935 lines
110 KiB
XML
935 lines
110 KiB
XML
<document id="1977EB5254B4E86D53E1CBBB34C48122" ID-DOI="10.11646/zootaxa.5319.1.5" ID-GBIF-Dataset="6bad9ed2-56d4-4e36-90d3-2802e325a57c" ID-ISSN="1175-5326" ID-Zenodo-Dep="10124970" ID-ZooBank="C15698EF-A99E-431F-9453-FC82FEF0ECC4" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690276967882" checkinUser="plazi" docAuthor="Xu, Yu-Hao, Yang, Dian-Cheng, Gong, Yan-An, Wu, Jia-Xiang, Huang, Ru-Yi, Liu, Yong-Jin, Liang, Sheng-Ming, Huang, Tian-Qi & Huang, Song" docDate="2023" docId="2166A771FFF8C978CFADFE17FA032B88" docLanguage="en" docName="zootaxa.5319.1.5.pdf" docOrigin="Zootaxa 5319 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hebius youjiangensis Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang 2023, sp. nov." docType="treatment" docVersion="11" lastPageNumber="88" masterDocId="DD5FDF09FFFFC974CF3AFF85FF942A25" masterDocTitle="A new species of the Genus Hebius Thompson, 1913 (Squamata: Colubridae) from Baise, Guangxi, China" masterLastPageNumber="90" masterPageNumber="76" pageNumber="83" updateTime="1700051107319" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FE7754959803416E0EB776A38A7D5C73">A new species of the Genus Hebius Thompson, 1913 (Squamata: Colubridae) from Baise, Guangxi, China</mods:title>
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<mods:namePart id="94C63FE03233FB1C30DC593D3E877CE4">Xu, Yu-Hao</mods:namePart>
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<mods:affiliation id="BC26A7CB21DAB9D6EAFBB6744ED6950E">Anhui Province Key Laboratory of the Conservation and Exploitation of Biological Resource, College of Life Sciences, Anhui Normal University, Wuhu 241000, China & Huangshan Noah Biodiversity Institute, Huangshan 245000, China</mods:affiliation>
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<mods:title id="45E46FA7828509F729C60D1E1AE9B015">Zootaxa</mods:title>
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<treatment id="2166A771FFF8C978CFADFE17FA032B88" ID-GBIF-Taxon="212421066" LSID="urn:lsid:plazi:treatment:2166A771FFF8C978CFADFE17FA032B88" httpUri="http://treatment.plazi.org/id/2166A771FFF8C978CFADFE17FA032B88" lastPageId="12" lastPageNumber="88" pageId="7" pageNumber="83">
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<heading id="F238A10BFFF8C973CFADFE17FBE52B89" bold="true" box="[151,1137,402,429]" fontSize="11" level="1" pageId="7" pageNumber="83" reason="1">
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<taxonomicName id="6ECF6DE4FFF8C973CFADFE17FE0F2B88" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[151,411,402,429]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="7" pageNumber="83" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
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<emphasis id="9BBBCA75FFF8C973CFADFE17FE0F2B88" bold="true" box="[151,411,402,429]" italics="true" pageId="7" pageNumber="83">Hebius youjiangensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8088770EFFF8C973CE98FE16FD942B88" box="[418,512,403,429]" pageId="7" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
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Yang, Xu, Wu, Gong, Huang & Huang (
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<figureCitation id="31F40AE2FFF8C973CCCDFE17FBFE2B89" box="[1015,1130,402,428]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="7.[152,255,1739,1763]" captionStartId-1="8.[152,255,1110,1134]" captionStartId-2="9.[152,255,1497,1521]" captionTargetBox-0="[151,1435,993,1716]" captionTargetBox-1="[151,1435,181,1086]" captionTargetBox-2="[151,1436,181,1473]" captionTargetId-0="figure-331@7.[151,1435,993,1716]" captionTargetId-1="figure-19@8.[151,1435,181,1086]" captionTargetId-2="figure-23@9.[151,1436,181,1473]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 3. General view of the holotype (ANU20220010) of Hebius youjiangensis sp. nov. in life. Photo by Dian-Cheng Yang & Yan-An Gong." captionText-1="FIGURE 4. Dorsal (A) and ventral (B) views of the holotype (ANU20220010) of Hebius youjiangensis sp. nov. in preservative. Photos by Dian-Cheng Yang." captionText-2="FIGURE 5. Right (A), left (B), ventral (C) and dorsal (D) views of the head, the anterior part of venter (E), and lateral view of dorsal body (F) of the holotype (ANU20220010) of Hebius youjiangensis sp. nov. in life. Photos by Dian-Cheng Yang & Yan-An Gong." figureDoi-0="http://doi.org/10.5281/zenodo.10135905" figureDoi-1="http://doi.org/10.5281/zenodo.10135907" figureDoi-2="http://doi.org/10.5281/zenodo.10135909" httpUri-0="https://zenodo.org/record/10135905/files/figure.png" httpUri-1="https://zenodo.org/record/10135907/files/figure.png" httpUri-2="https://zenodo.org/record/10135909/files/figure.png" pageId="7" pageNumber="83">Figs. 3–5</figureCitation>
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)
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="E1D545ECFFF8C973CFADFE5EFF792844" pageId="7" pageNumber="83" type="materials_examined">
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<materialsCitation id="19A71C3AFFF8C973CFADFE5EFF792844" ID-GBIF-Occurrence="4164457301" collectingDate="2022-11-28" collectionCode="ANU, HSR" collectorName="Jiaxiang Wu & Yongjin Liu" country="China" county="Autonomous Region" elevation="780" latitude="23.735281" location="Daleng Village" longLatPrecision="1" longitude="106.39533" municipality="Baise City" pageId="7" pageNumber="83" specimenCode="ANU20220010, HSR22184" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
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<typeStatus id="7674A8C5FFF8C973CFADFE5EFE922BD0" box="[151,262,475,501]" pageId="7" pageNumber="83" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="F969BE1CFFF8C973CE2EFE5EFE542BD0" box="[276,448,475,501]" collectionCode="ANU" country="Australia" httpUri="http://grbio.org/cool/xny3-ga2w" name="Australian National University" pageId="7" pageNumber="83">ANU20220010</specimenCode>
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(collection number:
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<specimenCode id="F969BE1CFFF8C973CD8EFE5EFCA22BD0" box="[692,822,475,501]" collectionCode="HSR" pageId="7" pageNumber="83">HSR22184</specimenCode>
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), an adult female, was collected by
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<collectorName id="043A73B1FFF8C973CBE1FE5EFAFD2BD0" box="[1243,1385,475,501]" pageId="7" pageNumber="83">Jiaxiang Wu</collectorName>
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and
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<collectorName id="043A73B1FFF8C973CFADFE7AFEB7283C" box="[151,291,511,537]" pageId="7" pageNumber="83">Yongjin Liu</collectorName>
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, on
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<date id="DD7130A7FFF8C973CE6FFE7AFDAF283C" box="[341,571,511,537]" pageId="7" pageNumber="83" value="2022-11-28">
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<collectingDate id="CD35C94FFFF8C973CE6FFE7AFDAF283C" box="[341,571,511,537]" pageId="7" pageNumber="83" value="2022-11-28">November 28, 2022</collectingDate>
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</date>
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, at night, in
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,
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|
||
,
|
||
<collectingMunicipality id="49148C1DFFF8C973CB62FE7AFB44283C" box="[1112,1232,511,537]" pageId="7" pageNumber="83">Baise City</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="6B0BD885FFF8C973CBE7FE7AFA09283C" box="[1245,1437,511,537]" country="China" name="Guangxi" pageId="7" pageNumber="83">Guangxi Zhuang</collectingRegion>
|
||
<collectingCounty id="40116EEBFFF8C973CFADFDA6FE122818" box="[151,390,547,573]" pageId="7" pageNumber="83">Autonomous Region</collectingCounty>
|
||
,
|
||
<collectingCountry id="D1D856F7FFF8C973CEA9FDA6FE4D2818" box="[403,473,547,573]" name="China" pageId="7" pageNumber="83">China</collectingCountry>
|
||
, (
|
||
<geoCoordinate id="CCFB70A0FFF8C973CED5FDA1FD032818" box="[495,663,547,575]" degrees="23" direction="north" minutes="44" orientation="latitude" pageId="7" pageNumber="83" precision="1" seconds="7.01" value="23.735281">23°44′7.01″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CCFB70A0FFF8C973CD9FFDA1FCF72818" box="[677,867,547,575]" degrees="106" direction="east" minutes="23" orientation="longitude" pageId="7" pageNumber="83" precision="1" seconds="43.21" value="106.39533">106°23′43.21″ E</geoCoordinate>
|
||
,
|
||
<elevation id="22E2F154FFF8C973CC48FDA6FB992818" box="[882,1037,547,573]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="7" pageNumber="83" unit="m" value="780.0">
|
||
<quantity id="6E37BB82FFF8C973CC48FDA6FC282818" box="[882,956,547,573]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="7" pageNumber="83" unit="m" value="780.0">780 m</quantity>
|
||
a. s. l.
|
||
</elevation>
|
||
), with an air temperature of about 17.6°C.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E1D545ECFFF8C973CFFDFDEFFF4F29EC" pageId="7" pageNumber="83" type="diagnosis">
|
||
<paragraph id="A9701667FFF8C973CFFDFDEFFF4F29EC" blockId="7.[151,1437,475,969]" pageId="7" pageNumber="83">
|
||
<emphasis id="9BBBCA75FFF8C973CFFDFDEFFED628A1" bold="true" box="[199,322,618,644]" pageId="7" pageNumber="83">Diagnosis.</emphasis>
|
||
<taxonomicName id="6ECF6DE4FFF8C973CE7DFDEEFDA228A1" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[327,566,619,645]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="7" pageNumber="83" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF8C973CE7DFDEEFDA228A1" box="[327,566,619,645]" italics="true" pageId="7" pageNumber="83">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF8C973CD06FDEEFD0028A0" bold="true" box="[572,660,619,645]" pageId="7" pageNumber="83">
|
||
<taxonomicNameLabel id="8088770EFFF8C973CD06FDEEFD0028A0" box="[572,660,619,645]" pageId="7" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from its congeners by the following combination of characters: (1) dorsal scale rows 19-17-17, feebly keeled except the outermost rows; (2) tail length comparatively long, TAL/TL ratio
|
||
<quantity id="6E37BB82FFF8C973CE4EFD36FE5528E8" box="[372,449,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="83" unit="in" value="0.3">0.30 in</quantity>
|
||
females; (3) ventrals 3+160; (4) subcaudals 112; (5) supralabials 9, the fourth to sixth in contact with the eye, the seventh to ninth largest; (6) infralabials 10, the first 5 touching the first pair of chin shields; (7) preocular 1; (8) postoculars 2; (9) temporals 4, arranged in three rows (1+1+2); (10) maxillary teeth 30, the last 3 enlarged, without diastema; (11) postocular streak presence; (12) background color of dorsal brownish black, a continuous beige stripe extending from behind the eye to the end of the tail on 5
|
||
<superScript id="5EBABB2FFFF8C973CB34FCC7FB8F2975" attach="left" box="[1038,1051,834,848]" fontSize="6" pageId="7" pageNumber="83">th</superScript>
|
||
to 7
|
||
<superScript id="5EBABB2FFFF8C973CB71FCC7FBCC2975" attach="left" box="[1099,1112,834,848]" fontSize="6" pageId="7" pageNumber="83">th</superScript>
|
||
scale rows in anterior part of body, and 4
|
||
<superScript id="5EBABB2FFFF8C973CE22FCE3FEB12951" attach="left" box="[280,293,870,884]" fontSize="6" pageId="7" pageNumber="83">th</superScript>
|
||
to 6
|
||
<superScript id="5EBABB2FFFF8C973CE6FFCE3FEF62951" attach="left" box="[341,354,870,884]" fontSize="6" pageId="7" pageNumber="83">th</superScript>
|
||
in middle and posterior part of body; (13) anterior venter creamish-yellow, gradually fades to the rear, with irregular black blotches in the middle and outer quarter of ventrals, the posterior part almost completely black.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="FDB046EFFFF8C973CFA2F94EFED12D22" ID-DOI="http://doi.org/10.5281/zenodo.10135905" ID-Zenodo-Dep="10135905" httpUri="https://zenodo.org/record/10135905/files/figure.png" pageId="7" pageNumber="83" startId="7.[152,255,1739,1763]" targetBox="[151,1435,993,1716]" targetPageId="7" targetType="figure">
|
||
<paragraph id="A9701667FFF8C973CFA2F94EFED12D22" blockId="7.[151,1436,1739,1799]" pageId="7" pageNumber="83">
|
||
<emphasis id="9BBBCA75FFF8C973CFA2F94EFE832CC6" bold="true" box="[152,279,1739,1763]" pageId="7" pageNumber="83">FIGURE 3.</emphasis>
|
||
General view of the holotype (ANU20220010) of
|
||
<taxonomicName id="6ECF6DE4FFF8C973CC2BF94EFC7D2CC6" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[785,1001,1739,1763]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="7" pageNumber="83" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF8C973CC2BF94EFC7D2CC6" box="[785,1001,1739,1763]" italics="true" pageId="7" pageNumber="83">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF8C973CCD4F94EFBA92CC6" bold="true" box="[1006,1085,1739,1763]" pageId="7" pageNumber="83">
|
||
<taxonomicNameLabel id="8088770EFFF8C973CCD4F94EFBA92CC6" box="[1006,1085,1739,1763]" pageId="7" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life. Photo by Dian-Cheng Yang & Yan-An Gong.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E1D545ECFFF8C97ECFFDF8DCFC632AF0" lastPageId="10" lastPageNumber="86" pageId="7" pageNumber="83" type="description">
|
||
<paragraph id="A9701667FFF8C973CFFDF8DCFCFA2DFA" blockId="7.[151,1437,1881,2016]" pageId="7" pageNumber="83">
|
||
<emphasis id="9BBBCA75FFF8C973CFFDF8DCFE712D56" bold="true" box="[199,485,1881,1907]" pageId="7" pageNumber="83">
|
||
Description of
|
||
<typeStatus id="7674A8C5FFF8C973CE42F8DCFE752D56" box="[376,481,1881,1907]" pageId="7" pageNumber="83" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
The adult female specimen has the following characters: TL
|
||
<quantity id="6E37BB82FFF8C973CBACF8DCFB602D51" box="[1174,1268,1881,1908]" metricMagnitude="-1" metricUnit="m" metricValue="5.82" pageId="7" pageNumber="83" unit="mm" value="582.0">582 mm</quantity>
|
||
(SVL
|
||
<quantity id="6E37BB82FFF8C973CA05F8DCFA092D51" box="[1343,1437,1881,1908]" metricMagnitude="-1" metricUnit="m" metricValue="4.09" pageId="7" pageNumber="83" unit="mm" value="409.0">409 mm</quantity>
|
||
and TAL
|
||
<quantity id="6E37BB82FFF8C973CE3CF8F8FEFD2DB2" box="[262,361,1917,1943]" metricMagnitude="-1" metricUnit="m" metricValue="1.73" pageId="7" pageNumber="83" unit="mm" value="173.0">173 mm</quantity>
|
||
), tail complete, TAL/TL ratio 0.30. Body elongated and cylindrical; head flattened anteriorly, distinct from the neck, HL
|
||
<quantity id="6E37BB82FFF8C973CEFFF824FDAE2D99" box="[453,570,1953,1980]" metricMagnitude="-2" metricUnit="m" metricValue="1.6949999999999998" pageId="7" pageNumber="83" unit="mm" value="16.95">16.95 mm</quantity>
|
||
, HW
|
||
<quantity id="6E37BB82FFF8C973CD40F824FD752D99" box="[634,737,1953,1980]" metricMagnitude="-3" metricUnit="m" metricValue="8.82" pageId="7" pageNumber="83" unit="mm" value="8.82">8.82 mm</quantity>
|
||
. Nostril lateral, round, piercing in the middle of the nasal. Eye large, ED
|
||
<quantity id="6E37BB82FFF8C973CE33F840FEF52DC5" box="[265,353,1989,2016]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="83" unit="mm" value="2.3">2.3 mm</quantity>
|
||
, pupil round. Tail complete, long and tapering.
|
||
</paragraph>
|
||
<caption id="FDB046EFFFF7C97CCFA2FBD3FE2B2EB6" ID-DOI="http://doi.org/10.5281/zenodo.10135907" ID-Zenodo-Dep="10135907" httpUri="https://zenodo.org/record/10135907/files/figure.png" pageId="8" pageNumber="84" startId="8.[152,255,1110,1134]" targetBox="[151,1435,181,1086]" targetPageId="8" targetType="figure">
|
||
<paragraph id="A9701667FFF7C97CCFA2FBD3FE2B2EB6" blockId="8.[151,1437,1110,1171]" pageId="8" pageNumber="84">
|
||
<emphasis id="9BBBCA75FFF7C97CCFA2FBD3FE832E4B" bold="true" box="[152,279,1110,1134]" pageId="8" pageNumber="84">FIGURE 4.</emphasis>
|
||
Dorsal (A) and ventral (B) views of the holotype (ANU20220010) of
|
||
<taxonomicName id="6ECF6DE4FFF7C97CCCEAFBD3FB3C2E4B" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[976,1192,1110,1134]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="8" pageNumber="84" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF7C97CCCEAFBD3FB3C2E4B" box="[976,1192,1110,1134]" italics="true" pageId="8" pageNumber="84">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF7C97CCB97FBD3FB682E4B" bold="true" box="[1197,1276,1110,1134]" pageId="8" pageNumber="84">
|
||
<taxonomicNameLabel id="8088770EFFF7C97CCB97FBD3FB682E4B" box="[1197,1276,1110,1134]" pageId="8" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in preservative. Photos by Dian-Cheng Yang.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A9701667FFF7C97CCFFDFB53FAC42F36" blockId="8.[151,1437,1238,2040]" pageId="8" pageNumber="84">
|
||
<emphasis id="9BBBCA75FFF7C97CCFFDFB53FEE32ED5" box="[199,375,1238,1264]" italics="true" pageId="8" pageNumber="84">Body scalation.</emphasis>
|
||
DSR 19-17-17, feebly keeled posterior to nape, and outermost dorsal scale row on both sides smooth, not notched at their posterior extremity. VEN 160 (+ 3 preventrals); SC 112, all paired; CP divided.
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF7C97CCFFDFA98FD942F12" blockId="8.[151,1437,1238,2040]" box="[199,512,1309,1335]" pageId="8" pageNumber="84">
|
||
<emphasis id="9BBBCA75FFF7C97CCFFDFA98FD942F12" box="[199,512,1309,1335]" italics="true" pageId="8" pageNumber="84">Dorsal scale row reduction.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF7C97CCFF4FAC5FE5A2F7F" blockId="8.[151,1437,1238,2040]" box="[206,462,1344,1370]" pageId="8" pageNumber="84">3+4→3 (76-77) (right)</paragraph>
|
||
<paragraph id="A9701667FFF7C97CCFADFAE6FD972F58" blockId="8.[151,1437,1238,2040]" box="[151,515,1379,1405]" pageId="8" pageNumber="84">19———————————17</paragraph>
|
||
<paragraph id="A9701667FFF7C97CCFF4FA03FE2A2F85" blockId="8.[151,1437,1238,2040]" box="[206,446,1414,1441]" pageId="8" pageNumber="84">3+4→3 (75-76) (left)</paragraph>
|
||
<paragraph id="A9701667FFF7C97CCFFDFA2FFDB62FC2" blockId="8.[151,1437,1238,2040]" pageId="8" pageNumber="84">
|
||
<emphasis id="9BBBCA75FFF7C97CCFFDFA2FFEA52FE6" box="[199,305,1450,1475]" italics="true" pageId="8" pageNumber="84">Dentition</emphasis>
|
||
. Maxillary teeth 30, gradually enlarging, the last 3 moderately enlarged, without diastema between the last three and the anterior teeth.
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF7C97CCFFDFA75FE752D62" blockId="8.[151,1437,1238,2040]" pageId="8" pageNumber="84">
|
||
<emphasis id="9BBBCA75FFF7C97CCFFDFA75FEED2C2F" box="[199,377,1520,1546]" italics="true" pageId="8" pageNumber="84">Head scalation.</emphasis>
|
||
Rostral visible dorsally, trapezoidal, wider than high; nasals nearly rectangular, about twice as wide as high, divided into two parts; nostril in the anterior part of the nasal, posterior margin of nostril with a distinct nostril cleft, the posterior section of nasal vertically subrectangle, about half as wide as the anterior part; internasals small, subtriangular, in broad contact with each other, narrowing anteriorly; prefrontals 2, subpentagonal, wider than long, in contact with loreal; frontal narrow, hexagonal, longer than wide, shield-like; supraocular 1/1, much wider than high; LOR 1/1, subrectangular, wider than long; PRO 1/1; PO 2/2, upper one larger than lower one; SL 9/9, the first
|
||
<quantity id="6E37BB82FFF7C97CCE10F946FEC32CF8" box="[298,343,1731,1757]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="84" unit="in" value="2.0">2 in</quantity>
|
||
contact with nasal, the 2
|
||
<superScript id="5EBABB2FFFF7C97CCD4EF946FD102CF4" attach="left" box="[628,644,1731,1745]" fontSize="6" pageId="8" pageNumber="84">nd</superScript>
|
||
to 4
|
||
<superScript id="5EBABB2FFFF7C97CCD83F946FD522CF4" attach="left" box="[697,710,1731,1745]" fontSize="6" pageId="8" pageNumber="84">th</superScript>
|
||
in contact with the loreal, 4
|
||
<superScript id="5EBABB2FFFF7C97CCB32F946FB812CF4" attach="left" box="[1032,1045,1731,1745]" fontSize="6" pageId="8" pageNumber="84">th</superScript>
|
||
to 6
|
||
<superScript id="5EBABB2FFFF7C97CCB73F946FBC22CF4" attach="left" box="[1097,1110,1731,1745]" fontSize="6" pageId="8" pageNumber="84">th</superScript>
|
||
entering orbit, the 7
|
||
<superScript id="5EBABB2FFFF7C97CCA05F946FAD82CF4" attach="left" box="[1343,1356,1731,1745]" fontSize="6" pageId="8" pageNumber="84">th</superScript>
|
||
and 9
|
||
<superScript id="5EBABB2FFFF7C97CCAAAF946FA092CF4" attach="none" box="[1424,1437,1731,1745]" fontSize="6" pageId="8" pageNumber="84">th</superScript>
|
||
largest; TEMP 4/4, small, slightly rectangular, arranged in three rows (1+1+2); chin shields in 2 pairs, posterior pair longer than anterior ones; IL 10/10, first pair in contact behind the mental, 1
|
||
<superScript id="5EBABB2FFFF7C97CCCC1F88FFB922D3D" attach="left" box="[1019,1030,1802,1816]" fontSize="6" pageId="8" pageNumber="84">st</superScript>
|
||
to 5
|
||
<superScript id="5EBABB2FFFF7C97CCB00F88CFBD32D32" attach="left" box="[1082,1095,1801,1815]" fontSize="6" pageId="8" pageNumber="84">th</superScript>
|
||
touching the first pair of chin shields, the 5
|
||
<superScript id="5EBABB2FFFF7C97CCE10F8A9FEA32D1F" attach="left" box="[298,311,1836,1850]" fontSize="6" pageId="8" pageNumber="84">th</superScript>
|
||
and 6
|
||
<superScript id="5EBABB2FFFF7C97CCE41F8A9FE1C2D1F" attach="left" box="[379,392,1836,1850]" fontSize="6" pageId="8" pageNumber="84">th</superScript>
|
||
largest.
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF7C97CCFFDF8D5FDB32DD2" blockId="8.[151,1437,1238,2040]" pageId="8" pageNumber="84">
|
||
<emphasis id="9BBBCA75FFF7C97CCFFDF8D5FE022D4F" bold="true" box="[199,406,1872,1898]" pageId="8" pageNumber="84">Coloration in life.</emphasis>
|
||
In life, dorsal surface of head is brownish-black, densely covered by irregular and vermiculate brownish yellow or light brown stripes. A thick, obvious beige postocular streak extends backwards to meet with the dorsolateral stripe on each side. A short, pale yellowish-brown streak on the outside of parietals and the vertebral scales behind the parietals. Supralabials ochre yellow, the first seven trailing edges brownish-black, the 8
|
||
<superScript id="5EBABB2FFFF7C97CCA7DF83CFAC02DE2" attach="left" box="[1351,1364,1977,1991]" fontSize="6" pageId="8" pageNumber="84">th</superScript>
|
||
upper half and 9
|
||
<superScript id="5EBABB2FFFF7C97CCE3DF858FE802DCE" attach="left" box="[263,276,2013,2027]" fontSize="6" pageId="8" pageNumber="84">th</superScript>
|
||
bottom brownish-black.
|
||
</paragraph>
|
||
<caption id="FDB046EFFFF6C97DCFA2FA5CFEB82C1F" ID-DOI="http://doi.org/10.5281/zenodo.10135909" ID-Zenodo-Dep="10135909" httpUri="https://zenodo.org/record/10135909/files/figure.png" pageId="9" pageNumber="85" startId="9.[152,255,1497,1521]" targetBox="[151,1436,181,1473]" targetPageId="9" targetType="figure">
|
||
<paragraph id="A9701667FFF6C97DCFA2FA5CFEB82C1F" blockId="9.[151,1437,1497,1594]" pageId="9" pageNumber="85">
|
||
<emphasis id="9BBBCA75FFF6C97DCFA2FA5CFE8D2FD7" bold="true" box="[152,281,1497,1522]" pageId="9" pageNumber="85">FIGURE 5.</emphasis>
|
||
Right (A), left (B), ventral (C) and dorsal (D) views of the head, the anterior part of venter (E), and lateral view of dorsal body (F) of the holotype (ANU20220010) of
|
||
<taxonomicName id="6ECF6DE4FFF6C97DCDE8FA78FC392C30" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[722,941,1533,1557]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="9" pageNumber="85" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF6C97DCDE8FA78FC392C30" box="[722,941,1533,1557]" italics="true" pageId="9" pageNumber="85">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF6C97DCC8FFA7BFB932C33" bold="true" box="[949,1031,1534,1558]" pageId="9" pageNumber="85">
|
||
<taxonomicNameLabel id="8088770EFFF6C97DCC8FFA7BFB932C33" box="[949,1031,1534,1558]" pageId="9" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life. Photos by Dian-Cheng Yang & Yan-An Gong.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A9701667FFF6C97DCFFDF909FD132D13" blockId="9.[151,1437,1676,2026]" pageId="9" pageNumber="85">
|
||
Body brownish-black, middle darker than the sides. Dorsal scales of the body brownish-black on both sides, and pale yellowish-brown in the middle. A conspicuous, continuous beige stripe extending from behind the eye to the end of the tail on the upper part of 5
|
||
<superScript id="5EBABB2FFFF6C97DCD17F956FDAE2CC4" attach="left" box="[557,570,1747,1761]" fontSize="6" pageId="9" pageNumber="85">th</superScript>
|
||
to the lower part of 7
|
||
<superScript id="5EBABB2FFFF6C97DCC14F956FCAF2CC4" attach="left" box="[814,827,1747,1761]" fontSize="6" pageId="9" pageNumber="85">th</superScript>
|
||
scale rows in anterior part of body, and the upper part of 4
|
||
<superScript id="5EBABB2FFFF6C97DCFFEF972FF452D20" attach="left" box="[196,209,1783,1797]" fontSize="6" pageId="9" pageNumber="85">th</superScript>
|
||
to 6
|
||
<superScript id="5EBABB2FFFF6C97DCE38F972FE9B2D20" attach="left" box="[258,271,1783,1797]" fontSize="6" pageId="9" pageNumber="85">th</superScript>
|
||
in middle and posterior part of body. The upper part of the 1
|
||
<superScript id="5EBABB2FFFF6C97DCC8DF97DFC562D23" attach="left" box="[951,962,1784,1798]" fontSize="6" pageId="9" pageNumber="85">st</superScript>
|
||
DSR in anterior part of body is brownish-black and the lower part is creamish-yellow.
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF6C97DCFFDF8C5FB572DCF" blockId="9.[151,1437,1676,2026]" pageId="9" pageNumber="85">Venter anterior creamish-yellow, gradually fades to the rear, speckled with dark or blackish-brown spots along its middle. The outermost edge of the ventrals is black. Outer quarter of ventrals with subrectangular, black blotches on each side, producing an irregular, continuous ventrolateral stripe. The black blotches gradually deepen and diffuse to the rear, making the posterior part of the venter almost completely black. Ventral surface of the tail glossy, uniform black, free margins of the posterior part of venter scales and subcaudals grayish-white.</paragraph>
|
||
<paragraph id="A9701667FFF5C97ECFFDFF13FC632AF0" blockId="10.[151,1437,150,1365]" pageId="10" pageNumber="86">
|
||
<emphasis id="9BBBCA75FFF5C97ECFFDFF13FD982A95" bold="true" box="[199,524,150,176]" pageId="10" pageNumber="86">Coloration in preservation.</emphasis>
|
||
After two months of preservation, coloration still resembles the specimen in life, except that the background color of the anterior venter becomes ivory cream.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E1D545ECFFF5C97ECFFDFF5AFBB02B64" pageId="10" pageNumber="86" type="etymology">
|
||
<paragraph id="A9701667FFF5C97ECFFDFF5AFBB02B64" blockId="10.[151,1437,150,1365]" pageId="10" pageNumber="86">
|
||
<emphasis id="9BBBCA75FFF5C97ECFFDFF5AFEDA2ADC" bold="true" box="[199,334,223,249]" pageId="10" pageNumber="86">Etymology.</emphasis>
|
||
We named this species based on its
|
||
<typeStatus id="7674A8C5FFF5C97ECDDCFF5AFC822ADC" box="[742,790,223,249]" pageId="10" pageNumber="86">type</typeStatus>
|
||
locality Daleng Village is adjacent to the Youjiang River,
|
||
<collectingRegion id="6B0BD885FFF5C97ECFADFE86FEC32B38" box="[151,343,259,285]" country="China" name="Guangxi" pageId="10" pageNumber="86">Guangxi Zhuang</collectingRegion>
|
||
Autonomous Region,
|
||
<collectingCountry id="D1D856F7FFF5C97ECD63FE86FD342B38" box="[601,672,259,285]" name="China" pageId="10" pageNumber="86">China</collectingCountry>
|
||
. The name we suggest is Yòu Jiāng Fù Liàn Shé (
|
||
<emphasis id="9BBBCA75FFF5C97ECBDCFE81FAE62B3E" bold="true" box="[1254,1394,260,283]" pageId="10" pageNumber="86">Ůĭkńfi</emphasis>
|
||
) as Chinese common name and Youjiang Keelback Snake as English common name.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E1D545ECFFF5C978CFFDFECFFA032B88" lastPageId="12" lastPageNumber="88" pageId="10" pageNumber="86" type="distribution">
|
||
<paragraph id="A9701667FFF5C97ECFFDFECFFC192BF4" blockId="10.[151,1437,150,1365]" pageId="10" pageNumber="86">
|
||
<emphasis id="9BBBCA75FFF5C97ECFFDFECFFE682B41" bold="true" box="[199,508,330,357]" pageId="10" pageNumber="86">Distribution and Habitat.</emphasis>
|
||
At present, the new species is only found at the
|
||
<typeStatus id="7674A8C5FFF5C97ECB0EFECEFBF02B40" box="[1076,1124,331,357]" pageId="10" pageNumber="86">type</typeStatus>
|
||
locality of Daleng Village, Youjiang District, Baise City,
|
||
<collectingRegion id="6B0BD885FFF5C97ECED7FEEAFD382BAC" box="[493,684,367,393]" country="China" name="Guangxi" pageId="10" pageNumber="86">Guangxi Zhuang</collectingRegion>
|
||
Autonomous Region,
|
||
<collectingCountry id="D1D856F7FFF5C97ECC91FEEAFC662BAC" box="[939,1010,367,393]" name="China" pageId="10" pageNumber="86">China</collectingCountry>
|
||
. We found the snake between 1:00 to 2:00 am after a light rain, swimming on the hill stream water surface. The habitat environment was well preserved subtropical evergreen broad-leaved forest, at an elevation of
|
||
<specimenCount id="BFC9DDEEFFF5C97ECC05FE32FC192BF4" box="[831,909,439,465]" pageId="10" pageNumber="86" type="generic">
|
||
<quantity id="6E37BB82FFF5C97ECC05FE32FC1D2BF4" box="[831,905,439,465]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="10" pageNumber="86" unit="m" value="780.0">780 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF5C97ECFFDFE5FFD5E2844" blockId="10.[151,1437,150,1365]" pageId="10" pageNumber="86">
|
||
<emphasis id="9BBBCA75FFF5C97ECFFDFE5FFEF92BD1" bold="true" box="[199,365,474,500]" pageId="10" pageNumber="86">Comparisons.</emphasis>
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECE4EFE5EFDF12BD1" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[372,613,475,501]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF5C97ECE4EFE5EFDF12BD1" box="[372,613,475,501]" italics="true" pageId="10" pageNumber="86">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF5C97ECD57FE5EFD532BD0" bold="true" box="[621,711,475,501]" pageId="10" pageNumber="86">
|
||
<taxonomicNameLabel id="8088770EFFF5C97ECD57FE5EFD532BD0" box="[621,711,475,501]" pageId="10" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from all other species of
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECBECFE5EFAB12BD0" authorityName="Thompson" authorityYear="1913" box="[1238,1317,475,501]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="genus">
|
||
<emphasis id="9BBBCA75FFF5C97ECBECFE5EFAB12BD0" box="[1238,1317,475,501]" italics="true" pageId="10" pageNumber="86">Hebius</emphasis>
|
||
</taxonomicName>
|
||
by having the following characters: preoculars, postoculars, supralabials, infralabials, prefrontal, ventrals, subcaudals, and DSR counts; a conspicuous, beige dorsolateral stripe extending from behind the eye to the end of the tail on each side; and the posterior part of venter almost black.
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF5C97ECFFDFDEEFBF92E10" blockId="10.[151,1437,150,1365]" pageId="10" pageNumber="86">
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECFFDFDEEFE2C28A1" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[199,440,619,645]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF5C97ECFFDFDEEFE2C28A1" box="[199,440,619,645]" italics="true" pageId="10" pageNumber="86">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF5C97ECEFAFDEEFD8E28A0" bold="true" box="[448,538,619,645]" pageId="10" pageNumber="86">
|
||
<taxonomicNameLabel id="8088770EFFF5C97ECEFAFDEEFD8E28A0" box="[448,538,619,645]" pageId="10" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be easily distinguished from
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECCAFFDEEFA0C28A0" authority="(Ziegler & Le Khac Quyet, 2006)" baseAuthorityName="Ziegler & Le Khac Quyet" baseAuthorityYear="2006" box="[917,1432,619,645]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="andreae">
|
||
<emphasis id="9BBBCA75FFF5C97ECCAFFDEEFB8728A0" box="[917,1043,619,645]" italics="true" pageId="10" pageNumber="86">H. andreae</emphasis>
|
||
(Ziegler & Le Khac Quyet, 2006)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECFADFD0AFE79288C" authority="(Gunther, 1864)" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[151,493,655,681]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="beddomei">
|
||
<emphasis id="9BBBCA75FFF5C97ECFADFD0AFF3F288D" box="[151,171,655,680]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECF87FD0AFEBE288C" box="[189,298,655,681]" italics="true" pageId="10" pageNumber="86">beddomei</emphasis>
|
||
(Günther, 1864)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECEC6FD0AFCAE288C" authority="(Wall, 1925)" baseAuthorityName="Wall" baseAuthorityYear="1925" box="[508,826,654,681]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="bitaeniatus">
|
||
<emphasis id="9BBBCA75FFF5C97ECEC6FD0AFD84288D" box="[508,528,655,680]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECD1BFD0AFD09288C" box="[545,669,655,681]" italics="true" pageId="10" pageNumber="86">bitaeniatus</emphasis>
|
||
(
|
||
<bibRefCitation id="CD5E6B96FFF5C97ECD8AFD0AFCA6288C" author="Wall, F." box="[688,818,654,681]" pageId="10" pageNumber="86" pagination="805 - 821" refId="ref9885" refString="Wall, F. (1925) Notes on snakes collected in Burma in 1924. Journal of the Bombay Natural History Society, 30 (4), 805 - 821." type="journal article" year="1925">Wall, 1925</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECC73FD0AFB33288C" authority="(Gressitt, 1937)" baseAuthorityName="Gressitt" baseAuthorityYear="1937" box="[841,1191,655,681]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="boulengeri">
|
||
<emphasis id="9BBBCA75FFF5C97ECC73FD0AFC7C288C" box="[841,1000,655,681]" italics="true" pageId="10" pageNumber="86">H. boulengeri</emphasis>
|
||
(
|
||
<bibRefCitation id="CD5E6B96FFF5C97ECCC0FD0AFB34288C" author="Gressitt, J. L." box="[1018,1184,655,681]" pageId="10" pageNumber="86" pagination="125 - 128" refId="ref8645" refString="Gressitt, J. L. (1937) A new snake from southeastern China. Proceedings of the Biological Society of Washington, 50, 125 - 128." type="journal article" year="1937">Gressitt, 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECB8DFD0AFED328E8" authority="(Peters & Doris, 1878)" baseAuthorityName="Peters & Doris" baseAuthorityYear="1878" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="celebicus">
|
||
<emphasis id="9BBBCA75FFF5C97ECB8DFD0AFB5F288D" box="[1207,1227,655,680]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECBE6FD0AFAD0288C" box="[1244,1348,655,681]" italics="true" pageId="10" pageNumber="86">celebicus</emphasis>
|
||
(Peters & Doris, 1878)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECE6EFD36FDC528E8" authority="(Wall, 1925)" baseAuthorityName="Wall" baseAuthorityYear="1925" box="[340,593,690,717]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="clerki">
|
||
<emphasis id="9BBBCA75FFF5C97ECE6EFD36FEFC28E9" box="[340,360,691,716]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECE42FD36FE2C28E8" box="[376,440,691,717]" italics="true" pageId="10" pageNumber="86">clerki</emphasis>
|
||
(
|
||
<bibRefCitation id="CD5E6B96FFF5C97ECEF2FD36FDDC28E8" author="Wall, F." box="[456,584,690,717]" pageId="10" pageNumber="86" pagination="805 - 821" refId="ref9885" refString="Wall, F. (1925) Notes on snakes collected in Burma in 1924. Journal of the Bombay Natural History Society, 30 (4), 805 - 821." type="journal article" year="1925">Wall, 1925</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECD64FD36FC2928E8" authority="(Malnate, 1963)" baseAuthorityName="Malnate" baseAuthorityYear="1963" box="[606,957,691,717]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="concelarus">
|
||
<emphasis id="9BBBCA75FFF5C97ECD64FD36FD6928E8" box="[606,765,691,717]" italics="true" pageId="10" pageNumber="86">H. concelarus</emphasis>
|
||
(Malnate, 1963)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECCF1FD36FAE028E8" authority="(Boulenger, 1899)" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[971,1396,691,717]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="craspedogaster">
|
||
<emphasis id="9BBBCA75FFF5C97ECCF1FD36FB0828E8" box="[971,1180,691,717]" italics="true" pageId="10" pageNumber="86">H. craspedogaster</emphasis>
|
||
(Boulenger, 1899)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECABBFD36FE4128D4" authority="(Boulenger, 1887)" baseAuthorityName="Boulenger" baseAuthorityYear="1887" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="flavifrons">
|
||
<emphasis id="9BBBCA75FFF5C97ECABBFD36FA0128E9" box="[1409,1429,691,716]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECFADFD52FE9528D4" box="[151,257,727,753]" italics="true" pageId="10" pageNumber="86">flavifrons</emphasis>
|
||
(Boulenger, 1887)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECEE5FD52FD7C28D4" authority="(Laidlaw, 1901)" baseAuthorityName="Laidlaw" baseAuthorityYear="1901" box="[479,744,727,753]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="inas">
|
||
<emphasis id="9BBBCA75FFF5C97ECEE5FD52FDA428D5" box="[479,560,727,752]" italics="true" pageId="10" pageNumber="86">H. inas</emphasis>
|
||
(Laidlaw, 1901)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECDCEFD52FA8428D4" authority="(Malnate & Munsterman, 1960)" baseAuthorityName="Malnate & Munsterman" baseAuthorityYear="1960" box="[756,1296,727,753]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="ishigakiensis">
|
||
<emphasis id="9BBBCA75FFF5C97ECDCEFD52FC9C28D5" box="[756,776,727,752]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECC2FFD52FC3228D4" box="[789,934,727,753]" italics="true" pageId="10" pageNumber="86">ishigakiensis</emphasis>
|
||
(Malnate & Munsterman, 1960)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECA21FD52FEF02930" authority="(Boulenger, 1908)" baseAuthorityName="Boulenger" baseAuthorityYear="1908" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="johannis">
|
||
<emphasis id="9BBBCA75FFF5C97ECA21FD52FABB28D5" box="[1307,1327,727,752]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECA06FD52FA0828D4" box="[1340,1436,727,753]" italics="true" pageId="10" pageNumber="86">johannis</emphasis>
|
||
(Boulenger, 1908)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECE54FD7EFC972930" authority="(David & Das, 2003)" baseAuthorityName="David & Das" baseAuthorityYear="2003" box="[366,771,763,789]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="kerinciensis">
|
||
<emphasis id="9BBBCA75FFF5C97ECE54FD7EFE162931" box="[366,386,763,788]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECEB4FD7EFD802930" box="[398,532,763,789]" italics="true" pageId="10" pageNumber="86">kerinciensis</emphasis>
|
||
(
|
||
<bibRefCitation id="CD5E6B96FFF5C97ECD1BFD7EFD6F2930" author="David, P. & Das, I." box="[545,763,763,789]" pageId="10" pageNumber="86" pagination="413 - 419" refId="ref8190" refString="David, P. & Das, I. (2003) A new species of the snake genus Amphiesma (Serpentes: Colubridae: Natricinae) from western Sumatra, Indonesia. Raffles Bulletin of Zoology, 51 (2), 413 - 419." type="journal article" year="2003">David & Das, 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECC36FD7EFBE62930" authority="(Boulenger, 1890)" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[780,1138,763,789]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="khasiensis">
|
||
<emphasis id="9BBBCA75FFF5C97ECC36FD7EFCB42931" box="[780,800,763,788]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECC17FD7EFC342930" box="[813,928,763,789]" italics="true" pageId="10" pageNumber="86">khasiensis</emphasis>
|
||
(Boulenger, 1890)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECB46FD7EFEB0291C" authority="Purkayastha & David, 2019" authorityName="Purkayastha & David" authorityYear="2019" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="lacrima">
|
||
<emphasis id="9BBBCA75FFF5C97ECB46FD7EFB042931" box="[1148,1168,763,788]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECBA6FD7EFB672930" box="[1180,1267,763,789]" italics="true" pageId="10" pageNumber="86">lacrima</emphasis>
|
||
<bibRefCitation id="CD5E6B96FFF5C97ECBC2FD7EFEB0291C" author="Purkayastha, J. & David, P." pageId="10" pageNumber="86" pagination="79 - 90" refId="ref9336" refString="Purkayastha, J. & David, P. (2019) A new species of the snake genus Hebius Thompson from Northeast India (Squamata: Natricidae). Zootaxa, 4555 (1), 79 - 90. https: // doi. org / 10.11646 / zootaxa. 4555.1.6" type="journal article" year="2019">Purkayastha & David, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECE15FC9AFA8C291C" authority="(David, Bain, Quang Truong, Orlov, Vogel, Ngoc Thanh & Ziegler, 2007)" baseAuthorityName="David, Bain, Quang Truong, Orlov, Vogel, Ngoc Thanh & Ziegler" baseAuthorityYear="2007" box="[303,1304,799,825]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="leucomystax">
|
||
<emphasis id="9BBBCA75FFF5C97ECE15FC9AFED7291D" box="[303,323,799,824]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECE6BFC9AFE4F291C" box="[337,475,799,825]" italics="true" pageId="10" pageNumber="86">leucomystax</emphasis>
|
||
(David, Bain, Quang Truong, Orlov, Vogel, Ngoc Thanh & Ziegler, 2007)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECA19FC9AFDB12978" authority="(Inger, Zhao, Shaffer & Wu, 1990)" baseAuthorityName="Inger, Zhao, Shaffer & Wu" baseAuthorityYear="1990" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="metusia">
|
||
<emphasis id="9BBBCA75FFF5C97ECA19FC9AFAA3291D" box="[1315,1335,799,824]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECA7FFC9AFA08291D" box="[1349,1436,799,824]" italics="true" pageId="10" pageNumber="86">metusia</emphasis>
|
||
(Inger, Zhao, Shaffer & Wu, 1990)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECD08FCC6FCFD2978" authority="(Maki, 1931)" baseAuthorityName="Maki" baseAuthorityYear="1931" box="[562,873,835,861]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="miyajimae">
|
||
<emphasis id="9BBBCA75FFF5C97ECD08FCC6FD5F2979" box="[562,715,835,860]" italics="true" pageId="10" pageNumber="86">H. miyajimae</emphasis>
|
||
(Maki, 1931)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECC4DFCC6FB762978" authority="(Sclater, 1891)" baseAuthorityName="Sclater" baseAuthorityYear="1891" box="[887,1250,835,861]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="nicobariensis">
|
||
<emphasis id="9BBBCA75FFF5C97ECC4DFCC6FC1F2979" box="[887,907,835,860]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECCA1FCC6FBA62978" box="[923,1074,835,861]" italics="true" pageId="10" pageNumber="86">nicobariensis</emphasis>
|
||
(Sclater, 1891)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECBCAFCC6FEF229A4" authority="(Boulenger, 1904)" baseAuthorityName="Boulenger" baseAuthorityYear="1904" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="octolineatus">
|
||
<emphasis id="9BBBCA75FFF5C97ECBCAFCC6FA902979" box="[1264,1284,835,860]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECA29FCC6FA082978" box="[1299,1436,835,861]" italics="true" pageId="10" pageNumber="86">octolineatus</emphasis>
|
||
(Boulenger, 1904)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECE48FCE2FD5D29A4" authority="(Hu & Zhao, 1966)" baseAuthorityName="Hu & Zhao" baseAuthorityYear="1966" box="[370,713,871,897]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="optatus">
|
||
<emphasis id="9BBBCA75FFF5C97ECE48FCE2FE7229A5" box="[370,486,871,896]" italics="true" pageId="10" pageNumber="86">H. optatus</emphasis>
|
||
(Hu & Zhao, 1966)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECDEFFCE2FBAB29A4" authority="(Boulenger, 1890)" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[725,1087,871,897]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="parallelus">
|
||
<emphasis id="9BBBCA75FFF5C97ECDEFFCE2FD7D29A5" box="[725,745,871,896]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECDC2FCE2FCFE29A4" box="[760,874,871,897]" italics="true" pageId="10" pageNumber="86">parallelus</emphasis>
|
||
(Boulenger, 1890)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECB71FCE2FA0329A4" authority="(Boulenger, 1893)" baseAuthorityName="Boulenger" baseAuthorityYear="1893" box="[1099,1431,871,897]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="petersii">
|
||
<emphasis id="9BBBCA75FFF5C97ECB71FCE2FB5629A5" box="[1099,1218,871,896]" italics="true" pageId="10" pageNumber="86">H. petersii</emphasis>
|
||
(Boulenger, 1893)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECFADFC0EFE232980" authority="(Schmidt, 1925)" baseAuthorityName="Schmidt" baseAuthorityYear="1925" box="[151,439,906,933]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="popei">
|
||
<emphasis id="9BBBCA75FFF5C97ECFADFC0EFF3F2981" box="[151,171,907,932]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECF83FC0EFF632981" box="[185,247,907,932]" italics="true" pageId="10" pageNumber="86">popei</emphasis>
|
||
(Schmidt, 1925)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECEF9FC0EFD6B2980" authority="(Boulenger, 1887)" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[451,767,907,933]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="pryeri">
|
||
<emphasis id="9BBBCA75FFF5C97ECEF9FC0EFE432981" box="[451,471,907,932]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECEDFFC0EFDBD2981" box="[485,553,907,932]" italics="true" pageId="10" pageNumber="86">pryeri</emphasis>
|
||
(Boulenger, 1887)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECC31FC0EFBF92980" authority="(Smedley, 1932)" baseAuthorityName="Smedley" baseAuthorityYear="1932" box="[779,1133,907,933]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="sanguineus">
|
||
<emphasis id="9BBBCA75FFF5C97ECC31FC0EFC8B2981" box="[779,799,907,932]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECC17FC0EFC3F2981" box="[813,939,907,932]" italics="true" pageId="10" pageNumber="86">sanguineus</emphasis>
|
||
(Smedley, 1932)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECB43FC0EFE1429EC" authority="Zhou, Qi, Lu, Lyu & Li, 2019" authorityName="Zhou, Qi, Lu, Lyu & Li" authorityYear="2019" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="sangzhiensis">
|
||
<emphasis id="9BBBCA75FFF5C97ECB43FC0EFABD2980" box="[1145,1321,907,933]" italics="true" pageId="10" pageNumber="86">H. sangzhiensis</emphasis>
|
||
<bibRefCitation id="CD5E6B96FFF5C97ECA0AFC0EFE1429EC" author="Zhou, Z. Y. & Sun, Z. & Qi, S. & Lu, Y. & Lyu, Z. & Wang, Y. & Li, P. & Ma, J." pageId="10" pageNumber="86" pagination="1" refId="ref9946" refString="Zhou, Z. Y., Sun, Z., Qi, S., Lu, Y., Lyu, Z., Wang, Y., Li, P. & Ma, J. (2019) A new species of the genus Hebius (Squamata: Colubridae: Natricinae) from Hunan Province, China. Zootaxa, 4674 (1), 068 - 082. https: // doi. org / 10.11646 / zootaxa. 4674.1.3" type="journal article" year="2019">Zhou, Qi, Lu, Lyu & Li, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECEB7FC2AFC8E29EC" authority="(Boulenger, 1896)" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[397,794,943,969]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="sarasinorum">
|
||
<emphasis id="9BBBCA75FFF5C97ECEB7FC2AFE3529ED" box="[397,417,943,968]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECE88FC2AFDD429ED" box="[434,576,943,968]" italics="true" pageId="10" pageNumber="86">sarasinorum</emphasis>
|
||
(Boulenger, 1896)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECC12FC2AFB2329EC" authority="(Schmidt, 1925)" baseAuthorityName="Schmidt" baseAuthorityYear="1925" box="[808,1207,942,969]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="septemlineatus">
|
||
<emphasis id="9BBBCA75FFF5C97ECC12FC2AFC6729EC" box="[808,1011,943,969]" italics="true" pageId="10" pageNumber="86">H. septemlineatus</emphasis>
|
||
(Schmidt, 1925)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECBFCFC2AFE7D29C8" authority="Hauser, Smits & David, 2022" authorityName="Hauser, Smits & David" authorityYear="2022" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="terrakarenorum">
|
||
<emphasis id="9BBBCA75FFF5C97ECBFCFC2AFA0829EC" box="[1222,1436,943,969]" italics="true" pageId="10" pageNumber="86">H. terrakarenorum</emphasis>
|
||
<bibRefCitation id="CD5E6B96FFF5C97ECFADFC56FE7D29C8" author="Hauser, J. & Smits, T. & David, P." box="[151,489,979,1005]" pageId="10" pageNumber="86" refId="ref8715" refString="Hauser, J., Smits, T. & David, P. (2022) On the distribution of the species of the genus Hebius Thompson, 1913 (Squamata: Natricidae) in northern Thailand, including the description of a new species and a discussion on snake diversity of this region. Zootaxa 5116 (1), 001 - 039 https: // doi. org / 10.11646 / zootaxa. 5116.1.1" type="book" year="2022">Hauser, Smits & David, 2022</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECECFFC56FC9B29C8" authority="(Boie, 1826)" baseAuthorityName="Boie" baseAuthorityYear="1826" box="[501,783,979,1005]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="vibakari">
|
||
<emphasis id="9BBBCA75FFF5C97ECECFFC56FDE129C8" box="[501,629,979,1005]" italics="true" pageId="10" pageNumber="86">H. vibakari</emphasis>
|
||
(Boie, 1826)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECC26FC56FBE629C8" authority="(Schenkel, 1901)" baseAuthorityName="Schenkel" baseAuthorityYear="1901" box="[796,1138,979,1005]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="viperinus">
|
||
<emphasis id="9BBBCA75FFF5C97ECC26FC56FCA429C9" box="[796,816,979,1004]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECC05FC56FC3329C9" box="[831,935,979,1004]" italics="true" pageId="10" pageNumber="86">viperinus</emphasis>
|
||
(Schenkel, 1901)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECB45FC56FD762E34" authority="Hou, Yuan, Wei, Zhao, Liu, Wu, Shen, Chen, Guo & Che, 2021" authorityName="Hou, Yuan, Wei, Zhao, Liu, Wu, Shen, Chen, Guo & Che" authorityYear="2021" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="weixiensis">
|
||
<emphasis id="9BBBCA75FFF5C97ECB45FC56FB0729C9" box="[1151,1171,979,1004]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECB98FC56FA8129C9" box="[1186,1301,979,1004]" italics="true" pageId="10" pageNumber="86">weixiensis</emphasis>
|
||
<bibRefCitation id="CD5E6B96FFF5C97ECA27FC56FD762E34" author="Hou, S. B. & Yuan, Z. Y. & Wei, P. F. & Zhao, G. G. & Liu, G. H. & Wu, Y. H. & Shen, W. - J. & Chen, J. - M. & Guo, P. & Che, J." pageId="10" pageNumber="86" pagination="620 - 625" refId="ref8792" refString="Hou, S. B., Yuan, Z. Y., Wei, P. F., Zhao, G. G., Liu, G. H., Wu, Y. H., Shen, W. - J., Chen, J. - M., Guo, P. & Che, J. (2021) Molecular phylogeny and morphological comparisons of the genus Hebius Thompson, 1913 (Reptilia: Squamata: Colubridae) uncover a new taxon from Yunnan Province, China, and support revalidation of Hebius septemlineatus (Schmidt, 1925). Zoological Research, 42 (5), 620 - 625. https: // doi. org / 10.24272 / j. issn. 2095 - 8137.2021.093" type="journal article" year="2021">Hou, Yuan, Wei, Zhao, Liu, Wu, Shen, Chen, Guo & Che, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECC20FC72FAEC2E34" authority="Liu, Zhong, Wang, Liu & Guo, 2018" authorityName="Liu, Zhong, Wang, Liu & Guo" authorityYear="2018" box="[794,1400,1015,1041]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="yanbianensis">
|
||
<emphasis id="9BBBCA75FFF5C97ECC20FC72FCBA2E35" box="[794,814,1015,1040]" italics="true" pageId="10" pageNumber="86">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF5C97ECC07FC72FC5B2E34" box="[829,975,1015,1041]" italics="true" pageId="10" pageNumber="86">yanbianensis</emphasis>
|
||
<bibRefCitation id="CD5E6B96FFF5C97ECCEDFC72FAEC2E34" author="Liu, Q. & Zhong, G. H. & Liu, Y. & Guo, P." box="[983,1400,1015,1041]" pageId="10" pageNumber="86" pagination="385 - 394" refId="ref9267" refString="Liu, Q., Zhong, G. H., W, P., Liu, Y. & Guo, P. (2018) A new species of the genus Hebius (Squamata: Colubridae) from Sichuan, China. Zootaxa, 4483 (2), 385 - 394. https: // doi. org / 10.11646 / zootaxa. 4483.2.10" type="journal article" year="2018">Liu, Zhong, Wang, Liu & Guo, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
by having 17 DSR at midbody (vs. 19 or 15), and the posterior part of venter almost black.
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF5C97ECFFDFBBAFD8C2F70" blockId="10.[151,1437,150,1365]" pageId="10" pageNumber="86">
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECFFDFBBAFE2C2E7D" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[199,440,1087,1113]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF5C97ECFFDFBBAFE2C2E7D" box="[199,440,1087,1113]" italics="true" pageId="10" pageNumber="86">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF5C97ECEFAFBBAFD8F2E7C" bold="true" box="[448,539,1087,1113]" pageId="10" pageNumber="86">
|
||
<taxonomicNameLabel id="8088770EFFF5C97ECEFAFBBAFD8F2E7C" box="[448,539,1087,1113]" pageId="10" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar to its sister species
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECC97FBBAFBAC2E7C" baseAuthorityName="Gunther" baseAuthorityYear="1875" box="[941,1080,1087,1113]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="9BBBCA75FFF5C97ECC97FBBAFBAC2E7C" box="[941,1080,1087,1113]" italics="true" pageId="10" pageNumber="86">H. modestus</emphasis>
|
||
</taxonomicName>
|
||
. However, the new species can be distinguished from
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECEABFBE6FD8E2E58" baseAuthorityName="Gunther" baseAuthorityYear="1875" box="[401,538,1123,1149]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="9BBBCA75FFF5C97ECEABFBE6FD8E2E58" box="[401,538,1123,1149]" italics="true" pageId="10" pageNumber="86">H. modestus</emphasis>
|
||
</taxonomicName>
|
||
by having 17 DSR at midbody (vs. 19), maxillary teeth 30, the last 3 moderately enlarged (vs. 27–29 maxillary teeth, the last 2 moderately enlarged), postocular streak presence (vs. absent), and dorsal brownish-black, a conspicuous, continuous beige stripe extending from behind the eye to the end of the tail on each side of body (vs. dorsal usually scattered with blackish-brown or black spots or blotches, a ochre-yellow, yellowish-brown, ochre-red, orange-brown or reddish-brown stripe often reduced to a succession of elongate blotches on the anterior part of the body on each side). For more detailed information and visual comparisons, please refer to
|
||
<tableCitation id="E44D23DCFFF5C97ECE06FABEFE1B2F70" box="[316,399,1339,1365]" captionStart="TABLE 3" captionStartId="10.[151,239,1412,1436]" captionTargetPageId="10" captionText="TABLE 3. Comparison of main morphological characters between H. youjiangensis sp. nov. and H. modestus." httpUri="http://table.plazi.org/id/FDB046EFFFF5C97ECFADFA01FAA22FB9" pageId="10" pageNumber="86" tableUuid="FDB046EFFFF5C97ECFADFA01FAA22FB9">Table 3</tableCitation>
|
||
&
|
||
<figureCitation id="31F40AE2FFF5C97ECE89FABEFD802F70" box="[435,532,1339,1365]" captionStart="FIGURE 6" captionStartId="11.[152,255,936,960]" captionTargetBox="[488,1099,182,913]" captionTargetId="figure-23@11.[488,1099,181,913]" captionTargetPageId="11" captionText="FIGURE 6. Comparisons of (1) dorsal overview, (2) ventral overview, (3) dorsal head, and (4) lateral head among H. youjiangensis sp. nov. (ANU20220010; holotype) (A), H. modestus (BMNH 1946.1.13.41; syntype) (B). Photos of the syntype of H. modestus are obtained from David et al. (2021), remaining photos by Yu-Hao Xu." figureDoi="http://doi.org/10.5281/zenodo.10135912" httpUri="https://zenodo.org/record/10135912/files/figure.png" pageId="10" pageNumber="86">Figure 6</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="FDB046EFFFF5C97ECFADFA01FAA22FB9" ID-Table-UUID="FDB046EFFFF5C97ECFADFA01FAA22FB9" box="[151,1334,1412,1437]" httpUri="http://table.plazi.org/id/FDB046EFFFF5C97ECFADFA01FAA22FB9" pageId="10" pageNumber="86" startId="10.[151,239,1412,1436]" targetBox="[163,1418,1453,1587]" targetIsTable="true" targetPageId="10" targetType="table">
|
||
<paragraph id="A9701667FFF5C97ECFADFA01FAA22FB9" blockId="10.[151,1334,1412,1437]" box="[151,1334,1412,1437]" pageId="10" pageNumber="86">
|
||
<emphasis id="9BBBCA75FFF5C97ECFADFA01FE9E2FB9" bold="true" box="[151,266,1412,1436]" pageId="10" pageNumber="86">TABLE 3.</emphasis>
|
||
Comparison of main morphological characters between
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECC5DFA00FB8F2FB9" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[871,1051,1413,1436]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF5C97ECC5DFA00FB8F2FB9" box="[871,1051,1413,1436]" italics="true" pageId="10" pageNumber="86">H. youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF5C97ECB1BFA00FBE32FB8" bold="true" box="[1057,1143,1413,1437]" pageId="10" pageNumber="86">
|
||
<taxonomicNameLabel id="8088770EFFF5C97ECB1BFA00FBE32FB8" box="[1057,1143,1413,1437]" pageId="10" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECB90FA00FAA62FB9" baseAuthorityName="Gunther" baseAuthorityYear="1875" box="[1194,1330,1412,1436]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="modestus">H. modestus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A9701667FFF5C97ECF99FA28FA1E2C16" pageId="10" pageNumber="86">
|
||
<table id="DBCFE4C7FFF5368BCF99FA28FA1E2C16" box="[163,1418,1453,1587]" gridcols="11" gridrows="3" pageId="10" pageNumber="86">
|
||
<tr id="17FF1425FFF5368BCF99FA28FA1E2FE0" box="[163,1418,1453,1477]" gridrow="0" pageId="10" pageNumber="86">
|
||
<th id="542E7D59FFF5368BCF99FA28FED82FE0" box="[163,332,1453,1477]" gridcol="0" gridrow="0" pageId="10" pageNumber="86">Species</th>
|
||
<th id="542E7D59FFF5368BCEB8FA28FE732FE0" box="[386,487,1453,1477]" gridcol="1" gridrow="0" pageId="10" pageNumber="86">TAL/TL</th>
|
||
<th id="542E7D59FFF5368BCD32FA28FDAD2FE0" box="[520,569,1453,1477]" gridcol="2" gridrow="0" pageId="10" pageNumber="86">PRO</th>
|
||
<th id="542E7D59FFF5368BCD6DFA28FD5D2FE0" box="[599,713,1453,1477]" gridcol="3" gridrow="0" pageId="10" pageNumber="86">PO</th>
|
||
<th id="542E7D59FFF5368BCDD0FA28FC9E2FE0" box="[746,778,1453,1477]" gridcol="4" gridrow="0" pageId="10" pageNumber="86">SL</th>
|
||
<th id="542E7D59FFF5368BCC0AFA28FCFE2FE0" box="[816,874,1453,1477]" gridcol="5" gridrow="0" pageId="10" pageNumber="86">IL</th>
|
||
<th id="542E7D59FFF5368BCCB0FA28FC5A2FE0" box="[906,974,1453,1477]" gridcol="6" gridrow="0" pageId="10" pageNumber="86">TEMP</th>
|
||
<th id="542E7D59FFF5368BCCD1FA28FBDC2FE0" box="[1003,1096,1453,1477]" gridcol="7" gridrow="0" pageId="10" pageNumber="86">DSR</th>
|
||
<th id="542E7D59FFF5368BCB58FA28FB2F2FE0" box="[1122,1211,1453,1477]" gridcol="8" gridrow="0" pageId="10" pageNumber="86">VEN</th>
|
||
<th id="542E7D59FFF5368BCBEEFA28FAB92FE0" box="[1236,1325,1453,1477]" gridcol="9" gridrow="0" pageId="10" pageNumber="86">SC</th>
|
||
<th id="542E7D59FFF5368BCA71FA28FA1E2FE0" box="[1355,1418,1453,1477]" gridcol="10" gridrow="0" pageId="10" pageNumber="86">MT</th>
|
||
</tr>
|
||
<tr id="17FF1425FFF5368BCF99FA50FA1E2C2F" box="[163,1418,1493,1546]" gridrow="1" pageId="10" pageNumber="86">
|
||
<th id="542E7D59FFF5368BCF99FA50FED82C2F" box="[163,332,1493,1546]" gridcol="0" gridrow="1" pageId="10" pageNumber="86">
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECF99FA53FED82FC8" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[163,332,1494,1517]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF5C97ECF99FA53FED82FC8" box="[163,332,1494,1517]" italics="true" pageId="10" pageNumber="86">H. youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF5C97ECF99FA77FF612C2F" bold="true" box="[163,245,1522,1546]" pageId="10" pageNumber="86">
|
||
<taxonomicNameLabel id="8088770EFFF5C97ECF99FA77FF612C2F" box="[163,245,1522,1546]" pageId="10" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<td id="542E7D59FFF5368BCEB8FA50FE732C2F" box="[386,487,1493,1546]" gridcol="1" gridrow="1" pageId="10" pageNumber="86">0.30</td>
|
||
<td id="542E7D59FFF5368BCD32FA50FDAD2C2F" box="[520,569,1493,1546]" gridcol="2" gridrow="1" pageId="10" pageNumber="86">1</td>
|
||
<td id="542E7D59FFF5368BCD6DFA50FD5D2C2F" box="[599,713,1493,1546]" gridcol="3" gridrow="1" pageId="10" pageNumber="86">2</td>
|
||
<td id="542E7D59FFF5368BCDD0FA50FC9E2C2F" box="[746,778,1493,1546]" gridcol="4" gridrow="1" pageId="10" pageNumber="86">9/9</td>
|
||
<td id="542E7D59FFF5368BCC0AFA50FCFE2C2F" box="[816,874,1493,1546]" gridcol="5" gridrow="1" pageId="10" pageNumber="86">10/10</td>
|
||
<td id="542E7D59FFF5368BCCB0FA50FC5A2C2F" box="[906,974,1493,1546]" gridcol="6" gridrow="1" pageId="10" pageNumber="86">1+1+2</td>
|
||
<td id="542E7D59FFF5368BCCD1FA50FBDC2C2F" box="[1003,1096,1493,1546]" gridcol="7" gridrow="1" pageId="10" pageNumber="86">19-17-17</td>
|
||
<td id="542E7D59FFF5368BCB58FA50FB2F2C2F" box="[1122,1211,1493,1546]" gridcol="8" gridrow="1" pageId="10" pageNumber="86">160+3</td>
|
||
<td id="542E7D59FFF5368BCBEEFA50FAB92C2F" box="[1236,1325,1493,1546]" gridcol="9" gridrow="1" pageId="10" pageNumber="86">112</td>
|
||
<td id="542E7D59FFF5368BCA71FA50FA1E2C2F" box="[1355,1418,1493,1546]" gridcol="10" gridrow="1" pageId="10" pageNumber="86">30</td>
|
||
</tr>
|
||
<tr id="17FF1425FFF5368BCF99F99FFA1E2C16" box="[163,1418,1562,1587]" gridrow="2" pageId="10" pageNumber="86">
|
||
<th id="542E7D59FFF5368BCF99F99FFED82C16" box="[163,332,1562,1587]" gridcol="0" gridrow="2" pageId="10" pageNumber="86">
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECF99F99EFE8B2C17" baseAuthorityName="Gunther" baseAuthorityYear="1875" box="[163,287,1562,1586]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="9BBBCA75FFF5C97ECF99F99EFE8B2C17" box="[163,287,1562,1586]" italics="true" pageId="10" pageNumber="86">H. modestus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="542E7D59FFF5368BCEB8F99FFE732C16" box="[386,487,1562,1587]" gridcol="1" gridrow="2" pageId="10" pageNumber="86">0.29–0.33</td>
|
||
<td id="542E7D59FFF5368BCD32F99FFDAD2C16" box="[520,569,1562,1587]" gridcol="2" gridrow="2" pageId="10" pageNumber="86">2 (1)</td>
|
||
<td id="542E7D59FFF5368BCD6DF99FFD5D2C16" box="[599,713,1562,1587]" gridcol="3" gridrow="2" pageId="10" pageNumber="86">3 (rarely 2)</td>
|
||
<td id="542E7D59FFF5368BCDD0F99FFC9E2C16" box="[746,778,1562,1587]" gridcol="4" gridrow="2" pageId="10" pageNumber="86">9/9</td>
|
||
<td id="542E7D59FFF5368BCC0AF99FFCFE2C16" box="[816,874,1562,1587]" gridcol="5" gridrow="2" pageId="10" pageNumber="86">10/10</td>
|
||
<td id="542E7D59FFF5368BCCB0F99FFC5A2C16" box="[906,974,1562,1587]" gridcol="6" gridrow="2" pageId="10" pageNumber="86">1+1</td>
|
||
<td id="542E7D59FFF5368BCCD1F99FFBDC2C16" box="[1003,1096,1562,1587]" gridcol="7" gridrow="2" pageId="10" pageNumber="86">19-19-17</td>
|
||
<td id="542E7D59FFF5368BCB58F99FFB2F2C16" box="[1122,1211,1562,1587]" gridcol="8" gridrow="2" pageId="10" pageNumber="86">143–163</td>
|
||
<td id="542E7D59FFF5368BCBEEF99FFAB92C16" box="[1236,1325,1562,1587]" gridcol="9" gridrow="2" pageId="10" pageNumber="86">104–122</td>
|
||
<td id="542E7D59FFF5368BCA71F99FFA1E2C16" box="[1355,1418,1562,1587]" gridcol="10" gridrow="2" pageId="10" pageNumber="86">27–29</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF5C97ECFFDF903FD802D15" blockId="10.[151,1437,1670,2020]" pageId="10" pageNumber="86">
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECFFDF903FE222C85" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[199,438,1670,1696]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF5C97ECFFDF903FE222C85" box="[199,438,1670,1696]" italics="true" pageId="10" pageNumber="86">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF5C97ECE86F903FD802C85" bold="true" box="[444,532,1670,1696]" pageId="10" pageNumber="86">
|
||
<taxonomicNameLabel id="8088770EFFF5C97ECE86F903FD802C85" box="[444,532,1670,1696]" pageId="10" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECD9EF902FCF32C85" baseAuthorityName="Taylor" baseAuthorityYear="1934" box="[676,871,1670,1696]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="deschauenseei">
|
||
<emphasis id="9BBBCA75FFF5C97ECD9EF902FCF32C85" box="[676,871,1670,1696]" italics="true" pageId="10" pageNumber="86">H. deschauenseei</emphasis>
|
||
</taxonomicName>
|
||
by having 17 DSR at midbody (vs. 19), subcaudals 112 (vs. 115–141), preoculars 1 (vs. 2), temporals 1+1+2 (vs. 1+2), and a conspicuous, continuous beige stripe extending from behind the eye to the end of the tail (vs. dorsal surface with dark blotches, not producing a chequered pattern, a dorsolateral series of large, elongate blotches, yellow-ochre, rusty red or yellowish-brown, forming a discontinuous dorsolateral stripe).
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF5C97ECFFDF8BFFEB02DC1" blockId="10.[151,1437,1670,2020]" pageId="10" pageNumber="86">
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECFFDF8BFFE2F2D71" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[199,443,1850,1876]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF5C97ECFFDF8BFFE2F2D71" box="[199,443,1850,1876]" italics="true" pageId="10" pageNumber="86">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF5C97ECEFCF8BFFDB72D71" bold="true" box="[454,547,1850,1876]" pageId="10" pageNumber="86">
|
||
<taxonomicNameLabel id="8088770EFFF5C97ECEFCF8BFFDB72D71" box="[454,547,1850,1876]" pageId="10" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECDFBF8BEFCF72D71" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[705,867,1850,1876]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="chapaensis">
|
||
<emphasis id="9BBBCA75FFF5C97ECDFBF8BEFCF72D71" box="[705,867,1850,1876]" italics="true" pageId="10" pageNumber="86">H. chapaensis</emphasis>
|
||
</taxonomicName>
|
||
by having 19-17-17 DSR counts (vs. 17-17-17), dorsal scales feebly keeled except the outermost rows (vs. dorsal scales weakly keeled anteriorly, strongly keeled posteriorly), preoculars 1 (vs. 2), and having a conspicuous, beige dorsolateral stripe extending from behind the eye to the end of the tail on each side (vs. pale orange blotches anteriorly and longitudinal rodlike pattern in
|
||
<taxonomicName id="6ECF6DE4FFF5C97ECABBF822FE872DC1" baseAuthorityName="Bourret" baseAuthorityYear="1934" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="10" pageNumber="86" phylum="Chordata" rank="species" species="chapaensis">
|
||
<emphasis id="9BBBCA75FFF5C97ECABBF822FE872DC1" italics="true" pageId="10" pageNumber="86">H. chapaensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="FDB046EFFFF4C97FCFA2FC2DFB9B2E2D" ID-DOI="http://doi.org/10.5281/zenodo.10135912" ID-Zenodo-Dep="10135912" httpUri="https://zenodo.org/record/10135912/files/figure.png" pageId="11" pageNumber="87" startId="11.[152,255,936,960]" targetBox="[488,1099,182,913]" targetPageId="11" targetType="figure">
|
||
<paragraph id="A9701667FFF4C97FCFA2FC2DFB9B2E2D" blockId="11.[151,1436,936,1033]" pageId="11" pageNumber="87">
|
||
<emphasis id="9BBBCA75FFF4C97FCFA2FC2DFE8929E5" bold="true" box="[152,285,936,960]" pageId="11" pageNumber="87">FIGURE 6.</emphasis>
|
||
Comparisons of (1) dorsal overview, (2) ventral overview, (3) dorsal head, and (4) lateral head among
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCABEFC2CFEB629C1" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF4C97FCABEFC2CFA0229E5" box="[1412,1430,937,960]" italics="true" pageId="11" pageNumber="87">H</emphasis>
|
||
.
|
||
<emphasis id="9BBBCA75FFF4C97FCFADFC48FEB629C1" box="[151,290,973,996]" italics="true" pageId="11" pageNumber="87">youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF4C97FCE13FC48FEED29C0" bold="true" box="[297,377,973,997]" pageId="11" pageNumber="87">
|
||
<taxonomicNameLabel id="8088770EFFF4C97FCE13FC48FEED29C0" box="[297,377,973,997]" pageId="11" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ANU20220010; holotype) (A),
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCDF2FC48FCD029C1" baseAuthorityName="Gunther" baseAuthorityYear="1875" box="[712,836,972,996]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="9BBBCA75FFF4C97FCDF2FC48FCD029C1" box="[712,836,972,996]" italics="true" pageId="11" pageNumber="87">H. modestus</emphasis>
|
||
</taxonomicName>
|
||
(BMNH 1946.1.13.41; syntype) (B). Photos of the syntype of
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCF89FC74FEA42E2D" baseAuthorityName="Gunther" baseAuthorityYear="1875" box="[179,304,1008,1032]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="9BBBCA75FFF4C97FCF89FC74FEA42E2D" box="[179,304,1008,1032]" italics="true" pageId="11" pageNumber="87">H. modestus</emphasis>
|
||
</taxonomicName>
|
||
are obtained from
|
||
<bibRefCitation id="CD5E6B96FFF4C97FCECBFC75FD202E2C" author="David, P. & Vogel, G. & Nguyen, T. Q. & Orlov, N. L. & Pauwels, O. S. G. & Teynie, A. & Ziegler, T." box="[497,692,1008,1033]" pageId="11" pageNumber="87" pagination="1 - 61" refId="ref8280" refString="David, P., Vogel, G., Nguyen, T. Q., Orlov, N. L., Pauwels, O. S. G., Teynie, A. & Ziegler, T. (2021) A revision of the dark-bellied, stream-dwelling snakes of the genus Hebius (Reptilia: Squamata: Natricidae) with the description of a new species from China, Vietnam and Thailand. Zootaxa, 4911 (1), 1 - 61. https: // doi. org / 10.11646 / zootaxa. 4911.1.1" type="journal article" year="2021">
|
||
David
|
||
<emphasis id="9BBBCA75FFF4C97FCD0FFC74FDF52E2D" box="[565,609,1008,1032]" italics="true" pageId="11" pageNumber="87">et al</emphasis>
|
||
. (2021)
|
||
</bibRefCitation>
|
||
, remaining photos by Yu-Hao Xu.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A9701667FFF4C97FCFFDFBDEFAE92EC4" blockId="11.[151,1436,1115,2041]" pageId="11" pageNumber="87">
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCFFDFBDEFE2D2E50" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[199,441,1115,1141]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF4C97FCFFDFBDEFE2D2E50" box="[199,441,1115,1141]" italics="true" pageId="11" pageNumber="87">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF4C97FCEFBFBDEFD8F2E50" bold="true" box="[449,539,1115,1141]" pageId="11" pageNumber="87">
|
||
<taxonomicNameLabel id="8088770EFFF4C97FCEFBFBDEFD8F2E50" box="[449,539,1115,1141]" pageId="11" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCD89FBD9FC8B2E50" box="[691,799,1116,1141]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="igneus">
|
||
<emphasis id="9BBBCA75FFF4C97FCD89FBD9FC8B2E50" box="[691,799,1116,1141]" italics="true" pageId="11" pageNumber="87">H. igneus</emphasis>
|
||
</taxonomicName>
|
||
by having 17 DSR at midbody (vs. 19), subcaudals 112 (vs. 115–129), preoculars 1 (vs. 2), a conspicuous, beige dorsolateral stripe extending from behind the eye to the end of the tail on each side (vs. a dorsolateral series of bright orange, coral or rusty red blotches), and the background color of the anterior venter creamish-yellow (vs. venter overall dark, i.e., dark brown, blackish-brown or black).
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF4C97FCFFDFB6EFC432F9C" blockId="11.[151,1436,1115,2041]" pageId="11" pageNumber="87">
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCFFDFB6EFE2F2F20" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[199,443,1259,1285]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF4C97FCFFDFB6EFE2F2F20" box="[199,443,1259,1285]" italics="true" pageId="11" pageNumber="87">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF4C97FCEFCFB6EFDB72F20" bold="true" box="[454,547,1259,1285]" pageId="11" pageNumber="87">
|
||
<taxonomicNameLabel id="8088770EFFF4C97FCEFCFB6EFDB72F20" box="[454,547,1259,1285]" pageId="11" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCDF8FB69FCF82F20" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[706,876,1260,1285]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="9BBBCA75FFF4C97FCDF8FB69FCF82F20" box="[706,876,1260,1285]" italics="true" pageId="11" pageNumber="87">H. annamensis</emphasis>
|
||
</taxonomicName>
|
||
by having DSR 19-17-17, feebly keeled except the outermost rows (vs. 15 or 17 dorsal scale rows at midbody, moderately keeled at midbody, strongly keeled posteriorly, usually smooth on the 1st dorsal scale row), prefrontals 2 (vs. prefrontal single), subcaudals 112 (vs. 116–146), preoculars 1 (vs. 2), and a conspicuous, beige dorsolateral stripe extending from behind the eye to the end of the tail on each side (vs. dorsal pattern made of bright orange or rusty-red blotches, usually transversally elongate as crossbars anteriorly, then becoming irregular, broken spots posteriorly).
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF4C97FCFFDFA46FAB52CB4" blockId="11.[151,1436,1115,2041]" pageId="11" pageNumber="87">
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCFFDFA46FE232FF8" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[199,439,1475,1501]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF4C97FCFFDFA46FE232FF8" box="[199,439,1475,1501]" italics="true" pageId="11" pageNumber="87">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF4C97FCE84FA46FD8C2FF8" bold="true" box="[446,536,1475,1501]" pageId="11" pageNumber="87">
|
||
<taxonomicNameLabel id="8088770EFFF4C97FCE84FA46FD8C2FF8" box="[446,536,1475,1501]" pageId="11" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCD96FA41FC562FF8" authority="(Wall, 1910)" baseAuthorityName="Wall" baseAuthorityYear="1910" box="[684,962,1475,1501]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="venningi">
|
||
<emphasis id="9BBBCA75FFF4C97FCD96FA41FCBA2FF8" box="[684,814,1476,1501]" italics="true" pageId="11" pageNumber="87">H. venningi</emphasis>
|
||
(
|
||
<bibRefCitation id="CD5E6B96FFF4C97FCC07FA46FC2E2FF8" author="Wall, F." box="[829,954,1475,1501]" pageId="11" pageNumber="87" pagination="345 - 346" refId="ref9854" refString="Wall, F. (1910) A new Tropidonotus from the Chin Hills. Journal of the Bombay Natural History Society, 20 (2), 345 - 346" type="journal article" year="1910">Wall, 1910</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by having 19 dorsal scale rows at anterior body, dorsal scales feebly keeled except the outermost rows (vs. 17 dorsal scale rows at anterior body, dorsal scales moderately keeled at midbody, strongly keeled posteriorly but 1st DSR smooth), subcaudals 112 (vs. 115–129), postoculars 2 (vs. 3), and a conspicuous, beige dorsolateral stripe extending from behind the eye to the end of the tail on each side (vs. a dorsolateral series of irregular blotches, yellow-ochre or yellowish-brown, enlarged and forming a chain on the first quarter to third of the body, progressively smaller, usually vanishing after midbody).
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF4C97FCFFDF91EFC6B2D4C" blockId="11.[151,1436,1115,2041]" pageId="11" pageNumber="87">
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCFFDF91EFE2D2C90" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[199,441,1691,1717]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF4C97FCFFDF91EFE2D2C90" box="[199,441,1691,1717]" italics="true" pageId="11" pageNumber="87">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF4C97FCEF8F91EFD8A2C90" bold="true" box="[450,542,1691,1717]" pageId="11" pageNumber="87">
|
||
<taxonomicNameLabel id="8088770EFFF4C97FCEF8F91EFD8A2C90" box="[450,542,1691,1717]" pageId="11" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCD8DF919FC7B2C93" authority="(Wall, 1925)" baseAuthorityName="Wall" baseAuthorityYear="1925" box="[695,1007,1691,1718]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="nigriventer">
|
||
<emphasis id="9BBBCA75FFF4C97FCD8DF919FCC32C90" box="[695,855,1692,1717]" italics="true" pageId="11" pageNumber="87">H. nigriventer</emphasis>
|
||
(
|
||
<bibRefCitation id="CD5E6B96FFF4C97FCC52F91EFC732C93" author="Wall, F." box="[872,999,1691,1718]" pageId="11" pageNumber="87" pagination="805 - 821" refId="ref9885" refString="Wall, F. (1925) Notes on snakes collected in Burma in 1924. Journal of the Bombay Natural History Society, 30 (4), 805 - 821." type="journal article" year="1925">Wall, 1925</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by having 19 DSR counts at anterior body (vs. 17), preoculars 1 (vs. 2), anterior venter creamish-yellow, with irregular black blotches in the middle and outer quarter of ventrals, the posterior part almost completely black (vs. venter entirely very dark, sometimes with two, narrow longitudinal paler stripes), and a conspicuous, beige dorsolateral stripe extending from behind the eye to the end of the tail on each side (vs. a bright orange, rusty-red or reddish-brown, rectangular blotches,, becoming smaller posteriorly and turning to an irregular stripe posteriorly on each side).
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF4C97FCFFDF8F6FAF32DDC" blockId="11.[151,1436,1115,2041]" pageId="11" pageNumber="87">
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCFFDF8F6FE2D2DA8" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[199,441,1907,1933]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF4C97FCFFDF8F6FE2D2DA8" box="[199,441,1907,1933]" italics="true" pageId="11" pageNumber="87">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF4C97FCEFBF8F6FD882DA8" bold="true" box="[449,540,1907,1933]" pageId="11" pageNumber="87">
|
||
<taxonomicNameLabel id="8088770EFFF4C97FCEFBF8F6FD882DA8" box="[449,540,1907,1933]" pageId="11" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCD8FF8F1FC662DA8" authority="(Smith, 1940)" baseAuthorityName="Smith" baseAuthorityYear="1940" box="[693,1010,1907,1933]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="taronensis">
|
||
<emphasis id="9BBBCA75FFF4C97FCD8FF8F1FCD92DA8" box="[693,845,1908,1933]" italics="true" pageId="11" pageNumber="87">H. taronensis</emphasis>
|
||
(
|
||
<bibRefCitation id="CD5E6B96FFF4C97FCC66F8F6FC7D2DA8" author="Smith, M. A." box="[860,1001,1907,1933]" pageId="11" pageNumber="87" pagination="465 - 486" refId="ref9593" refString="Smith, M. A. (1940) The amphibians and reptiles obtained by Mr. Ronald Kaulback in Upper Burma. Records of the Indian Museum, 43 (3), 465 - 486, pl. 7. https: // doi. org / 10.26515 / rzsi / v 42 / i 3 / 1940 / 162431" type="journal article" year="1940">Smith, 1940</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by having 19-17-17 DSR counts (vs. 17-17-17), relatively longer tail length (TAL/TL ratio 0.30 vs. TAL/TL <0.29), PRO 1, PO 2 (vs. PRO 2, PO 3), and a conspicuous, beige dorsolateral stripe extending from behind the eye to the end of the tail on each side (vs. a series of small, elongate, dorsolateral blotches, pale yellowish-brown or ochre-brown on each side in
|
||
<taxonomicName id="6ECF6DE4FFF4C97FCBFAF865FAC32DDC" baseAuthorityName="Smith" baseAuthorityYear="1940" box="[1216,1367,2016,2041]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="11" pageNumber="87" phylum="Chordata" rank="species" species="taronensis">
|
||
<emphasis id="9BBBCA75FFF4C97FCBFAF865FAC32DDC" box="[1216,1367,2016,2041]" italics="true" pageId="11" pageNumber="87">H. taronensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A9701667FFF3C978CFFDFF12FA032B88" blockId="12.[151,1437,151,429]" pageId="12" pageNumber="88">
|
||
In addition, with other seven congeners that have 17 DSR at midbody.
|
||
<taxonomicName id="6ECF6DE4FFF3C978CCCEFF12FB722A95" authority="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang, 2023" authorityName="Xu & Yang & Gong & Wu & Huang & Liu & Liang & Huang & Huang" authorityYear="2023" box="[1012,1254,151,177]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="youjiangensis" status="sp. nov.">
|
||
<emphasis id="9BBBCA75FFF3C978CCCEFF12FB722A95" box="[1012,1254,151,177]" italics="true" pageId="12" pageNumber="88">Hebius youjiangensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9BBBCA75FFF3C978CBD4FF12FADE2A94" bold="true" box="[1262,1354,151,177]" pageId="12" pageNumber="88">
|
||
<taxonomicNameLabel id="8088770EFFF3C978CBD4FF12FADE2A94" box="[1262,1354,151,177]" pageId="12" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated from
|
||
<taxonomicName id="6ECF6DE4FFF3C978CE70FF3EFD5A2AF0" authority="(David & Vogel, 2010)" baseAuthorityName="David & Vogel" baseAuthorityYear="2010" box="[330,718,187,213]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="arquus">
|
||
<emphasis id="9BBBCA75FFF3C978CE70FF3EFE2F2AF1" box="[330,443,187,212]" italics="true" pageId="12" pageNumber="88">H. arquus</emphasis>
|
||
(
|
||
<bibRefCitation id="CD5E6B96FFF3C978CEF1FF3EFD522AF0" author="David, P. & Vogel, G." box="[459,710,187,213]" pageId="12" pageNumber="88" pagination="121 - 127" refId="ref8237" refString="David, P. & Vogel, G. (2010) A new species of the natricine snake genus Amphiesma from Borneo (Squamata: Natricidae). Russian Journal of Herpetology, 17 (2), 121 - 127." type="journal article" year="2010">David & Vogel, 2010</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF3C978CDE6FF3EFBD62AF0" authority="(Bourret, 1934)" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[732,1090,187,213]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="atemporalis">
|
||
<emphasis id="9BBBCA75FFF3C978CDE6FF3EFC122AF0" box="[732,902,187,213]" italics="true" pageId="12" pageNumber="88">H. atemporalis</emphasis>
|
||
(
|
||
<bibRefCitation id="CD5E6B96FFF3C978CCADFF3EFBAE2AF0" author="Bourret, R." box="[919,1082,187,213]" pageId="12" pageNumber="88" pagination="3 - 12" refId="ref8021" refString="Bourret, R. (1934) Notes herpetologiques sur l'Indochine francaise. III. Ophidiens d'Annam et du Moyen Laos. Bulletin general de l'Instruction Publique, 13 e Annee (1933 - 1934), 9 Mai 1934, 167 - 176. [Separate reprint: 3 - 12]" type="book chapter" year="1934">Bourret, 1934</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF3C978CB6AFF3EFAE72AF0" authority="(Dunn, 1923)" baseAuthorityName="Dunn" baseAuthorityYear="1923" box="[1104,1395,187,213]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="frenatus">
|
||
<emphasis id="9BBBCA75FFF3C978CB6AFF3EFB5B2AF0" box="[1104,1231,187,213]" italics="true" pageId="12" pageNumber="88">H. frenatus</emphasis>
|
||
(Dunn, 1923)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF3C978CABBFF3EFE4B2ADC" authority="(Gunther, 1872)" baseAuthorityName="Gunther" baseAuthorityYear="1872" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="sarwacensis">
|
||
<emphasis id="9BBBCA75FFF3C978CABBFF3EFEB42ADD" italics="true" pageId="12" pageNumber="88">H. sarwacensis</emphasis>
|
||
(Günther, 1872)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF3C978CED1FF5AFCA02ADC" authority="(Boulenger, 1909)" baseAuthorityName="Boulenger" baseAuthorityYear="1909" box="[491,820,223,249]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="sauteri">
|
||
<emphasis id="9BBBCA75FFF3C978CED1FF5AFDC92ADD" box="[491,605,223,248]" italics="true" pageId="12" pageNumber="88">H. sauteri</emphasis>
|
||
(Boulenger, 1909)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="6ECF6DE4FFF3C978CC4BFF5AFB212ADC" authority="(Malnate, 1962)" baseAuthorityName="Malnate" baseAuthorityYear="1962" box="[881,1205,223,249]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="maximus">
|
||
<emphasis id="9BBBCA75FFF3C978CC4BFF5AFC6C2ADD" box="[881,1016,223,248]" italics="true" pageId="12" pageNumber="88">H. maximus</emphasis>
|
||
(Malnate, 1962)
|
||
</taxonomicName>
|
||
by having 19-17-17 DSR counts (vs. 17-17-
|
||
<quantity id="6E37BB82FFF3C978CEA4FE87FE432B38" box="[414,471,258,285]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="12" pageNumber="88" unit="in" value="15.0">15 in</quantity>
|
||
<taxonomicName id="6ECF6DE4FFF3C978CEE4FE86FDD92B39" baseAuthorityName="David & Vogel" baseAuthorityYear="2010" box="[478,589,259,284]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="arquus">
|
||
<emphasis id="9BBBCA75FFF3C978CEE4FE86FDD92B39" box="[478,589,259,284]" italics="true" pageId="12" pageNumber="88">H. arquus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6ECF6DE4FFF3C978CDB8FE86FD6B2B38" baseAuthorityName="Dunn" baseAuthorityYear="1923" box="[642,767,259,285]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="frenatus">
|
||
<emphasis id="9BBBCA75FFF3C978CDB8FE86FD6B2B38" box="[642,767,259,285]" italics="true" pageId="12" pageNumber="88">H. frenatus</emphasis>
|
||
</taxonomicName>
|
||
, and 17-17-
|
||
<quantity id="6E37BB82FFF3C978CCBCFE86FC2B2B38" box="[902,959,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="12" pageNumber="88" unit="in" value="17.0">17 in</quantity>
|
||
<taxonomicName id="6ECF6DE4FFF3C978CCFCFE86FBFA2B38" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[966,1134,259,285]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="atemporalis">
|
||
<emphasis id="9BBBCA75FFF3C978CCFCFE86FBFA2B38" box="[966,1134,259,285]" italics="true" pageId="12" pageNumber="88">H. atemporalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="9BBBCA75FFF3C978CB41FE86FA082B39" box="[1147,1436,259,284]" italics="true" pageId="12" pageNumber="88">
|
||
<taxonomicName id="6ECF6DE4FFF3C978CB41FE86FAB22B39" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[1147,1318,259,284]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="sarwacensis">H. sarwacensis</taxonomicName>
|
||
<taxonomicName id="6ECF6DE4FFF3C978CA17FE86FA082B39" baseAuthorityName="Boulenger" baseAuthorityYear="1909" box="[1325,1436,259,284]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="sauteri">H. sauteri</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="6ECF6DE4FFF3C978CFFCFEA2FED82B65" baseAuthorityName="Malnate" baseAuthorityYear="1962" box="[198,332,295,320]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="maximus">
|
||
<emphasis id="9BBBCA75FFF3C978CFFCFEA2FED82B65" box="[198,332,295,320]" italics="true" pageId="12" pageNumber="88">H. maximus</emphasis>
|
||
</taxonomicName>
|
||
), from
|
||
<taxonomicName id="6ECF6DE4FFF3C978CE9AFEA2FD6F2B64" authority="(Smith, 1922)" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[416,763,295,321]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="groundwateri">
|
||
<emphasis id="9BBBCA75FFF3C978CE9AFEA2FDCD2B64" box="[416,601,295,321]" italics="true" pageId="12" pageNumber="88">H. groundwateri</emphasis>
|
||
(Smith, 1922)
|
||
</taxonomicName>
|
||
by having divided cloacal plate (vs. CP entire), preoculars 1 (vs. 2), ventrals 3+160 (vs. 147), and subcaudals 112 (vs. 120). Furthermore, the new species differs from
|
||
<taxonomicName id="6ECF6DE4FFF3C978CA14FECEFA082B41" baseAuthorityName="David & Vogel" baseAuthorityYear="2010" box="[1326,1436,331,356]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="arquus">
|
||
<emphasis id="9BBBCA75FFF3C978CA14FECEFA082B41" box="[1326,1436,331,356]" italics="true" pageId="12" pageNumber="88">H. arquus</emphasis>
|
||
</taxonomicName>
|
||
by having a single loreal (vs. the absence of a loreal). It differs from
|
||
<taxonomicName id="6ECF6DE4FFF3C978CC90FEEAFBC72BAC" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[938,1107,367,393]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="atemporalis">
|
||
<emphasis id="9BBBCA75FFF3C978CC90FEEAFBC72BAC" box="[938,1107,367,393]" italics="true" pageId="12" pageNumber="88">H. atemporalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF3C978CB58FEEAFB752BAC" baseAuthorityName="Dunn" baseAuthorityYear="1923" box="[1122,1249,367,393]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="frenatus">
|
||
<emphasis id="9BBBCA75FFF3C978CB58FEEAFB752BAC" box="[1122,1249,367,393]" italics="true" pageId="12" pageNumber="88">H. frenatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6ECF6DE4FFF3C978CBCAFEEAFA082BAD" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[1264,1436,367,392]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="sarwacensis">
|
||
<emphasis id="9BBBCA75FFF3C978CBCAFEEAFA082BAD" box="[1264,1436,367,392]" italics="true" pageId="12" pageNumber="88">H. sarwacensis</emphasis>
|
||
</taxonomicName>
|
||
also by having 9 supralabials (vs.
|
||
<quantity id="6E37BB82FFF3C978CD29FE16FDAA2B88" box="[531,574,403,429]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="12" pageNumber="88" unit="in" value="6.0">6 in</quantity>
|
||
<taxonomicName id="6ECF6DE4FFF3C978CD7FFE16FD792B88" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[581,749,403,429]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="atemporalis">
|
||
<emphasis id="9BBBCA75FFF3C978CD7FFE16FD792B88" box="[581,749,403,429]" italics="true" pageId="12" pageNumber="88">H. atemporalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="6E37BB82FFF3C978CDC1FE17FCD62B88" box="[763,834,402,429]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.032" metricValueMin="1.27" pageId="12" pageNumber="88" unit="in" value="6.5" valueMax="8.0" valueMin="5.0">5–8 in</quantity>
|
||
<taxonomicName id="6ECF6DE4FFF3C978CC73FE16FC2D2B89" baseAuthorityName="Boulenger" baseAuthorityYear="1909" box="[841,953,403,428]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="sauteri">
|
||
<emphasis id="9BBBCA75FFF3C978CC73FE16FC2D2B89" box="[841,953,403,428]" italics="true" pageId="12" pageNumber="88">H. sauteri</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity id="6E37BB82FFF3C978CCCCFE16FBB52B88" box="[1014,1057,403,429]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="12" pageNumber="88" unit="in" value="8.0">8 in</quantity>
|
||
<taxonomicName id="6ECF6DE4FFF3C978CB12FE16FB312B88" baseAuthorityName="Dunn" baseAuthorityYear="1923" box="[1064,1189,403,429]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="frenatus">
|
||
<emphasis id="9BBBCA75FFF3C978CB12FE16FB312B88" box="[1064,1189,403,429]" italics="true" pageId="12" pageNumber="88">H. frenatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6ECF6DE4FFF3C978CBE6FE16FA132B89" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[1244,1415,403,428]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="12" pageNumber="88" phylum="Chordata" rank="species" species="sarwacensis">
|
||
<emphasis id="9BBBCA75FFF3C978CBE6FE16FA132B89" box="[1244,1415,403,428]" italics="true" pageId="12" pageNumber="88">H. sarwacensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |