242 lines
23 KiB
XML
242 lines
23 KiB
XML
<document ID-DOI="10.1590/S0073-47212003000400008" ID-GBIF-Dataset="32bd22ce-7d3f-4eac-8203-e9f61a19c7f0" ID-ISSN="1678-4766" ID-Zenodo-Dep="3719467" approvedBy="tatiana" checkinTime="1584724827204" checkinUser="tatiana" docAuthor="Barcellos, Aline & Grazia, Jocélia" docDate="2003" docId="217887A7991A80284B1A5D51FC0666D2" docLanguage="en" docName="Iheringia.93.4.413-446.pdf.imf" docOrigin="Iheringia, Sér. Zool. 93 (4)" docStyle="DocumentStyle{}" docTitle="Brachystethus coxalis Breddin. 1904" docType="treatment" docVersion="7" lastPageNumber="438" masterDocId="DD41FFDF990D80314A705E42FF826415" masterDocTitle="Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae)" masterLastPageNumber="446" masterPageNumber="413" pageNumber="436" updateTime="1668426789504" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Barcellos, Aline</mods:namePart>
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<mods:affiliation>Museu de Ciências Naturais, Fundação Zoobotânica do Rio Grande do Sul, Caixa Postal 1188, 90001 - 970, Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:nameIdentifier type="email">alinebar@ig.com.br</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grazia, Jocélia</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves, 9500, Prédio 43435.2, 91501 - 970, Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:nameIdentifier type="email">jocelia@ufrgs.br</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Iheringia, Sér. Zool.</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2003</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2003-12-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>93</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>413</mods:start>
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<mods:end>446</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1590/S0073-47212003000400008</mods:identifier>
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<mods:identifier type="GBIF-Dataset">32bd22ce-7d3f-4eac-8203-e9f61a19c7f0</mods:identifier>
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<mods:identifier type="ISSN">1678-4766</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3719467</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3719831" ID-GBIF-Taxon="162653799" ID-Zenodo-Dep="3719831" LSID="urn:lsid:plazi:treatment:217887A7991A80284B1A5D51FC0666D2" httpUri="http://treatment.plazi.org/id/217887A7991A80284B1A5D51FC0666D2" lastPageId="25" lastPageNumber="438" pageId="23" pageNumber="436">
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<subSubSection box="[362,770,787,811]" pageId="23" pageNumber="436" type="nomenclature">
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<paragraph blockId="23.[362,770,787,836]" box="[362,770,787,811]" pageId="23" pageNumber="436">
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<heading bold="true" box="[362,770,787,811]" centered="true" fontSize="10" level="1" pageId="23" pageNumber="436" reason="0">
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<taxonomicName authority="Breddin, 1904" authorityName="Breddin." authorityYear="1904" box="[362,770,787,811]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="436" phylum="Arthropoda" rank="species" species="coxalis">
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<emphasis bold="true" box="[362,770,787,811]" pageId="23" pageNumber="436">
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<emphasis bold="true" box="[362,603,787,811]" italics="true" pageId="23" pageNumber="436">Brachystethus coxalis</emphasis>
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Breddin, 1904
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[447,685,817,836]" pageId="23" pageNumber="436" type="description">
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<paragraph blockId="23.[362,770,787,836]" box="[447,685,817,836]" pageId="23" pageNumber="436">
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<heading box="[447,685,817,836]" pageId="23" pageNumber="436">
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(
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<figureCitation box="[454,532,817,836]" captionStart="Figs" captionStartId="5.[101,144,1441,1460]" captionTargetBox="[161,975,188,1422]" captionTargetId="figure@5.[161,975,188,1422]" captionText="Figs. 2-13. 2, 3, Brachystethus rubromaculatus Dallas, 1851; 4, 5, B. signoreti Stål; 1872; 6, B. cribrus (Fabricius, 1781); 7, B. schuhi sp. nov.; 8, B. tricolor Bolívar, 1879; 9, B. vexillum Breddin, 1903; 10, B. geniculatus (Fabricius, 1787); 11, B. coxalis Breddin, 1904; 12, B. improvisus Breddin, 1905; 13, B. vicinus Signoret, 1851. Magnification 1.63X." figureDoi="http://doi.org/10.5281/zenodo.3719471" httpUri="https://zenodo.org/record/3719471/files/figure.png" pageId="23" pageNumber="436">Figs. 11</figureCitation>
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,
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<figureCitation box="[545,569,817,836]" captionStart="Figs" captionStartId="9.[101,142,1044,1063]" captionText="Figs. 20-23. Pygophore, dorsal: 20, Brachystethus geniculatus; 21, B. coxalis; 22, B. improvisus; 23, B. vicinus (dor, dorsal rim; gcp, genital cup process; ldor, lobe of dorsal rim; pa, paramere; vr, ventral rim; X, tenth segment). Scale, 1 mm." pageId="23" pageNumber="436">21</figureCitation>
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,
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<figureCitation box="[582,606,817,836]" captionStart="Figs" captionStartId="14.[139,183,884,903]" captionTargetBox="[184,1025,180,849]" captionTargetId="figure@14.[179,1030,179,859]" captionTargetPageId="14" captionText="Figs. 30-33. Pygophore, caudal (tenth segment omitted): 30, Brachystethus geniculatus; 31, B. coxalis; 32, B. improvisus; 33, B. vicinus (dor, dorsal rim; pa, paramere; vr, ventral rim). Scale, 1mm." figureDoi="http://doi.org/10.5281/zenodo.3719475" httpUri="https://zenodo.org/record/3719475/files/figure.png" pageId="23" pageNumber="436">31</figureCitation>
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,
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<figureCitation box="[619,643,817,836]" captionStart="Figs" captionStartId="17.[101,144,1245,1264]" captionTargetBox="[157,966,181,1208]" captionTargetId="figure@17.[152,974,177,1222]" captionTargetPageId="17" captionText="Figs. 34-43. Pygophore, ventral rim, ventral: 34, Brachystethus rubromaculatus; 35, B. signoreti; 36, B. cribrus; 37, B. schuhi; 38, B. tricolor; 39, B. vexillum; 40, B. geniculatus; 41, B. coxalis; 42, B. improvisus; 43, B. vicinus. Scale, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3719477" httpUri="https://zenodo.org/record/3719477/files/figure.png" pageId="23" pageNumber="436">41</figureCitation>
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,
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<figureCitation box="[656,680,817,836]" captionStart="Figs" captionStartId="26.[139,180,930,949]" captionTargetBox="[144,1054,182,912]" captionTargetId="figure@26.[139,1062,177,921]" captionTargetPageId="26" captionText="Figs. 61-64. Genital plates, ventral: 61, Brachystethus geniculatus; 62, B. coxalis; 63, B. improvisus; 64, B. vicinus (gc8, gonocoxite 8; gc9, gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; VII, seventh segment; X, tenth segment). Scale, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3719485" httpUri="https://zenodo.org/record/3719485/files/figure.png" pageId="23" pageNumber="436">62</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[101,940,865,884]" pageId="23" pageNumber="436" type="reference_group">
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<paragraph blockId="23.[101,940,865,884]" box="[101,940,865,884]" pageId="23" pageNumber="436">
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<treatmentCitationGroup box="[101,940,865,884]" pageId="23" pageNumber="436">
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<treatmentCitation author="BREDDIN" box="[101,479,865,884]" page="178" pageId="23" pageNumber="436" year="1904">
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<taxonomicName authority="BREDDIN, 1904" authorityName="Breddin." authorityYear="1904" box="[101,443,865,884]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="436" phylum="Arthropoda" rank="species" species="coxalis">
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<emphasis box="[101,296,865,884]" italics="true" pageId="23" pageNumber="436">Brachystethus coxalis</emphasis>
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BREDDIN, 1904
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</taxonomicName>
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:178
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</treatmentCitation>
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;
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<treatmentCitation author="BERGROTH" box="[494,680,865,884]" page="178" pageId="23" pageNumber="436" year="1908">
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<bibRefCitation author="BERGROTH, E." box="[494,680,865,884]" journalOrPublisher="Mem. Soc. r. ent. Belg., Bruxelles" pageId="23" pageNumber="436" pagination="131 - 200" part="15" refId="ref19790" refString="BERGROTH, E. 1908. Enumeratio Pentatomidarum post catalogum Bruxellensem descriptarum. Mem. Soc. r. ent. Belg., Bruxelles, 15: 131 - 200." title="Enumeratio Pentatomidarum post catalogum Bruxellensem descriptarum" type="journal article" year="1908">BERGROTH, 1908:178</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="KIRKALDY" box="[695,878,865,884]" page="152" pageId="23" pageNumber="436" year="1909">
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<bibRefCitation author="KIRKALDY, G. W." bookContentInfo="392 p." box="[695,878,865,884]" journalOrPublisher="Berlin, Felix L. Dames" pageId="23" pageNumber="436" refId="ref20822" refString="KIRKALDY, G. W. 1909. Catalogue of the Hemiptera (Heteroptera). 1. Cimicidae. Berlin, Felix L. Dames. 392 p." title="Catalogue of the Hemiptera (Heteroptera). 1. Cimicidae" type="book" year="1909">KIRKALDY, 1909:152</bibRefCitation>
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</treatmentCitation>
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(cat.).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="436" type="materials_examined">
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<paragraph blockId="23.[101,1032,908,1517]" pageId="23" pageNumber="436">
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Typematerial.
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<materialsCitation ID-GBIF-Occurrence="2579424918" country="Peru" location="Amazonas" pageId="23" pageNumber="436" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="lectotype">
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<typeStatus box="[337,446,908,931]" pageId="23" pageNumber="436">Lectotype</typeStatus>
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, heredesignated, labeled: a)
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<collectingRegion box="[808,926,908,931]" country="Peru" name="Amazonas" pageId="23" pageNumber="436">Amazonas</collectingRegion>
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,
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<collectingCountry box="[940,994,908,931]" name="Peru" pageId="23" pageNumber="436" value="Peru">Peru</collectingCountry>
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; b)
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<taxonomicName authority="Bredd." authorityName="Bredd." box="[101,401,937,960]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="436" phylum="Arthropoda" rank="species" species="coxalis">
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<emphasis box="[101,323,937,960]" italics="true" pageId="23" pageNumber="436">Brachystethus coxalis</emphasis>
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Bredd.
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</taxonomicName>
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; coll. Breddin
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2579424955" collectionCode="DEIC" country="Peru" location="Chiriqui" municipality="Chiriqui" pageId="23" pageNumber="436" specimenCount="1" stateProvince="Amazonas" typeStatus="syntipus">
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d)
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<typeStatus box="[578,671,937,960]" pageId="23" pageNumber="436">Syntipus</typeStatus>
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; e) DEI Eberswalde; f) 1 ST
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<emphasis box="[959,1031,937,960]" italics="true" pageId="23" pageNumber="436">coxalis</emphasis>
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(
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<collectionCode box="[109,174,966,989]" collectionName="Germany, Muencheberg, Deutsches Entomologisches Institut im ZALF" pageId="23" pageNumber="436">DEIC</collectionCode>
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). Breddin described the species based on two specimens, one male and one female. Only the male syntype was found.
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</materialsCitation>
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Type locality:
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<collectingCountry box="[595,642,995,1018]" name="Peru" pageId="23" pageNumber="436" value="Peru">Peru</collectingCountry>
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(
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<collectingRegion box="[654,766,995,1018]" country="Peru" name="Amazonas" pageId="23" pageNumber="436">Amazonas</collectingRegion>
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) and
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<collectingCountry box="[820,902,995,1018]" name="Panama" pageId="23" pageNumber="436">Panama</collectingCountry>
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(Chiriquí).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="25" lastPageNumber="438" pageId="23" pageNumber="436" type="description">
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<paragraph blockId="23.[101,1032,908,1517]" lastBlockId="24.[139,1069,1247,1533]" lastPageId="24" lastPageNumber="437" pageId="23" pageNumber="436">
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Description. Dorsal surface dark-castaneous to black, with a marginal yellow stripe (
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<figureCitation box="[109,178,1054,1077]" pageId="23" pageNumber="436">fig. 11</figureCitation>
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). Head dark-castaneous, with well-marked wrinkles. Puncturesdensely distributed. Outer margins of jugae with yellowish stripe, wider at base. Antennae dark-castaneous to black, except the first segment, yellowish. Fourth segment cylindrical. First antennal segment a little more than 3/4 as long as second, third almost twice as long as second, fourth slightly less than one and a half as long as third, fifth lacking. Pronotum darkcastaneous to black, with a broad yellow stripe along anterolateral margins. Punctures fine regularly distributed, with sparse coarse punctures restrict tothe area after cicatrices. Before cicatrices, smallerpunctures more concentrated alonganterior marginof pronotum. Scutellum dark-castaneous to black, with fine punctures regularly distributed. Coarse punctures on basal third, concentrated in 1+1 groups along lateral margins. Punctures slightly smaller, sparse, on basal half. Corium dark-castaneous, with yellow border on basal third of costal margin and with a small yellowish callus near and entad the apex of radial vein. Legs yellowish on coxae, trochanters, base and apex of femora, ventral face and apex of tibiae and tarsi; remaining segments dark-castaneous, almost black. Ventral surface of thorax dark-castaneous to black, with castaneous to black medium-sized punctures. Ostiolar rugae reaching 2/3 of distance between inner margin of ostiolumand lateral margin of metapleura. Metasternal carina flat in profile. Connexivum with a subcaloused yellow stripe, scarcely punctured, over outer lateral third of each segment; remaining of connexival segments dark-castaneous, almost black, with punctures minutely and densely distributed. Venter dark-castaneous, with a marginal, subcaloused yellowish stripe. Punctures fine regularly distributed, except on the middle, where they are less frequent.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3719483" ID-Zenodo-Dep="3719483" httpUri="https://zenodo.org/record/3719483/files/figure.png" pageId="24" pageNumber="437" startId="24.[139,184,1161,1180]" targetBox="[176,1022,181,1137]" targetPageId="24">
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<paragraph blockId="24.[139,1071,1161,1228]" pageId="24" pageNumber="437">
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Figs.56-60. Genital plates, ventral: 56,
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<taxonomicName authorityName="Dallas" authorityYear="1851" box="[530,824,1161,1180]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="437" phylum="Arthropoda" rank="species" species="rubromaculatus">
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<emphasis box="[530,824,1161,1180]" italics="true" pageId="24" pageNumber="437">Brachystethus rubromaculatus</emphasis>
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</taxonomicName>
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; 57,
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<taxonomicName authorityName="Stal" authorityYear="1872" box="[879,993,1161,1180]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="437" phylum="Arthropoda" rank="species" species="signoreti">
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<emphasis box="[879,993,1161,1180]" italics="true" pageId="24" pageNumber="437">B. signoreti</emphasis>
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</taxonomicName>
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; 58,
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1781" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="437" phylum="Arthropoda" rank="species" species="cribrus">
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<emphasis italics="true" pageId="24" pageNumber="437">B. cribrus</emphasis>
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</taxonomicName>
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; 59,
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<taxonomicName authorityName="Bolívar" authorityYear="1879" box="[255,352,1185,1204]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="437" phylum="Arthropoda" rank="species" species="tricolor">
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<emphasis box="[255,352,1185,1204]" italics="true" pageId="24" pageNumber="437">B. tricolor</emphasis>
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</taxonomicName>
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; 60,
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<taxonomicName authorityName="Breddin" authorityYear="1903" box="[403,507,1185,1204]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="437" phylum="Arthropoda" rank="species" species="vexillum">
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<emphasis box="[403,507,1185,1204]" italics="true" pageId="24" pageNumber="437">B. vexillum</emphasis>
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</taxonomicName>
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(gc8, gonocoxite 8; gc9, gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; VII, seventhsegment; X, tenthsegment). Scale, 1 mm.
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</paragraph>
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</caption>
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<paragraph blockId="24.[139,1069,1247,1533]" pageId="24" pageNumber="437">
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Male genitalia. Pygophore quadrangular. Lobes of dorsal rim conspicuous, moderately projected posterad, rounded at apex. Genital cup processes not visible in dorsal view. Parameres withacute apex, turned anterad (
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<figureCitation box="[731,853,1480,1503]" pageId="24" pageNumber="437">figs. 21, 31</figureCitation>
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). Ventral rim openly V-like (
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<figureCitation box="[214,281,1510,1533]" pageId="24" pageNumber="437">fig. 41</figureCitation>
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).
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</paragraph>
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<paragraph blockId="25.[101,1032,190,711]" pageId="25" pageNumber="438">Measurements (n=1). Headlength 2.03, width 3.00; interoculardistance 1.64; anteocularlength 0.74; antennalsegments - lacking; pronotallength 4.34, width 10.02; scutellarlength 6.40, width 6.24; corium length, not measured; abdominalwidth 11.19; totallength 15.87.</paragraph>
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<paragraph blockId="25.[101,1032,190,711]" pageId="25" pageNumber="438">
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Female genitalia. Posteriorborders of gonocoxites 8 forming an almost rightangle over laterotergites 9. Sutural borders of gonocoxites 8 parallel (
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<figureCitation box="[770,843,336,359]" pageId="25" pageNumber="438">fig. 62</figureCitation>
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).
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</paragraph>
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<paragraph blockId="25.[101,1032,190,711]" pageId="25" pageNumber="438">Measurements (n=1). Headlength 2.00, width 3.04; interoculardistance 1.60; anteocularlength 0.72; lengthof antennalsegments: I - 0.88; II - 1.12; III - 2.00; IV - 2.80; V - lacking; pronotallength 4.18, width 10.02; scutellarlength 6.68, width 6.51; corium length 9.52; abdominalwidth 11.11; totallength 15.70.</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="438" type="materials_examined">
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<paragraph blockId="25.[101,1032,190,711]" pageId="25" pageNumber="438">
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<materialsCitation ID-GBIF-Occurrence="2579424885" collectingDate="1984-02-11" collectionCode="AMNH, ZMUC" country="Venezuela" elevation="1880" location="Cerro de la Neblina" municipality="Cerro de la Neblina" pageId="25" pageNumber="438" specimenCount="1" stateProvince="Amazonas">
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Material examined.
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<collectingCountry box="[381,542,483,506]" name="Venezuela" pageId="25" pageNumber="438" value="Venezuela">VENEZUELA</collectingCountry>
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,
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<emphasis bold="true" box="[554,672,483,507]" pageId="25" pageNumber="438">
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<collectingRegion box="[554,672,483,507]" country="Venezuela" name="Amazonas" pageId="25" pageNumber="438">Amazonas</collectingRegion>
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</emphasis>
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:
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<collectingMunicipality box="[687,901,483,506]" pageId="25" pageNumber="438">Cerro de la Neblina</collectingMunicipality>
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(
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<quantity box="[916,995,483,506]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="25" pageNumber="438" unit="m" value="1880.0">
|
|
<elevation box="[916,995,483,506]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="25" pageNumber="438" unit="m" value="1880.0">1880m</elevation>
|
|
</quantity>
|
|
),,
|
|
<date box="[101,212,512,535]" pageId="25" pageNumber="438" value="1984-02-11">
|
|
<collectingDate box="[101,212,512,535]" pageId="25" pageNumber="438" value="1984-02-11">11.II.1984</collectingDate>
|
|
</date>
|
|
, Stupakoff (
|
|
<collectionCode box="[337,423,512,535]" collectionName="USA, New York, New York, American Museum of Natural History" pageId="25" pageNumber="438">AMNH</collectionCode>
|
|
). No data, (
|
|
<collectionCode box="[557,639,512,535]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" pageId="25" pageNumber="438">ZMUC</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection box="[165,956,542,565]" pageId="25" pageNumber="438" type="distribution">
|
|
<paragraph blockId="25.[101,1032,190,711]" box="[165,956,542,565]" pageId="25" pageNumber="438">Distribution. Panama (Chiriquí), Venezuela (Amazonas), Peru (Amazonas).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="25" pageNumber="438" type="discussion">
|
|
<paragraph blockId="25.[101,1032,190,711]" pageId="25" pageNumber="438">
|
|
Comments.
|
|
<taxonomicName authorityName="Bredd." box="[290,517,571,594]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="438" phylum="Arthropoda" rank="species" species="coxalis">
|
|
<emphasis box="[290,517,571,594]" italics="true" pageId="25" pageNumber="438">Brachystethus coxalis</emphasis>
|
|
</taxonomicName>
|
|
is the sister group of
|
|
<taxonomicName authorityName="Signoret" authorityYear="1851" box="[733,835,571,594]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="438" phylum="Arthropoda" rank="species" species="vicinus">
|
|
<emphasis box="[733,835,571,594]" italics="true" pageId="25" pageNumber="438">B. vicinus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authorityName="Breddin" authorityYear="1905" box="[882,1025,571,594]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="438" phylum="Arthropoda" rank="species" species="improvisus">
|
|
<emphasis box="[882,1025,571,594]" italics="true" pageId="25" pageNumber="438">B. improvisus</emphasis>
|
|
</taxonomicName>
|
|
, sharing with them one synapomorphy, the yellowish tarsi, and two homoplastic characters, coxae, and apex of femora and base of tibiae yellowish (
|
|
<bibRefCitation author="BARCELLOS, A. & GRAZIA, J." journalOrPublisher="Zootaxa, Auckland" pageId="25" pageNumber="438" pagination="1 - 14" part="256" refId="ref19723" refString="BARCELLOS, A. & GRAZIA, J. 2003. Cladistic analysis and biogeography of Brachystethus Laporte (Heteroptera, Pentatomidae, Edessinae). Zootaxa, Auckland, 256: 1 - 14." title="Cladistic analysis and biogeography of Brachystethus Laporte (Heteroptera, Pentatomidae, Edessinae)" type="journal article" year="2003">BARCELLOS & GRAZIA, 2003</bibRefCitation>
|
|
). It can be distinguished from
|
|
<taxonomicName authorityName="Signoret" authorityYear="1851" box="[483,588,659,682]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="438" phylum="Arthropoda" rank="species" species="vicinus">
|
|
<emphasis box="[483,588,659,682]" italics="true" pageId="25" pageNumber="438">B. vicinus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authorityName="Breddin" authorityYear="1905" box="[639,786,659,682]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="438" phylum="Arthropoda" rank="species" species="improvisus">
|
|
<emphasis box="[639,786,659,682]" italics="true" pageId="25" pageNumber="438">B. improvisus</emphasis>
|
|
</taxonomicName>
|
|
by the flat metasternal carina, and from the latter also by the shape of parameres, with acute apex.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |