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<document id="9F019060F1F4801B6CADD80ADF56D752" ID-CLB-Dataset="231091" ID-DOI="10.1080/00222933.2022.2152750" ID-GBIF-Dataset="53db727e-224e-4dbc-82fa-67760d26f0e3" ID-ISSN="1464-5262" ID-Zenodo-Dep="7560931" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1674200384013" checkinUser="plazi" docAuthor="Páll-Gergely, Barna &amp; White, Tom S." docDate="2023" docId="2050F30FBA46FFCFFE5E255FFDC8FC90" docLanguage="en" docName="JNATHIST.56.45-48.2011-2029.pdf" docOrigin="Journal of Natural History 56 (45 - 48)" docSource="http://dx.doi.org/10.1080/00222933.2022.2152750" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Bensonella lakainguta Hwang 2014" docType="treatment" docVersion="7" lastPageNumber="2026" masterDocId="DC698B77BA48FFC0FFF9214FFF93FFB7" masterDocTitle="Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae)" masterLastPageNumber="2029" masterPageNumber="2011" pageNumber="2025" updateTime="1701999375362" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B0FC94EE9AB20580ED56AE1FE06B3909">Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae)</mods:title>
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<mods:namePart id="7FD979A8565E8C346148597DBD0CF428">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="9BDE7E44875A89D5F37C917ABC2C58E6">White, Tom S.</mods:namePart>
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<mods:title id="A170DB7EA704702141432CBD5DF09AD4">Journal of Natural History</mods:title>
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<mods:date id="35A4529F8369E06F32B44529C19EA41F">2023</mods:date>
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<mods:number id="F0C1852B086AFFABBA453CFC4EFB1862">2023-01-18</mods:number>
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<mods:number id="6EF03E3700327BDCE86E348B566838BB">56</mods:number>
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<mods:number id="ED4CCD32A55C911F1D99AF0ABC1BB643">45 - 48</mods:number>
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<mods:start id="ABA286504D95AAED342989BCCC50E5C2">2011</mods:start>
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<treatment id="2050F30FBA46FFCFFE5E255FFDC8FC90" ID-DOI="http://doi.org/10.5281/zenodo.7560947" ID-GBIF-Taxon="209100203" ID-Zenodo-Dep="7560947" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90" httpUri="http://treatment.plazi.org/id/2050F30FBA46FFCFFE5E255FFDC8FC90" lastPageId="15" lastPageNumber="2026" pageId="14" pageNumber="2025">
<subSubSection id="E0E31192BA46FFCEFE5E255FFCEFFB9C" box="[423,892,1040,1067]" pageId="14" pageNumber="2025" type="nomenclature">
<paragraph id="A8464219BA46FFCEFE5E255FFCEFFB9C" blockId="14.[423,892,1040,1067]" box="[423,892,1040,1067]" pageId="14" pageNumber="2025">
<heading id="F30EF575BA46FFCEFE5E255FFCEFFB9C" bold="true" box="[423,892,1040,1067]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="2025" reason="3">
<taxonomicName id="6FF9399ABA46FFCEFE5E255FFCEFFB9C" ID-CoL="2fa6593c-4457-43aa-8fe7-efdd86638755" authority="Hwang, 2014" authorityName="Hwang" authorityYear="2014" box="[423,892,1040,1067]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="2025" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA46FFCEFE5E255FFCEFFB9C" bold="true" box="[423,892,1040,1067]" pageId="14" pageNumber="2025">
<emphasis id="9A8D9E0BBA46FFCEFE5E255FFD55FB9C" bold="true" box="[423,710,1040,1067]" italics="true" pageId="14" pageNumber="2025">Bensonella lakainguta</emphasis>
<bibRefCitation id="CC683FE8BA46FFCEFD36255FFCEFFB9C" author="Hwang C-C" box="[719,892,1040,1067]" pageId="14" pageNumber="2025" pagination="15 - 26" refId="ref6907" refString="Hwang C-C. 2014. A new subspecies of land snail Bensonella plicidens lakainguta (Gastropoda, Vertiginidae) from southern Taiwan. Bull Malacol Taiwan 貝類學報. 37: 15 - 26." type="journal article" year="2014">Hwang, 2014</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="E0E31192BA46FFCEFDED250CFED3FB02" pageId="14" pageNumber="2025" type="description">
<paragraph id="A8464219BA46FFCEFDED250CFC9CFBE9" blockId="14.[532,783,1091,1118]" box="[532,783,1091,1118]" pageId="14" pageNumber="2025">
(
<figureCitation id="30C25E9CBA46FFCEFDE5250CFD3FFBE9" box="[540,684,1091,1118]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStartId-0="9.[160,225,1102,1125]" captionStartId-1="10.[160,225,1556,1579]" captionTargetBox-0="[167,1147,145,1068]" captionTargetBox-1="[168,1147,145,1522]" captionTargetId-0="figure-77@9.[166,1149,144,1070]" captionTargetId-1="figure-3@10.[166,1149,144,1524]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figure 7. Bensonella lakainguta Hwang, 2014. a–c: Holotype of Bensonella lakainguta (NMNS-7244-00); D: Bensonella lakainguta Hwang, 2014, Japan,Kochi Prefecture, Kami-shi,Ryugado Prefectural National Park, above the cave, 500 m along a side-road, 290 m, 33.601917°N, 133.746217°E, Leg. A. Hunyadi, D. Murányi and B. Páll-Gergely, 8 October 2016 (HNHM 105323); e–f: Holotype of Boysidia (Bensonella) qingliangfengensis F. Fang, J. Wang and Y. Chen, 2015. Photos: a–c and e–f: images from the original descriptions, d: B. Páll-Gergely. Figs e and f are not to scale." captionText-1="Figure 8. Bensonella lakainguta Hwang, 2014, Japan, Omi, NHMUK 201901108, specimen 1. All photos: B. Páll-Gergely." figureDoi-0="http://doi.org/10.5281/zenodo.7560951" figureDoi-1="http://doi.org/10.5281/zenodo.7560955" httpUri-0="https://zenodo.org/record/7560951/files/figure.png" httpUri-1="https://zenodo.org/record/7560955/files/figure.png" pageId="14" pageNumber="2025">Figures 7–8</figureCitation>
,
<figureCitation id="30C25E9CBA46FFCEFD48250CFD53FBE9" box="[689,704,1091,1118]" captionStart="Figure 9" captionStartId="11.[160,225,749,772]" captionTargetBox="[167,1146,145,715]" captionTargetId="figure-246@11.[166,1148,144,717]" captionTargetPageId="11" captionText="Figure 9. a, b: Bensonella plicidens (Benson, 1849), NHMUK 56.9.15.69A, specimen 1 (same as Figure 4); c,d: Bensonella hooki sp. nov., holotype, NHMUK 20191114 (same as Figure 6); e,f: Bensonella lakainguta Hwang, 2014, HNHM 105323 (same as Figure 7(d)); g, h: Bensonella lakainguta, NHMUK 201901108 (same as Figure 8). All images: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7560957" httpUri="https://zenodo.org/record/7560957/files/figure.png" pageId="14" pageNumber="2025">9</figureCitation>
(e–h))
</paragraph>
<paragraph id="A8464219BA46FFCEFF592536FED3FB02" blockId="14.[160,1156,1145,1447]" pageId="14" pageNumber="2025">
<taxonomicName id="6FF9399ABA46FFCEFF592536FE7FFB25" box="[160,492,1145,1170]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="2025" phylum="Mollusca" rank="species" species="plicidens" subGenus="Bensonella">
<emphasis id="9A8D9E0BBA46FFCEFF592536FF69FB25" box="[160,250,1145,1170]" italics="true" pageId="14" pageNumber="2025">Boysidia</emphasis>
(
<emphasis id="9A8D9E0BBA46FFCEFEF02536FEEEFB25" box="[265,381,1145,1170]" italics="true" pageId="14" pageNumber="2025">Bensonella</emphasis>
)
<emphasis id="9A8D9E0BBA46FFCEFE772536FE7FFB25" box="[398,492,1145,1170]" italics="true" pageId="14" pageNumber="2025">plicidens</emphasis>
</taxonomicName>
– Pilsbry, 1917 (in
<bibRefCitation id="CC683FE8BA46FFCEFD3C2536FCDCFB25" author="Pilsbry HA" box="[709,847,1145,1170]" pageId="14" pageNumber="2025" refId="ref7592" refString="Pilsbry HA. 1916 - 1918. Manual of conchology, structural and systematic, with illustrations of the species. Ser. 2, Pulmonata. Vol. 24: Pupillidae (Gastrocoptinae). pp 1 - 380, pls 1 - 49. Philadelphia, published by the Conchological Section, Academy of Natural Sciences. [pp. 1 - 112, Pl. 1 - 13, 18 Dec 1916; pp. 113 - 176, pl. 14 - 29, 18 July 1917; pp. 177 - 256, pl. 30 - 38, 09 Nov 1917; pp. 257 - 380, pl. 39 - 49, 24 April 1918]." type="book" year="1916">Pilsbry 1916</bibRefCitation>
–1918): 198 (partim), pl. 34, figs 3, 9–10
</paragraph>
</subSubSection>
<subSubSection id="E0E31192BA46FFCEFF5925F1FEBEFA10" pageId="14" pageNumber="2025" type="reference_group">
<paragraph id="A8464219BA46FFCEFF5925F1FCBBFB60" blockId="14.[160,1156,1145,1447]" box="[160,808,1214,1239]" pageId="14" pageNumber="2025">
<treatmentCitationGroup id="88E96537BA46FFCEFF5925F1FCBBFB60" box="[160,808,1214,1239]" pageId="14" pageNumber="2025">
<taxonomicName id="6FF9399ABA46FFCEFF5925F1FE79FB60" ID-CoL="7QD4P" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,490,1214,1239]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="2025" phylum="Mollusca" rank="species" species="plicidens" subGenus="Bensonella">
<emphasis id="9A8D9E0BBA46FFCEFF5925F1FF69FB60" box="[160,250,1214,1239]" italics="true" pageId="14" pageNumber="2025">Boysidia</emphasis>
(
<emphasis id="9A8D9E0BBA46FFCEFEF025F1FEEEFB60" box="[265,381,1214,1239]" italics="true" pageId="14" pageNumber="2025">Bensonella</emphasis>
)
<emphasis id="9A8D9E0BBA46FFCEFE7525F1FE79FB60" box="[396,490,1214,1239]" italics="true" pageId="14" pageNumber="2025">plicidens</emphasis>
</taxonomicName>
–
<treatmentCitation id="29586408BA46FFCEFDFF25F1FD2BFB60" author="Habe T" box="[518,696,1214,1239]" page="109" pageId="14" pageNumber="2025" year="1956">
<bibRefCitation id="CC683FE8BA46FFCEFDFF25F1FD2BFB60" author="Habe T" box="[518,696,1214,1239]" pageId="14" pageNumber="2025" pagination="109 - 117" refId="ref6841" refString="Habe T. 1956. Anatomical studies on the Japanese Land Snails (7). Venus (Japanese Journal of Malacology). 19 (2): 109 - 117." type="journal article" year="1956">Habe 1956: 109</bibRefCitation>
</treatmentCitation>
, figs 9–11
</treatmentCitationGroup>
</paragraph>
<paragraph id="A8464219BA46FFCEFF5925AEFDD8FB4D" blockId="14.[160,1156,1145,1447]" box="[160,587,1249,1274]" pageId="14" pageNumber="2025">
<treatmentCitationGroup id="88E96537BA46FFCEFF5925AEFDD8FB4D" box="[160,587,1249,1274]" pageId="14" pageNumber="2025">
<taxonomicName id="6FF9399ABA46FFCEFF5925AEFEE9FB4D" ID-CoL="LHH3" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,378,1249,1274]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="2025" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA46FFCEFF5925AEFEE9FB4D" box="[160,378,1249,1274]" italics="true" pageId="14" pageNumber="2025">Bensonella plicidens</emphasis>
</taxonomicName>
–
<treatmentCitation id="29586408BA46FFCEFE6F25AEFDD8FB4D" author="Minato H" box="[406,587,1249,1274]" page="41" pageId="14" pageNumber="2025" year="1988">
<bibRefCitation id="CC683FE8BA46FFCEFE6F25AEFDD8FB4D" author="Minato H" box="[406,587,1249,1274]" pageId="14" pageNumber="2025" refId="ref7132" refString="Minato H. 1988. A systematic and bibliographic list of the Japanese land snails. Shirahama: H. Minato." type="book" year="1988">Minato 1988: 41</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A8464219BA46FFCEFF59244BFD47FAAA" blockId="14.[160,1156,1145,1447]" box="[160,724,1284,1309]" pageId="14" pageNumber="2025">
<treatmentCitationGroup id="88E96537BA46FFCEFF59244BFD47FAAA" box="[160,724,1284,1309]" pageId="14" pageNumber="2025">
<taxonomicName id="6FF9399ABA46FFCEFF59244BFEE9FAAA" ID-CoL="LHH3" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,378,1284,1309]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="2025" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA46FFCEFF59244BFEE9FAAA" box="[160,378,1284,1309]" italics="true" pageId="14" pageNumber="2025">Bensonella plicidens</emphasis>
</taxonomicName>
–
<treatmentCitation id="29586408BA46FFCEFE6F244BFDE4FAAA" author="Schileyko AA" box="[406,631,1284,1309]" page="139" pageId="14" pageNumber="2025" year="1998">
<bibRefCitation id="CC683FE8BA46FFCEFE6F244BFDE4FAAA" author="Schileyko AA" box="[406,631,1284,1309]" pageId="14" pageNumber="2025" pagination="129 - 261" refId="ref7802" refString="Schileyko AA. 1998. Treatise on recent terrestrial pulmonate molluscs. Part 2. Gastrocoptidae, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachnodidae, Enidae, Sagdidae. Ruthenica. Supplement 2: 129 - 261." type="journal article" year="1998">Schileyko 1998: 139</bibRefCitation>
</treatmentCitation>
, fig. 159
</treatmentCitationGroup>
</paragraph>
<paragraph id="A8464219BA46FFCEFF592469FDF8FA88" blockId="14.[160,1156,1145,1447]" box="[160,619,1318,1343]" pageId="14" pageNumber="2025">
<treatmentCitationGroup id="88E96537BA46FFCEFF592469FDF8FA88" box="[160,619,1318,1343]" pageId="14" pageNumber="2025">
<taxonomicName id="6FF9399ABA46FFCEFF592469FEE9FA88" ID-CoL="LHH3" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,378,1318,1343]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="2025" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA46FFCEFF592469FEE9FA88" box="[160,378,1318,1343]" italics="true" pageId="14" pageNumber="2025">Bensonella plicidens</emphasis>
</taxonomicName>
–
<treatmentCitation id="29586408BA46FFCEFE6F2469FDF8FA88" author="Maassen WJM" box="[406,619,1318,1343]" page="126" pageId="14" pageNumber="2025" year="1999">
<bibRefCitation id="CC683FE8BA46FFCEFE6F2469FDF8FA88" author="Maassen WJM" box="[406,619,1318,1343]" pageId="14" pageNumber="2025" pagination="121 - 126" refId="ref7100" refString="Maassen WJM. 1999. Some new records of Streptaxidae and Hypselostomatidae from Sumatra, Indonesia, with descriptions of three new species. Basteria. 63: 121 - 126." type="journal article" year="1999">Maassen 1999: 126</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A8464219BA46FFCEFF592406FC8EFAD5" blockId="14.[160,1156,1145,1447]" box="[160,797,1353,1378]" pageId="14" pageNumber="2025">
<treatmentCitationGroup id="88E96537BA46FFCEFF592406FC8EFAD5" box="[160,797,1353,1378]" pageId="14" pageNumber="2025">
<treatmentCitation id="29586408BA46FFCEFF592406FD2FFAD5" author="Hwang C-C" box="[160,700,1353,1378]" page="18" pageId="14" pageNumber="2025" year="2014">
<taxonomicName id="6FF9399ABA46FFCEFF592406FD2FFAD5" ID-CoL="5DMZD" authority="Hwang, 2014: 18" authorityName="Hwang" authorityPageNumber="18" authorityYear="2014" box="[160,700,1353,1378]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="2025" phylum="Mollusca" rank="subSpecies" species="plicidens" subSpecies="lakainguta">
<emphasis id="9A8D9E0BBA46FFCEFF592406FE65FAD5" box="[160,502,1353,1378]" italics="true" pageId="14" pageNumber="2025">Bensonella plicidens lakainguta</emphasis>
<bibRefCitation id="CC683FE8BA46FFCEFE042406FD2FFAD5" author="Hwang C-C" box="[509,700,1353,1378]" pageId="14" pageNumber="2025" pagination="15 - 26" refId="ref6907" refString="Hwang C-C. 2014. A new subspecies of land snail Bensonella plicidens lakainguta (Gastropoda, Vertiginidae) from southern Taiwan. Bull Malacol Taiwan 貝類學報. 37: 15 - 26." type="journal article" year="2014">Hwang, 2014: 18</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 2–4
</treatmentCitationGroup>
</paragraph>
<paragraph id="A8464219BA46FFCEFF592423FEBEFA10" blockId="14.[160,1156,1145,1447]" pageId="14" pageNumber="2025">
<taxonomicName id="6FF9399ABA46FFCEFF592423FB9DFA32" ID-CoL="993CJ" authority="F. Fang, J. Wang and Y. Chen, 2015: 692" authorityName="F. Fang, J. Wang and Y. Chen" authorityPageNumber="692" authorityYear="2015" box="[160,1038,1388,1413]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="2025" phylum="Mollusca" rank="species" species="qingliangfengensis" subGenus="Bensonella">
<emphasis id="9A8D9E0BBA46FFCEFF592423FF69FA32" box="[160,250,1388,1413]" italics="true" pageId="14" pageNumber="2025">Boysidia</emphasis>
(
<emphasis id="9A8D9E0BBA46FFCEFEFE2423FEE8FA32" box="[263,379,1388,1413]" italics="true" pageId="14" pageNumber="2025">Bensonella</emphasis>
)
<emphasis id="9A8D9E0BBA46FFCEFE702423FDC7FA32" box="[393,596,1388,1413]" italics="true" pageId="14" pageNumber="2025">qingliangfengensis</emphasis>
F. Fang, J. Wang and Y. Chen, 2015: 692
</taxonomicName>
, fig. 1
<emphasis id="9A8D9E0BBA46FFCEFBA82423FEBEFA10" bold="true" pageId="14" pageNumber="2025">new synonym</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="E0E31192BA46FFCFFF592499FBCDFF5A" lastPageId="15" lastPageNumber="2026" pageId="14" pageNumber="2025" type="materials_examined">
<paragraph id="A8464219BA46FFCEFF592499FEECFA47" blockId="14.[160,1157,1494,1728]" box="[160,383,1494,1520]" pageId="14" pageNumber="2025">
<emphasis id="9A8D9E0BBA46FFCEFF592499FEECFA47" bold="true" box="[160,383,1494,1520]" italics="true" pageId="14" pageNumber="2025">Material examined</emphasis>
</paragraph>
<paragraph id="A8464219BA46FFCFFF5924B5FBCDFF5A" blockId="14.[160,1157,1494,1728]" lastBlockId="15.[160,1157,142,237]" lastPageId="15" lastPageNumber="2026" pageId="14" pageNumber="2025">
<materialsCitation id="18914844BA46FFCEFF5924B5FCC7F9A4" ID-GBIF-Occurrence="4018076304" box="[160,852,1530,1555]" collectionCode="NHMUK" collectorName="Omi &amp; Hirase" lastPageId="14" lastPageNumber="2026" pageId="14" pageNumber="2025" specimenCode="NHMUK 201901108, 12">
<specimenCode id="F85FEA62BA46FFCEFF5924B5FE16F9A4" box="[160,389,1530,1555]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="2025" type="Museum">NHMUK 201901108</specimenCode>
,
<specimenCode id="F85FEA62BA46FFCEFE6A24B5FE3CF9A4" box="[403,431,1530,1555]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="2025" type="Museum">12</specimenCode>
shells,
<collectorName id="050C27CFBA46FFCEFDF324B5FDAFF9A4" box="[522,572,1530,1555]" pageId="14" pageNumber="2025">Omi</collectorName>
,
<collectingCountry id="D0EE0289BA46FFCEFDB324B5FD03F9A4" box="[586,656,1530,1555]" name="Japan" pageId="14" pageNumber="2025">Japan</collectingCountry>
,
<collectorName id="050C27CFBA46FFCEFD6724B5FD77F9A4" box="[670,740,1530,1555]" pageId="14" pageNumber="2025">Hirase</collectorName>
coll. 457
</materialsCitation>
;
<materialsCitation id="18914844BA46FFCEFC9A24B5FE4DF981" ID-GBIF-Occurrence="4018076321" collectionCode="NHMUK" collectorName="Omi" lastPageId="14" lastPageNumber="2026" pageId="14" pageNumber="2025" specimenCode="NHMUK 20191128">
<specimenCode id="F85FEA62BA46FFCEFC9A24B5FBAAF9A4" box="[867,1081,1530,1555]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="2025" type="Museum">NHMUK 20191128</specimenCode>
, three shells,
<location id="AD2614C2BA46FFCEFF0B2752FEDBF981" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA46FFCEFF0B2752FEDBF981" box="[242,328,1565,1590]" country="Japan" name="Rijozen" pageId="14" pageNumber="2025">Rijozen</location>
,
<collectorName id="050C27CFBA46FFCEFEAE2752FE1AF981" box="[343,393,1565,1590]" pageId="14" pageNumber="2025">Omi</collectorName>
,
<collectingCountry id="D0EE0289BA46FFCEFE612752FE4DF981" box="[408,478,1565,1590]" name="Japan" pageId="14" pageNumber="2025">Japan</collectingCountry>
</materialsCitation>
;
<materialsCitation id="18914844BA46FFCEFE142752FD4BF9EF" ID-GBIF-Occurrence="4018076307" collectingDate="1951-09-24" collectorName="Mt" elevation="300.0" lastPageId="14" lastPageNumber="2026" pageId="14" pageNumber="2025">
NHMW-MO-38313 (1 shell),
<collectorName id="050C27CFBA46FFCEFCC02752FCC9F981" box="[825,858,1565,1590]" pageId="14" pageNumber="2025">Mt</collectorName>
. Fujiwara,
<elevation id="23D4A52ABA46FFCEFC252752FBB8F981" box="[988,1067,1565,1590]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="2025" unit="m" value="300.0">
<quantity id="6F01EFFCBA46FFCEFC252752FBB8F981" box="[988,1067,1565,1590]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="2025" unit="m" value="300.0">300 m</quantity>
</elevation>
,
<collectingCountry id="D0EE0289BA46FFCEFBC22752FB12F981" box="[1083,1153,1565,1590]" name="Japan" pageId="14" pageNumber="2025">Japan</collectingCountry>
, coll.
<location id="AD2614C2BA46FFCEFF232770FEB1F9EF" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA46FFCEFF232770FEB1F9EF" box="[218,290,1599,1624]" name="Blume" pageId="14" pageNumber="2025">Blume</location>
ex coll.
<location id="AD2614C2BA46FFCEFE742770FE4CF9EF" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA46FFCEFE742770FE4CF9EF" box="[397,479,1599,1624]" country="Japan" name="Azuma" pageId="14" pageNumber="2025">Azuma</location>
,
<date id="DC4764D9BA46FFCEFE142770FD4BF9EF" box="[493,728,1599,1624]" pageId="14" pageNumber="2025" value="1951-09-24">
<collectingDate id="CC039D31BA46FFCEFE142770FD4BF9EF" box="[493,728,1599,1624]" pageId="14" pageNumber="2025" value="1951-09-24">24 September 1951</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="18914844BA46FFCEFD1E2770FC12F9CC" ID-GBIF-Occurrence="4018076325" elevation="300.0" lastPageId="14" lastPageNumber="2026" pageId="14" pageNumber="2025">
NHMW-MO-55810 (5 shells),
<collectingCountry id="D0EE0289BA46FFCEFBC22770FB12F9EF" box="[1083,1153,1599,1624]" name="Japan" pageId="14" pageNumber="2025">Japan</collectingCountry>
, Mie Pref.,
<location id="AD2614C2BA46FFCEFEE7272DFE52F9CC" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA46FFCEFEE7272DFE52F9CC" box="[286,449,1634,1659]" country="Japan" name="Fujiwara-dake" pageId="14" pageNumber="2025">Fujiwara-dake</location>
,
<elevation id="23D4A52ABA46FFCEFE28272DFDB3F9CC" box="[465,544,1634,1659]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="2025" unit="m" value="300.0">
<quantity id="6F01EFFCBA46FFCEFE28272DFDB3F9CC" box="[465,544,1634,1659]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="2025" unit="m" value="300.0">300 m</quantity>
</elevation>
, coll.
<location id="AD2614C2BA46FFCEFD94272DFD50F9CC" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA46FFCEFD94272DFD50F9CC" box="[621,707,1634,1659]" name="Edlauer" pageId="14" pageNumber="2025">Edlauer</location>
ex coll.
<location id="AD2614C2BA46FFCEFCCD272DFC12F9CC" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA46FFCEFCCD272DFC12F9CC" box="[820,897,1634,1659]" country="Japan" county="Mino" municipality="Yoro" name="Kuiper" pageId="14" pageNumber="2025">Kuiper</location>
</materialsCitation>
;
<materialsCitation id="18914844BA46FFCEFC68272DFEE4F929" ID-GBIF-Occurrence="4018076312" lastPageId="14" lastPageNumber="2026" pageId="14" pageNumber="2025">
NHMW-MO-38645 (4 shells),
<collectingMunicipality id="4822D863BA46FFCEFF0F27CAFEBDF929" box="[246,302,1669,1694]" pageId="14" pageNumber="2025">Yoro</collectingMunicipality>
,
<collectingCounty id="41273A95BA46FFCEFEC327CAFEE4F929" box="[314,375,1669,1694]" pageId="14" pageNumber="2025">Mino</collectingCounty>
</materialsCitation>
;
<materialsCitation id="18914844BA46FFCEFE7B27CAFEF8F977" ID-GBIF-Occurrence="4018076301" collectorName="Riozen &amp; Omi" lastPageId="14" lastPageNumber="2026" pageId="14" pageNumber="2025">
NHMW-MO-69480 (2 shells),
<collectorName id="050C27CFBA46FFCEFD3427CAFC8FF929" box="[717,796,1669,1694]" pageId="14" pageNumber="2025">Riozen</collectorName>
,
<collectorName id="050C27CFBA46FFCEFCD127CAFCC9F929" box="[808,858,1669,1694]" pageId="14" pageNumber="2025">Omi</collectorName>
,
<collectingCountry id="D0EE0289BA46FFCEFC9C27CAFC38F929" box="[869,939,1669,1694]" name="Japan" pageId="14" pageNumber="2025">Japan</collectingCountry>
, coll.
<location id="AD2614C2BA46FFCEFC1427CAFB10F929" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA46FFCEFC1427CAFB10F929" box="[1005,1155,1669,1694]" name="Oberwimmer" pageId="14" pageNumber="2025">Oberwimmer</location>
ex coll.
<location id="AD2614C2BA46FFCEFEF027E8FEF8F977" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA46FFCEFEF027E8FEF8F977" box="[265,363,1703,1728]" country="Japan" name="Jetschin" pageId="14" pageNumber="2025">Jetschin</location>
</materialsCitation>
;
<materialsCitation id="18914844BA46FFCEFE8427E8FBB5F977" ID-GBIF-Occurrence="4018076331" box="[381,1062,1703,1728]" collectorName="Omi" lastPageId="14" lastPageNumber="2026" pageId="14" pageNumber="2025">
NHMW-MO-113726 (3 shells),
<collectorName id="050C27CFBA46FFCEFD0827E8FCB7F977" box="[753,804,1703,1728]" pageId="14" pageNumber="2025">Omi</collectorName>
,
<collectingCountry id="D0EE0289BA46FFCEFCCF27E8FCEEF977" box="[822,893,1703,1728]" name="Japan" pageId="14" pageNumber="2025">Japan</collectingCountry>
, coll.
<location id="AD2614C2BA46FFCEFC3727E8FBB5F977" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA46FFCEFC3727E8FBB5F977" box="[974,1062,1703,1728]" country="Japan" name="Rusnov" pageId="14" pageNumber="2025">Rušnov</location>
</materialsCitation>
;
<materialsCitation id="18914844BA46FFCFFBC127E8FBCDFF5A" ID-GBIF-Occurrence="4018076329" collectingDate="2016-08-08" collectorName="Leg" elevation="290.0" lastPageId="15" lastPageNumber="2026" latitude="33.601917" longitude="133.74622" pageId="14" pageNumber="2025">
<collectionCode id="CEE8DADCBA46FFCEFBC127E8FB17F977" box="[1080,1156,1703,1728]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="14" pageNumber="2025">HNHM</collectionCode>
105323 (figured shell),
<collectingCountry id="D0EE0289BA47FFCFFE3421C1FD86FF10" box="[461,533,142,167]" name="Japan" pageId="15" pageNumber="2026">Japan</collectingCountry>
,
<collectingRegion id="6A3D8CFBBA47FFCFFDD321C1FD6AFF10" box="[554,761,142,167]" country="Japan" name="Kochi" pageId="15" pageNumber="2026">Kochi Prefecture</collectingRegion>
,
<collectingMunicipality id="4822D863BA47FFCFFCF721C1FCE6FF10" box="[782,885,142,167]" pageId="15" pageNumber="2026">Kami-shi</collectingMunicipality>
,
<location id="AD2614C2BA47FFCFFC7221C1FEDAFF7D" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA47FFCFFC7221C1FEDAFF7D" country="Japan" latitude="33.601917" longLatPrecision="1" longitude="133.74622" municipality="Kami-shi" name="Ryugado Prefectural National Park" pageId="15" pageNumber="2026" stateProvince="Kochi">Ryugado Prefectural National Park</location>
, above the cave,
<quantity id="6F01EFFCBA47FFCFFDCA21FEFD10FF7D" box="[563,643,177,202]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="2026" unit="m" value="500.0">
<elevation id="23D4A52ABA47FFCFFDCA21FEFD10FF7D" box="[563,643,177,202]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="2026" unit="m" value="500.0">500 m</elevation>
</quantity>
along a side-road, 290 m,
<geoCoordinate id="CDCD24DEBA47FFCFFC1321FEFB12FF7D" box="[1002,1153,177,202]" degrees="33.601917" direction="north" orientation="latitude" pageId="15" pageNumber="2026" precision="1" value="33.601917">33.601917°N</geoCoordinate>
,
<geoCoordinate id="CDCD24DEBA47FFCFFF58219BFEAEFF5A" box="[161,317,212,237]" degrees="133.746217" direction="east" orientation="longitude" pageId="15" pageNumber="2026" precision="1" value="133.74622">133.746217°E</geoCoordinate>
,
<collectorName id="050C27CFBA47FFCFFEB3219BFEE5FF5A" box="[330,374,212,237]" pageId="15" pageNumber="2026">Leg</collectorName>
. Hunyadi, A., Murányi, D. and Páll-Gergely, B.,
<date id="DC4764D9BA47FFCFFC67219BFBC9FF5A" box="[926,1114,212,237]" pageId="15" pageNumber="2026" value="2016-08-08">08 August 2016</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E0E31192BA47FFCFFF592067FDC8FC90" pageId="15" pageNumber="2026" type="discussion">
<paragraph id="A8464219BA47FFCFFF592067FE95FEF5" blockId="15.[160,1156,296,807]" box="[160,262,296,322]" pageId="15" pageNumber="2026">
<emphasis id="9A8D9E0BBA47FFCFFF592067FE95FEF5" bold="true" box="[160,262,296,322]" italics="true" pageId="15" pageNumber="2026">Remarks</emphasis>
</paragraph>
<paragraph id="A8464219BA47FFCFFF592003FD78FDA5" blockId="15.[160,1156,296,807]" pageId="15" pageNumber="2026">
In the original description,
<bibRefCitation id="CC683FE8BA47FFCFFE202003FDEAFED2" author="Hwang C-C" box="[473,633,332,357]" pageId="15" pageNumber="2026" pagination="15 - 26" refId="ref6907" refString="Hwang C-C. 2014. A new subspecies of land snail Bensonella plicidens lakainguta (Gastropoda, Vertiginidae) from southern Taiwan. Bull Malacol Taiwan 貝類學報. 37: 15 - 26." type="journal article" year="2014">Hwang (2014)</bibRefCitation>
mentioned three differences between Indian
<taxonomicName id="6FF9399ABA47FFCFFF592021FE8FFE30" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,284,366,391]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA47FFCFFF592021FE8FFE30" box="[160,284,366,391]" italics="true" pageId="15" pageNumber="2026">B. plicidens</emphasis>
</taxonomicName>
and the subspecies
<taxonomicName id="6FF9399ABA47FFCFFDF62021FCF8FE30" authorityName="Hwang" authorityYear="2014" box="[527,875,366,391]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="subSpecies" species="plicidens" subSpecies="lakainguta">
<emphasis id="9A8D9E0BBA47FFCFFDF62021FCF8FE30" box="[527,875,366,391]" italics="true" pageId="15" pageNumber="2026">Bensonella plicidens lakainguta</emphasis>
</taxonomicName>
. The most important of them was the submarginal elevations of columellar, basal and palatal barriers, i.e. anterior to the hooks those barriers become more elevated. However, this trait can also be seen in Indian
<taxonomicName id="6FF9399ABA47FFCFFF142099FED0FE58" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[237,323,470,495]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA47FFCFFF142099FED0FE58" box="[237,323,470,495]" italics="true" pageId="15" pageNumber="2026">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA47FFCFFEB32099FE32FE58" box="[330,417,470,495]" pageId="15" pageNumber="2026" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="30C25E9CBA47FFCFFE562099FD9FFE58" box="[431,524,470,495]" captionStart="Figure 9" captionStartId="11.[160,225,749,772]" captionTargetBox="[167,1146,145,715]" captionTargetId="figure-246@11.[166,1148,144,717]" captionTargetPageId="11" captionText="Figure 9. a, b: Bensonella plicidens (Benson, 1849), NHMUK 56.9.15.69A, specimen 1 (same as Figure 4); c,d: Bensonella hooki sp. nov., holotype, NHMUK 20191114 (same as Figure 6); e,f: Bensonella lakainguta Hwang, 2014, HNHM 105323 (same as Figure 7(d)); g, h: Bensonella lakainguta, NHMUK 201901108 (same as Figure 8). All images: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7560957" httpUri="https://zenodo.org/record/7560957/files/figure.png" pageId="15" pageNumber="2026">Figure 9</figureCitation>
(c,d)); see differences between
<taxonomicName id="6FF9399ABA47FFCFFC9C2099FC67FE58" authorityName="Hwang" authorityYear="2014" box="[869,1012,470,495]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA47FFCFFC9C2099FC67FE58" box="[869,1012,470,495]" italics="true" pageId="15" pageNumber="2026">B. lakainguta</emphasis>
</taxonomicName>
and
<taxonomicName id="6FF9399ABA47FFCFFBD42099FB10FE58" authority="Páll-Gergely &amp; White, 2023" authorityName="Páll-Gergely &amp; White" authorityYear="2023" box="[1069,1155,470,495]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
<emphasis id="9A8D9E0BBA47FFCFFBD42099FB10FE58" box="[1069,1155,470,495]" italics="true" pageId="15" pageNumber="2026">B. hooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81BE2370BA47FFCFFF5920B6FF64FDA5" box="[160,247,505,530]" pageId="15" pageNumber="2026" rank="species">sp. nov.</taxonomicNameLabel>
outlined under the description of the latter.
</paragraph>
<paragraph id="A8464219BA47FFCFFF392353FF4EFD08" blockId="15.[160,1156,296,807]" pageId="15" pageNumber="2026">
<taxonomicName id="6FF9399ABA47FFCFFF392353FDE9FD82" authorityName="F. Fang, J. Wang and Y. Chen" authorityYear="2015" box="[192,634,540,565]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="qingliangfengensis" subGenus="Bensonella">
<emphasis id="9A8D9E0BBA47FFCFFF392353FE89FD82" box="[192,282,540,565]" italics="true" pageId="15" pageNumber="2026">Boysidia</emphasis>
(
<emphasis id="9A8D9E0BBA47FFCFFED32353FE0DFD82" box="[298,414,540,565]" italics="true" pageId="15" pageNumber="2026">Bensonella</emphasis>
)
<emphasis id="9A8D9E0BBA47FFCFFE562353FDE9FD82" box="[431,634,540,565]" italics="true" pageId="15" pageNumber="2026">qingliangfengensis</emphasis>
</taxonomicName>
(
<figureCitation id="30C25E9CBA47FFCFFD732353FD7AFD82" box="[650,745,540,565]" captionStart="Figure 7" captionStartId="9.[160,225,1102,1125]" captionTargetBox="[167,1147,145,1068]" captionTargetId="figure-77@9.[166,1149,144,1070]" captionTargetPageId="9" captionText="Figure 7. Bensonella lakainguta Hwang, 2014. a–c: Holotype of Bensonella lakainguta (NMNS-7244-00); D: Bensonella lakainguta Hwang, 2014, Japan,Kochi Prefecture, Kami-shi,Ryugado Prefectural National Park, above the cave, 500 m along a side-road, 290 m, 33.601917°N, 133.746217°E, Leg. A. Hunyadi, D. Murányi and B. Páll-Gergely, 8 October 2016 (HNHM 105323); e–f: Holotype of Boysidia (Bensonella) qingliangfengensis F. Fang, J. Wang and Y. Chen, 2015. Photos: a–c and e–f: images from the original descriptions, d: B. Páll-Gergely. Figs e and f are not to scale." figureDoi="http://doi.org/10.5281/zenodo.7560951" httpUri="https://zenodo.org/record/7560951/files/figure.png" pageId="15" pageNumber="2026">Figure 7</figureCitation>
(e,f))
<materialsCitation id="18914844BA47FFCFFCDB2353FD62FDCD" ID-GBIF-Occurrence="4018076315" collectorName="Hangzhou" location="Provinces" municipality="Chinese" pageId="15" pageNumber="2026" specimenCount="1" stateProvince="Anhui and Zhejiang">
was described from the border region of the
<collectingMunicipality id="4822D863BA47FFCFFEC02371FE01FDE0" box="[313,402,574,599]" pageId="15" pageNumber="2026">Chinese</collectingMunicipality>
<collectingRegion id="6A3D8CFBBA47FFCFFE602371FDE4FDE0" box="[409,631,574,599]" isEnumeration="true" pageId="15" pageNumber="2026">Anhui and Zhejiang</collectingRegion>
<location id="AD2614C2BA47FFCFFD872371FD7AFDE0" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA47FFCFFD872371FD7AFDE0" box="[638,745,574,599]" municipality="Chinese" name="Provinces" pageId="15" pageNumber="2026" stateProvince="Anhui and Zhejiang">Provinces</location>
(very close to Pilsbry̾s (1917) record from
<collectorName id="050C27CFBA47FFCFFF27232EFEC5FDCD" box="[222,342,609,634]" pageId="15" pageNumber="2026">Hangzhou</collectorName>
), and is claimed to be larger than
<taxonomicName id="6FF9399ABA47FFCFFD18232EFD62FDCD" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[737,753,609,634]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA47FFCFFD18232EFD62FDCD" box="[737,753,609,634]" italics="true" pageId="15" pageNumber="2026">B</emphasis>
</taxonomicName>
</materialsCitation>
<emphasis id="9A8D9E0BBA47FFCFFD0B232EFCCFFDCD" box="[754,860,609,634]" italics="true" pageId="15" pageNumber="2026">. plicidens</emphasis>
, yet looks identical for all other characters.
<materialsCitation id="18914844BA47FFCFFE9823CBFC3AFD2A" ID-GBIF-Occurrence="4018076313" box="[353,937,644,669]" location="Therefore" pageId="15" pageNumber="2026" specimenCount="1" stateProvince="Anhui and Zhejiang">
<location id="AD2614C2BA47FFCFFE9823CBFE43FD2A" LSID="urn:lsid:plazi:treatment:2050F30FBA46FFCFFE5E255FFDC8FC90:AD2614C2BA47FFCFFE9823CBFE43FD2A" box="[353,464,644,669]" name="Therefore" pageId="15" pageNumber="2026" stateProvince="Anhui and Zhejiang">Therefore</location>
, it is treated here as a junior synonym of
<taxonomicName id="6FF9399ABA47FFCFFC6023CBFC3AFD2A" authority="Hwang, 2014" authorityName="Hwang" authorityYear="2014" box="[921,937,644,669]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA47FFCFFC6023CBFC3AFD2A" box="[921,937,644,669]" italics="true" pageId="15" pageNumber="2026">B</emphasis>
</taxonomicName>
</materialsCitation>
<emphasis id="9A8D9E0BBA47FFCFFC5023CBFBBBFD2A" box="[937,1064,644,669]" italics="true" pageId="15" pageNumber="2026">. lakainguta</emphasis>
<bibRefCitation id="CC683FE8BA47FFCFFBD723CBFF44FD08" author="Hwang C-C" pageId="15" pageNumber="2026" pagination="15 - 26" refId="ref6907" refString="Hwang C-C. 2014. A new subspecies of land snail Bensonella plicidens lakainguta (Gastropoda, Vertiginidae) from southern Taiwan. Bull Malacol Taiwan 貝類學報. 37: 15 - 26." type="journal article" year="2014">Hwang, 2014</bibRefCitation>
.
</paragraph>
<paragraph id="A8464219BA47FFCFFF392386FDC8FC90" blockId="15.[160,1156,296,807]" pageId="15" pageNumber="2026">
<bibRefCitation id="CC683FE8BA47FFCFFF392386FEFFFD55" author="Maassen WJM" box="[192,364,713,738]" pageId="15" pageNumber="2026" pagination="121 - 126" refId="ref7100" refString="Maassen WJM. 1999. Some new records of Streptaxidae and Hypselostomatidae from Sumatra, Indonesia, with descriptions of three new species. Basteria. 63: 121 - 126." type="journal article" year="1999">Maassen (1999)</bibRefCitation>
mentioned that he compared
<taxonomicName id="6FF9399ABA47FFCFFD472386FC0EFD55" authorityName="Maassen" authorityYear="1999" box="[702,925,713,738]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="karoensis">
<emphasis id="9A8D9E0BBA47FFCFFD472386FC0EFD55" box="[702,925,713,738]" italics="true" pageId="15" pageNumber="2026">Bensonella karoensis</emphasis>
</taxonomicName>
with Japanese shells of
<taxonomicName id="6FF9399ABA47FFCFFF4423A3FE04FCB2" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[189,407,748,773]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA47FFCFFF4423A3FE04FCB2" box="[189,407,748,773]" italics="true" pageId="15" pageNumber="2026">Bensonella plicidens</emphasis>
</taxonomicName>
(=
<taxonomicName id="6FF9399ABA47FFCFFE4523A3FD3EFCB2" authorityName="Hwang" authorityYear="2014" box="[444,685,748,773]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="lakainguta">
<emphasis id="9A8D9E0BBA47FFCFFE4523A3FD3EFCB2" box="[444,685,748,773]" italics="true" pageId="15" pageNumber="2026">Bensonella lakainguta</emphasis>
</taxonomicName>
). Furthermore, he erroneously stated that
<taxonomicName id="6FF9399ABA47FFCFFF592241FEE9FC90" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,378,782,807]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="2026" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="9A8D9E0BBA47FFCFFF592241FEE9FC90" box="[160,378,782,807]" italics="true" pageId="15" pageNumber="2026">Bensonella plicidens</emphasis>
</taxonomicName>
inhabited
<collectingCountry id="D0EE0289BA47FFCFFE0D2241FDC4FC90" box="[500,599,782,807]" name="Thailand" pageId="15" pageNumber="2026">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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