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<mods:title id="9D85CC11480F20F30B6E8B0008EE2783">A review of Cryphocricos Signoret, 1850 (Naucoridae: Cryphocricinae) with descriptions of three new species</mods:title>
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<treatment id="207387E3E071FF9602CBFA1C52CE9E69" ID-DOI="http://doi.org/10.5281/zenodo.4693086" ID-GBIF-Taxon="181600675" ID-Zenodo-Dep="4693086" LSID="urn:lsid:plazi:treatment:207387E3E071FF9602CBFA1C52CE9E69" httpUri="http://treatment.plazi.org/id/207387E3E071FF9602CBFA1C52CE9E69" lastPageId="6" lastPageNumber="77" pageId="5" pageNumber="77">
<subSubSection id="E0C0657EE071FF9502CBFA1C532B9A9D" box="[151,558,1436,1463]" pageId="5" pageNumber="77" type="nomenclature">
<paragraph id="A86536F5E071FF9502CBFA1C532B9A9D" blockId="5.[151,558,1436,1498]" box="[151,558,1436,1463]" pageId="5" pageNumber="77">
<heading id="F32D8199E071FF9502CBFA1C532B9A9D" bold="true" box="[151,558,1436,1463]" fontSize="11" level="1" pageId="5" pageNumber="77" reason="1">
<taxonomicName id="6FDA4D76E071FF9502CBFA1C532B9A9D" ID-CoL="ZT2W" authority="De Carlo" authorityName="De Carlo" authorityYear="1940" box="[151,558,1436,1463]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="77" phylum="Arthropoda" rank="species" species="daguerrei">
<emphasis id="9AAEEAE7E071FF9502CBFA1C532B9A9D" bold="true" box="[151,558,1436,1463]" pageId="5" pageNumber="77">
<emphasis id="9AAEEAE7E071FF9502CBFA1C50B09A9C" bold="true" box="[151,437,1436,1463]" italics="true" pageId="5" pageNumber="77">Cryphocricos daguerrei</emphasis>
De Carlo
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E0C0657EE071FF9502CBFA4050179AF1" box="[151,274,1472,1498]" pageId="5" pageNumber="77" type="description">
<paragraph id="A86536F5E071FF9502CBFA4050179AF1" blockId="5.[151,558,1436,1498]" box="[151,274,1472,1498]" pageId="5" pageNumber="77">
(
<figureCitation id="30E12A70E071FF9502C3FA40500F9AF1" box="[159,266,1472,1498]" captionStart="FIGURES 78" captionStartId="5.[151,264,1330,1354]" captionTargetBox="[151,1435,417,1307]" captionTargetId="figure-126@5.[151,1435,417,1307]" captionTargetPageId="5" captionText="FIGURES 78. Cryphocricos daguerrei holotype (7) dorsal and (8) ventral. Photos courtesy of H.D.D. Rodrigues." figureDoi="http://doi.org/10.5281/zenodo.4693084" httpUri="https://zenodo.org/record/4693084/files/figure.png" pageId="5" pageNumber="77">Figs. 78</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E0C0657EE071FF9502CBF98555D59936" box="[151,1232,1541,1565]" pageId="5" pageNumber="77" type="reference_group">
<paragraph id="A86536F5E071FF9502CBF98555D59936" blockId="5.[151,1232,1541,1565]" box="[151,1232,1541,1565]" pageId="5" pageNumber="77">
<taxonomicName id="6FDA4D76E071FF9502CBF985532C9936" authority="De Carlo 1940" authorityName="De Carlo" authorityYear="1940" box="[151,553,1541,1565]" class="Insecta" family="Naucoridae" genus="Cryphocricus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="77" phylum="Arthropoda" rank="species" species="daguerrei">
<emphasis id="9AAEEAE7E071FF9502CBF985508F9936" box="[151,394,1541,1565]" italics="true" pageId="5" pageNumber="77">Cryphocricus daguerrei</emphasis>
<bibRefCitation id="CC4B4B04E071FF9503CDF985532C9936" author="De Carlo, J. A." box="[401,553,1541,1565]" pageId="5" pageNumber="77" pagination="426 - 433" refId="ref12276" refString="De Carlo, J. A. (1940) I) Descripcion de tres especies nuevas del genero Cryphocricus Signoret. II). Una nueva especie del genero Heleocoris Stal (Hemiptera-Naucoridae). Revista de la Sociedad Entomologica Argentina, 10, 426 - 433." type="journal article" year="1940">De Carlo 1940</bibRefCitation>
</taxonomicName>
:
<journalOrPublisher id="721B3C39E071FF95006FF98552649936" box="[563,865,1541,1565]" pageId="5" pageNumber="77">
Rev. Soc. Entomol.
<collectingCountry id="D0CD7665E071FF9500A0F98552649936" box="[764,865,1541,1565]" name="Argentina" pageId="5" pageNumber="77">Argentina</collectingCountry>
</journalOrPublisher>
11:430431 (misspelling of genus).
</paragraph>
</subSubSection>
<subSubSection id="E0C0657EE071FF9502CBF9CB525C99A2" pageId="5" pageNumber="77" type="materials_examined">
<paragraph id="A86536F5E071FF9502CBF9CB546E994D" blockId="5.[151,1437,1611,2033]" box="[151,1387,1611,1638]" pageId="5" pageNumber="77">
<materialsCitation id="18B23CA8E071FF9502CBF9CB5462994D" ID-GBIF-Occurrence="3091107321" box="[151,1383,1611,1638]" country="Argentina" location="Museo Argentino de Ciencias Naturales" pageId="5" pageNumber="77" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires" typeStatus="holotype">
<emphasis id="9AAEEAE7E071FF9502CBF9CB5058994E" bold="true" box="[151,349,1611,1637]" pageId="5" pageNumber="77">
<typeStatus id="77618857E071FF9502CBF9CB51D7994E" box="[151,210,1611,1637]" pageId="5" pageNumber="77">Type</typeStatus>
repository:
</emphasis>
<location id="AD05602EE071FF95033FF9CB5224994E" LSID="urn:lsid:plazi:treatment:207387E3E071FF9602CBFA1C52CE9E69:AD05602EE071FF95033FF9CB5224994E" box="[355,801,1611,1637]" country="Argentina" name="Museo Argentino de Ciencias Naturales" pageId="5" pageNumber="77" stateProvince="Ciudad Autonoma de Buenos Aires">Museo Argentino de Ciencias Naturales</location>
“Bernardino Rivadavia” (
<collectingRegion id="6A1EF817E071FF95061AF9CB55E5994E" box="[1094,1248,1611,1637]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="5" pageNumber="77">Buenos Aires</collectingRegion>
,
<collectingCountry id="D0CD7665E071FF9506B6F9CB545A994D" box="[1258,1375,1611,1638]" name="Argentina" pageId="5" pageNumber="77">Argentina</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A86536F5E071FF95029BF9EF525C99A2" blockId="5.[151,1437,1611,2033]" box="[199,857,1647,1673]" pageId="5" pageNumber="77">
<materialsCitation id="18B23CA8E071FF95029BF9EF525199A2" ID-GBIF-Occurrence="3091107308" box="[199,852,1647,1673]" country="Argentina" location="Concordia" pageId="5" pageNumber="77" specimenCount="1" stateProvince="Entre Rios" typeStatus="holotype">
<emphasis id="9AAEEAE7E071FF95029BF9EF506C99A2" bold="true" box="[199,361,1647,1673]" pageId="5" pageNumber="77">
<typeStatus id="77618857E071FF95029BF9EF500799A2" box="[199,258,1647,1673]" pageId="5" pageNumber="77">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="D0CD7665E071FF950333F9EF50E699A2" box="[367,483,1647,1673]" name="Argentina" pageId="5" pageNumber="77">Argentina</collectingCountry>
:
<collectingRegion id="6A1EF817E071FF9503B1F9EF53D499A2" box="[493,721,1647,1673]" country="Argentina" name="Entre Rios" pageId="5" pageNumber="77">Entre Ríos Province</collectingRegion>
,
<location id="AD05602EE071FF950080F9EF525199A2" LSID="urn:lsid:plazi:treatment:207387E3E071FF9602CBFA1C52CE9E69:AD05602EE071FF950080F9EF525199A2" box="[732,852,1647,1673]" country="Argentina" name="Concordia" pageId="5" pageNumber="77" stateProvince="Entre Rios">Concordia</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E0C0657EE071FF95029BF91352FC98AE" pageId="5" pageNumber="77" type="discussion">
<paragraph id="A86536F5E071FF95029BF91352FC98AE" blockId="5.[151,1437,1611,2033]" pageId="5" pageNumber="77">
<emphasis id="9AAEEAE7E071FF95029BF913504A9986" bold="true" box="[199,335,1683,1709]" pageId="5" pageNumber="77">Discussion:</emphasis>
The
<typeStatus id="77618857E071FF9503D5F91350EE9986" box="[393,491,1683,1709]" pageId="5" pageNumber="77" type="holotype">holotype</typeStatus>
is a brachypterous male with only the right foreleg intact. The left foreleg and right hindleg are complete only through the femur, the left middle leg is complete through the tibia, and the right middle leg and left hindleg are missing (
<figureCitation id="30E12A70E071FF950055F95B537399DE" box="[521,630,1755,1781]" captionStart="FIGURES 78" captionStartId="5.[151,264,1330,1354]" captionTargetBox="[151,1435,417,1307]" captionTargetId="figure-126@5.[151,1435,417,1307]" captionTargetPageId="5" captionText="FIGURES 78. Cryphocricos daguerrei holotype (7) dorsal and (8) ventral. Photos courtesy of H.D.D. Rodrigues." figureDoi="http://doi.org/10.5281/zenodo.4693084" httpUri="https://zenodo.org/record/4693084/files/figure.png" pageId="5" pageNumber="77">Figs. 78</figureCitation>
). The rivers in Concordia are wide, flat, and muddy, which are not suitable conditions for
<taxonomicName id="6FDA4D76E071FF950329F97F53089832" authorityName="Signoret" authorityYear="1850" box="[373,525,1791,1817]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="9AAEEAE7E071FF950329F97F53089832" box="[373,525,1791,1817]" italics="true" pageId="5" pageNumber="77">Cryphocricos</emphasis>
</taxonomicName>
; thus, if indeed the type specimen was collected there, river dynamics probably have changed since its collection in 1931.
<bibRefCitation id="CC4B4B04E071FF950031F8A352269815" author="De Carlo, J. A." box="[621,803,1827,1854]" pageId="5" pageNumber="77" pagination="99 - 104" refId="ref12526" refString="De Carlo, J. A. (1968) Descripcion de especies nuevas de los generos Ambrysus, Cryphocricos y Heleocoris. Allotypus de Cryphocricos daguerrei (Hemiptera - Naucoridae). Revista de la Sociedad Entomologica Argentina, 30, 99 - 104. [1967]" type="journal article" year="1968">De Carlo (1968)</bibRefCitation>
proposed an
<typeStatus id="77618857E071FF9501EBF8A355149816" box="[951,1041,1827,1853]" pageId="5" pageNumber="77" type="allotype">allotype</typeStatus>
from
<collectingCountry id="D0CD7665E071FF950608F8A355BF9816" box="[1108,1210,1827,1853]" name="Uruguay" pageId="5" pageNumber="77">Uruguay</collectingCountry>
; however, this is not permissible under the rules of the ICZN, and
<bibRefCitation id="CC4B4B04E071FF9500C8F8C75259984A" author="Lopez Ruf, M. L." box="[660,860,1863,1889]" pageId="5" pageNumber="77" pagination="103 - 120" refId="ref13109" refString="Lopez Ruf, M. L. (1991) El genero Cryphocricos en la Argentina (Hemiptera-Limnocoridae). Revista de la Sociedad Entomologica Argentina, 49 (1 - 4), 103 - 120." type="journal article" year="1991">Lopez Ruf (1991)</bibRefCitation>
challenged the accuracy of the identification based on differences in color, pronotum, and head shape with those of the
<typeStatus id="77618857E071FF9501CCF8EB52F398AE" box="[912,1014,1899,1925]" pageId="5" pageNumber="77" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E0C0657EE071FF96029BF80F52CE9E69" lastPageId="6" lastPageNumber="78" pageId="5" pageNumber="77" type="diagnosis">
<paragraph id="A86536F5E071FF96029BF80F52CE9E69" blockId="5.[151,1437,1611,2033]" lastBlockId="6.[151,1437,151,322]" lastPageId="6" lastPageNumber="78" pageId="5" pageNumber="77">
<emphasis id="9AAEEAE7E071FF95029BF80F50419882" bold="true" box="[199,324,1935,1961]" pageId="5" pageNumber="77">Diagnosis:</emphasis>
<bibRefCitation id="CC4B4B04E071FF950311F80F530C9882" author="De Carlo, J. A." box="[333,521,1935,1961]" pageId="5" pageNumber="77" pagination="37 - 41" refId="ref12325" refString="De Carlo, J. A. (1941) I) Descripcion de dos especies nuevas del genero Limnocoris Stal. II) nuevas consideraciones sobre Cryphocricus daguerrei y Cryphocricus rufus De Carlo (Hemiptera-Naucoridae). Revista de la Sociedad Entomologica Argentina, 11, 37 - 41." type="journal article" year="1941">De Carlo (1941)</bibRefCitation>
offered the diagnostic character of abdominal segments II and III (he probably meant III and IV) lateral margins individually slightly curved; thus, creating a non-uniform curvature of the entire abdominal lateral margin. This species differs from
<taxonomicName id="6FDA4D76E071FF9500B5F85752FB98DA" authority="De Carlo, 1940" authorityName="De Carlo" authorityYear="1940" box="[745,1022,2007,2033]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="77" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="9AAEEAE7E071FF9500B5F857524098DA" box="[745,837,2007,2033]" italics="true" pageId="5" pageNumber="77">C. rufus</emphasis>
<bibRefCitation id="CC4B4B04E071FF950112F85752FB98DA" author="De Carlo, J. A." box="[846,1022,2007,2033]" pageId="5" pageNumber="77" pagination="426 - 433" refId="ref12276" refString="De Carlo, J. A. (1940) I) Descripcion de tres especies nuevas del genero Cryphocricus Signoret. II). Una nueva especie del genero Heleocoris Stal (Hemiptera-Naucoridae). Revista de la Sociedad Entomologica Argentina, 10, 426 - 433." type="journal article" year="1940">De Carlo, 1940</bibRefCitation>
</taxonomicName>
by its smaller size, the shape of the hemelytra, the concavity in the prothoracic anterior margin, the shape of the head, and fewer granulations (=tubercles) (
<bibRefCitation id="CC4B4B04E072FF9602A3FF3B50AC9FFE" author="De Carlo, J. A." box="[255,425,187,213]" pageId="6" pageNumber="78" pagination="426 - 433" refId="ref12276" refString="De Carlo, J. A. (1940) I) Descripcion de tres especies nuevas del genero Cryphocricus Signoret. II). Una nueva especie del genero Heleocoris Stal (Hemiptera-Naucoridae). Revista de la Sociedad Entomologica Argentina, 10, 426 - 433." type="journal article" year="1940">De Carlo 1940</bibRefCitation>
). This species differs from
<taxonomicName id="6FDA4D76E072FF960086FF3B524F9FFE" baseAuthorityName="De Carlo" baseAuthorityYear="1968" box="[730,842,187,213]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="78" phylum="Arthropoda" rank="species" species="fittkaui">
<emphasis id="9AAEEAE7E072FF960086FF3B524F9FFE" box="[730,842,187,213]" italics="true" pageId="6" pageNumber="78">C. fittkaui</emphasis>
</taxonomicName>
by its darker color and the female subgenital plate is very similar, although it has its differences (
<bibRefCitation id="CC4B4B04E072FF9600D2FF5F523E9FD1" author="De Carlo, J. A." box="[654,827,223,250]" pageId="6" pageNumber="78" pagination="189 - 200" refId="ref12493" refString="De Carlo, J. A. (1967) Una nueva especie del genero Ranatra y nuevas especies de Naucoridae. Amazoniana, 1 (2), 189 - 200." type="journal article" year="1967">De Carlo 1967</bibRefCitation>
). The female subgenital plate was illustrated with the lateral margins sinuate and the posterior margin to have three distinct lobes, with the middle lobe half the length of the lateral lobes, and is comparable to that of
<taxonomicName id="6FDA4D76E072FF9600C9FEA752C29E69" authority="(De Carlo 1968)" baseAuthorityName="De Carlo" baseAuthorityYear="1968" box="[661,967,295,322]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="78" phylum="Arthropoda" rank="species" species="fittkaui">
<emphasis id="9AAEEAE7E072FF9600C9FEA752009E6A" box="[661,773,295,321]" italics="true" pageId="6" pageNumber="78">C. fittkaui</emphasis>
(
<bibRefCitation id="CC4B4B04E072FF960148FEA752BB9E69" author="De Carlo, J. A." box="[788,958,295,322]" pageId="6" pageNumber="78" pagination="99 - 104" refId="ref12526" refString="De Carlo, J. A. (1968) Descripcion de especies nuevas de los generos Ambrysus, Cryphocricos y Heleocoris. Allotypus de Cryphocricos daguerrei (Hemiptera - Naucoridae). Revista de la Sociedad Entomologica Argentina, 30, 99 - 104. [1967]" type="journal article" year="1968">De Carlo 1968</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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