Santarém">Santarem</normalizedToken>
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, northern Brazil (Fig. 41). Only one female was available for examination. The series is very small, but species sufficiently distinct to allow description. Differs from
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<taxonomicName id="49371169C635813260AD9B62A39525AA" genus="Falsocis" lsidName="Falsocis aquilonius" pageId="7" pageNumber="66" rank="species" species="aquilonius">Falsocis aquilonius</taxonomicName>
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sp. n. and
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<taxonomicName id="91B688C9E1625DDCBE1A79D19933BE78" genus="Falsocis" lsidName="Falsocis opacus" pageId="7" pageNumber="66" rank="species" species="opacus">Falsocis opacus</taxonomicName>
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in having the epipleura narrow posteriorly.
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<taxonomicName id="707681DFF3AF2D1BE31FA83A2077B357" genus="Falsocis" lsidName="Falsocis occultus" pageId="7" pageNumber="66" rank="species" species="occultus">Falsocis occultus</taxonomicName>
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sp. n. has deeper and closer pronotal punctation. Very similar to
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<taxonomicName id="2D27B1FF1B0B070D630199D336A9BA49" genus="Falsocis" lsidName="Falsocis brasiliensis" pageId="7" pageNumber="66" rank="species" species="brasiliensis">Falsocis brasiliensis</taxonomicName>
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, but differing mainly in features mentioned in the diagnosis and in the conspicuous basal piece of male terminalia. In
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<taxonomicName id="FB556F10E27E6DE0740A136936A9B384" genus="Falsocis" lsidName="Falsocis brasiliensis" pageId="7" pageNumber="66" rank="species" species="brasiliensis">Falsocis brasiliensis</taxonomicName>
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, the eighth sternite has the anterior margin only slightly curved inward; the basal piece was not observed in the available slide preparations and is possibly membranous (
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<bibRefCitation id="0697733CA1D98514361462047E9E5847" author="Lopes-Andrade, C" journalOrPublisher="Zootaxa" pageId="16" pageNumber="75" pagination="41 - 58" title="Notes on Falsocis Pic (Coleoptera: Tenebrionoidea: Ciidae), with the description of an endangered Brazilian species." volume="1544" year="2007">Lopes-Andrade 2007</bibRefCitation>
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); the tegmen has a subtriangular posterior portion, subparallel lateral margins and anterior V-shape emargination is only one-fourth to one-third deep.
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</paragraph>
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<caption id="24B139BD6EA0DF17E97EB914195A7673" pageId="8" pageNumber="67">
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<paragraph id="92AA61B9531F1E291BA3C53DC0689606" pageId="8" pageNumber="67">
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<pageBreakToken id="61D1C437C0BFE5A20DB6C0B3CBD7255F" pageId="8" pageNumber="67" start="start">Figures</pageBreakToken>
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10-16.
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<taxonomicName id="79C12A1F7284D87EDB1E9629E4B9895E" genus="Falsocis" lsidName="Falsocis egregius" pageId="8" pageNumber="67" rank="species" species="egregius">Falsocis egregius</taxonomicName>
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Lopes-Andrade & Lawrence, sp. n., male holotype 10-13 and female paratype 14 shown in the same scale, and slide preparation of male terminalia of a paratype 15-16. 10 Dorsal view (pronotal tuft of long setae, arrow) 11 Lateral view 12 Ventral view (pronotal tuft of long setae, arrow) 13 Head and pronotum view from above (triangular plate behind eyes, arrows) 14 Female 15 Eighth sternite 16 Aedeagus (left) and gross outline of aedeagus (right) showing tegmen (continuous line), penis (dashed line) and basal piece (dotted line).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |