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<document ID-CLB-Dataset="9487" ID-DOI="http://dx.doi.org/10.3897/phytokeys.168.51904" ID-GBIF-Dataset="e7345e6b-fa55-474e-9b6e-d6b185121541" ID-PMC="PMC7718216" ID-Pensoft-Pub="1314-2003-168-1" ID-Pensoft-UUID="5F39D34A0DEF5952A2C4E9090C14B498" ID-PubMed="33335445" ModsDocID="1314-2003-168-1" checkinTime="1606572050664" checkinUser="pensoft" docAuthor="Dean, Ellen, Poore, Jennifer, Anguiano-Constante, Marco Antonio, Nee, Michael H., Kang, Hannah, Starbuck, Thomas, Rodrigues, Annamarie &amp; Conner, Matthew" docDate="2020" docId="20EC8BBAAA9763BB87EEE2852AE99B88" docLanguage="en" docName="PhytoKeys 168: 1-333" docOrigin="PhytoKeys 168" docSource="http://dx.doi.org/10.3897/phytokeys.168.51904" docTitle="Lycianthes inconspicua Bitter, Repert. Spec. Nov. Regni Veg. 20: 368 1924" docType="treatment" docVersion="6" id="5F39D34A0DEF5952A2C4E9090C14B498" lastPageNumber="1" masterDocId="5F39D34A0DEF5952A2C4E9090C14B498" masterDocTitle="The genus Lycianthes (Solanaceae, Capsiceae) in Mexico and Guatemala" masterLastPageNumber="333" masterPageNumber="1" pageNumber="1" updateTime="1732941180293" updateUser="ExternalLinkService">
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<mods:title id="72C8F982231171CF89F15DBCBA24B39C">The genus Lycianthes (Solanaceae, Capsiceae) in Mexico and Guatemala</mods:title>
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<mods:namePart id="27CA9E34E76866533E564EE032461818">Dean, Ellen</mods:namePart>
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<mods:namePart id="A133A7E43EDBAEB8349DC55F0C88D50E">Poore, Jennifer</mods:namePart>
<mods:affiliation id="C5A69457D8C5E97D0C0AA4223F30B2EC">UC Davis Center for Plant Diversity, Plant Sciences M. S. 7, One Shields Ave., Davis, CA 95616, USA</mods:affiliation>
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<mods:namePart id="F8A4FA384D36E9E04C632BF6A03EA5F1">Anguiano-Constante, Marco Antonio</mods:namePart>
<mods:affiliation id="37C6F655567DCC9845CAD8D9EE0C0EA6">Laboratorio Nacional de Identificacion y Caracterizacion Vegetal (LaniVeg), Consejo Nacional de Ciencia y Tecnologia (CONACyT), Centro Universitario de Ciencias Biologicas y Agropecuarias, Universidad de Guadalajara, Camino Ramon Padilla Sanchez 2100, 45110 Nextipac, Zapopan, Jalisco, Mexico</mods:affiliation>
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<mods:namePart id="3A65C06C153582019D954B72D64CC281">Nee, Michael H.</mods:namePart>
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<mods:namePart id="5C349EDAA9CB7BDF94E200E4990FCE0C">Kang, Hannah</mods:namePart>
<mods:affiliation id="40948B6CD4C281F82B4E48CC10CF3FD0">UC Davis Center for Plant Diversity, Plant Sciences M. S. 7, One Shields Ave., Davis, CA 95616, USA</mods:affiliation>
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<mods:namePart id="3015786C877B8307CA2F93E92E9500C0">Starbuck, Thomas</mods:namePart>
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<mods:namePart id="D6E68D5656B16DAEFA783D69BA46DEEC">Rodrigues, Annamarie</mods:namePart>
<mods:affiliation id="8131589B911DBE2E8B24F2A3A02BB76F">UC Davis Center for Plant Diversity, Plant Sciences M. S. 7, One Shields Ave., Davis, CA 95616, USA</mods:affiliation>
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<mods:namePart id="5CE45F0995F0BA3D3FC211A42254D9DF">Conner, Matthew</mods:namePart>
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<mods:date id="EB90EC29C6D5F03E6E52522B65905D12">2020</mods:date>
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<mods:classification id="E54AF020FFA7106B9F0E9B9C5981286E">journal article</mods:classification>
<mods:identifier id="4FB5006750B8F4608E41999A385A47E7" type="DOI">http://dx.doi.org/10.3897/phytokeys.168.51904</mods:identifier>
<mods:identifier id="04F3C4243B0CA329599C5D426FDBF183" type="Pensoft-Pub">1314-2003-168-1</mods:identifier>
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<treatment id="20EC8BBAAA9763BB87EEE2852AE99B88" ID-GBIF-Taxon="170596552" LSID="urn:lsid:plazi:treatment:20EC8BBAAA9763BB87EEE2852AE99B88" httpUri="http://treatment.plazi.org/id/20EC8BBAAA9763BB87EEE2852AE99B88" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Solanaceae">
<subSubSection id="A5A7F44AC4B53AB3337BE2878703C8B8" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="C2D35ECA11D60BACE2B7F382A849D133" pageId="0" pageNumber="1">
24
<taxonomicName id="31BDBB8E18B4443070A74D8EBFB1DE2A" ID-CoL="3WK69" authority="Bitter, Repert. Spec. Nov. Regni Veg. 20: 368, 1924" authorityName="Bitter, Repert. Spec. Nov. Regni Veg. 20: 368" authorityYear="1924" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes inconspicua" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="inconspicua">Lycianthes inconspicua Bitter, Repert. Spec. Nov. Regni Veg. 20: 368, 1924</taxonomicName>
<figureCitation id="974A949824801679F7B834636555ACCB" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Image of herbarium specimen of L. inconspicua in Guatemala, Croat 40950 (MO). Specimen used with permission from the Missouri Botanical Garden (http: // www. tropicos. org)." figureDoi="10.3897/phytokeys.168.51904.figure54" httpUri="https://binary.pensoft.net/fig/480066" pageId="0" pageNumber="1">Fig. 54</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A7278EC2352765FD5C682855152AB882" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="2AEFE5CDCF1F19DF642FED1AE1C2B675" pageId="0" pageNumber="1">
<taxonomicName id="BBE6214A3C05E56A9AA16BDCDC85BF01" authorityName="Morton &amp; Standl" authorityYear="1938" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes storkii" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="storkii">Lycianthes storkii</taxonomicName>
C.V.Morton &amp; Standl., Publ. Field Mus. Nat. Hist., Bot. Ser. 18: 1061. 1938. Type: Costa Rica, 0.5 mile south of Santa
<normalizedToken id="A3A4341485A22B98F17E28BED6FD58D7" originalValue="María">Maria</normalizedToken>
, 5700 ft, 8 Aug 1932,
<emphasis id="E4883EE4A7A181D190B22662D816FE37" italics="true" pageId="0" pageNumber="1">H. Stork 3138</emphasis>
(holotype: F [0072922F, acc. # 672865]).
</paragraph>
</subSubSection>
<subSubSection id="FAC9535B10921E15C55FA2696EB707B9" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="55E977203BEA21851F38220B27606EA7" pageId="0" pageNumber="1">Type.</paragraph>
<paragraph id="D9E08851E1AEBB9EF1451B68D923C7E7" pageId="0" pageNumber="1">
Guatemala. [Quetzaltenango]: [
<normalizedToken id="C9959985E3EB518B1DF71B90B8126732" originalValue="Volcán">Volcan</normalizedToken>
] Santa
<normalizedToken id="65DF93AFC0CD7256DD243EA8AC196BCF" originalValue="María">Maria</normalizedToken>
, [
<geoCoordinate id="040950BB91538A12AE8E141C334DDC9E" degrees="14.7559" direction="north" orientation="latitude" precision="5" value="14.7559">14.7559</geoCoordinate>
,
<geoCoordinate id="F47A027DC1A6ACF754A72CDA6903D46E" degrees="91.5537" direction="west" orientation="longitude" precision="5" value="-91.5537">-91.5537</geoCoordinate>
], Dec 1877,
<emphasis id="3B24464BE6FDFCF5449AD6CB516F74B5" italics="true" pageId="0" pageNumber="1">K. Bernoulli &amp; O. Cario 2373</emphasis>
(holotype: GOET [GOET003447]).
</paragraph>
<caption id="75BFA9B0E370FDFDFBE3B259B5B7BD70" doi="10.3897/phytokeys.168.51904.figure54" httpUri="https://binary.pensoft.net/fig/480066" pageId="0" pageNumber="1" start="Figure 54" startId="F54">
<paragraph id="98490D044A8DE5ED5EC4DBE69C7B4346" pageId="0" pageNumber="1">
<emphasis id="36B5690159511DD8380158968F130EEC" bold="true" pageId="0" pageNumber="1">Figure 54.</emphasis>
Image of herbarium specimen of
<taxonomicName id="5F54A1FABD316901B12249751DA2FA17" lsidName="L. inconspicua" pageId="0" pageNumber="1" rank="species" species="inconspicua">
<emphasis id="96AA4879938C7A0C904D617D4418E094" italics="true" pageId="0" pageNumber="1">L. inconspicua</emphasis>
</taxonomicName>
in Guatemala,
<emphasis id="36620F7907324FD42784854223E97638" italics="true" pageId="0" pageNumber="1">Croat 40950</emphasis>
(MO). Specimen used with permission from the Missouri Botanical Garden (http://www.tropicos.org).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="ADA9E3628D2D6C4ECCF5B34A21AF988A" pageId="0" pageNumber="1" type="description">
<paragraph id="7FE8D1750190F1C41985E1C3036BC235" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="F72D6BA47E14E664C7B9B13151E3483B" pageId="0" pageNumber="1">
Subshrub, shrub or treelet, erect, 0.3-6 m tall. Indument pale yellow, uniseriate, multicellular, simple, eglandular, ascending-appressed to ascending trichomes 0.3-1.5 mm long, these usually remaining cylindrical and acute upon drying, the trichomes sometimes becoming less appressed on older stems. Stems green and slender when young, moderately to densely pubescent, sometimes glabrate with age, often compressed upon drying in a plant press, becoming woody with age; upper sympodial branching points usually monochasial with a few dichasial branching points, the branching angles not particularly apparent. Leaves simple, the leaves of the upper sympodia usually paired, usually conspicuously different in size and shape, the larger ones with blades 6-19.5
<normalizedToken id="0D0E39D36960955A8326ACB61E1E4015" originalValue="×">x</normalizedToken>
2-7 cm, narrowly elliptic, lanceolate, or oblanceolate, sometimes appearing slightly falcate, the smaller ones with blades 0.6-3.5
<normalizedToken id="6D5B427490DF2090A627027CDCBEE6E4" originalValue="×">x</normalizedToken>
0.4-2.1 cm, ovate, the blades of both the large and small leaves chartaceous, moderately pubescent, the pubescence densest along the veins of the abaxial side, the trichomes along the midvein of the abaxial side appressed, the base of large blades cuneate, the base of small blades rounded, oblique, the margin entire, usually undulate, the apex acute to acuminate, the petiole to 0.6 (1.1) cm long, sometimes absent, the large leaf blades with (6) 8-12 primary veins on each side of the midvein. Flowers solitary or in groups of 2-3, axillary, nodding; peduncles absent; pedicels very slender, (11) 15-30 mm and straight to arching or deflexed in flower, to 36 mm long, arching or deflexed in fruit, moderately pubescent; 1.5-2.5 mm long, 2-3 mm in diameter, obconic to campanulate, moderately pubescent, the margin truncate, with 5-10 narrow, erect appendages 0.5-3 mm long, emerging ca. 0.25 mm below the calyx rim; fruiting calyx usually enlarged, widely campanulate to bowl-shaped, 1-2 mm long, 3.5-7 mm in diameter, the appendages 1-4 mm long, erect, sometimes withering; corolla 0.5-1 cm long, campanulate to rotate in orientation, stellate in outline, divided 1/2 to nearly all the way to the base, interpetalar tissue present near base, white, adaxial markings unknown, sparsely pubescent abaxially especially near the lobe tips; stamens equal, straight, the filaments 1-2 mm long, glabrous, the anthers 2-3 mm long, ovate, somewhat narrowed at the tip (the narrowed portion ca. 0.25 mm long), free of one another, yellow, glabrous, poricidal at the tips, the pores round, dehiscing distally, not opening into longitudinal slits; pistil with glabrous ovary, the style 3.5-5 mm long, linear, straight, glabrous, the stigma capitate. Fruit a berry, 3.5-9 mm long, 3.5-11 mm in diameter, globose to ovoid, orange to red at maturity, glabrous, lacking sclerotic granules. Seeds 50-100 per fruit, 0.9-1.5
<normalizedToken id="49072A83A958787E3C6F95EC96610E7C" originalValue="×">x</normalizedToken>
1 mm, compressed, but not flat, ridged, rhombic to triangular in outline, tan to orange, the surface reticulum with minute serpentine pattern and shallow luminae.
</paragraph>
</subSubSection>
<subSubSection id="26C263205239D9A04C1879512F30D34D" pageId="0" pageNumber="1" type="chromosome number">
<paragraph id="B8A5DC4C6B607EBA91659C2059713ADE" pageId="0" pageNumber="1">Chromosome number.</paragraph>
<paragraph id="47661C8B5C6CC5AB1E68997CF3398DA7" pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection id="8C0B6A836704FA278C7D73102C6C2BA5" pageId="0" pageNumber="1" type="distribution">
<paragraph id="40C9B99943ECCAFE2367748FB59A93C7" pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph id="1AB941C8EB1F7E29C41440555F2ECDE4" pageId="0" pageNumber="1">
Guatemala (Quetzaltenango, San Marcos,
<normalizedToken id="DBB9A71BA9F2A1D08708FB73F4A65B13" originalValue="Suchitepéquez">Suchitepequez</normalizedToken>
), Costa Rica, and Panama, in cloud forest, montane forest, and oak forest, sometimes on slopes in canyons and drainages, often near streams, 1250-1870 m in elevation (Fig.
<figureCitation id="A5D37514478D0026096014F964EC27A3" captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Map of geographic distribution of L. inconspicua based on herbarium specimen data." figureDoi="10.3897/phytokeys.168.51904.figure55" httpUri="https://binary.pensoft.net/fig/480067" pageId="0" pageNumber="1">55</figureCitation>
).
</paragraph>
<caption id="A81A44782741401BBC1039B9B580C665" doi="10.3897/phytokeys.168.51904.figure55" httpUri="https://binary.pensoft.net/fig/480067" pageId="0" pageNumber="1" start="Figure 55" startId="F55">
<paragraph id="1FDEDC3D95ABEF93AAFE9C6FAD8B5162" pageId="0" pageNumber="1">
<emphasis id="3ACC503D6A1C921A541DB653F334D1C2" bold="true" pageId="0" pageNumber="1">Figure 55.</emphasis>
Map of geographic distribution of
<taxonomicName id="2E176C32FDF4E9C690BF4A3D4A54C21B" lsidName="L. inconspicua" pageId="0" pageNumber="1" rank="species" species="inconspicua">
<emphasis id="F8FFED8E5A90E6A58E6264CB423C6FCE" italics="true" pageId="0" pageNumber="1">L. inconspicua</emphasis>
</taxonomicName>
based on herbarium specimen data.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="75E073599219DEA5B89BE187E3DD943C" pageId="0" pageNumber="1" type="common names and uses">
<paragraph id="F4FEC0EF7463684A6A906A03790EA6CD" pageId="0" pageNumber="1">Common names and uses.</paragraph>
<paragraph id="69EDCC21900302995F8EB846AF341514" pageId="0" pageNumber="1">
Guatemala. Chiltepe de
<normalizedToken id="E140621811AA3A3FCAF04CE1EA6747E6" originalValue="montaña">montana</normalizedToken>
(
<bibRefCitation id="2DD525376AE734624CD5DD54CBA1DCC4" author="Gentry, Jr JL" journalOrPublisher="Taxon" pageId="0" pageNumber="1" refId="B56" refString="Gentry, Jr JL, Standley, PC, 1974. Flora of Guatemala. Fieldiana: Botany 24 (part 10) no. 1-2. Field Museum of Natural History, Chicago." title="Flora of Guatemala. Fieldiana: Botany 24 (part 10) no. 1 - 2. Field Museum of Natural History, Chicago." year="1974">Gentry and Standley 1974</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8B322649C5040C999CA92A7C6F447C74" pageId="0" pageNumber="1" type="phenology">
<paragraph id="D18E7402DE842040240D1CD83E3E5211" pageId="0" pageNumber="1">Phenology.</paragraph>
<paragraph id="15CC78B84DAD04D4CF6BAE338CA0632E" pageId="0" pageNumber="1">Flowering specimens have been collected in January, February, June, and July. Specimens with mature fruits have been collected in July, August, and February. Information about the diurnal movements of the corolla of this species has not been determined; the flowers on specimens range from fully open to somewhat closed (campanulate).</paragraph>
</subSubSection>
<subSubSection id="54F8BF0C1928EF8003D5EF8325951BA7" pageId="0" pageNumber="1" type="preliminary conservation status">
<paragraph id="990FE13AF6890BC4AFD56564E177DD6B" pageId="0" pageNumber="1">Preliminary conservation status.</paragraph>
<paragraph id="904E5ACBE0530E7AAC0F7B5206B01258" pageId="0" pageNumber="1">
In Guatemala,
<taxonomicName id="24255A201A5E5CF0DDC53DB38CFDFF65" authorityName="Bitter, Repert. Spec. Nov. Regni Veg. 20: 368" authorityYear="1924" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes inconspicua" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="inconspicua">
<emphasis id="C0E5EA73B1CC8447192B8686605CCF85" italics="true" pageId="0" pageNumber="1">Lycianthes inconspicua</emphasis>
</taxonomicName>
is represented by only three collections, although it is more common in Costa Rica and Panama, and present in one protected area in Panama (Cerros de La Carpintera, Panama). The EOO is 27,148.801 km2, and the AOO is 32 km2. Based on the
<bibRefCitation id="C68E32F876A4C8EB7E959935306A5571" author="IUCN" journalOrPublisher="The Linnean Society of London, Academic Press, London" pageId="0" pageNumber="1" publicationUrl="http://jr.iucnredlist.org/documents/redlistGuidelines" refId="B67" refString="IUCN, 2019. Guidelines for using the IUCN red list categories and criteria. version 12. Prepared by the Standards and Petitions Subcommittee in February 2019. http://jr.iucnredlist.org/documents/redlistGuidelines" title="Guidelines for using the IUCN red list categories and criteria. version 12. Prepared by the Standards and Petitions Subcommittee in February 2019." url="http://jr.iucnredlist.org/documents/redlistGuidelines" year="2019">IUCN (2019)</bibRefCitation>
criteria, the preliminary assessment category for Guatemala is Endangered (EN).
</paragraph>
</subSubSection>
<subSubSection id="4DBFA24B2DC74EC5B4B70A5053568D21" pageId="0" pageNumber="1" type="discussion">
<paragraph id="76430C1238153B49AA5E2D379FCBF7BF" pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph id="AC5530146E9FE35C7134B1B89533638C" pageId="0" pageNumber="1">
<taxonomicName id="15B7DFA3CD78AD7A07106981EC5CAAE3" authorityName="Bitter, Repert. Spec. Nov. Regni Veg. 20: 368" authorityYear="1924" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes inconspicua" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="inconspicua">
<emphasis id="FA5FB2C19F2032A6160CA841D2323B1E" italics="true" pageId="0" pageNumber="1">Lycianthes inconspicua</emphasis>
</taxonomicName>
is an uncommon species of high elevation areas in Central America. It is uncommon in Guatemala, but more common in high elevation areas of Costa Rica and Panama. It is morphologically similar to three other Mexican and Central American species:
<taxonomicName id="2BE6CAD2414CC197DD83C89B05962153" lsidName="L. glabripetala" pageId="0" pageNumber="1" rank="species" species="glabripetala">
<emphasis id="39473D2DFE7EB8A61348F2E302D0E514" italics="true" pageId="0" pageNumber="1">L. glabripetala</emphasis>
</taxonomicName>
,
<taxonomicName id="BB64F2666EC9F651360916932AE1A67D" lsidName="L. amatitlanensis" pageId="0" pageNumber="1" rank="species" species="amatitlanensis">
<emphasis id="8FAAAD93DCF6C1C8F515C9BF6C5692B7" italics="true" pageId="0" pageNumber="1">L. amatitlanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="2EE83123E60F952E73DB0D7377F77192" lsidName="L. inaequilatera" pageId="0" pageNumber="1" rank="species" species="inaequilatera">
<emphasis id="74766D533965D5F818062418181E83C6" italics="true" pageId="0" pageNumber="1">L. inaequilatera</emphasis>
</taxonomicName>
.
<taxonomicName id="2A1BB554ABBFFAA2546E1CC4D6FED7AC" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes amatitlanensis" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="amatitlanensis">
<emphasis id="99446AEA27F6A8932E33B601CE0E06E0" italics="true" pageId="0" pageNumber="1">Lycianthes amatitlanensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="37F9191F3FE5C3B9E96C750C4C0D718F" lsidName="L. inconspicua" pageId="0" pageNumber="1" rank="species" species="inconspicua">
<emphasis id="CF6B0200CDD1FBC81BF4078DE0E7F108" italics="true" pageId="0" pageNumber="1">L. inconspicua</emphasis>
</taxonomicName>
in having shorter pedicels (4-12 mm in flower and 6-16 mm in fruit), long, coarse trichomes that spread away from the midvein on the abaxial side of the leaf (usually with some trichomes at an angle close to ninety degrees), and lanceolate anthers with a longer attenuate portion at the tip (ca. 0.5 mm long).
<taxonomicName id="AE250BD561E374B217ADB433CDB74A6B" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes inaequilatera" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="inaequilatera">
<emphasis id="821F548079BEDCFDB9970D05E78AFE20" italics="true" pageId="0" pageNumber="1">Lycianthes inaequilatera</emphasis>
</taxonomicName>
and
<taxonomicName id="45470C47115C18A41B32B2FE2A98979C" lsidName="L. glabripetala" pageId="0" pageNumber="1" rank="species" species="glabripetala">
<emphasis id="458794A359D3BA2DFA1FC016E3469A74" italics="true" pageId="0" pageNumber="1">L. glabripetala</emphasis>
</taxonomicName>
are more similar to
<taxonomicName id="E664B4134C1B5A20901EF3807BDC7971" lsidName="L. inconspicua" pageId="0" pageNumber="1" rank="species" species="inconspicua">
<emphasis id="FADBC0074E1DD04EF6F83A06531F9534" italics="true" pageId="0" pageNumber="1">L. inconspicua</emphasis>
</taxonomicName>
in terms of having appressed trichomes along the leaf blade midvein, but they tend to have shorter pedicels (15 mm or less in flower); in addition,
<taxonomicName id="2D3913FC08FF94D380B43088A4CA44E8" lsidName="L. glabripetala" pageId="0" pageNumber="1" rank="species" species="glabripetala">
<emphasis id="097C4BC0584B72B160B8CEE3B41C3BFA" italics="true" pageId="0" pageNumber="1">L. glabripetala</emphasis>
</taxonomicName>
has larger corollas (1-1.2 cm long).
</paragraph>
</subSubSection>
<subSubSection id="2D0D41692857AB64D4D2A9BAEE8B5F4D" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="1F95A138EFE66028F8498E145F4ED675" pageId="0" pageNumber="1">Representative specimens examined.</paragraph>
<paragraph id="3FC241CB252FD2D3A18C3739DA68B75D" pageId="0" pageNumber="1">
<emphasis id="E08AFD5A742A47AF75C8EC9E3A673AA6" bold="true" pageId="0" pageNumber="1">Guatemala. [Quetzaltenango</emphasis>
]: [
<normalizedToken id="2566FA2228C870A6600DE1A3EF48A2B7" originalValue="Volcán">Volcan</normalizedToken>
] Santa
<normalizedToken id="1D8AFEEFDE5DFE638A67AC40B2BD640F" originalValue="María">Maria</normalizedToken>
, [
<geoCoordinate id="48583E3FD7C6698F8EC72C712D955145" degrees="14.7559" direction="north" orientation="latitude" precision="5" value="14.7559">14.7559</geoCoordinate>
,
<geoCoordinate id="F1B0604A36FF7F8EBCCB4261D6538D88" degrees="91.5537" direction="west" orientation="longitude" precision="5" value="-91.5537">-91.5537</geoCoordinate>
], Dec 1877,
<emphasis id="99B033C24E9412E79B48EAC955826DC0" italics="true" pageId="0" pageNumber="1">K. Bernoulli &amp; O. Cario 2373</emphasis>
(GOET).
<emphasis id="114CF41619CB23912153D2F99DB19116" bold="true" pageId="0" pageNumber="1">San Marcos</emphasis>
: Finca Armenia, Rafael de Cuesta, San Marcos, 5000 ft, 6-7 Jul 1977,
<emphasis id="67A81301681907A257214FF91ADC325E" italics="true" pageId="0" pageNumber="1">J.D. Dwyer 14408</emphasis>
(LL, MO).
<emphasis id="2E773D0A5861CAEB37A44096B4306656" bold="true" pageId="0" pageNumber="1">
<normalizedToken id="8E422AE32284866EE7593D4547D7ECB2" originalValue="Suchitepéquez">Suchitepequez</normalizedToken>
</emphasis>
: Mpio. San Francisco
<normalizedToken id="A83B6B4B494528C2F5B9801DEA717F10" originalValue="Zapotitlán">Zapotitlan</normalizedToken>
, Reserva Natural Privada Las Nubes,
<geoCoordinate id="51F9A8DE37B6B08BEAEF435B3089D7B9" degrees="14.7061" direction="north" orientation="latitude" precision="5" value="14.7061">14.7061</geoCoordinate>
,
<geoCoordinate id="CFF112D1CC509D777F7640E7DAEA0E61" degrees="90.9744" direction="west" orientation="longitude" precision="5" value="-90.9744">-90.9744</geoCoordinate>
, 1600 m, 3 Jul 2014,
<emphasis id="560EA413ED075656F5E0BD69B1DD04FB" italics="true" pageId="0" pageNumber="1">B. Escobar 171</emphasis>
(BIGU).
</paragraph>
</subSubSection>
</treatment>
</document>