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<mods:title id="E9D9467245A92FD1BC794B763416F696">Description of two new Australian species of Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) and females of two further species with notes on their biogeography</mods:title>
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<mods:namePart id="72E6B9D7768A62FCF868B8576A0FB6AC">Moore, Michael D.</mods:namePart>
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<mods:namePart id="231876622F5E7D64D804C0B990CDF367">Beaver, Ethan P.</mods:namePart>
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<mods:affiliation id="356CC81D54A4BD8E2AEA3C59273A0F39">Biological and Earth Sciences, South Australian Museum, SA 5000, Australia &amp; ethan. beaver @ live. com. au; https: // orcid. org / 0000 - 0002 - 0613 - 7046</mods:affiliation>
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<mods:namePart id="8A8CC17967E2AE3B04B5BD6B4F857FF2">Velasco-Castrillón, Alejandro</mods:namePart>
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<mods:affiliation id="B1926340DDD6439A00D41293D6AFF44E">Biological and Earth Sciences, South Australian Museum, SA 5000, Australia &amp; a. velascocastrillon @ gmail. com; https: // orcid. org / 0000 - 0002 - 3516 - 6655</mods:affiliation>
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<mods:namePart id="4D278BFED15C88D8EA6A11B1BDBFAFE8">Stevens, Mark I.</mods:namePart>
<mods:affiliation id="DA0A761A100C587E3EEFC400A86A3B35">Biological and Earth Sciences, South Australian Museum, SA 5000, Australia &amp; University of South Australia, Clinical and Health Sciences, SA 5000, Australia.</mods:affiliation>
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<treatment id="216587E00D23FF86EFC4F51F3472FF06" ID-DOI="http://doi.org/10.5281/zenodo.4401272" ID-GBIF-Taxon="177010364" ID-Zenodo-Dep="4401272" LSID="urn:lsid:plazi:treatment:216587E00D23FF86EFC4F51F3472FF06" httpUri="http://treatment.plazi.org/id/216587E00D23FF86EFC4F51F3472FF06" lastPageId="19" lastPageNumber="89" pageId="15" pageNumber="86" scope_family="Hepialidae" scope_order="Lepidoptera">
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<taxonomicName id="6ECC4D750D23FF9AEFC4F51F3220FB82" ID-CoL="99VVK" authority="Simonsen, 2018" authorityName="Simonsen" authorityYear="2018" box="[151,579,1122,1149]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D23FF9AEFC4F51F311AFB83" bold="true" box="[151,377,1122,1149]" italics="true" pageId="15" pageNumber="86">Abantiades pallida</emphasis>
<bibRefCitation id="CD5D4B070D23FF9AEED3F51F3220FB82" author="Simonsen, T. J." box="[384,579,1122,1148]" pageId="15" pageNumber="86" pagination="1 - 300" refId="ref12387" refString="Simonsen, T. J. (2018) Splendid Ghost Moths and Their Allies. Monographs on Australian Lepidoptera, 12, i-xii + 1 - 300. https: // doi. org / 10.1071 / 9781486307487" type="journal article" year="2018">Simonsen, 2018</bibRefCitation>
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(female)
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</paragraph>
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<figureCitation id="31F72A730D23FF9AEFC4F5FA30B8FB5F" box="[151,219,1159,1185]" captionStart="FIGURE 8" captionStartId="16.[151,260,1596,1622]" captionTargetBox="[162,1420,360,1556]" captionTargetId="figure-117@16.[151,1436,342,1571]" captionTargetPageId="16" captionText="FIGURE 8. Abantiades pallida. (a) Adult male. SAMA. West. Aust.; Wheatbelt, Wave Rock, Hyden, 32o2631”S, 118o5349”E, M+M Moore; 12 May 17. Specimen No 17162, Leg removed for DNA analysis; Sample designation H036. SAMA No. 31-020257. (b) Adult female. West.Aust. Wheatbelt, Koorda Caravan Park, Koorda, 30o4918”S, 117o2912”E, M+M Moore 14 May 17. Spec. No., 17185, leg removed, for tissue, storage. M.D. Moore. Sample designation No H 043. SAMA No. 31-020314." figureDoi="http://doi.org/10.5281/zenodo.4401286" httpUri="https://zenodo.org/record/4401286/files/figure.png" pageId="15" pageNumber="86">Figs 8</figureCitation>
ab, 9ab.
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<paragraph id="A97336F60D23FF9AEFC4F5B23114FB17" blockId="15.[151,1437,1231,1437]" box="[151,375,1231,1257]" pageId="15" pageNumber="86">Material Examined:</paragraph>
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;
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<collectingMunicipality id="4917AC8C0D23FF9AED08F58E332CFAF3" box="[603,847,1267,1293]" pageId="15" pageNumber="86">Koorda Caravan Park</collectingMunicipality>
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4918”S
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2912”E
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, M+
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14
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17.
<collectorName id="043953200D23FF9AEFAAF46A310BFACF" box="[249,360,1303,1329]" pageId="15" pageNumber="86">Spec. No.</collectorName>
, 17185, leg removed, for tissue, storage.
<collectorName id="043953200D23FF9AEC7EF46A3422FACF" box="[813,1089,1303,1329]" pageId="15" pageNumber="86">M.D. Moore. Sample No</collectorName>
H 043
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.
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<collectionCode id="CFDDAE330D23FF9AEBCBF46A3484FACF" box="[1176,1255,1303,1329]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="15" pageNumber="86" type="Museum">SAMA</collectionCode>
<location id="AC13602D0D23FF9AEBBEF46A3575FACF" LSID="urn:lsid:plazi:treatment:216587E00D23FF86EFC4F51F3472FF06:AC13602D0D23FF9AEBBEF46A3575FACF" box="[1261,1302,1303,1329]" name="No." pageId="15" pageNumber="86">No.</location>
31-020314
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;
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3819”S
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0007”E
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<date id="DD7210360D23FF9AEAD1F446316EFA87" pageId="15" pageNumber="86" value="2018-05-07">
<collectingDate id="CD36E9DE0D23FF9AEAD1F446316EFA87" pageId="15" pageNumber="86" value="2018-05-07">
7
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<collectorName id="043953200D23FF9AEE45F42231E7FA87" box="[278,388,1375,1401]" pageId="15" pageNumber="86">Spec. No.</collectorName>
18122, leg removed, for tissue, storage.
<collectorName id="043953200D23FF9AEC14F422343EFA87" box="[839,1117,1375,1401]" pageId="15" pageNumber="86">M.D. Moore. Sample No</collectorName>
H 122.
<collectingMunicipality id="4917AC8C0D23FF9AEBE6F4223541FA87" box="[1205,1314,1375,1401]" pageId="15" pageNumber="86">Dissected</collectingMunicipality>
E.
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,
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16
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March 2020
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</materialsCitation>
.
<materialsCitation id="19A43CAB0D23FF9AEE0EF4FE3203FA63" ID-GBIF-Occurrence="3019881309" box="[349,608,1411,1437]" collectionCode="SAMA" location="No." pageId="15" pageNumber="86" specimenCount="1">
<collectionCode id="CFDDAE330D23FF9AEE0EF4FE31CFFA63" box="[349,428,1411,1437]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="15" pageNumber="86" type="Museum">SAMA</collectionCode>
<location id="AC13602D0D23FF9AEEE0F4FE31BEFA63" LSID="urn:lsid:plazi:treatment:216587E00D23FF86EFC4F51F3472FF06:AC13602D0D23FF9AEEE0F4FE31BEFA63" box="[435,477,1411,1437]" name="No." pageId="15" pageNumber="86">No.</location>
31-020414
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E1D6657D0D23FF9AEFC4F4933355F967" pageId="15" pageNumber="86" type="diagnosis">
<paragraph id="A97336F60D23FF9AEFC4F493317AF9F6" blockId="15.[151,281,1518,1544]" box="[151,281,1518,1544]" pageId="15" pageNumber="86">
<heading id="F23B819A0D23FF9AEFC4F493317AF9F6" bold="true" box="[151,281,1518,1544]" fontSize="11" level="1" pageId="15" pageNumber="86" reason="1">
<emphasis id="9BB8EAE40D23FF9AEFC4F493317AF9F6" bold="true" box="[151,281,1518,1544]" pageId="15" pageNumber="86">Diagnosis:</emphasis>
</heading>
</paragraph>
<paragraph id="A97336F60D23FF9AEFC4F74A3355F967" blockId="15.[151,1437,1591,1689]" pageId="15" pageNumber="86">
A large grey moth with a distinct white submarginal line but only an indistinct brown/grey coloured discoidal mark. Specifically, the presence of setose covered nipple shaped projections on the lateral lobes of the antevaginal lamella is unique in all described triforked females (
<figureCitation id="31F72A730D23FF9AEDD5F70232ABF967" box="[646,712,1663,1689]" captionStart="FIGURE 9" captionStartId="17.[151,260,1702,1728]" captionTargetBox="[182,1411,1189,1660]" captionTargetId="figure-385@17.[151,1437,1170,1678]" captionTargetPageId="17" captionText="FIGURE 9. Abantiades pallida. (West.Aust. Nullarbor, Quarantine Station, Border Village, 31o3819”S, 129o07”E, R. Penning 7 May 18. Spec. No. 18122, leg removed, for tissue, storage. M.D. Moore. Sample designation No. H 122. SAMA No. 31-020414. Dissected E.P. Beaver.) (left) posterior view female genitalia, (right) lateral view female genitalia. Morphological structures: Dp: dorsal plate; Lal: lateral lobe of the antevaginal lamella; Mal: medial lobe of the antevaginal lamella; Sub: sub-anal plate. Arrow: Setose nipple." figureDoi="http://doi.org/10.5281/zenodo.4401288" httpUri="https://zenodo.org/record/4401288/files/figure.png" pageId="15" pageNumber="86">Fig. 9</figureCitation>
arrow).
</paragraph>
</subSubSection>
<subSubSection id="E1D6657D0D23FF84EFC4F79731DBFCC7" lastPageId="17" lastPageNumber="88" pageId="15" pageNumber="86" type="description">
<paragraph id="A97336F60D23FF9AEFC4F7973151F8FA" blockId="15.[151,306,1770,1796]" box="[151,306,1770,1796]" pageId="15" pageNumber="86">
<heading id="F23B819A0D23FF9AEFC4F7973151F8FA" bold="true" box="[151,306,1770,1796]" fontSize="11" level="1" pageId="15" pageNumber="86" reason="1">
<emphasis id="9BB8EAE40D23FF9AEFC4F7973151F8FA" bold="true" box="[151,306,1770,1796]" pageId="15" pageNumber="86">Description:</emphasis>
</heading>
</paragraph>
<paragraph id="A97336F60D23FF9AEFC4F64E308AF8B3" blockId="15.[151,1437,1843,2013]" box="[151,233,1843,1869]" pageId="15" pageNumber="86">Female</paragraph>
<paragraph id="A97336F60D23FF85EF94F62A3151FEBF" blockId="15.[151,1437,1843,2013]" lastBlockId="16.[151,1436,151,321]" lastPageId="16" lastPageNumber="87" pageId="15" pageNumber="86">
<emphasis id="9BB8EAE40D23FF9AEF94F62A316EF88F" box="[199,269,1879,1905]" italics="true" pageId="15" pageNumber="86">Head:</emphasis>
Dorsally covered in thick, elongate light grey scales, fronto-clypeal region dark grey. Each eye only slightly smaller than head capsule. Labial palps approximately straight, three segmented; basal palpomere thickly covered in long brown/grey scales, some whitish at base; second palpomere longest, thickly covered in light brown scales, darker ventrally; third palpomere, smallest, sub-spherical, densely covered in brown scales. Antennae: total 60 segments,
<quantity id="6E349B130D3CFF85EE63F1EA31CBFF4F" box="[304,424,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="16" pageNumber="87" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">1213 mm</quantity>
long, less than 20% forewing length; monopectinate, basal rami lamellar transitioning distally to spatulate then with lateral forks; rami cruciform on basal half then fully triple branched before spade shaped near the tip. Scattered robust dark setae and more numerous long colourless setae on central fork. Colourless setae present on upper surface of lateral fork and on faces. Pectinations lean slightly towards distal end and central fork a little more so.
</paragraph>
<caption id="FDB3667E0D3CFF85EFC4F74132C0F918" ID-DOI="http://doi.org/10.5281/zenodo.4401286" ID-Zenodo-Dep="4401286" httpUri="https://zenodo.org/record/4401286/files/figure.png" pageId="16" pageNumber="87" startId="16.[151,260,1596,1622]" targetBox="[162,1420,360,1556]" targetPageId="16">
<paragraph id="A97336F60D3CFF85EFC4F74132C0F918" blockId="16.[151,1436,1596,1767]" pageId="16" pageNumber="87">
<emphasis id="9BB8EAE40D3CFF85EFC4F7413142F9A8" bold="true" box="[151,289,1596,1622]" pageId="16" pageNumber="87">FIGURE 8.</emphasis>
<emphasis id="9BB8EAE40D3CFF85EE7BF7413267F9A8" box="[296,516,1596,1622]" italics="true" pageId="16" pageNumber="87">
<taxonomicName id="6ECC4D750D3CFF85EE7BF7413263F9A8" authorityName="Simonsen" authorityYear="2018" box="[296,512,1596,1622]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="pallida">Abantiades pallida</taxonomicName>
.
</emphasis>
<emphasis id="9BB8EAE40D3CFF85ED58F741324FF9A8" bold="true" box="[523,556,1596,1622]" pageId="16" pageNumber="87">(a)</emphasis>
Adult male. SAMA. West. Aust.; Wheatbelt, Wave Rock, Hyden, 32
<superScript id="5EB99BBE0D3CFF85EA63F741355BF9B4" attach="none" box="[1328,1336,1596,1610]" fontSize="6" pageId="16" pageNumber="87">o</superScript>
2631”S, 118
<superScript id="5EB99BBE0D3CFF85EF93F71D30ABF990" attach="none" box="[192,200,1632,1646]" fontSize="6" pageId="16" pageNumber="87">o</superScript>
5349”E, M+M Moore; 12 May 17. Specimen No 17162, Leg removed for DNA analysis; Sample designation H036. SAMA No. 31-020257.
<emphasis id="9BB8EAE40D3CFF85EEBEF7F9326CF960" bold="true" box="[493,527,1668,1694]" pageId="16" pageNumber="87">(b)</emphasis>
Adult female. West.Aust. Wheatbelt, Koorda Caravan Park, Koorda, 30
<superScript id="5EB99BBE0D3CFF85EA63F7F9355BF96C" attach="none" box="[1328,1336,1668,1682]" fontSize="6" pageId="16" pageNumber="87">o</superScript>
4918”S, 117
<superScript id="5EB99BBE0D3CFF85EF93F7D530ABF948" attach="none" box="[192,200,1704,1718]" fontSize="6" pageId="16" pageNumber="87">o</superScript>
2912”E, M+M Moore 14 May 17. Spec. No., 17185, leg removed, for tissue, storage. M.D. Moore. Sample designation No H 043. SAMA No. 31-020314.
</paragraph>
</caption>
<paragraph id="A97336F60D3CFF84EF94F66933A9FDE7" blockId="16.[151,1437,1812,2019]" lastBlockId="17.[151,1437,151,825]" lastPageId="17" lastPageNumber="88" pageId="16" pageNumber="87">
<emphasis id="9BB8EAE40D3CFF85EF94F6693142F8D0" box="[199,289,1812,1838]" italics="true" pageId="16" pageNumber="87">Thorax:</emphasis>
Dorsal surface densely clothed by medium grey, elongate, robust scales, slightly paler than head. Ventral surface light white grey. Forewing length
<quantity id="6E349B130D3CFF85EDF0F6453343F8AD" box="[675,800,1848,1875]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="6.8" metricValueMin="6.2" pageId="16" pageNumber="87" unit="mm" value="65.0" valueMax="68.0" valueMin="62.0">6268 mm</quantity>
, membrane transparent, veins whitish, browner towards base, covered in small dark grey scales; dorsal costa covered in small grey/brown scales; costa to Rs and apically Rs1, sparsely covered in dark grey scales. Two large marks on forewing present, an indistinct discoidal mark, medium brown basally, whiter distally and a distinct dull white submarginal band that extends between Rs2 to CuA1, linear but wider centrally and highlighted by narrow border of dark grey scales. The anterior portion of the discoidal mark has three or four light grey scroll lines present then out to
<collectionCode id="CFDDAE330D3DFF84EC34F1EA3319FF4F" box="[871,890,151,177]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="88">R</collectionCode>
and Rs1+2 a thin band of dark brown highlighting scales. Irregular obscure, white scaled scroll lines present over forewing, apically scroll marks elliptical, along termen, linear, with pale grey scales between lines. Dorsally and basally wing scales become longer and more red-brown, sparse throughout. Ventral surface: dorsal markings visible through wing; costa covered in small yellow-grey scales with longer grey-brown scales towards Sc. Apex with a few, thinly spread dark grey scales, rest of wing sparsely covered in white-grey piliform scales that are longer and more golden towards base. Hindwing length
<quantity id="6E349B130D3DFF84EFC4F0133170FE77" box="[151,275,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" metricValueMax="5.3" metricValueMin="4.5" pageId="17" pageNumber="88" unit="mm" value="49.0" valueMax="53.0" valueMin="45.0">4553 mm</quantity>
, veins white-yellow, membrane transparent. Dorsal surface, sparsely covered in long light grey to reddish-brown scales, basally piliform golden scales. Ventral surface, as for ventral forewing. Legs: femur dorsal surface covered by short yellow sandy-brown scales, lateral scales light grey; inner lateral surface light grey; ventrally whitish grey. Tibia dorsally sandy-grey, ventrally sandy yellow. Tarsus, dorsally sandy-grey, ventrally sandy-yellow, increasingly golden towards distal tarsus. Arolium and epiphysis present.
</paragraph>
<paragraph id="A97336F60D3DFF84EF94F35E3451FD9F" blockId="17.[151,1437,151,825]" pageId="17" pageNumber="88">
<emphasis id="9BB8EAE40D3DFF84EF94F35E3159FDC3" box="[199,314,547,573]" italics="true" pageId="17" pageNumber="88">Abdomen:</emphasis>
Dorsal surface thickly covered in long, dark grey piliform scales. Ventrally thickly covered in piliform scales, sternites IIII light white grey, IVV yellow-grey, VIVIII yellowish.
</paragraph>
<paragraph id="A97336F60D3DFF84EF94F31631DBFCC7" blockId="17.[151,1437,151,825]" pageId="17" pageNumber="88">
<emphasis id="9BB8EAE40D3DFF84EF94F3163159FD7B" box="[199,314,619,645]" italics="true" pageId="17" pageNumber="88">Genitalia:</emphasis>
(
<figureCitation id="31F72A730D3DFF84EE19F31631F3FD7B" box="[330,400,619,645]" captionStart="FIGURE 9" captionStartId="17.[151,260,1702,1728]" captionTargetBox="[182,1411,1189,1660]" captionTargetId="figure-385@17.[151,1437,1170,1678]" captionTargetPageId="17" captionText="FIGURE 9. Abantiades pallida. (West.Aust. Nullarbor, Quarantine Station, Border Village, 31o3819”S, 129o07”E, R. Penning 7 May 18. Spec. No. 18122, leg removed, for tissue, storage. M.D. Moore. Sample designation No. H 122. SAMA No. 31-020414. Dissected E.P. Beaver.) (left) posterior view female genitalia, (right) lateral view female genitalia. Morphological structures: Dp: dorsal plate; Lal: lateral lobe of the antevaginal lamella; Mal: medial lobe of the antevaginal lamella; Sub: sub-anal plate. Arrow: Setose nipple." figureDoi="http://doi.org/10.5281/zenodo.4401288" httpUri="https://zenodo.org/record/4401288/files/figure.png" pageId="17" pageNumber="88">Fig. 9</figureCitation>
). Dorsal plate lightly sclerotized, bilobed, each hemispherical, central “V shaped invagination, inner margin setose. Antevaginal lamella trilobed; medial lobe more heavily sclerotized, setose, inner margin continuously curved with wide, blunt ended lateral projections and small concave depression centrally. Inner margin lightly sclerotized and membranous in some places. Lateral lobes with setose nipple like projection which is quite distinct in the more lateral view (see
<figureCitation id="31F72A730D3DFF84ED7FF3863219FCEB" box="[556,634,763,789]" captionStart="FIGURE 9" captionStartId="17.[151,260,1702,1728]" captionTargetBox="[182,1411,1189,1660]" captionTargetId="figure-385@17.[151,1437,1170,1678]" captionTargetPageId="17" captionText="FIGURE 9. Abantiades pallida. (West.Aust. Nullarbor, Quarantine Station, Border Village, 31o3819”S, 129o07”E, R. Penning 7 May 18. Spec. No. 18122, leg removed, for tissue, storage. M.D. Moore. Sample designation No. H 122. SAMA No. 31-020414. Dissected E.P. Beaver.) (left) posterior view female genitalia, (right) lateral view female genitalia. Morphological structures: Dp: dorsal plate; Lal: lateral lobe of the antevaginal lamella; Mal: medial lobe of the antevaginal lamella; Sub: sub-anal plate. Arrow: Setose nipple." figureDoi="http://doi.org/10.5281/zenodo.4401288" httpUri="https://zenodo.org/record/4401288/files/figure.png" pageId="17" pageNumber="88">Fig. 9b</figureCitation>
arrow). Subanal plates with heavily sclerotized distal margin. No bursa copulatrix was recovered.
</paragraph>
</subSubSection>
<subSubSection id="E1D6657D0D3DFF84EFC4F2F73190FB83" pageId="17" pageNumber="88" type="distribution">
<paragraph id="A97336F60D3DFF84EFC4F2F731B5FC5A" blockId="17.[151,470,906,933]" box="[151,470,906,933]" pageId="17" pageNumber="88">
<heading id="F23B819A0D3DFF84EFC4F2F731B5FC5A" bold="true" box="[151,470,906,933]" fontSize="11" level="1" pageId="17" pageNumber="88" reason="1">
<emphasis id="9BB8EAE40D3DFF84EFC4F2F731B5FC5A" bold="true" box="[151,470,906,933]" pageId="17" pageNumber="88">Distribution and Biology:</emphasis>
</heading>
</paragraph>
<paragraph id="A97336F60D3DFF84EFC4F2AE3495FBEF" blockId="17.[151,1437,979,1149]" pageId="17" pageNumber="88">
<emphasis id="9BB8EAE40D3DFF84EFC4F2AE31E2FC13" box="[151,385,979,1005]" italics="true" pageId="17" pageNumber="88">Recorded Localities:</emphasis>
(
<figureCitation id="31F72A730D3DFF84EEC2F2AE3182FC13" box="[401,481,979,1005]" captionStart="FIGURE 11" captionStartId="19.[151,260,1460,1486]" captionTargetBox="[173,1428,279,1434]" captionTargetId="figure-70@19.[151,1436,270,1436]" captionTargetPageId="19" captionText="FIGURE 11. Distribution Map of Australia for Abantiades pallida and A. tembyi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4401292" httpUri="https://zenodo.org/record/4401292/files/figure.png" pageId="17" pageNumber="88">Fig. 11</figureCitation>
) Previously known only from Border Village,
<collectionCode id="CFDDAE330D3DFF84ECA1F2AE347BFC13" box="[1010,1048,979,1005]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="17" pageNumber="88">SA</collectionCode>
, and Eucla, WA (
<bibRefCitation id="CD5D4B070D3DFF84EB8FF2AE35F2FC13" author="Simonsen, T. J." box="[1244,1425,979,1005]" pageId="17" pageNumber="88" pagination="1 - 300" refId="ref12387" refString="Simonsen, T. J. (2018) Splendid Ghost Moths and Their Allies. Monographs on Australian Lepidoptera, 12, i-xii + 1 - 300. https: // doi. org / 10.1071 / 9781486307487" type="journal article" year="2018">Simonsen 2018</bibRefCitation>
). Now also from Coolgardie; Bullabulling; Hyden; Koorda; and Kellerberrin all in
<collectingRegion id="6B08F8140D3DFF84EB74F28A3492FBEF" box="[1063,1265,1015,1041]" country="Australia" name="Western Australia" pageId="17" pageNumber="88">Western Australia</collectingRegion>
.
</paragraph>
<paragraph id="A97336F60D3DFF84EF94F566337DFBA7" blockId="17.[151,1437,979,1149]" pageId="17" pageNumber="88">
<emphasis id="9BB8EAE40D3DFF84EF94F566313AFBCA" box="[199,345,1051,1077]" italics="true" pageId="17" pageNumber="88">Flight times:</emphasis>
<bibRefCitation id="CD5D4B070D3DFF84EE32F5663241FBCB" author="Simonsen, T. J." box="[353,546,1051,1077]" pageId="17" pageNumber="88" pagination="1 - 300" refId="ref12387" refString="Simonsen, T. J. (2018) Splendid Ghost Moths and Their Allies. Monographs on Australian Lepidoptera, 12, i-xii + 1 - 300. https: // doi. org / 10.1071 / 9781486307487" type="journal article" year="2018">Simonsen (2018)</bibRefCitation>
lists the flight period as between 28
<superScript id="5EB99BBE0D3DFF84EC92F56733ADFBD6" attach="left" box="[961,974,1050,1064]" fontSize="6" pageId="17" pageNumber="88">th</superScript>
of April to the 11
<superScript id="5EB99BBE0D3DFF84EBCFF56734CAFBD6" attach="left" box="[1180,1193,1050,1064]" fontSize="6" pageId="17" pageNumber="88">th</superScript>
of May. The females examined here were collected on the 04
<superScript id="5EB99BBE0D3DFF84ED07F5433202FBB2" attach="left" box="[596,609,1086,1100]" fontSize="6" pageId="17" pageNumber="88">th</superScript>
and 14
<superScript id="5EB99BBE0D3DFF84EDE0F54332A3FBB2" attach="left" box="[691,704,1086,1100]" fontSize="6" pageId="17" pageNumber="88">th</superScript>
of May.
</paragraph>
<paragraph id="A97336F60D3DFF84EF94F51E3190FB83" blockId="17.[151,1437,979,1149]" box="[199,499,1123,1149]" pageId="17" pageNumber="88">
<emphasis id="9BB8EAE40D3DFF84EF94F51E3115FB83" box="[199,374,1123,1149]" italics="true" pageId="17" pageNumber="88">Larval biology:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="FDB3667E0D3DFF84EFC4F7DB3306F88A" ID-DOI="http://doi.org/10.5281/zenodo.4401288" ID-Zenodo-Dep="4401288" httpUri="https://zenodo.org/record/4401288/files/figure.png" pageId="17" pageNumber="88" startId="17.[151,260,1702,1728]" targetBox="[182,1411,1189,1660]" targetPageId="17">
<paragraph id="A97336F60D3DFF84EFC4F7DB315EF8D2" blockId="17.[151,1436,1702,1909]" pageId="17" pageNumber="88">
<emphasis id="9BB8EAE40D3DFF84EFC4F7DB317DF93E" bold="true" box="[151,286,1702,1728]" pageId="17" pageNumber="88">FIGURE 9.</emphasis>
<emphasis id="9BB8EAE40D3DFF84EE70F7DB319DF93E" box="[291,510,1702,1728]" italics="true" pageId="17" pageNumber="88">
<taxonomicName id="6ECC4D750D3DFF84EE70F7DB3199F93E" authorityName="Simonsen" authorityYear="2018" box="[291,506,1702,1728]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="88" phylum="Arthropoda" rank="species" species="pallida">Abantiades pallida</taxonomicName>
.
</emphasis>
(West.Aust. Nullarbor, Quarantine Station, Border Village, 31
<superScript id="5EB99BBE0D3DFF84EBFCF7DB34D4F94A" attach="none" box="[1199,1207,1702,1716]" fontSize="6" pageId="17" pageNumber="88">o</superScript>
3819”S, 129
<superScript id="5EB99BBE0D3DFF84EA19F7DB3531F94A" attach="none" box="[1354,1362,1702,1716]" fontSize="6" pageId="17" pageNumber="88">o</superScript>
07”E, R. Penning 7 May 18. Spec. No. 18122, leg removed, for tissue, storage. M.D. Moore. Sample designation No. H 122. SAMA No. 31-020414. Dissected E.P. Beaver.)
<emphasis id="9BB8EAE40D3DFF84EDBDF793334BF8F6" bold="true" box="[750,808,1774,1800]" pageId="17" pageNumber="88">(left)</emphasis>
posterior view female genitalia,
<emphasis id="9BB8EAE40D3DFF84EBCFF7933489F8F6" bold="true" box="[1180,1258,1774,1800]" pageId="17" pageNumber="88">(right)</emphasis>
lateral view female genitalia.
</paragraph>
<paragraph id="A97336F60D3DFF84EFC4F64B3306F88A" blockId="17.[151,1436,1702,1909]" pageId="17" pageNumber="88">
Morphological structures:
<emphasis id="9BB8EAE40D3DFF84EE97F64A3192F8AF" bold="true" box="[452,497,1847,1873]" pageId="17" pageNumber="88">Dp:</emphasis>
dorsal plate;
<emphasis id="9BB8EAE40D3DFF84EDDDF64A32A3F8AF" bold="true" box="[654,704,1847,1873]" pageId="17" pageNumber="88">Lal:</emphasis>
lateral lobe of the antevaginal lamella;
<emphasis id="9BB8EAE40D3DFF84EBDAF64A34A0F8AF" bold="true" box="[1161,1219,1847,1873]" pageId="17" pageNumber="88">Mal:</emphasis>
medial lobe of the antevaginal lamella;
<emphasis id="9BB8EAE40D3DFF84EED3F62731DBF88A" bold="true" box="[384,440,1882,1908]" pageId="17" pageNumber="88">Sub:</emphasis>
sub-anal plate.
<emphasis id="9BB8EAE40D3DFF84ED3BF62732A3F88A" bold="true" box="[616,704,1882,1908]" pageId="17" pageNumber="88">Arrow:</emphasis>
Setose nipple.
</paragraph>
</caption>
<caption id="FDB3667E0D3EFF87EFC4F42631D9FA67" ID-DOI="http://doi.org/10.5281/zenodo.4401290" ID-Zenodo-Dep="4401290" httpUri="https://zenodo.org/record/4401290/files/figure.png" pageId="18" pageNumber="89" startId="18.[151,260,1371,1397]" targetBox="[172,1427,185,1340]" targetPageId="18">
<paragraph id="A97336F60D3EFF87EFC4F42631D9FA67" blockId="18.[151,1436,1371,1434]" pageId="18" pageNumber="89">
<emphasis id="9BB8EAE40D3EFF87EFC4F426314CFA8B" bold="true" box="[151,303,1371,1397]" pageId="18" pageNumber="89">FIGURE 10.</emphasis>
Distribution Map of Australia for
<taxonomicName id="6ECC4D750D3EFF87EDE9F42633D9FA8B" authorityName="Simonsen" authorityYear="2018" box="[698,954,1371,1397]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="kristenseni">
<emphasis id="9BB8EAE40D3EFF87EDE9F42633D9FA8B" box="[698,954,1371,1397]" italics="true" pageId="18" pageNumber="89">Abantiades kristenseni</emphasis>
</taxonomicName>
,
<taxonomicName id="6ECC4D750D3EFF87EC9AF4213436FA8B" authorityName="Simonsen" authorityYear="2018" box="[969,1109,1372,1398]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="inexpecta">
<emphasis id="9BB8EAE40D3EFF87EC9AF4213436FA8B" box="[969,1109,1372,1398]" italics="true" pageId="18" pageNumber="89">A. inexpecta</emphasis>
</taxonomicName>
,
<emphasis id="9BB8EAE40D3EFF87EB36F4213508FA8B" box="[1125,1387,1371,1398]" italics="true" pageId="18" pageNumber="89">
<taxonomicName id="6ECC4D750D3EFF87EB36F4213504FA8B" authorityName="Simonsen" authorityYear="2018" box="[1125,1383,1371,1398]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="macropusinsulariae">A. macropusinsulariae</taxonomicName>
,
</emphasis>
and
<taxonomicName id="6ECC4D750D3EFF87EFC4F4FD313AFA67" authority="Moore &amp; Beaver &amp; Velasco-Castrillón &amp; Stevens, 2020" authorityName="Moore &amp; Beaver &amp; Velasco-Castrillón &amp; Stevens" authorityYear="2020" box="[151,345,1407,1434]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="cephalocorvus" status="sp. nov.">
<emphasis id="9BB8EAE40D3EFF87EFC4F4FD313AFA67" box="[151,345,1407,1434]" italics="true" pageId="18" pageNumber="89">A. cephalocorvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="808B579F0D3EFF87EE32F40231D9FA67" box="[353,442,1407,1433]" pageId="18" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="E1D6657D0D3EFF86EFC4F4963472FF06" lastPageId="19" lastPageNumber="90" pageId="18" pageNumber="89" type="discussion">
<paragraph id="A97336F60D3EFF87EFC4F4963170F9FB" blockId="18.[151,275,1515,1541]" box="[151,275,1515,1541]" pageId="18" pageNumber="89">
<heading id="F23B819A0D3EFF87EFC4F4963170F9FB" bold="true" box="[151,275,1515,1541]" fontSize="11" level="1" pageId="18" pageNumber="89" reason="1">
<emphasis id="9BB8EAE40D3EFF87EFC4F4963170F9FB" bold="true" box="[151,275,1515,1541]" pageId="18" pageNumber="89">Remarks:</emphasis>
</heading>
</paragraph>
<paragraph id="A97336F60D3EFF87EFC4F74E348FF9B3" blockId="18.[151,1437,1587,2010]" box="[151,1260,1587,1614]" pageId="18" pageNumber="89">
See above for intra-cladal differences and comparison to
<taxonomicName id="6ECC4D750D3EFF87EC44F7493476F9B3" authorityName="Simonsen" authorityYear="2018" box="[791,1045,1587,1614]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="macropusinsulariae">
<emphasis id="9BB8EAE40D3EFF87EC44F7493476F9B3" box="[791,1045,1587,1614]" italics="true" pageId="18" pageNumber="89">A. macropusinsulariae</emphasis>
</taxonomicName>
and
<emphasis id="9BB8EAE40D3EFF87EB1FF749348FF9B3" box="[1100,1260,1587,1614]" italics="true" pageId="18" pageNumber="89">
<taxonomicName id="6ECC4D750D3EFF87EB1FF749348BF9B3" authorityName="Simonsen" authorityYear="2018" box="[1100,1256,1587,1614]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="kristenseni">A. kristenseni</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="A97336F60D3EFF87EF94F72A31B2F920" blockId="18.[151,1437,1587,2010]" pageId="18" pageNumber="89">
The females (
<accessionNumber id="B69FAB150D3EFF87EE34F72A3185F98C" box="[359,486,1623,1650]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT052733" pageId="18" pageNumber="89">MT052733</accessionNumber>
and
<accessionNumber id="B69FAB150D3EFF87ED76F72A32C7F98C" box="[549,676,1623,1650]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT052735" pageId="18" pageNumber="89">MT052735</accessionNumber>
) were identified as belonging to
<taxonomicName id="6ECC4D750D3EFF87EB78F72534FEF98F" authorityName="Simonsen" authorityYear="2018" box="[1067,1181,1623,1650]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D3EFF87EB78F72534FEF98F" box="[1067,1181,1623,1650]" italics="true" pageId="18" pageNumber="89">A. pallida</emphasis>
</taxonomicName>
by sequencing of the mtDNA (COI) gene, and are clustered within a clade that includes three male specimens (
<figureCitation id="31F72A730D3EFF87EBF8F7063493F96B" box="[1195,1264,1659,1685]" captionStart="FIGURE 1" captionStartId="3.[151,260,1815,1841]" captionTargetBox="[151,1427,283,1786]" captionTargetId="figure-67@3.[151,1436,270,1791]" captionTargetPageId="3" captionText="FIGURE 1. Maximum credibility tree from Bayesian phylogenetic analysis of the mitochondrial DNA COI gene of all 42 Abantiades sequences used and the ♀ specimens for A. macropusinsulariae and A. pallida. Posterior probabilities greater than 0.7 are shown at each node. Details for all sequences including GenBank accessions are available in Table S1." figureDoi="http://doi.org/10.5281/zenodo.4401270" httpUri="https://zenodo.org/record/4401270/files/figure.png" pageId="18" pageNumber="89">Fig. 1</figureCitation>
). Even though separated by hundreds of kilometres, neither of the two females had an obvious discoidal mark on their forewings, which is rare in
<taxonomicName id="6ECC4D750D3EFF87EE1FF7BE31A9F923" authorityName="Herrich-Schaffer" authorityYear="1855" box="[332,458,1731,1757]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="9BB8EAE40D3EFF87EE1FF7BE31A9F923" box="[332,458,1731,1757]" italics="true" pageId="18" pageNumber="89">Abantiades</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A97336F60D3EFF86EF94F79A3472FF06" blockId="18.[151,1437,1587,2010]" lastBlockId="19.[151,1437,151,249]" lastPageId="19" lastPageNumber="90" pageId="18" pageNumber="89">
<taxonomicName id="6ECC4D750D3EFF87EF94F79A31F8F8FF" authorityName="Simonsen" authorityYear="2018" box="[199,411,1767,1793]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D3EFF87EF94F79A31F8F8FF" box="[199,411,1767,1793]" italics="true" pageId="18" pageNumber="89">Abantiades pallida</emphasis>
</taxonomicName>
has a wide distribution that significantly overlaps with both
<taxonomicName id="6ECC4D750D3EFF87EB16F79534BEF8FF" authorityName="Simonsen" authorityYear="2018" box="[1093,1245,1767,1794]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="kristenseni">
<emphasis id="9BB8EAE40D3EFF87EB16F79534BEF8FF" box="[1093,1245,1767,1794]" italics="true" pageId="18" pageNumber="89">A. kristenseni</emphasis>
</taxonomicName>
and
<taxonomicName id="6ECC4D750D3EFF87EA46F79531AEF8DB" authority="Moore &amp; Beaver, 2020" authorityName="Moore &amp; Beaver" authorityYear="2020" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="zonatriticum">
<emphasis id="9BB8EAE40D3EFF87EA46F79530A5F8DB" italics="true" pageId="18" pageNumber="89">A. zonatriticum</emphasis>
<bibRefCitation id="CD5D4B070D3EFF87EF9FF67131AEF8DB" author="Beaver, E. P. &amp; Moore, M. D. &amp; Velasco-Castrillon, A. &amp; Stevens, M. I." box="[204,461,1803,1830]" pageId="18" pageNumber="89" pagination="351 - 374" refId="ref11553" refString="Beaver, E. P., Moore, M. D., Velasco-Castrillon, A. &amp; Stevens, M. I. (2020) Three new ghost moths of the genus Oxycanus Walker, 1856 from Australia (Lepidoptera: Hepialidae). Zootaxa, 4732 (3), 351 - 374. https: // doi. org / 10.11646 / zootaxa. 4732.3.1" type="journal article" year="2020">Moore &amp; Beaver, 2020</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="CD5D4B070D3EFF87EE88F67632CBF8DB" author="Moore, M. D. &amp; Beaver, E. P. &amp; Velasco-Castrillon, A. &amp; Stevens, M. I." box="[475,680,1803,1829]" pageId="18" pageNumber="89" pagination="115 - 141" refId="ref12260" refString="Moore, M. D., Beaver, E. P., Velasco-Castrillon, A. &amp; Stevens, M. I. (2020) Four new species of triforked Abantiades Herrich- Schaffer (Lepidoptera: Hepialidae) from the &quot; dark obscura clade &quot;. Zootaxa, 4801 (1), 115 - 141. https: // doi. org / 10.11646 / zootaxa. 4801.1.5" type="journal article" year="2020">
Moore
<emphasis id="9BB8EAE40D3EFF87ED7DF6713206F8DB" box="[558,613,1803,1829]" italics="true" pageId="18" pageNumber="89">et al.</emphasis>
2020
</bibRefCitation>
). The females of the triforked
<taxonomicName id="6ECC4D750D3EFF87ECABF6763415F8DB" authorityName="Herrich-Schaffer" authorityYear="1855" box="[1016,1142,1803,1829]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="9BB8EAE40D3EFF87ECABF6763415F8DB" box="[1016,1142,1803,1829]" italics="true" pageId="18" pageNumber="89">Abantiades</emphasis>
</taxonomicName>
species are inherently difficult to identify without dissection. However,
<taxonomicName id="6ECC4D750D3EFF87EDCFF64D336AF8B7" authorityName="Simonsen" authorityYear="2018" box="[668,777,1839,1866]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D3EFF87EDCFF64D336AF8B7" box="[668,777,1839,1866]" italics="true" pageId="18" pageNumber="89">A. pallida</emphasis>
</taxonomicName>
has one obvious white submarginal line and only an almost obscure discoidal one, while the other two species have two obvious marks (either white or brown). Antennal structure is useful;
<taxonomicName id="6ECC4D750D3EFF87EE64F6053186F86F" authorityName="Moore &amp; Beaver" authorityYear="2020" box="[311,485,1912,1938]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="zonatriticum">
<emphasis id="9BB8EAE40D3EFF87EE64F6053186F86F" box="[311,485,1912,1938]" italics="true" pageId="18" pageNumber="89">A. zonatriticum</emphasis>
</taxonomicName>
has triangular shaped lateral processes on the antenna whereas in
<taxonomicName id="6ECC4D750D3EFF87EB9EF6053558F86F" authorityName="Simonsen" authorityYear="2018" box="[1229,1339,1911,1938]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D3EFF87EB9EF6053558F86F" box="[1229,1339,1911,1938]" italics="true" pageId="18" pageNumber="89">A. pallida</emphasis>
</taxonomicName>
they are linear. Upon dissection of the genitalia,
<taxonomicName id="6ECC4D750D3EFF87ED0AF6E13364F84B" authorityName="Moore &amp; Beaver" authorityYear="2020" box="[601,775,1948,1974]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="89" phylum="Arthropoda" rank="species" species="zonatriticum">
<emphasis id="9BB8EAE40D3EFF87ED0AF6E13364F84B" box="[601,775,1948,1974]" italics="true" pageId="18" pageNumber="89">A. zonatriticum</emphasis>
</taxonomicName>
has a process which seems like an extension to the medial lobe (rather than part of it) of the antevaginal lamella and projects more into the central space (has triangular shaped lateral processes), whereas with
<taxonomicName id="6ECC4D750D3FFF86ED55F1EA3216FF4F" authorityName="Simonsen" authorityYear="2018" box="[518,629,151,177]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="90" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D3FFF86ED55F1EA3216FF4F" box="[518,629,151,177]" italics="true" pageId="19" pageNumber="90">A. pallida</emphasis>
</taxonomicName>
the projection is more part of the medial lobe, has a smaller projection into the central space with blunt ended lateral processes trapezoidal in shape. The lateral lobes of the antevaginal lamella have a nipple like protrusion in
<taxonomicName id="6ECC4D750D3FFF86ED06F1A232A0FF07" authorityName="Simonsen" authorityYear="2018" box="[597,707,223,249]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="90" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="9BB8EAE40D3FFF86ED06F1A232A0FF07" box="[597,707,223,249]" italics="true" pageId="19" pageNumber="90">A. pallida</emphasis>
</taxonomicName>
but not so in
<emphasis id="9BB8EAE40D3FFF86EC0FF1A23472FF06" box="[860,1041,223,249]" italics="true" pageId="19" pageNumber="90">
<taxonomicName id="6ECC4D750D3FFF86EC0FF1A2346EFF06" authorityName="Moore &amp; Beaver" authorityYear="2020" box="[860,1037,223,249]" class="Insecta" family="Hepialidae" genus="Abantiades" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="90" phylum="Arthropoda" rank="species" species="zonatriticum">A. zonatriticum</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>