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<mods:title id="FFA00AE90D0B6FA9993A1A62A53138D8">Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae)</mods:title>
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<emphasis id="9A8D9E0BBA4FFFC7FE5021C1FD3EFF1E" bold="true" box="[425,685,142,169]" italics="true" pageId="7" pageNumber="2018">Bensonella plicidens</emphasis>
(
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)
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(
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(ae), 4, 5, 9(a,b)).
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<emphasis id="9A8D9E0BBA4FFFC7FF592046FF44FE95" box="[160,215,265,290]" italics="true" pageId="7" pageNumber="2018">Pupa</emphasis>
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<emphasis id="9A8D9E0BBA4FFFC7FF262046FEAEFE95" box="[223,317,265,290]" italics="true" pageId="7" pageNumber="2018">plicidens</emphasis>
<bibRefCitation id="CC683FE8BA4FFFC7FEBD2046FD82FE95" author="Benson WH" box="[324,529,265,290]" pageId="7" pageNumber="2018" pagination="125 - 128" refId="ref6320" refString="Benson WH. 1849. Descriptions of four new Asiatic species of the genus Pupa of Draparnaud. Ann Mag Nat Hist. 4 (20): 125 - 128. doi: 10.1080 / 03745486009496158." type="journal article" year="1849">Benson, 1849: 126</bibRefCitation>
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<emphasis id="9A8D9E0BBA4FFFC7FF592063FEAEFEF2" box="[160,317,300,325]" italics="true" pageId="7" pageNumber="2018">
Pupa
<taxonomicName id="6FF9399ABA4FFFC7FF262063FEAEFEF2" ID-CoL="LHH3" authorityName="Benson" authorityYear="1849" box="[223,317,300,325]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">plicidens</taxonomicName>
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<bibRefCitation id="CC683FE8BA4FFFC7FEA02063FE4FFEF2" author="Kuster HC" box="[345,476,300,325]" pageId="7" pageNumber="2018" refId="ref7017" refString="Kuster HC. 1841 - 1855. Die Gattungen Pupa, Megaspira, Balea und Tornatellina. In Abbildungen nach der Natur mit Beschreibungen. In: Systematisches ConchylienCabinet von Martini und Chemnitz. Nurnberg: Bauer &amp; Raspe; p. 194." type="journal volume" year="1841">Küster 1841</bibRefCitation>
1855: 136, pl. 17, figs 2324
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<emphasis id="9A8D9E0BBA4FFFC7FF592000FECCFEDF" box="[160,351,335,360]" italics="true" pageId="7" pageNumber="2018">Boysidia plicidens</emphasis>
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<treatmentCitation id="29586408BA4FFFC7FE822000FDACFEDF" author="Pfeiffer L" box="[379,575,335,360]" page="553" pageId="7" pageNumber="2018" year="1853">
<bibRefCitation id="CC683FE8BA4FFFC7FE822000FDACFEDF" author="Pfeiffer L" box="[379,575,335,360]" pageId="7" pageNumber="2018" refId="ref7434" refString="Pfeiffer L. 1853. Monographia heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum. Vol. 3, Volumen tertium. Lipsiae: Brockhaus." type="book" year="1853">Pfeiffer 1853: 553</bibRefCitation>
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<emphasis id="9A8D9E0BBA4FFFC7FF59203DFF7EFE3C" box="[160,237,370,395]" italics="true" pageId="7" pageNumber="2018">Vertigo</emphasis>
(
<emphasis id="9A8D9E0BBA4FFFC7FF05203DFE1CFE3C" box="[252,399,370,395]" italics="true" pageId="7" pageNumber="2018">Odontocyclas</emphasis>
)
<emphasis id="9A8D9E0BBA4FFFC7FE67203DFE6FFE3C" box="[414,508,370,395]" italics="true" pageId="7" pageNumber="2018">plicidens</emphasis>
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<treatmentCitation id="29586408BA4FFFC7FDE1203DFCF3FE3C" author="Adams H &amp; Adams A" box="[536,864,370,395]" page="173" pageId="7" pageNumber="2018" year="1858">
<bibRefCitation id="CC683FE8BA4FFFC7FDE1203DFCF3FE3C" author="Adams H &amp; Adams A" box="[536,864,370,395]" pageId="7" pageNumber="2018" refId="ref6179" refString="Adams H, Adams A. 1858. The genera of recent mollusca, arranged according to their organization. Vol. 2. London: John van Voorst." type="book" year="1858">Adams and Adams 1858: 173</bibRefCitation>
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<emphasis id="9A8D9E0BBA4FFFC7FF5920DBFF44FE1A" box="[160,215,404,429]" italics="true" pageId="7" pageNumber="2018">Pupa</emphasis>
(
<emphasis id="9A8D9E0BBA4FFFC7FF1E20DBFEE2FE1A" box="[231,369,404,429]" italics="true" pageId="7" pageNumber="2018">Scopelophila</emphasis>
)
<emphasis id="9A8D9E0BBA4FFFC7FE7920DBFE4DFE1A" box="[384,478,404,429]" italics="true" pageId="7" pageNumber="2018">plicidens</emphasis>
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<treatmentCitation id="29586408BA4FFFC7FE0320DBFCDDFE1A" author="Albers JC &amp; von Martens E" box="[506,846,404,429]" page="296" pageId="7" pageNumber="2018" year="1860">
<bibRefCitation id="CC683FE8BA4FFFC7FE0320DBFCDDFE1A" author="Albers JC &amp; von Martens E" box="[506,846,404,429]" pageId="7" pageNumber="2018" refId="ref6209" refString="Albers JC, von Martens E. 1860. Die Heliceen nach naturlicher Verwandtschaft systematisch geordnet von Joh. Christ. Albers. 2 nd ed. Leipzig: Engelman." type="book" year="1860">Albers and Martens, 1860: 296</bibRefCitation>
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<taxonomicName id="6FF9399ABA4FFFC7FF5920F8FECCFE67" ID-CoL="7QD4P" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,351,439,464]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA4FFFC7FF5920F8FECCFE67" box="[160,351,439,464]" italics="true" pageId="7" pageNumber="2018">Boysidia plicidens</emphasis>
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Hanley and Theobald, 1874 (18701876): pl. 100, fig. 8
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<taxonomicName id="6FF9399ABA4FFFC7FF592096FECCFE45" box="[160,351,473,498]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA4FFFC7FF592096FECCFE45" box="[160,351,473,498]" italics="true" pageId="7" pageNumber="2018">Boysidia plicidens</emphasis>
</taxonomicName>
<bibRefCitation id="CC683FE8BA4FFFC7FE822096FDB3FE45" author="Sowerby GB II." box="[379,544,473,498]" pageId="7" pageNumber="2018" pagination="1 - 10" refId="ref7840" refString="Sowerby GB II. 1876. Monograph of the genus Pupinidae. In: Reeve L, editor. Conchologia iconica, or illustrations of the shells of molluscous animals. Vol. 20. London: L. Reeve; p. 1 - 10. unpaginated text." type="book chapter" year="1876">Sowerby, 1876</bibRefCitation>
: pl. 16, fig. 151
</paragraph>
<paragraph id="A8464219BA4FFFC7FF5920B3FCCEFDA2" blockId="7.[160,1128,265,845]" box="[160,861,508,533]" pageId="7" pageNumber="2018">
<emphasis id="9A8D9E0BBA4FFFC7FF5920B3FF44FDA2" box="[160,215,508,533]" italics="true" pageId="7" pageNumber="2018">Pupa</emphasis>
<taxonomicName id="6FF9399ABA4FFFC7FF2620B3FE75FDA2" box="[223,486,508,533]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens" subGenus="Odontocyclas">
(
<emphasis id="9A8D9E0BBA4FFFC7FF1E20B3FEE9FDA2" box="[231,378,508,533]" italics="true" pageId="7" pageNumber="2018">Odontocyclas</emphasis>
)
<emphasis id="9A8D9E0BBA4FFFC7FE7120B3FE75FDA2" box="[392,486,508,533]" italics="true" pageId="7" pageNumber="2018">plicidens</emphasis>
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Pfeiffer, 1879 (18781881): 350
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<emphasis id="9A8D9E0BBA4FFFC7FF592350FECCFD8F" box="[160,351,543,568]" italics="true" pageId="7" pageNumber="2018">Boysidia plicidens</emphasis>
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</paragraph>
<paragraph id="A8464219BA4FFFC7FF59230EFCC7FDED" blockId="7.[160,1128,265,845]" box="[160,852,577,602]" pageId="7" pageNumber="2018">
<treatmentCitationGroup id="88E96537BA4FFFC7FF59230EFCC7FDED" box="[160,852,577,602]" pageId="7" pageNumber="2018">
<taxonomicName id="6FF9399ABA4FFFC7FF59230EFE79FDED" ID-CoL="8V3K7" baseAuthorityName="W. H. Benson" baseAuthorityYear="1849" box="[160,490,577,602]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens" subGenus="Bensonella">
<emphasis id="9A8D9E0BBA4FFFC7FF59230EFF69FDED" box="[160,250,577,602]" italics="true" pageId="7" pageNumber="2018">Bifidaria</emphasis>
(
<emphasis id="9A8D9E0BBA4FFFC7FEF0230EFEEEFDED" box="[265,381,577,602]" italics="true" pageId="7" pageNumber="2018">Bensonella</emphasis>
)
<emphasis id="9A8D9E0BBA4FFFC7FE75230EFE79FDED" box="[396,490,577,602]" italics="true" pageId="7" pageNumber="2018">plicidens</emphasis>
</taxonomicName>
<treatmentCitation id="29586408BA4FFFC7FDFF230EFCC7FDED" author="Pilsbry HA &amp; Vanatta EG" box="[518,852,577,602]" page="591" pageId="7" pageNumber="2018" year="1900">
<bibRefCitation id="CC683FE8BA4FFFC7FDFF230EFCC7FDED" author="Pilsbry HA &amp; Vanatta EG" box="[518,852,577,602]" pageId="7" pageNumber="2018" pagination="582 - 611" refId="ref7771" refString="Pilsbry HA, Vanatta EG. 1900. A partial revision of the Pupae of the United States. Proc Acad Nat Sci Phila. 52: 582 - 611." type="journal article" year="1900">Pilsbry and Vanatta 1900: 591</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A8464219BA4FFFC7FF59232BFDBFFDCA" blockId="7.[160,1128,265,845]" box="[160,556,612,637]" pageId="7" pageNumber="2018">
<treatmentCitationGroup id="88E96537BA4FFFC7FF59232BFDBFFDCA" box="[160,556,612,637]" pageId="7" pageNumber="2018">
<taxonomicName id="6FF9399ABA4FFFC7FF59232BFECCFDCA" ID-CoL="7QD4P" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,351,612,637]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA4FFFC7FF59232BFECCFDCA" box="[160,351,612,637]" italics="true" pageId="7" pageNumber="2018">Boysidia plicidens</emphasis>
</taxonomicName>
<treatmentCitation id="29586408BA4FFFC7FE82232BFDBFFDCA" author="Gude GK" box="[379,556,612,637]" page="294" pageId="7" pageNumber="2018" year="1914">
<bibRefCitation id="CC683FE8BA4FFFC7FE82232BFDBFFDCA" author="Gude GK" box="[379,556,612,637]" pageId="7" pageNumber="2018" pagination="520" refId="ref6760" refString="Gude GK. 1914. Mollusca-II. Trochomorphidae-Janellidae. In: Shipley AE, Marshall GAK, editors. The Fauna of British India, including Ceylon and Burma. London: Taylor &amp; Francis; p. XII + 520." type="book chapter" year="1914">Gude 1914: 294</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A8464219BA4FFFC7FF5923C8FC5AFD17" blockId="7.[160,1128,265,845]" box="[160,969,647,672]" pageId="7" pageNumber="2018">
<treatmentCitationGroup id="88E96537BA4FFFC7FF5923C8FC5AFD17" box="[160,969,647,672]" pageId="7" pageNumber="2018">
<taxonomicName id="6FF9399ABA4FFFC7FF5923C8FD94FD17" ID-CoL="7QBW8" authorityName="Pilsbry" authorityYear="1915" box="[160,519,647,672]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="landurensis" subGenus="Bensonella">
<emphasis id="9A8D9E0BBA4FFFC7FF5923C8FF69FD17" box="[160,250,647,672]" italics="true" pageId="7" pageNumber="2018">Bifidaria</emphasis>
(
<emphasis id="9A8D9E0BBA4FFFC7FEF023C8FEEEFD17" box="[265,381,647,672]" italics="true" pageId="7" pageNumber="2018">Bensonella</emphasis>
)
<emphasis id="9A8D9E0BBA4FFFC7FE7523C8FD94FD17" box="[396,519,647,672]" italics="true" pageId="7" pageNumber="2018">landurensis</emphasis>
</taxonomicName>
[sic]
<treatmentCitation id="29586408BA4FFFC7FDBB23C8FC88FD17" author="Pilsbry HA" box="[578,795,647,672]" page="73" pageId="7" pageNumber="2018" year="1915">
<bibRefCitation id="CC683FE8BA4FFFC7FDBB23C8FC88FD17" author="Pilsbry HA" box="[578,795,647,672]" pageId="7" pageNumber="2018" pagination="73 - 74" refId="ref7566" refString="Pilsbry HA. 1915. A new Indian species of Pupillidae. Nautilus (Philadelphia). 29 (7): 73 - 74." type="journal article" year="1915">Pilsbry 1915: 7374</bibRefCitation>
</treatmentCitation>
<emphasis id="9A8D9E0BBA4FFFC7FCDA23C8FC5AFD17" bold="true" box="[803,969,647,672]" pageId="7" pageNumber="2018">new synonym</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A8464219BA4FFFC7FF5923E6FC78FD75" blockId="7.[160,1128,265,845]" box="[160,1003,681,706]" pageId="7" pageNumber="2018">
<taxonomicName id="6FF9399ABA4FFFC7FF5923E6FEEFFD75" ID-CoL="7QD4G" baseAuthorityName="Pilsbry" baseAuthorityYear="1915" box="[160,380,681,706]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="landurensis">
<emphasis id="9A8D9E0BBA4FFFC7FF5923E6FEEFFD75" box="[160,380,681,706]" italics="true" pageId="7" pageNumber="2018">Boysidia landurensis</emphasis>
</taxonomicName>
Pilsbry, 1917 (in
<bibRefCitation id="CC683FE8BA4FFFC7FDA823E6FD48FD75" author="Pilsbry HA" box="[593,731,681,706]" pageId="7" pageNumber="2018" refId="ref7592" refString="Pilsbry HA. 1916 - 1918. Manual of conchology, structural and systematic, with illustrations of the species. Ser. 2, Pulmonata. Vol. 24: Pupillidae (Gastrocoptinae). pp 1 - 380, pls 1 - 49. Philadelphia, published by the Conchological Section, Academy of Natural Sciences. [pp. 1 - 112, Pl. 1 - 13, 18 Dec 1916; pp. 113 - 176, pl. 14 - 29, 18 July 1917; pp. 177 - 256, pl. 30 - 38, 09 Nov 1917; pp. 257 - 380, pl. 39 - 49, 24 April 1918]." type="book" year="1916">Pilsbry 1916</bibRefCitation>
1918): 204, pl. 35, fig. 9
</paragraph>
<paragraph id="A8464219BA4FFFC7FF592383FD6CFD52" blockId="7.[160,1128,265,845]" box="[160,767,716,741]" pageId="7" pageNumber="2018">
<treatmentCitationGroup id="88E96537BA4FFFC7FF592383FD6CFD52" box="[160,767,716,741]" pageId="7" pageNumber="2018">
<taxonomicName id="6FF9399ABA4FFFC7FF592383FE2FFD52" ID-CoL="8VQFW" authorityName="Pilsbry" authorityYear="1915" box="[160,444,716,741]" class="Gastropoda" family="Gastrocoptidae" genus="Paraboysidia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA4FFFC7FF592383FE2FFD52" box="[160,444,716,741]" italics="true" pageId="7" pageNumber="2018">Paraboysidia landourensis</emphasis>
</taxonomicName>
<treatmentCitation id="29586408BA4FFFC7FE212383FD6CFD52" author="van Benthem Jutting WSS" box="[472,767,716,741]" page="39" pageId="7" pageNumber="2018" year="1950">
<bibRefCitation id="CC683FE8BA4FFFC7FE212383FD6CFD52" author="van Benthem Jutting WSS" box="[472,767,716,741]" pageId="7" pageNumber="2018" pagination="5 - 47" refId="ref7993" refString="van Benthem Jutting WSS. 1950. The Malayan species of Boysidia, Paraboysidia, Hypselostoma and Gyliotrachela (Gastropoda, Pulmonata, Vertiginidae) with a catalogue of all the species hitherto described. Bulletin of the Raffles Museum. 21: 5 - 47." type="journal article" year="1950">Benthem Jutting, 1950: 39</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A8464219BA4FFFC7FF5923A0FC4FFCBF" blockId="7.[160,1128,265,845]" box="[160,988,751,776]" pageId="7" pageNumber="2018">
<taxonomicName id="6FF9399ABA4FFFC7FF5923A0FEE9FCBF" ID-CoL="LHH3" authorityName="Benson" authorityYear="1849" box="[160,378,751,776]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA4FFFC7FF5923A0FEE9FCBF" box="[160,378,751,776]" italics="true" pageId="7" pageNumber="2018">Bensonella plicidens</emphasis>
</taxonomicName>
<bibRefCitation id="CC683FE8BA4FFFC7FE6F23A0FDB7FCBF" author="Hwang C-C" box="[406,548,751,776]" pageId="7" pageNumber="2018" pagination="15 - 26" refId="ref6907" refString="Hwang C-C. 2014. A new subspecies of land snail Bensonella plicidens lakainguta (Gastropoda, Vertiginidae) from southern Taiwan. Bull Malacol Taiwan 貝類學報. 37: 15 - 26." type="journal article" year="2014">Hwang 2014</bibRefCitation>
: fig. 5 (partim: fig. 5 shows
<taxonomicName id="6FF9399ABA4FFFC7FCA223A0FC47FCBF" ID-CoL="LHH3" authorityName="Benson" authorityYear="1849" box="[859,980,751,776]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA4FFFC7FCA223A0FC47FCBF" box="[859,980,751,776]" italics="true" pageId="7" pageNumber="2018">B. plicidens</emphasis>
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="E0E31192BA4FFFC7FF59225EFEFCFBF4" pageId="7" pageNumber="2018" type="description">
<paragraph id="A8464219BA4FFFC7FF59225EFE4BFCFA" blockId="7.[160,1128,265,845]" pageId="7" pageNumber="2018">
<taxonomicName id="6FF9399ABA4FFFC7FF59225EFE36FC9D" box="[160,421,785,810]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA4FFFC7FF59225EFE36FC9D" box="[160,421,785,810]" italics="true" pageId="7" pageNumber="2018">Bensonella landourensis</emphasis>
</taxonomicName>
and
<taxonomicName id="6FF9399ABA4FFFC7FE26225EFDAEFC9D" authorityName="Benson" authorityYear="1849" box="[479,573,785,810]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA4FFFC7FE26225EFDAEFC9D" box="[479,573,785,810]" italics="true" pageId="7" pageNumber="2018">plicidens</emphasis>
</taxonomicName>
<bibRefCitation id="CC683FE8BA4FFFC7FDA0225EFC11FC9D" author="Budha PB &amp; Backeljau T" box="[601,898,785,810]" pageId="7" pageNumber="2018" pagination="189 - 201" refId="ref6448" refString="Budha PB, Backeljau T. 2017. First report of Paraboysidia landourensis (Pilsbry, 1915) from Nepal, with description of a putative new species and a note on Bensonella plicidens (Benson, 1849) (Stylommatophora: family unclear). In: Budha PB, editor. Taxonomy of terrestrial molluscs of Nepal PhD dissertation. Antwerp (Belgium): Evolutionary Ecology Group, University of Antwerp; p. 189 - 201." type="book chapter" year="2017">Budha and Backeljau 2017</bibRefCitation>
(partim: figs 2B, 3, 5 A and F show
<taxonomicName id="6FF9399ABA4FFFC7FEAE227BFE43FCFA" authorityName="Benson" authorityYear="1849" box="[343,464,820,845]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA4FFFC7FEAE227BFE43FCFA" box="[343,464,820,845]" italics="true" pageId="7" pageNumber="2018">B. plicidens</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="A8464219BA4FFFC7FF592233FECCFC21" blockId="7.[160,1156,892,1091]" box="[160,351,892,918]" pageId="7" pageNumber="2018">
<emphasis id="9A8D9E0BBA4FFFC7FF592233FECCFC21" bold="true" box="[160,351,892,918]" italics="true" pageId="7" pageNumber="2018">
<typeStatus id="7742FCBBBA4FFFC7FF592233FF70FC21" box="[160,227,892,918]" pageId="7" pageNumber="2018">Types</typeStatus>
examined
</emphasis>
</paragraph>
<paragraph id="A8464219BA4FFFC7FF5922D0FEFCFBF4" blockId="7.[160,1156,892,1091]" pageId="7" pageNumber="2018">
<materialsCitation id="18914844BA4FFFC7FF5922D0FDCCFC0F" ID-GBIF-Occurrence="4018076311" box="[160,607,927,952]" pageId="7" pageNumber="2018" typeStatus="lectotype">
UMZC I.103325.A,
<typeStatus id="7742FCBBBA4FFFC7FE9C22D0FE5FFC0F" box="[357,460,927,952]" pageId="7" pageNumber="2018" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="6FF9399ABA4FFFC7FE1222D0FDCCFC0F" authorityName="Benson" authorityYear="1849" box="[491,607,927,952]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA4FFFC7FE1222D0FDCCFC0F" box="[491,607,927,952]" italics="true" pageId="7" pageNumber="2018">P. plicidens</emphasis>
</taxonomicName>
</materialsCitation>
(selected herein from a series of
<specimenCount id="BEFF8990BA4FFFC7FC4622D0FBD3FC0F" box="[959,1088,927,952]" pageId="7" pageNumber="2018" type="generic" typeStatus="syntypes">15 syntypes</specimenCount>
in the UMZC), R. McAndrew coll. (ex-
<collectorName id="050C27CFBA4FFFC7FDFB228DFD03FC6C" box="[514,656,962,987]" pageId="7" pageNumber="2018">W.H. Benson</collectorName>
coll.),
<materialsCitation id="18914844BA4FFFC7FD25228DFB13FC49" ID-GBIF-Occurrence="4018076332" collectionCode="NHMUK" pageId="7" pageNumber="2018" specimenCode="NHMUK 1954.6" typeStatus="paralectotype">
<location id="AD2614C2BA4FFFC7FD25228DFCACFC6C" LSID="urn:lsid:plazi:treatment:2050F30FBA4FFFCCFE5021C1FCF3FA3F:AD2614C2BA4FFFC7FD25228DFCACFC6C" box="[732,831,962,987]" county="Himalaya" municipality="Landour" name="Rockville" pageId="7" pageNumber="2018">Rockville</location>
,
<collectingMunicipality id="4822D863BA4FFFC7FCB5228DFC39FC6C" box="[844,938,962,987]" pageId="7" pageNumber="2018">Landour</collectingMunicipality>
,
<collectingCounty id="41273A95BA4FFFC7FC4E228DFB8CFC6C" box="[951,1055,962,987]" pageId="7" pageNumber="2018">Himalaya</collectingCounty>
;
<specimenCode id="F85FEA62BA4FFFC7FBD5228DFF7EFC49" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="2018" type="Museum">NHMUK 1954.6</specimenCode>
.2.937,
<typeStatus id="7742FCBBBA4FFFC7FEB922AAFE4FFC49" box="[320,476,997,1022]" pageId="7" pageNumber="2018" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="6FF9399ABA4FFFC7FDF522AAFCB8FC49" authority=", J. S. Hawkins" authorityName="J. S. Hawkins" box="[524,811,997,1022]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA4FFFC7FDF522AAFD18FC49" box="[524,651,997,1022]" italics="true" pageId="7" pageNumber="2018">P. plicidens</emphasis>
, J.S. Hawkins
</taxonomicName>
coll. (ex- W.H. Benson coll.)
</materialsCitation>
,
<materialsCitation id="18914844BA4FFFC7FF592548FD3FFB97" ID-GBIF-Occurrence="4018076333" box="[160,684,1031,1056]" collectionCode="ANSP" collectorName="Himalaya" pageId="7" pageNumber="2018" specimenCode="ANSP 16721" typeStatus="paratype">
W
<collectorName id="050C27CFBA4FFFC7FF442548FEB1FB97" box="[189,290,1031,1056]" pageId="7" pageNumber="2018">Himalaya</collectorName>
(labelled as
<typeStatus id="7742FCBBBA4FFFC7FE542548FD81FB97" box="[429,530,1031,1056]" pageId="7" pageNumber="2018" type="paratype">paratype</typeStatus>
);
<specimenCode id="F85FEA62BA4FFFC7FDDB2548FD3FFB97" box="[546,684,1031,1056]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="7" pageNumber="2018">ANSP 16721</specimenCode>
</materialsCitation>
,
<typeStatus id="7742FCBBBA4FFFC7FD4C2548FC84FB97" box="[693,791,1031,1056]" pageId="7" pageNumber="2018" type="holotype">holotype</typeStatus>
of
<emphasis id="9A8D9E0BBA4FFFC7FCC52548FC4FFB97" box="[828,988,1031,1056]" italics="true" pageId="7" pageNumber="2018">
<taxonomicName id="6FF9399ABA4FFFC7FCC52548FCDFFB97" box="[828,844,1031,1056]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="landourensis">B</taxonomicName>
. landourensis
</emphasis>
(photos examined: see
<figureCitation id="30C25E9CBA4FFFC7FEF12565FEF0FBF4" box="[264,355,1066,1091]" captionStart="Figure 5" captionStartId="7.[160,227,1642,1665]" captionTargetBox="[167,1146,1149,1608]" captionTargetId="figure-258@7.[166,1148,1148,1610]" captionTargetPageId="7" captionText="Figure 5. Bensonella plicidens (holotype of Bifidaria landourensis Pilsbry, 1915, ANSP 16721). Downloaded from the website of the Academy of Natural Sciences, Philadelphia, USA (ANSP)." figureDoi="http://doi.org/10.5281/zenodo.7560945" httpUri="https://zenodo.org/record/7560945/files/figure.png" pageId="7" pageNumber="2018">Figure 5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="FC861291BA4FFFC7FF592725FBD2F929" ID-DOI="http://doi.org/10.5281/zenodo.7560945" ID-Zenodo-Dep="7560945" httpUri="https://zenodo.org/record/7560945/files/figure.png" pageId="7" pageNumber="2018" startId="7.[160,227,1642,1665]" targetBox="[167,1146,1149,1608]" targetPageId="7">
<paragraph id="A8464219BA4FFFC7FF592725FBD2F929" blockId="7.[160,1157,1642,1694]" pageId="7" pageNumber="2018">
<emphasis id="9A8D9E0BBA4FFFC7FF592725FE91F936" bold="true" box="[160,258,1642,1665]" pageId="7" pageNumber="2018">Figure 5.</emphasis>
<taxonomicName id="6FF9399ABA4FFFC7FEF62725FE48F936" box="[271,475,1642,1665]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA4FFFC7FEF62725FE48F936" box="[271,475,1642,1665]" italics="true" pageId="7" pageNumber="2018">Bensonella plicidens</emphasis>
</taxonomicName>
(holotype of
<taxonomicName id="6FF9399ABA4FFFC7FD8F2725FC70F936" authority="Pilsbry, 1915" authorityName="Pilsbry" authorityYear="1915" box="[630,995,1642,1665]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="2018" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA4FFFC7FD8F2725FCC2F936" box="[630,849,1642,1665]" italics="true" pageId="7" pageNumber="2018">Bifidaria landourensis</emphasis>
<bibRefCitation id="CC683FE8BA4FFFC7FCA72725FC70F936" author="Pilsbry HA" box="[862,995,1642,1665]" pageId="7" pageNumber="2018" pagination="73 - 74" refId="ref7566" refString="Pilsbry HA. 1915. A new Indian species of Pupillidae. Nautilus (Philadelphia). 29 (7): 73 - 74." type="journal article" year="1915">Pilsbry, 1915</bibRefCitation>
</taxonomicName>
, ANSP 16721). Downloaded from the website of the Academy of Natural Sciences, Philadelphia, USA (ANSP).
</paragraph>
</caption>
<caption id="FC861291BA40FFC8FF59275BFF61F9FE" ID-DOI="http://doi.org/10.5281/zenodo.7560949" ID-Zenodo-Dep="7560949" httpUri="https://zenodo.org/record/7560949/files/figure.png" pageId="8" pageNumber="2019" startId="8.[160,225,1556,1579]" targetBox="[167,1147,145,1523]" targetPageId="8">
<paragraph id="A8464219BA40FFC8FF59275BFF61F9FE" blockId="8.[160,1156,1556,1609]" pageId="8" pageNumber="2019">
<emphasis id="9A8D9E0BBA40FFC8FF59275BFF6EF99C" bold="true" box="[160,253,1556,1579]" pageId="8" pageNumber="2019">Figure 6.</emphasis>
<taxonomicName id="6FF9399ABA40FFC8FEFE275BFE3FF99C" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[263,428,1556,1579]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="2019" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA40FFC8FEFE275BFE3FF99C" box="[263,428,1556,1579]" italics="true" pageId="8" pageNumber="2019">Bensonella hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA40FFC8FE4F275BFD95F99C" box="[438,518,1556,1579]" pageId="8" pageNumber="2019" rank="species">sp. nov.</taxonomicNameLabel>
NHMUK 20191114, specimen 1 (holotype). All photos: B. PállGergely.
</paragraph>
</caption>
<caption id="FC861291BA41FFC9FF592501FD60FB40" ID-DOI="http://doi.org/10.5281/zenodo.7560951" ID-Zenodo-Dep="7560951" httpUri="https://zenodo.org/record/7560951/files/figure.png" pageId="9" pageNumber="2020" startId="9.[160,225,1102,1125]" targetBox="[167,1147,145,1068]" targetPageId="9">
<paragraph id="A8464219BA41FFC9FF592501FD60FB40" blockId="9.[160,1157,1102,1271]" pageId="9" pageNumber="2020">
<emphasis id="9A8D9E0BBA41FFC9FF592501FF6AFBD2" bold="true" box="[160,249,1102,1125]" pageId="9" pageNumber="2020">Figure 7.</emphasis>
<taxonomicName id="6FF9399ABA41FFC9FF062501FDC0FBD2" authority="Hwang, 2014" authorityName="Hwang" authorityYear="2014" box="[255,595,1102,1125]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="2020" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA41FFC9FF062501FE5FFBD2" box="[255,460,1102,1125]" italics="true" pageId="9" pageNumber="2020">Bensonella lakainguta</emphasis>
<bibRefCitation id="CC683FE8BA41FFC9FE2B2501FDC0FBD2" author="Hwang C-C" box="[466,595,1102,1125]" pageId="9" pageNumber="2020" pagination="15 - 26" refId="ref6907" refString="Hwang C-C. 2014. A new subspecies of land snail Bensonella plicidens lakainguta (Gastropoda, Vertiginidae) from southern Taiwan. Bull Malacol Taiwan 貝類學報. 37: 15 - 26." type="journal article" year="2014">Hwang, 2014</bibRefCitation>
</taxonomicName>
. ac: Holotype of
<taxonomicName id="6FF9399ABA41FFC9FCFD2501FC45FBD2" authorityName="Hwang" authorityYear="2014" box="[772,982,1102,1125]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="2020" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA41FFC9FCFD2501FC45FBD2" box="[772,982,1102,1125]" italics="true" pageId="9" pageNumber="2020">Bensonella lakainguta</emphasis>
</taxonomicName>
(NMNS-7244-00); D:
<taxonomicName id="6FF9399ABA41FFC9FF402524FD83FB35" authority="Hwang, 2014" authorityName="Hwang" authorityYear="2014" box="[185,528,1131,1154]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="2020" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA41FFC9FF402524FE18FB35" box="[185,395,1131,1154]" italics="true" pageId="9" pageNumber="2020">Bensonella lakainguta</emphasis>
<bibRefCitation id="CC683FE8BA41FFC9FE692524FD83FB35" author="Hwang C-C" box="[400,528,1131,1154]" pageId="9" pageNumber="2020" pagination="15 - 26" refId="ref6907" refString="Hwang C-C. 2014. A new subspecies of land snail Bensonella plicidens lakainguta (Gastropoda, Vertiginidae) from southern Taiwan. Bull Malacol Taiwan 貝類學報. 37: 15 - 26." type="journal article" year="2014">Hwang, 2014</bibRefCitation>
</taxonomicName>
, Japan, Kochi Prefecture, Kami-shi, Ryugado Prefectural National Park, above the cave, 500 m along a side-road, 290 m, 33.601917°N, 133.746217°E, Leg. A. Hunyadi, D. Murányi and B. Páll-Gergely, 8 October 2016 (HNHM 105323); ef: Holotype of
<taxonomicName id="6FF9399ABA41FFC9FC4325E9FD27FB6D" authority="F. Fang, J. Wang and Y. Chen, 2015" authorityName="F. Fang, J. Wang and Y. Chen" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="2020" phylum="Mollusca" rank="species" species="qingliangfengensis" subGenus="Bensonella">
<emphasis id="9A8D9E0BBA41FFC9FC4325E9FB9AFB0A" box="[954,1033,1190,1213]" italics="true" pageId="9" pageNumber="2020">Boysidia</emphasis>
(
<emphasis id="9A8D9E0BBA41FFC9FBEF25E9FBEFFB0A" box="[1046,1148,1190,1213]" italics="true" pageId="9" pageNumber="2020">Bensonella</emphasis>
)
<emphasis id="9A8D9E0BBA41FFC9FF59258CFEC1FB6D" box="[160,338,1219,1242]" italics="true" pageId="9" pageNumber="2020">qingliangfengensis</emphasis>
F. Fang, J. Wang and Y. Chen, 2015
</taxonomicName>
. Photos: ac and ef: images from the original descriptions, d: B. Páll-Gergely. Figs e and f are not to scale.
</paragraph>
</caption>
<subSubSection id="E0E31192BA41FFC9FF59242EFF64F9DA" pageId="9" pageNumber="2020" type="materials_examined">
<paragraph id="A8464219BA41FFC9FF59242EFD91FACC" blockId="9.[160,1156,1377,1645]" box="[160,514,1377,1403]" pageId="9" pageNumber="2020">
<emphasis id="9A8D9E0BBA41FFC9FF59242EFD91FACC" bold="true" box="[160,514,1377,1403]" italics="true" pageId="9" pageNumber="2020">Additional material examined</emphasis>
</paragraph>
<paragraph id="A8464219BA41FFC9FF5924CBFF64F9DA" blockId="9.[160,1156,1377,1645]" pageId="9" pageNumber="2020">
<collectionCode id="CEE8DADCBA41FFC9FF5924CBFF6BFA2A" box="[160,248,1412,1437]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="2020" type="Museum">NHMUK</collectionCode>
1856.9.15.69/A, 36 shells, Capt.
<collectionCode id="CEE8DADCBA41FFC9FDAF24CBFDF5FA2A" box="[598,614,1412,1437]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="2020">T</collectionCode>
. Hutton coll., Moussorie;
<collectionCode id="CEE8DADCBA41FFC9FC8124CBFC43FA2A" box="[888,976,1412,1437]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="2020" type="Museum">NHMUK</collectionCode>
20191111/A, 10 shells, Cherra,
<collectingCountry id="D0EE0289BA41FFC9FEA924E8FE1FFA77" box="[336,396,1447,1472]" name="India" pageId="9" pageNumber="2020">India</collectingCountry>
, mixed sample that includes
<collectionCode id="CEE8DADCBA41FFC9FD0B24E8FCD8FA77" box="[754,843,1447,1472]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="2020" type="Museum">NHMUK</collectionCode>
20191111/B, identified as
<taxonomicName id="6FF9399ABA41FFC9FF592485FF6FFA54" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[160,252,1482,1507]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="2020" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA41FFC9FF592485FF6FFA54" box="[160,252,1482,1507]" italics="true" pageId="9" pageNumber="2020">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA41FFC9FEFE2485FEF6FA54" box="[263,357,1482,1507]" pageId="9" pageNumber="2020" rank="species">sp. nov.</taxonomicNameLabel>
;
<collectionCode id="CEE8DADCBA41FFC9FE8A2485FE58FA54" box="[371,459,1482,1507]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="2020" type="Museum">NHMUK</collectionCode>
1903.7.1.2882/B, six shells, Godwin-Austen coll., Moussorie, NW Himalaya, mixed sample that includes
<collectionCode id="CEE8DADCBA41FFC9FD4324A3FC87F9B2" box="[698,788,1516,1541]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="2020" type="Museum">NHMUK</collectionCode>
1903.7.1.2882/C, identified as
<taxonomicName id="6FF9399ABA41FFC9FF592740FF6FF99F" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[160,252,1551,1576]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="2020" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA41FFC9FF592740FF6FF99F" box="[160,252,1551,1576]" italics="true" pageId="9" pageNumber="2020">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA41FFC9FEFE2740FEF5F99F" box="[263,358,1551,1576]" pageId="9" pageNumber="2020" rank="species">sp. nov.</taxonomicNameLabel>
;
<collectionCode id="CEE8DADCBA41FFC9FE8D2740FE5FF99F" box="[372,460,1551,1576]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="2020" type="Museum">NHMUK</collectionCode>
1906.2.2.190/B, 7 shells, W.
<collectionCode id="CEE8DADCBA41FFC9FCEB2740FCB1F99F" box="[786,802,1551,1576]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="2020">T</collectionCode>
. Blanford coll., Cherra Poonje, Khasi Hills, mixed sample that includes
<collectionCode id="CEE8DADCBA41FFC9FD8C277DFD5EF9FC" box="[629,717,1586,1611]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="2020" type="Museum">NHMUK</collectionCode>
1906.2.2.190/C, identified as
<taxonomicName id="6FF9399ABA41FFC9FBD1277DFB10F9FC" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[1064,1155,1586,1611]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="2020" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA41FFC9FBD1277DFB10F9FC" box="[1064,1155,1586,1611]" italics="true" pageId="9" pageNumber="2020">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA41FFC9FF59271BFF64F9DA" box="[160,247,1620,1645]" pageId="9" pageNumber="2020" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="FC861291BA42FFCAFF59275BFEA5F9FF" ID-DOI="http://doi.org/10.5281/zenodo.7560955" ID-Zenodo-Dep="7560955" httpUri="https://zenodo.org/record/7560955/files/figure.png" pageId="10" pageNumber="2021" startId="10.[160,225,1556,1579]" targetBox="[168,1147,145,1522]" targetPageId="10">
<paragraph id="A8464219BA42FFCAFF59275BFEA5F9FF" blockId="10.[160,1157,1556,1608]" pageId="10" pageNumber="2021">
<emphasis id="9A8D9E0BBA42FFCAFF59275BFF6AF99C" bold="true" box="[160,249,1556,1579]" pageId="10" pageNumber="2021">Figure 8.</emphasis>
<taxonomicName id="6FF9399ABA42FFCAFF07275BFDC5F99C" authority="Hwang, 2014" authorityName="Hwang" authorityYear="2014" box="[254,598,1556,1579]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="2021" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA42FFCAFF07275BFE43F99C" box="[254,464,1556,1579]" italics="true" pageId="10" pageNumber="2021">Bensonella lakainguta</emphasis>
<bibRefCitation id="CC683FE8BA42FFCAFE2F275BFDC5F99C" author="Hwang C-C" box="[470,598,1556,1579]" pageId="10" pageNumber="2021" pagination="15 - 26" refId="ref6907" refString="Hwang C-C. 2014. A new subspecies of land snail Bensonella plicidens lakainguta (Gastropoda, Vertiginidae) from southern Taiwan. Bull Malacol Taiwan 貝類學報. 37: 15 - 26." type="journal article" year="2014">Hwang, 2014</bibRefCitation>
</taxonomicName>
, Japan, Omi, NHMUK 201901108, specimen 1. All photos: B. Páll-Gergely.
</paragraph>
</caption>
<caption id="FC861291BA43FFCBFF5923A2FD4DFCEC" ID-DOI="http://doi.org/10.5281/zenodo.7560957" ID-Zenodo-Dep="7560957" httpUri="https://zenodo.org/record/7560957/files/figure.png" pageId="11" pageNumber="2022" startId="11.[160,225,749,772]" targetBox="[167,1146,145,715]" targetPageId="11">
<paragraph id="A8464219BA43FFCBFF5923A2FD4DFCEC" blockId="11.[160,1155,749,859]" pageId="11" pageNumber="2022">
<emphasis id="9A8D9E0BBA43FFCBFF5923A2FF6AFCB3" bold="true" box="[160,249,749,772]" pageId="11" pageNumber="2022">Figure 9.</emphasis>
a, b:
<taxonomicName id="6FF9399ABA43FFCBFED523A2FD12FCB3" authority="(Benson, 1849)" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[300,641,749,772]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA43FFCBFED523A2FE79FCB3" box="[300,490,749,772]" italics="true" pageId="11" pageNumber="2022">Bensonella plicidens</emphasis>
(
<bibRefCitation id="CC683FE8BA43FFCBFE0C23A2FDEAFCB3" author="Benson WH" box="[501,633,749,772]" pageId="11" pageNumber="2022" pagination="125 - 128" refId="ref6320" refString="Benson WH. 1849. Descriptions of four new Asiatic species of the genus Pupa of Draparnaud. Ann Mag Nat Hist. 4 (20): 125 - 128. doi: 10.1080 / 03745486009496158." type="journal article" year="1849">Benson, 1849</bibRefCitation>
)
</taxonomicName>
, NHMUK 56.9.15.69A, specimen 1 (same as Figure 4); c,d:
<taxonomicName id="6FF9399ABA43FFCBFF372245FEEAFC96" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[206,377,778,801]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA43FFCBFF372245FEEAFC96" box="[206,377,778,801]" italics="true" pageId="11" pageNumber="2022">Bensonella hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA43FFCBFE7D2245FE49FC96" box="[388,474,778,801]" pageId="11" pageNumber="2022" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, NHMUK 20191114 (same as Figure 6); e,f:
<taxonomicName id="6FF9399ABA43FFCBFBE32245FE00FC89" authority="Hwang, 2014" authorityName="Hwang" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA43FFCBFBE32245FE95FC89" italics="true" pageId="11" pageNumber="2022">Bensonella lakainguta</emphasis>
<bibRefCitation id="CC683FE8BA43FFCBFEF62268FE00FC89" author="Hwang C-C" box="[271,403,807,830]" pageId="11" pageNumber="2022" pagination="15 - 26" refId="ref6907" refString="Hwang C-C. 2014. A new subspecies of land snail Bensonella plicidens lakainguta (Gastropoda, Vertiginidae) from southern Taiwan. Bull Malacol Taiwan 貝類學報. 37: 15 - 26." type="journal article" year="2014">Hwang, 2014</bibRefCitation>
</taxonomicName>
, HNHM 105323 (same as Figure 7(d)); g, h:
<taxonomicName id="6FF9399ABA43FFCBFCAA2268FB10FC89" authority=", NHMUK" authorityName="NHMUK" box="[851,1155,807,830]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA43FFCBFCAA2268FBBBFC89" box="[851,1064,807,830]" italics="true" pageId="11" pageNumber="2022">Bensonella lakainguta</emphasis>
, NHMUK
</taxonomicName>
201901108 (same as Figure 8). All images: B. Páll-Gergely.
</paragraph>
</caption>
<subSubSection id="E0E31192BA43FFCBFF59228EFB98F917" pageId="11" pageNumber="2022" type="diagnosis">
<paragraph id="A8464219BA43FFCBFF59228EFE16FC6C" blockId="11.[160,1156,961,1368]" box="[160,389,961,987]" pageId="11" pageNumber="2022">
<emphasis id="9A8D9E0BBA43FFCBFF59228EFE16FC6C" bold="true" box="[160,389,961,987]" italics="true" pageId="11" pageNumber="2022">Extended diagnosis</emphasis>
</paragraph>
<paragraph id="A8464219BA43FFCBFF5922ABFDF8FAEF" blockId="11.[160,1156,961,1368]" pageId="11" pageNumber="2022">Shell triangular (pyramidal), slightly ovoid, height somewhat variable; shell colour greyishyellowish and corneous; sculpture with inconspicuous growth lines and very fine spiral striation (its strength variable, in some shells nearly absent); aperture with numerous barriers; parietal wall with a parietal and an angular lamella, both being lower at their middles, and a third, short, deeply situated one between parietal lamella and columellarparietal junction; palatal wall with three long main plicae with elevated central parts, usually (but not always) with additional short plicae between them; columellar lamella similar to palatal plicae; between the lowest palatal plicae and the columellar lamella there are two short plicae; an additional, blunt tubercle sits on the edge of the palatal lip, at the position where the palatal lip forms a little sinus; lower part of columellar lip with two very low, blunt denticles (swellings).</paragraph>
<paragraph id="A8464219BA43FFCBFF5924DBFE0EFA19" blockId="11.[160,1156,1428,1696]" box="[160,413,1428,1454]" pageId="11" pageNumber="2022">
<emphasis id="9A8D9E0BBA43FFCBFF5924DBFE0EFA19" bold="true" box="[160,413,1428,1454]" italics="true" pageId="11" pageNumber="2022">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="A8464219BA43FFCBFF5924F8FB98F917" blockId="11.[160,1156,1428,1696]" pageId="11" pageNumber="2022">
For differences with
<taxonomicName id="6FF9399ABA43FFCBFE7024F8FE72FA67" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[393,481,1463,1488]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA43FFCBFE7024F8FE72FA67" box="[393,481,1463,1488]" italics="true" pageId="11" pageNumber="2022">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA43FFCBFE1324F8FDD7FA67" box="[490,580,1463,1488]" pageId="11" pageNumber="2022" rank="species">sp. nov.</taxonomicNameLabel>
, see under that species.
<taxonomicName id="6FF9399ABA43FFCBFCA224F8FBA5FA67" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[859,1078,1463,1488]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA43FFCBFCA224F8FBA5FA67" box="[859,1078,1463,1488]" italics="true" pageId="11" pageNumber="2022">Bensonella plicidens</emphasis>
</taxonomicName>
differs from
<taxonomicName id="6FF9399ABA43FFCBFF272495FEE3FA44" box="[222,368,1498,1523]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA43FFCBFF272495FEE3FA44" box="[222,368,1498,1523]" italics="true" pageId="11" pageNumber="2022">B. lakainguta</emphasis>
</taxonomicName>
in the following traits: shell shape is similar, but
<taxonomicName id="6FF9399ABA43FFCBFC5E2495FBB2FA44" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[935,1057,1498,1523]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA43FFCBFC5E2495FBB2FA44" box="[935,1057,1498,1523]" italics="true" pageId="11" pageNumber="2022">B. plicidens</emphasis>
</taxonomicName>
is wider triangular; the apex is less pointed; the sculpture is clearly densely spirally striated, whereas
<taxonomicName id="6FF9399ABA43FFCBFEF12750FE0FF98F" box="[264,412,1567,1592]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA43FFCBFEF12750FE0FF98F" box="[264,412,1567,1592]" italics="true" pageId="11" pageNumber="2022">B. lakainguta</emphasis>
</taxonomicName>
is smooth; the arrangement of the apertural barriers is similar, but the barriers of
<taxonomicName id="6FF9399ABA43FFCBFE87270DFE6AF9EC" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[382,505,1602,1627]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA43FFCBFE87270DFE6AF9EC" box="[382,505,1602,1627]" italics="true" pageId="11" pageNumber="2022">B. plicidens</emphasis>
</taxonomicName>
are elongated ridges, which are highest at their middle parts, whereas in
<taxonomicName id="6FF9399ABA43FFCBFE98272AFE7CF9C9" box="[353,495,1637,1662]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="2022" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA43FFCBFE98272AFE7CF9C9" box="[353,495,1637,1662]" italics="true" pageId="11" pageNumber="2022">B. lakainguta</emphasis>
</taxonomicName>
the central part of each barrier is a hook pointing anteriorly (outside of the shell), with an anteriorly elongated low ridge before the hook.
</paragraph>
</subSubSection>
<subSubSection id="E0E31192BA44FFCCFF5921C1FCF3FA3F" pageId="12" pageNumber="2023" type="discussion">
<paragraph id="A8464219BA44FFCCFF5921C1FE95FF1F" blockId="12.[160,1156,142,1416]" box="[160,262,142,168]" pageId="12" pageNumber="2023">
<emphasis id="9A8D9E0BBA44FFCCFF5921C1FE95FF1F" bold="true" box="[160,262,142,168]" italics="true" pageId="12" pageNumber="2023">Remarks</emphasis>
</paragraph>
<paragraph id="A8464219BA44FFCCFF5921FDFD6FFE84" blockId="12.[160,1156,142,1416]" pageId="12" pageNumber="2023">
The original spelling of
<taxonomicName id="6FF9399ABA44FFCCFE5221FDFDC3FF7C" authorityName="Pilsbry" authorityYear="1915" box="[427,592,178,203]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFE5221FDFDC3FF7C" box="[427,592,178,203]" italics="true" pageId="12" pageNumber="2023">B. landourensis</emphasis>
</taxonomicName>
was
<taxonomicName id="6FF9399ABA44FFCCFD7321FDFCB3FF7C" baseAuthorityName="Pilsbry" baseAuthorityYear="1915" box="[650,800,178,203]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landurensis">
<emphasis id="9A8D9E0BBA44FFCCFD7321FDFCB3FF7C" box="[650,800,178,203]" italics="true" pageId="12" pageNumber="2023">B. landurensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="CC683FE8BA44FFCCFCB521FDFC76FF7C" author="Pilsbry HA" box="[844,997,178,203]" pageId="12" pageNumber="2023" pagination="73 - 74" refId="ref7566" refString="Pilsbry HA. 1915. A new Indian species of Pupillidae. Nautilus (Philadelphia). 29 (7): 73 - 74." type="journal article" year="1915">Pilsbry (1915)</bibRefCitation>
, although the
<typeStatus id="7742FCBBBA44FFCCFF59219BFF41FF5A" box="[160,210,212,237]" pageId="12" pageNumber="2023">type</typeStatus>
locality was spelled Landour (town in Mussoorie,
<collectingCountry id="D0EE0289BA44FFCCFCD2219BFCF4FF5A" box="[811,871,212,237]" name="India" pageId="12" pageNumber="2023">India</collectingCountry>
). Therefore, the spelling ̍
<taxonomicName id="6FF9399ABA44FFCCFF5C21B8FEBDFEA7" authorityName="Pilsbry" authorityYear="1915" box="[165,302,247,272]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFF5C21B8FEBDFEA7" box="[165,302,247,272]" italics="true" pageId="12" pageNumber="2023">landourensis</emphasis>
</taxonomicName>
̾ by
<bibRefCitation id="CC683FE8BA44FFCCFE9921B8FD0FFEA7" author="van Benthem Jutting WSS" box="[352,668,247,272]" pageId="12" pageNumber="2023" pagination="5 - 47" refId="ref7993" refString="van Benthem Jutting WSS. 1950. The Malayan species of Boysidia, Paraboysidia, Hypselostoma and Gyliotrachela (Gastropoda, Pulmonata, Vertiginidae) with a catalogue of all the species hitherto described. Bulletin of the Raffles Museum. 21: 5 - 47." type="journal article" year="1950">van Benthem Jutting (1950)</bibRefCitation>
is a justified emendation, and ̍
<taxonomicName id="6FF9399ABA44FFCCFBFA21B8FBEDFEA7" baseAuthorityName="Pilsbry" baseAuthorityYear="1915" box="[1027,1150,247,272]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landurensis">
<emphasis id="9A8D9E0BBA44FFCCFBFA21B8FBEDFEA7" box="[1027,1150,247,272]" italics="true" pageId="12" pageNumber="2023">landurensis</emphasis>
</taxonomicName>
̾ is an incorrect original spelling (under ICZN Art. 32.5).
</paragraph>
<paragraph id="A8464219BA44FFCCFF392073FC72FD64" blockId="12.[160,1156,142,1416]" pageId="12" pageNumber="2023">
<bibRefCitation id="CC683FE8BA44FFCCFF392073FECEFEE2" author="Pilsbry HA" box="[192,349,316,341]" pageId="12" pageNumber="2023" pagination="73 - 74" refId="ref7566" refString="Pilsbry HA. 1915. A new Indian species of Pupillidae. Nautilus (Philadelphia). 29 (7): 73 - 74." type="journal article" year="1915">Pilsbry (1915)</bibRefCitation>
distinguished
<taxonomicName id="6FF9399ABA44FFCCFDF52073FD25FEE2" authorityName="Pilsbry" authorityYear="1915" box="[524,694,316,341]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFDF52073FD25FEE2" box="[524,694,316,341]" italics="true" pageId="12" pageNumber="2023">B. landourensis</emphasis>
</taxonomicName>
from
<taxonomicName id="6FF9399ABA44FFCCFCFA2073FC12FEE2" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[771,897,316,341]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFCFA2073FC12FEE2" box="[771,897,316,341]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
by the uninterrupted angular lamella in the former. Additional differences were also mentioned, such as the length of the angular lamella, which does not emerge to the edge of the peristome (i.e. it stops before the peristome edge) in
<taxonomicName id="6FF9399ABA44FFCCFDAD20EBFD6FFE0A" authorityName="Pilsbry" authorityYear="1915" box="[596,764,420,445]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFDAD20EBFD6FFE0A" box="[596,764,420,445]" italics="true" pageId="12" pageNumber="2023">B. landourensis</emphasis>
</taxonomicName>
but does in
<taxonomicName id="6FF9399ABA44FFCCFC6220EBFB84FE0A" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[923,1047,420,445]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFC6220EBFB84FE0A" box="[923,1047,420,445]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
, and the parietal lamella, which is shorter in
<taxonomicName id="6FF9399ABA44FFCCFDB12088FD56FE57" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[584,709,455,480]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFDB12088FD56FE57" box="[584,709,455,480]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
than in
<taxonomicName id="6FF9399ABA44FFCCFCC92088FC4BFE57" authorityName="Pilsbry" authorityYear="1915" box="[816,984,455,480]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFCC92088FC4BFE57" box="[816,984,455,480]" italics="true" pageId="12" pageNumber="2023">B. landourensis</emphasis>
</taxonomicName>
. Although the original description of
<taxonomicName id="6FF9399ABA44FFCCFE5220A5FDC0FDB4" authorityName="Pilsbry" authorityYear="1915" box="[427,595,490,515]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFE5220A5FDC0FDB4" box="[427,595,490,515]" italics="true" pageId="12" pageNumber="2023">B. landourensis</emphasis>
</taxonomicName>
does not mention hooked apertural barriers in
<taxonomicName id="6FF9399ABA44FFCCFF592343FE89FD92" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,282,524,549]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFF592343FE89FD92" box="[160,282,524,549]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
, it is clear that Pilsbry̾s (1915) new species was not compared with correctly identified
<taxonomicName id="6FF9399ABA44FFCCFEE82360FE1BFDFF" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[273,392,559,584]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFEE82360FE1BFDFF" box="[273,392,559,584]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
but with the species that is described here as new (
<taxonomicName id="6FF9399ABA44FFCCFC3C2360FB88FDFF" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[965,1051,559,584]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA44FFCCFC3C2360FB88FDFF" box="[965,1051,559,584]" italics="true" pageId="12" pageNumber="2023">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA44FFCCFBD82360FBE9FDFF" box="[1057,1146,559,584]" pageId="12" pageNumber="2023" rank="species">sp. nov.</taxonomicNameLabel>
), because
<taxonomicName id="6FF9399ABA44FFCCFEFB231DFEC4FDDC" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[258,343,594,619]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA44FFCCFEFB231DFEC4FDDC" box="[258,343,594,619]" italics="true" pageId="12" pageNumber="2023">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA44FFCCFEA7231DFE20FDDC" box="[350,435,594,619]" pageId="12" pageNumber="2023" rank="species">sp. nov.</taxonomicNameLabel>
has an interrupted angular lamella, whereas that of
<taxonomicName id="6FF9399ABA44FFCCFC0C231DFBFEFDDC" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[1013,1133,594,619]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFC0C231DFBFEFDDC" box="[1013,1133,594,619]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
is not interrupted. Comparison of the
<typeStatus id="7742FCBBBA44FFCCFD99233BFD54FD3A" box="[608,711,628,653]" pageId="12" pageNumber="2023" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6FF9399ABA44FFCCFD03233BFC39FD3A" authorityName="Pilsbry" authorityYear="1915" box="[762,938,628,653]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFD03233BFC39FD3A" box="[762,938,628,653]" italics="true" pageId="12" pageNumber="2023">B. landourensis</emphasis>
</taxonomicName>
with
<typeStatus id="7742FCBBBA44FFCCFC00233BFBCDFD3A" box="[1017,1118,628,653]" pageId="12" pageNumber="2023" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="6FF9399ABA44FFCCFF5923D8FE8DFD07" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,286,663,688]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFF5923D8FE8DFD07" box="[160,286,663,688]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
reveal no significant differences, other than the somewhat more slender shell of
<taxonomicName id="6FF9399ABA44FFCCFF0023F5FE0EFD64" authorityName="Pilsbry" authorityYear="1915" box="[249,413,698,723]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFF0023F5FE0EFD64" box="[249,413,698,723]" italics="true" pageId="12" pageNumber="2023">B. landourensis</emphasis>
</taxonomicName>
, which can be explained by intraspecific variability.
</paragraph>
<paragraph id="A8464219BA44FFCCFF392393FE37FB9A" blockId="12.[160,1156,142,1416]" pageId="12" pageNumber="2023">
<bibRefCitation id="CC683FE8BA44FFCCFF392393FECAFD42" author="Pilsbry HA" box="[192,345,732,757]" pageId="12" pageNumber="2023" pagination="73 - 74" refId="ref7566" refString="Pilsbry HA. 1915. A new Indian species of Pupillidae. Nautilus (Philadelphia). 29 (7): 73 - 74." type="journal article" year="1915">Pilsbry (1915)</bibRefCitation>
mentioned that the single shell of
<taxonomicName id="6FF9399ABA44FFCCFD162393FC07FD42" authorityName="Pilsbry" authorityYear="1915" box="[751,916,732,757]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFD162393FC07FD42" box="[751,916,732,757]" italics="true" pageId="12" pageNumber="2023">B. landourensis</emphasis>
</taxonomicName>
(=
<taxonomicName id="6FF9399ABA44FFCCFC442393FBA4FD42" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[957,1079,732,757]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFC442393FBA4FD42" box="[957,1079,732,757]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
in our current understanding) was found amongst shells of
<taxonomicName id="6FF9399ABA44FFCCFD1723B0FCF5FCAF" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[750,870,767,792]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFD1723B0FCF5FCAF" box="[750,870,767,792]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
(=
<taxonomicName id="6FF9399ABA44FFCCFC7323B0FC73FCAF" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[906,992,767,792]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA44FFCCFC7323B0FC73FCAF" box="[906,992,767,792]" italics="true" pageId="12" pageNumber="2023">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA44FFCCFC1F23B0FBAFFCAF" box="[998,1084,767,792]" pageId="12" pageNumber="2023" rank="species">sp. nov.</taxonomicNameLabel>
in our current understanding), all of which had been received from Benson. We were unable to examine the shells studied by Pilsbry, other than the
<typeStatus id="7742FCBBBA44FFCCFCEA220BFCE4FCEA" box="[787,887,836,861]" pageId="12" pageNumber="2023" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6FF9399ABA44FFCCFC5B220BFBDAFCEA" authorityName="Pilsbry" authorityYear="1915" box="[930,1097,836,861]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFC5B220BFBDAFCEA" box="[930,1097,836,861]" italics="true" pageId="12" pageNumber="2023">B. landourensis</emphasis>
</taxonomicName>
. It is possible that Benson had both
<taxonomicName id="6FF9399ABA44FFCCFE032228FDE2FC37" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[506,625,871,896]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFE032228FDE2FC37" box="[506,625,871,896]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
(a species with normal barriers) and
<taxonomicName id="6FF9399ABA44FFCCFBF12228FBCDFC37" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[1032,1118,871,896]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA44FFCCFBF12228FBCDFC37" box="[1032,1118,871,896]" italics="true" pageId="12" pageNumber="2023">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA44FFCCFB9D2228FF42FC14" pageId="12" pageNumber="2023" rank="species">sp. nov.</taxonomicNameLabel>
(a species with hooked barriers) when he described
<taxonomicName id="6FF9399ABA44FFCCFCCF22C5FC22FC14" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[822,945,906,931]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFCCF22C5FC22FC14" box="[822,945,906,931]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
. However, neither
<bibRefCitation id="CC683FE8BA44FFCCFF5922E3FED3FC72" author="Benson WH" box="[160,320,940,965]" pageId="12" pageNumber="2023" pagination="125 - 128" refId="ref6320" refString="Benson WH. 1849. Descriptions of four new Asiatic species of the genus Pupa of Draparnaud. Ann Mag Nat Hist. 4 (20): 125 - 128. doi: 10.1080 / 03745486009496158." type="journal article" year="1849">Benson (1849)</bibRefCitation>
nor subsequent authors mentioned hooked barriers as a character of that species, and the
<typeStatus id="7742FCBBBA44FFCCFEA72280FE53FC5F" box="[350,448,975,1000]" pageId="12" pageNumber="2023" type="syntype">syntypes</typeStatus>
in the original Benson collection in the UMZC all have normal barriers (Preece et al., in litteris). Even the name ̍
<taxonomicName id="6FF9399ABA44FFCCFD2522BDFCA9FBBC" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[732,826,1010,1035]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFD2522BDFCA9FBBC" box="[732,826,1010,1035]" italics="true" pageId="12" pageNumber="2023">plicidens</emphasis>
</taxonomicName>
̾ indicates that the apertural barriers were plica-like.
</paragraph>
<paragraph id="A8464219BA44FFCCFF392578FBD2FA97" blockId="12.[160,1156,142,1416]" pageId="12" pageNumber="2023">
We do not know how Pilsbry received Benson̾s original specimens, or whether it is really true that he did. It is remarkable that all of the
<typeStatus id="7742FCBBBA44FFCCFD1C2515FCD4FBC4" box="[741,839,1114,1139]" pageId="12" pageNumber="2023" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="6FF9399ABA44FFCCFC902515FC72FBC4" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[873,993,1114,1139]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFC902515FC72FBC4" box="[873,993,1114,1139]" italics="true" pageId="12" pageNumber="2023">
B.
<materialsCitation id="18914844BA44FFCCFC7A2515FC72FBC4" ID-GBIF-Occurrence="4018076303" box="[899,993,1114,1139]" collectionCode="UMZC, NHM" collectorName="J. S. Hawkins" country="United Kingdom" location="Pilsbry" pageId="12" pageNumber="2023" specimenCount="1">plicidens</materialsCitation>
</emphasis>
</taxonomicName>
that remain in
<collectingCountry id="D0EE0289BA44FFCCFF592533FF6FFB22" box="[160,252,1148,1173]" name="United Kingdom" pageId="12" pageNumber="2023">England</collectingCountry>
(UMZC and the shell from the
<collectorName id="050C27CFBA44FFCCFDA72533FD75FB22" box="[606,742,1148,1173]" pageId="12" pageNumber="2023">J.S. Hawkins</collectorName>
collection in the NHM) have normal barriers, but the lot
<location id="AD2614C2BA44FFCCFE7825D0FE58FB0F" LSID="urn:lsid:plazi:treatment:2050F30FBA4FFFCCFE5021C1FCF3FA3F:AD2614C2BA44FFCCFE7825D0FE58FB0F" box="[385,459,1183,1208]" country="United Kingdom" name="Pilsbry" pageId="12" pageNumber="2023">Pilsbry</location>
received, according to this analysis, was entirely hooked with only one normal
<taxonomicName id="6FF9399ABA44FFCCFEA6258DFE44FB6C" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[351,471,1218,1243]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFEA6258DFE44FB6C" box="[351,471,1218,1243]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
that was redescribed as
<taxonomicName id="6FF9399ABA44FFCCFD14258DFC03FB6C" authorityName="Pilsbry" authorityYear="1915" box="[749,912,1218,1243]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="landourensis">
<emphasis id="9A8D9E0BBA44FFCCFD14258DFC03FB6C" box="[749,912,1218,1243]" italics="true" pageId="12" pageNumber="2023">B. landourensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="CC683FE8BA44FFCCFC40258DFBC2FB6C" author="Pilsbry HA" box="[953,1105,1218,1243]" pageId="12" pageNumber="2023" pagination="73 - 74" refId="ref7566" refString="Pilsbry HA. 1915. A new Indian species of Pupillidae. Nautilus (Philadelphia). 29 (7): 73 - 74." type="journal article" year="1915">Pilsbry (1915)</bibRefCitation>
. It is possible that the ̍
<taxonomicName id="6FF9399ABA44FFCCFE9F25ABFE4EFB4A" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[358,477,1252,1277]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFE9F25ABFE4EFB4A" box="[358,477,1252,1277]" italics="true" pageId="12" pageNumber="2023">
B.
<materialsCitation id="18914844BA44FFCCFE8625ABFE4EFB4A" ID-GBIF-Occurrence="4018076317" box="[383,477,1252,1277]" collectorName="Benson" country="United Kingdom" location="William Benson" pageId="12" pageNumber="2023" specimenCount="1">plicidens</materialsCitation>
</emphasis>
</taxonomicName>
̾ specimens that
<bibRefCitation id="CC683FE8BA44FFCCFD6125ABFCBEFB4A" author="Pilsbry HA" box="[664,813,1252,1277]" pageId="12" pageNumber="2023" pagination="73 - 74" refId="ref7566" refString="Pilsbry HA. 1915. A new Indian species of Pupillidae. Nautilus (Philadelphia). 29 (7): 73 - 74." type="journal article" year="1915">Pilsbry (1915)</bibRefCitation>
referred to as originating from
<collectorName id="050C27CFBA44FFCCFF592448FF60FA97" box="[160,243,1287,1312]" pageId="12" pageNumber="2023">Benson</collectorName>
were not part of the type series, or were not directly from
<location id="AD2614C2BA44FFCCFC742448FBAFFA97" LSID="urn:lsid:plazi:treatment:2050F30FBA4FFFCCFE5021C1FCF3FA3F:AD2614C2BA44FFCCFC742448FBAFFA97" box="[909,1084,1287,1312]" country="United Kingdom" name="William Benson" pageId="12" pageNumber="2023">William Benson</location>
.
</paragraph>
<paragraph id="A8464219BA44FFCCFF392465FCF3FA3F" blockId="12.[160,1156,142,1416]" pageId="12" pageNumber="2023">
<materialsCitation id="18914844BA44FFCCFF392465FE29FAD2" ID-GBIF-Occurrence="4018076316" collectionCode="UMZC" pageId="12" pageNumber="2023" specimenCount="2" typeStatus="lectotype">
Here we select the syntype in Cambridge (
<collectionCode id="CEE8DADCBA44FFCCFD472465FC94FAF4" box="[702,775,1322,1347]" pageId="12" pageNumber="2023">UMZC</collectionCode>
) as a
<typeStatus id="7742FCBBBA44FFCCFCAD2465FC2DFAF4" box="[852,958,1322,1347]" pageId="12" pageNumber="2023" type="lectotype">lectotype</typeStatus>
to avoid further nomenclatural problems
</materialsCitation>
. Consequently,
<taxonomicName id="6FF9399ABA44FFCCFD8B2403FD7DFAD2" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[626,750,1356,1381]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA44FFCCFD8B2403FD7DFAD2" box="[626,750,1356,1381]" italics="true" pageId="12" pageNumber="2023">B. plicidens</emphasis>
</taxonomicName>
is a species with normal apertural barriers and
<taxonomicName id="6FF9399ABA44FFCCFED72420FE16FA3F" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[302,389,1391,1416]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="2023" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA44FFCCFED72420FE16FA3F" box="[302,389,1391,1416]" italics="true" pageId="12" pageNumber="2023">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA44FFCCFE742420FE77FA3F" box="[397,484,1391,1416]" pageId="12" pageNumber="2023" rank="species">sp. nov.</taxonomicNameLabel>
is a species with hooked barriers.
</paragraph>
</subSubSection>
</treatment>
</document>